471 lines
63 KiB
XML
471 lines
63 KiB
XML
<document id="F6AAC608A9674B00E0EF4620A09735DB" ID-DOI="10.5281/zenodo.175024" ID-GBIF-Dataset="bcb5927c-502d-414e-bdaf-142164323e8a" ID-ISSN="1175-5326" ID-Zenodo-Dep="175024" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459794732622" checkinUser="plazi" docAuthor="Kok, Philippe J. R." docDate="2006" docId="03E2002DFF87FFD69537A29A74E1AE5E" docLanguage="en" docName="zt01378p035.pdf" docOrigin="Zootaxa 1378" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Atractus tamessari Kok, 2006, sp. nov." docType="treatment" docVersion="9" lastPageNumber="25" masterDocId="FFDB7855FF86FFD1943FA42C752CA821" masterDocTitle="A new snake of the genus Atractus Wagler, 1828 (Reptilia: Squamata: Colubridae) from Kaieteur National Park, Guyana, northeastern South America" masterLastPageNumber="35" masterPageNumber="19" pageNumber="20" updateTime="1698219557935" updateUser="plazi">
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<mods:title id="8529C8332C932271AA322AB452305392">A new snake of the genus Atractus Wagler, 1828 (Reptilia: Squamata: Colubridae) from Kaieteur National Park, Guyana, northeastern South America</mods:title>
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<mods:namePart id="F231C17440CB0B62C84F70200C632186">Kok, Philippe J. R.</mods:namePart>
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<mods:title id="87DA537A5E8EBDF9FDFB53F7E5F24C18">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="A3E53972FF1DCEB8A757BDCA0D47F6E6">
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<mods:date id="2AF3A7EEFD9912595B91851C7A1177F3">2006</mods:date>
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<mods:number id="A39B3F8E6281E0496ED3BBE4DA7DEBAF">1378</mods:number>
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<mods:identifier id="0BD08EBE9409C63A582369D85DA28FE1" type="DOI">10.5281/zenodo.175024</mods:identifier>
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<treatment id="03E2002DFF87FFD69537A29A74E1AE5E" ID-DOI="http://doi.org/10.5281/zenodo.5693696" ID-GBIF-Taxon="119350927" ID-Zenodo-Dep="5693696" LSID="urn:lsid:plazi:treatment:03E2002DFF87FFD69537A29A74E1AE5E" httpUri="http://treatment.plazi.org/id/03E2002DFF87FFD69537A29A74E1AE5E" lastPageId="7" lastPageNumber="25" pageId="1" pageNumber="20">
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<subSubSection id="C351E2B0FF87FFD09537A29A7441AED2" pageId="1" pageNumber="20" type="nomenclature">
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<paragraph id="8BF4B13BFF87FFD09537A29A7762AEF1" blockId="1.[264,590,1718,1779]" box="[264,590,1718,1744]" pageId="1" pageNumber="20">
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<heading id="D0BC0657FF87FFD09537A29A7762AEF1" bold="true" box="[264,590,1718,1744]" fontSize="11" level="1" pageId="1" pageNumber="20" reason="1">
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<emphasis id="B93F6D29FF87FFD09537A29A7762AEF1" bold="true" box="[264,590,1718,1744]" pageId="1" pageNumber="20">
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<taxonomicName id="4C4BCAB8FF87FFD09537A29A74C5AEF1" box="[264,489,1718,1744]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="20" phylum="Chordata" rank="species" species="tamessari" status="sp. nov.">
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<emphasis id="B93F6D29FF87FFD09537A29A74C5AEF1" bold="true" box="[264,489,1718,1744]" italics="true" pageId="1" pageNumber="20">Atractus tamessari</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20CD052FF87FFD095CEA29A7762AEF1" box="[497,590,1718,1744]" pageId="1" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BF4B13BFF87FFD09537A2F57441AED2" blockId="1.[264,590,1718,1779]" box="[264,365,1753,1779]" pageId="1" pageNumber="20">
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<figureCitation id="1370ADBEFF87FFD09537A2F57441AED2" box="[264,365,1753,1779]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="4.[264,367,1841,1865]" captionStartId-1="5.[264,367,1805,1829]" captionStartId-2="7.[264,367,1007,1031]" captionStartId-3="8.[264,367,1756,1780]" captionTargetBox-0="[340,1223,325,1778]" captionTargetBox-1="[439,1155,315,1772]" captionTargetBox-2="[264,1323,326,983]" captionTargetBox-3="[264,1322,846,1732]" captionTargetId-0="figure@4.[327,1250,279,1805]" captionTargetId-1="figure@5.[439,1157,302,1784]" captionTargetId-2="figure@7.[264,1323,326,983]" captionTargetId-3="figure@8.[264,1323,846,1732]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="7" captionTargetPageId-3="8" captionText-0="FIGURE 1. Atractus tamessari sp. nov. (holotype, IRSNB 2640): (A) dorsal, (B) lateral, and (C) ventral views of the head." captionText-1="FIGURE 2. Atractus tamessari sp. nov.: (A) the male holotype (IRSNB 2640) in life, (B) ventral view of the male holotype (IRSNB 2640) in life, and (C) a subadult female paratopotype (IRSNB 2641) in life." captionText-2="FIGURE 3. Almost fully everted right hemipenis of Atractus tamessari sp. nov. (holotype, IRSNB 2640): (left) sulcate, and (right) asulcate sides; scale = 5 mm." captionText-3="FIGURE 4. Atractus tamessari sp. nov.: (A) dorsal view of the preserved male holotype (IRSNB 2640), (B) dorsal view of a preserved subadult female paratopotype (IRSNB 2641), (C) ventral view of the preserved male holotype (IRSNB 2640), and (D) ventral view of a preserved subadult female paratopotype (IRSNB 2641)." httpUri-0="https://zenodo.org/record/175025/files/figure.png" httpUri-1="https://zenodo.org/record/175026/files/figure.png" httpUri-2="https://zenodo.org/record/175027/files/figure.png" httpUri-3="https://zenodo.org/record/175028/files/figure.png" pageId="1" pageNumber="20">Figs 1–4</figureCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C351E2B0FF87FFD39537A3057156A9F7" lastPageId="2" lastPageNumber="21" pageId="1" pageNumber="20" type="materials_examined">
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<paragraph id="8BF4B13BFF87FFD39537A30574C7A97F" blockId="1.[264,1323,1833,1899]" lastBlockId="2.[264,1323,284,1870]" lastPageId="2" lastPageNumber="21" pageId="1" pageNumber="20">
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<emphasis id="B93F6D29FF87FFD09537A30574ADAF62" bold="true" box="[264,385,1833,1859]" pageId="1" pageNumber="20">
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<typeStatus id="54F00F99FF87FFD09537A3057450AF62" box="[264,380,1833,1859]" pageId="1" pageNumber="20" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<collectionCode id="ED5A29FEFF87FFD095B4A30574CEAF62" box="[395,482,1833,1859]" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="1" pageNumber="20">IRSNB</collectionCode>
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2640 (field n° PK 1365), an adult male, collected along a tributary of Elinkwa River, ESE Kaieteur National Park,
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<emphasis id="B93F6D29FF87FFD09729A37D7615AF4B" box="[790,825,1873,1898]" italics="true" pageId="1" pageNumber="20">ca.</emphasis>
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<quantity id="4CB31CDEFF87FFD0977FA37D76A6AF4A" box="[832,906,1873,1899]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="20" unit="m" value="500.0">500 m</quantity>
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elevation, Potaro-Siparuni District,
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<collectingCountry id="F35CF1ABFF84FFD39537A530744FA917" box="[264,355,284,310]" name="Guyana" pageId="2" pageNumber="21">Guyana</collectingCountry>
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(
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<geoCoordinate id="EE7FD7FCFF84FFD3954CA53074DBA917" box="[371,503,284,310]" direction="north" orientation="latitude" pageId="2" pageNumber="21" precision="15" value="5.1358333">5°08’09”N</geoCoordinate>
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,
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<geoCoordinate id="EE7FD7FCFF84FFD3963CA53077B0A917" box="[515,668,284,310]" direction="west" orientation="longitude" pageId="2" pageNumber="21" precision="15" value="-59.424446">59°25’28”W</geoCoordinate>
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), on
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<date id="FFF597FBFF84FFD396E9A53076A4A917" box="[726,904,284,310]" pageId="2" pageNumber="21" value="2006-03-23">23 March 2006</date>
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by P. J. R. Kok, Paul Benjamin and Giuliano Seegobin.
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</paragraph>
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<paragraph id="8BF4B13BFF84FFD39507A5477156A9F7" blockId="2.[264,1323,284,1870]" pageId="2" pageNumber="21">
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<emphasis id="B93F6D29FF84FFD39507A5477490A9A4" bold="true" box="[312,444,363,389]" pageId="2" pageNumber="21">
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<typeStatus id="54F00F99FF84FFD39507A547749BA9A4" box="[312,439,363,389]" pageId="2" pageNumber="21" type="paratype">Paratypes</typeStatus>
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.
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</emphasis>
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<collectionCode id="ED5A29FEFF84FFD395FBA5407736A9A7" box="[452,538,364,390]" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="2" pageNumber="21">IRSNB</collectionCode>
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2641 (field n° PK 1372), a subadult female collected on
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<date id="FFF597FBFF84FFD39086A540746AA98F" pageId="2" pageNumber="21" value="2006-03-24">24 March 2006</date>
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, same data as the
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<typeStatus id="54F00F99FF84FFD39605A5B87788A98F" box="[570,676,404,430]" pageId="2" pageNumber="21" type="holotype">holotype</typeStatus>
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.
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<collectionCode id="ED5A29FEFF84FFD39689A5B87620A98F" box="[694,780,404,430]" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="2" pageNumber="21">IRSNB</collectionCode>
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2642 (field n° PK 1550), an adult female collected at the same locality as the
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<typeStatus id="54F00F99FF84FFD39691A5907634A9F7" box="[686,792,444,470]" pageId="2" pageNumber="21" type="holotype">holotype</typeStatus>
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, in
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<date id="FFF597FBFF84FFD3977EA5907695A9F7" box="[833,953,444,470]" pageId="2" pageNumber="21" value="2006-05">May 2006</date>
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by G. Seegobin.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C351E2B0FF84FFD39507A5CF7703AAE7" pageId="2" pageNumber="21" type="etymology">
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<paragraph id="8BF4B13BFF84FFD39507A5CF7703AAE7" blockId="2.[264,1323,284,1870]" pageId="2" pageNumber="21">
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<emphasis id="B93F6D29FF84FFD39507A5CF74E8A9DC" bold="true" box="[312,452,483,509]" pageId="2" pageNumber="21">Etymology.</emphasis>
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The specific name is a noun in the genitive case, honoring Michael Tamessar, retired Senior Scientific Officer of the Department of Biology at the University of
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<collectingCountry id="F35CF1ABFF84FFD39517A61874AFAA6F" box="[296,387,564,590]" name="Guyana" pageId="2" pageNumber="21">Guyana</collectingCountry>
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(UG). Mike Tamessar spent most of his life encouraging Guyanese students to pursue studies in the fields of ichthyology and herpetology. Mike also collected many of the amphibians and reptiles in the collection of the Centre for Study of the Biological Diversity (CSBD) at UG.
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</paragraph>
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</subSubSection>
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<subSubSection id="C351E2B0FF84FFD29507A6F877C2AB47" lastPageId="3" lastPageNumber="22" pageId="2" pageNumber="21" type="diagnosis">
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<paragraph id="8BF4B13BFF84FFD39507A6F874E8AC0F" blockId="2.[264,1323,284,1870]" pageId="2" pageNumber="21">
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<emphasis id="B93F6D29FF84FFD39507A6F8749BAACF" bold="true" box="[312,439,724,750]" pageId="2" pageNumber="21">Diagnosis.</emphasis>
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An
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<taxonomicName id="4C4BCAB8FF84FFD395CFA6F8777FAACC" box="[496,595,724,749]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="21" phylum="Chordata" rank="genus">
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<emphasis id="B93F6D29FF84FFD395CFA6F8777FAACC" box="[496,595,724,749]" italics="true" pageId="2" pageNumber="21">Atractus</emphasis>
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</taxonomicName>
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species with 15 dorsal scale rows that can be distinguished from all other congeners by the following combination of characters: (1) medium size, adults reaching about
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<quantity id="4CB31CDEFF84FFD39632A7087743AB1F" box="[525,623,804,830]" metricMagnitude="-1" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="21" unit="mm" value="400.0">400 mm</quantity>
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in TTL; (2) loreal much longer than high and touching the eye; (3) two postoculars; (4) eight supralabials, fourth and fifth touching the eye; (5) seven to eight infralabials, four in contact with chinshield; (6) six maxillary teeth; (7) 153–163 ventrals; (8) 28–31 subcaudals; (9) dorsal color pattern of irregular red or pale red markings, sometimes forming an incomplete broken dorsolateral stripe, on a medium brown to brownish black ground color; (10) venter yellowish cream, heavily mottled with brownish black.
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</paragraph>
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<paragraph id="8BF4B13BFF84FFD39507A010779EAD67" blockId="2.[264,1323,284,1870]" pageId="2" pageNumber="21">
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The new species can be referred to the genus
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<taxonomicName id="4C4BCAB8FF84FFD397B3A01076C3AC74" box="[908,1007,1084,1109]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="21" phylum="Chordata" rank="genus">
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<emphasis id="B93F6D29FF84FFD397B3A01076C3AC74" box="[908,1007,1084,1109]" italics="true" pageId="2" pageNumber="21">Atractus</emphasis>
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</taxonomicName>
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because it possesses the following combination of characters: medium size; head small, slightly distinct from neck; eye small, with semi-elliptical pupil; scales smooth; dorsal scales in 15 rows without reduction; loreal present, touching the eye and separating prefrontals and supralabials; nostril between two scales; single pair of chinshields; anal single; subcaudals divided; maxillary teeth decreasing in size posteriorly; occurring in northern South
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<collectingCountry id="F35CF1ABFF84FFD390FAA1287005AD3F" box="[1221,1321,1284,1310]" name="United States of America" pageId="2" pageNumber="21">America</collectingCountry>
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(
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<bibRefCitation id="EFDACCCAFF84FFD3952FA10074FCAD67" author="Boulenger" box="[272,464,1324,1350]" pageId="2" pageNumber="31" refString="Boulenger, G. A. (1894) Catalogue of the Snakes in the British Museum (Natural History), Volume II. Taylor and Francis, London, 382 pp." type="book" year="1894">Boulenger 1894</bibRefCitation>
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,
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<bibRefCitation id="EFDACCCAFF84FFD395E3A1007789AD67" author="Hoogmoed" box="[476,677,1324,1350]" pageId="2" pageNumber="32" refString="Hoogmoed, M. S. (1980) Revision of the genus Atractus in Surinam, with the resurrection of two species (Colubridae, Reptilia). Zoologische Verhandelingen, 175, 1 - 47." type="journal article" year="1980">Hoogmoed 1980</bibRefCitation>
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||
).
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</paragraph>
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<paragraph id="8BF4B13BFF84FFD29507A17877C2AB47" blockId="2.[264,1323,284,1870]" lastBlockId="3.[264,1323,284,1870]" lastPageId="3" lastPageNumber="22" pageId="2" pageNumber="21">
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By having 15 dorsal scale rows at midbody,
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<taxonomicName id="4C4BCAB8FF84FFD39746A1787171AD4C" box="[889,1117,1364,1389]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="tamessari">
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<emphasis id="B93F6D29FF84FFD39746A1787171AD4C" box="[889,1117,1364,1389]" italics="true" pageId="2" pageNumber="21">Atractus tamessari</emphasis>
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||
</taxonomicName>
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differs from all
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||
<taxonomicName id="4C4BCAB8FF84FFD39537A1507447ADB4" box="[264,363,1404,1429]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="21" phylum="Chordata" rank="genus">
|
||
<emphasis id="B93F6D29FF84FFD39537A1507447ADB4" box="[264,363,1404,1429]" italics="true" pageId="2" pageNumber="21">Atractus</emphasis>
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||
</taxonomicName>
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||
species having 17 scale rows in the
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<collectingCountry id="F35CF1ABFF84FFD39725A1507643ADB7" box="[794,879,1404,1430]" name="Guyana" pageId="2" pageNumber="21">Guiana</collectingCountry>
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||
Shield (
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||
<bibRefCitation id="EFDACCCAFF84FFD397EDA15071B7ADB7" author="Silva" box="[978,1179,1404,1430]" pageId="2" pageNumber="34" refString="Silva Haad, J. J. (2004) Las serpientes del genero Atractus Wagler, 1828 (Colubridae, Xenodontinae) en la Amazonia Colombiana. Revista de la Academia Colombiana de Ciencias Exactas, Fisicas y Naturales, 28, 409 - 446." type="journal article" year="2004">Silva Haad 2004</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFDACCCAFF84FFD39097A1507464AD9F" author="Avila-Pires" pageId="2" pageNumber="31" refString="Avila-Pires, T. C. S. (2005) Reptiles. In: Hollowell, T., and Reynolds, R. P. (Eds.), Checklist of the terrestrial vertebrates of the Guiana Shield. Bulletin of the Biological Society of Washington, 13, pp. 24 - 40." type="journal article" year="2005">Avila-Pires 2005</bibRefCitation>
|
||
):
|
||
<taxonomicName id="4C4BCAB8FF84FFD39562A1887706AD9C" box="[349,554,1444,1469]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="alphonsehogei">
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||
<emphasis id="B93F6D29FF84FFD39562A1887706AD9C" box="[349,554,1444,1469]" italics="true" pageId="2" pageNumber="21">A. alphonsehogei</emphasis>
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||
</taxonomicName>
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||
<bibRefCitation id="EFDACCCAFF84FFD3960DA188765BAD9F" author="Cunha" box="[562,887,1444,1470]" pageId="2" pageNumber="32" refString="Cunha, O. R. & Nascimento, F. P. (1983) Ofidios da Amazonia XX: as especies de Atractus Wagler, 1828, na Amazonia oriental e Maranhao (Ophidia, Colubridae). Boletim do Museu Paraense Emilio Goeldi, 123, 1 - 38." type="journal article" year="1983">
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||
<collectingCountry id="F35CF1ABFF84FFD3960DA1887752AD9F" box="[562,638,1444,1470]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="2" pageNumber="21">Cunha</collectingCountry>
|
||
& Nascimento 1983
|
||
</bibRefCitation>
|
||
,
|
||
<taxonomicName id="4C4BCAB8FF84FFD397BBA18871A0AD9F" authority="Boie 1827" authorityName="Boie" authorityYear="1827" box="[900,1164,1444,1470]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="badius">
|
||
<emphasis id="B93F6D29FF84FFD397BBA18876DFAD9C" box="[900,1011,1444,1469]" italics="true" pageId="2" pageNumber="21">A. badius</emphasis>
|
||
(
|
||
<bibRefCitation id="EFDACCCAFF84FFD3903BA18871AFAD9F" author="Boie" box="[1028,1155,1444,1470]" pageId="2" pageNumber="31" refString="Boie, F. (1827) Bemerkungen uber Merrem's Versuch eines Systems der Amphibien. Marburg. 1820. Erste Lieferung: Ophidier. Isis von Oken, 20, 508 - 566." type="journal article" year="1827">Boie 1827</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4BCAB8FF84FFD390A6A18874FEADC7" authority="Silva Haad 2004" authorityName="Silva Haad" authorityYear="2004" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="charitoae">
|
||
<emphasis id="B93F6D29FF84FFD390A6A1887006AD9C" box="[1177,1322,1444,1469]" italics="true" pageId="2" pageNumber="21">A. charitoae</emphasis>
|
||
<bibRefCitation id="EFDACCCAFF84FFD39537A1E074FEADC7" author="Silva" box="[264,466,1484,1510]" pageId="2" pageNumber="34" refString="Silva Haad, J. J. (2004) Las serpientes del genero Atractus Wagler, 1828 (Colubridae, Xenodontinae) en la Amazonia Colombiana. Revista de la Academia Colombiana de Ciencias Exactas, Fisicas y Naturales, 28, 409 - 446." type="journal article" year="2004">Silva Haad 2004</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4BCAB8FF84FFD395DFA1E077D0ADC7" authority="Roze 1961" authorityName="Roze" authorityYear="1961" box="[480,764,1484,1510]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="duidensis">
|
||
<emphasis id="B93F6D29FF84FFD395DFA1E0775DADC4" box="[480,625,1484,1509]" italics="true" pageId="2" pageNumber="21">A. duidensis</emphasis>
|
||
<bibRefCitation id="EFDACCCAFF84FFD39645A1E077D0ADC7" author="Roze" box="[634,764,1484,1510]" lastPageId="2" lastPageNumber="34" pageId="2" pageNumber="33" refString="Roze, J. A. (1961) El genero Atractus (Serpentes: Colubridae) en Venezuela. Acta Biologica Venezuelica, 3, 103 - 119." type="journal article" year="1961">Roze 1961</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4BCAB8FF84FFD39735A1E07132ADC7" authority="Filippi 1840" authorityName="Filippi" authorityYear="1840" box="[778,1054,1484,1510]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="favae">
|
||
<emphasis id="B93F6D29FF84FFD39735A1E07641ADC4" box="[778,877,1484,1509]" italics="true" pageId="2" pageNumber="21">A. favae</emphasis>
|
||
(
|
||
<bibRefCitation id="EFDACCCAFF84FFD39742A1E07139ADC7" author="Filippi" box="[893,1045,1484,1510]" pageId="2" pageNumber="32" refString="Filippi, F. De (1840) Catalogo ragionato e descrittivo della raccolta de serpenti del Museo dell' I. R. Universita di Pavia. Biblioteca Italiana, 99, 65 pp." type="book" year="1840">Filippi 1840</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4BCAB8FF84FFD39014A1E07463AE2F" authority="Boie 1827" authorityName="Boie" authorityYear="1827" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="flammigerus">
|
||
<emphasis id="B93F6D29FF84FFD39014A1E071CCADC4" box="[1067,1248,1484,1509]" italics="true" pageId="2" pageNumber="21">A. flammigerus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFDACCCAFF84FFD390CEA1E0746AAE2F" author="Boie" pageId="2" pageNumber="31" refString="Boie, F. (1827) Bemerkungen uber Merrem's Versuch eines Systems der Amphibien. Marburg. 1820. Erste Lieferung: Ophidier. Isis von Oken, 20, 508 - 566." type="journal article" year="1827">Boie 1827</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4BCAB8FF84FFD39563A1D8778CAE2F" authority="Gunther 1868" authorityName="Gunther" authorityYear="1868" box="[348,672,1524,1550]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="latifrons">
|
||
<emphasis id="B93F6D29FF84FFD39563A1D874F3AE2C" box="[348,479,1524,1549]" italics="true" pageId="2" pageNumber="21">A. latifrons</emphasis>
|
||
(
|
||
<bibRefCitation id="EFDACCCAFF84FFD395D1A1D877BAAE2F" author="Gunther" box="[494,662,1524,1550]" pageId="2" pageNumber="32" refString="Gunther, A. (1868) Sixth account of new species of snakes in the collection of the British Museum. Annals Magazine of Natural History, 4, 413 - 429." type="journal article" year="1868">Günther 1868</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4BCAB8FF84FFD39693A1D876C0AE2F" authority="Silva Haad 2004" authorityName="Silva Haad" authorityYear="2004" box="[684,1004,1524,1550]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="lucilae">
|
||
<emphasis id="B93F6D29FF84FFD39693A1D87631AE2C" box="[684,797,1524,1549]" italics="true" pageId="2" pageNumber="21">A. lucilae</emphasis>
|
||
<bibRefCitation id="EFDACCCAFF84FFD3971BA1D876C0AE2F" author="Silva" box="[804,1004,1524,1550]" pageId="2" pageNumber="34" refString="Silva Haad, J. J. (2004) Las serpientes del genero Atractus Wagler, 1828 (Colubridae, Xenodontinae) en la Amazonia Colombiana. Revista de la Academia Colombiana de Ciencias Exactas, Fisicas y Naturales, 28, 409 - 446." type="journal article" year="2004">Silva Haad 2004</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4BCAB8FF84FFD397C7A1D87009AE2F" authority="Boulenger 1894" authorityName="Boulenger" authorityYear="1894" box="[1016,1317,1524,1550]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="major">
|
||
<emphasis id="B93F6D29FF84FFD397C7A1D87173AE2C" box="[1016,1119,1524,1549]" italics="true" pageId="2" pageNumber="21">A. major</emphasis>
|
||
<bibRefCitation id="EFDACCCAFF84FFD39059A1D87009AE2F" author="Boulenger" box="[1126,1317,1524,1550]" pageId="2" pageNumber="31" refString="Boulenger, G. A. (1894) Catalogue of the Snakes in the British Museum (Natural History), Volume II. Taylor and Francis, London, 382 pp." type="book" year="1894">Boulenger 1894</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4BCAB8FF84FFD39537A230772FAE17" authority="Roze 1961" authorityName="Roze" authorityYear="1961" box="[264,515,1564,1590]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="riveroi">
|
||
<emphasis id="B93F6D29FF84FFD39537A2307454AE14" box="[264,376,1564,1589]" italics="true" pageId="2" pageNumber="21">A. riveroi</emphasis>
|
||
<bibRefCitation id="EFDACCCAFF84FFD395BEA230772FAE17" author="Roze" box="[385,515,1564,1590]" lastPageId="2" lastPageNumber="34" pageId="2" pageNumber="33" refString="Roze, J. A. (1961) El genero Atractus (Serpentes: Colubridae) en Venezuela. Acta Biologica Venezuelica, 3, 103 - 119." type="journal article" year="1961">Roze 1961</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4BCAB8FF84FFD3962FA2307631AE17" authority="Boie 1827" authorityName="Boie" authorityYear="1827" box="[528,797,1564,1590]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="schach">
|
||
<emphasis id="B93F6D29FF84FFD3962FA23077A8AE14" box="[528,644,1564,1589]" italics="true" pageId="2" pageNumber="21">A. schach</emphasis>
|
||
(
|
||
<bibRefCitation id="EFDACCCAFF84FFD396ABA2307638AE17" author="Boie" box="[660,788,1564,1590]" pageId="2" pageNumber="31" refString="Boie, F. (1827) Bemerkungen uber Merrem's Versuch eines Systems der Amphibien. Marburg. 1820. Erste Lieferung: Ophidier. Isis von Oken, 20, 508 - 566." type="journal article" year="1827">Boie 1827</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4BCAB8FF84FFD39715A23076FBAE14" box="[810,983,1564,1589]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="snethlageae">
|
||
<emphasis id="B93F6D29FF84FFD39715A23076FBAE14" box="[810,983,1564,1589]" italics="true" pageId="2" pageNumber="21">A. snethlageae</emphasis>
|
||
</taxonomicName>
|
||
<bibRefCitation id="EFDACCCAFF84FFD397DFA230700AAE17" author="Cunha" box="[992,1318,1564,1590]" pageId="2" pageNumber="32" refString="Cunha, O. R. & Nascimento, F. P. (1983) Ofidios da Amazonia XX: as especies de Atractus Wagler, 1828, na Amazonia oriental e Maranhao (Ophidia, Colubridae). Boletim do Museu Paraense Emilio Goeldi, 123, 1 - 38." type="journal article" year="1983">
|
||
<collectingCountry id="F35CF1ABFF84FFD397DFA2307100AE17" box="[992,1068,1564,1590]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="2" pageNumber="21">Cunha</collectingCountry>
|
||
& Nascimento 1983
|
||
</bibRefCitation>
|
||
,
|
||
<taxonomicName id="4C4BCAB8FF84FFD39537A268776DAE7F" authority="Roze 1958" authorityName="Roze" authorityYear="1958" box="[264,577,1604,1630]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="steyermarki">
|
||
<emphasis id="B93F6D29FF84FFD39537A2687499AE7C" box="[264,437,1604,1629]" italics="true" pageId="2" pageNumber="21">A. steyermarki</emphasis>
|
||
Roze 1958
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4BCAB8FF84FFD39670A26871ACAE7F" authority="Dumeril, Bibron & Dumeril 1854" authorityName="Dumeril, Bibron & Dumeril" authorityYear="1854" box="[591,1152,1604,1630]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="torquatus">
|
||
<emphasis id="B93F6D29FF84FFD39670A26877CDAE7C" box="[591,737,1604,1629]" italics="true" pageId="2" pageNumber="21">A. torquatus</emphasis>
|
||
Duméril, Bibron & Duméril 1854
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C4BCAB8FF84FFD390FDA268770AAEA7" authority="Gasc & Rodrigues 1979" authorityName="Gasc & Rodrigues" authorityYear="1979" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="zidoki">
|
||
<emphasis id="B93F6D29FF84FFD390FDA2687007AE7C" box="[1218,1323,1604,1629]" italics="true" pageId="2" pageNumber="21">A. zidoki</emphasis>
|
||
<bibRefCitation id="EFDACCCAFF84FFD39537A240770AAEA7" author="Gasc" box="[264,550,1644,1670]" pageId="2" pageNumber="32" refString="Gasc, J. - P. & Rodrigues, M. T. (1979) Une nouvelle espece du genre Atractus (Colubridae, Serpentes) de la Guyane francaise. Bulletin du Museum national d'Histoire naturelle de Paris, 1, 547 - 557." type="journal article" year="1979">Gasc & Rodrigues 1979</bibRefCitation>
|
||
</taxonomicName>
|
||
. Among the few
|
||
<taxonomicName id="4C4BCAB8FF84FFD396D0A240767EAEA4" box="[751,850,1644,1669]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="21" phylum="Chordata" rank="genus">
|
||
<emphasis id="B93F6D29FF84FFD396D0A240767EAEA4" box="[751,850,1644,1669]" italics="true" pageId="2" pageNumber="21">Atractus</emphasis>
|
||
</taxonomicName>
|
||
species with 15 dorsal scale rows found in the region,
|
||
<taxonomicName id="4C4BCAB8FF84FFD39586A2B877B7AE8C" box="[441,667,1684,1709]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="tamessari">
|
||
<emphasis id="B93F6D29FF84FFD39586A2B877B7AE8C" box="[441,667,1684,1709]" italics="true" pageId="2" pageNumber="21">Atractus tamessari</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C4BCAB8FF84FFD3977BA2B87140AE8F" authority="Gunther 1858" authorityName="Gunther" authorityYear="1858" box="[836,1132,1684,1710]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="elaps">
|
||
<emphasis id="B93F6D29FF84FFD3977BA2B8768BAE8C" box="[836,935,1684,1709]" italics="true" pageId="2" pageNumber="21">A. elaps</emphasis>
|
||
(
|
||
<bibRefCitation id="EFDACCCAFF84FFD39785A2B87149AE8F" author="Gunther" box="[954,1125,1684,1710]" pageId="2" pageNumber="32" refString="Gunther, A. (1858) Catalogue of the colubrine snakes in the collection of the British Museum. Trustees of British Museum, London, 281 pp." type="book" year="1858">Günther 1858</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
by having two postoculars (one in
|
||
<taxonomicName id="4C4BCAB8FF84FFD395CDA290777DAEF4" box="[498,593,1724,1749]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="elaps">
|
||
<emphasis id="B93F6D29FF84FFD395CDA290777DAEF4" box="[498,593,1724,1749]" italics="true" pageId="2" pageNumber="21">A. elaps</emphasis>
|
||
</taxonomicName>
|
||
), eight supralabials (six), and no red or yellow rings on body (present); from
|
||
<taxonomicName id="4C4BCAB8FF84FFD395F6A2C87652AEDF" authority="Silva Haad 2004" authorityName="Silva Haad" authorityYear="2004" box="[457,894,1764,1790]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="franciscopaivai">
|
||
<emphasis id="B93F6D29FF84FFD395F6A2C87789AEDC" box="[457,677,1764,1789]" italics="true" pageId="2" pageNumber="21">A. franciscopaivai</emphasis>
|
||
<bibRefCitation id="EFDACCCAFF84FFD3968EA2C87652AEDF" author="Silva" box="[689,894,1764,1790]" pageId="2" pageNumber="34" refString="Silva Haad, J. J. (2004) Las serpientes del genero Atractus Wagler, 1828 (Colubridae, Xenodontinae) en la Amazonia Colombiana. Revista de la Academia Colombiana de Ciencias Exactas, Fisicas y Naturales, 28, 409 - 446." type="journal article" year="2004">Silva Haad 2004</bibRefCitation>
|
||
</taxonomicName>
|
||
by having eight supralabials, three contacting the loreal (five supralabials, two contacting the loreal in
|
||
<taxonomicName id="4C4BCAB8FF84FFD39000A3207035AF04" box="[1087,1305,1804,1829]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="franciscopaivai">
|
||
<emphasis id="B93F6D29FF84FFD39000A3207035AF04" box="[1087,1305,1804,1829]" italics="true" pageId="2" pageNumber="21">A. franciscopaivai</emphasis>
|
||
</taxonomicName>
|
||
), seven to eight infralabials (five), two postoculars (one), 1+2 temporals (1+1), and a yellowish cream venter heavily mottled with brownish black blotches (banded black and orange or black and yellow); from
|
||
<taxonomicName id="4C4BCAB8FF85FFD2968AA568714BA97F" authority="Silva Haad 2004" authorityName="Silva Haad" authorityYear="2004" box="[693,1127,324,350]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="22" phylum="Chordata" rank="species" species="heliobelluomini">
|
||
<emphasis id="B93F6D29FF85FFD2968AA56876BDA97C" box="[693,913,324,349]" italics="true" pageId="3" pageNumber="22">A. heliobelluomini</emphasis>
|
||
<bibRefCitation id="EFDACCCAFF85FFD297A3A568714BA97F" author="Silva" box="[924,1127,324,350]" pageId="3" pageNumber="34" refString="Silva Haad, J. J. (2004) Las serpientes del genero Atractus Wagler, 1828 (Colubridae, Xenodontinae) en la Amazonia Colombiana. Revista de la Academia Colombiana de Ciencias Exactas, Fisicas y Naturales, 28, 409 - 446." type="journal article" year="2004">Silva Haad 2004</bibRefCitation>
|
||
</taxonomicName>
|
||
by having eight supralabials, three contacting the loreal (seven supralabials, two contacting the loreal in
|
||
<taxonomicName id="4C4BCAB8FF85FFD2912EA54074ECA98C" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="22" phylum="Chordata" rank="species" species="heliobelluomini">
|
||
<emphasis id="B93F6D29FF85FFD2912EA54074ECA98C" italics="true" pageId="3" pageNumber="22">A. heliobelluomini</emphasis>
|
||
</taxonomicName>
|
||
), two postoculars (one), 1+2 temporals (1+1), and a yellowish cream venter heavily mottled with brownish black blotches (immaculate cream); from
|
||
<taxonomicName id="4C4BCAB8FF85FFD2912EA590772AA9DF" authority="Roze 1961" authorityName="Roze" authorityYear="1961" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="22" phylum="Chordata" rank="species" species="insipidus">
|
||
<emphasis id="B93F6D29FF85FFD2912EA590745EA9DC" italics="true" pageId="3" pageNumber="22">A. insipidus</emphasis>
|
||
<bibRefCitation id="EFDACCCAFF85FFD295BFA5C8772AA9DF" author="Roze" box="[384,518,484,510]" lastPageId="3" lastPageNumber="34" pageId="3" pageNumber="33" refString="Roze, J. A. (1961) El genero Atractus (Serpentes: Colubridae) en Venezuela. Acta Biologica Venezuelica, 3, 103 - 119." type="journal article" year="1961">Roze 1961</bibRefCitation>
|
||
</taxonomicName>
|
||
by having eight supralabials, three contacting the loreal (seven supralabials, two contacting the loreal in
|
||
<taxonomicName id="4C4BCAB8FF85FFD29732A62076B2AA04" box="[781,926,524,549]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="22" phylum="Chordata" rank="species" species="insipidus">
|
||
<emphasis id="B93F6D29FF85FFD29732A62076B2AA04" box="[781,926,524,549]" italics="true" pageId="3" pageNumber="22">A. insipidus</emphasis>
|
||
</taxonomicName>
|
||
), and a yellowish cream venter heavily mottled with brownish black blotches (immaculate cream); from
|
||
<taxonomicName id="4C4BCAB8FF85FFD29056A6187460AA57" authority="Jan 1862" authorityName="Jan" authorityYear="1862" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="22" phylum="Chordata" rank="species" species="poeppigi">
|
||
<emphasis id="B93F6D29FF85FFD29056A61871DEAA6C" box="[1129,1266,564,589]" italics="true" pageId="3" pageNumber="22">A. poeppigi</emphasis>
|
||
(
|
||
<bibRefCitation id="EFDACCCAFF85FFD2913DA6187468AA57" author="Jan" lastPageId="3" lastPageNumber="33" pageId="3" pageNumber="32" refString="Jan, G. (1862) Enumerazione sistematica delle specie d'ofidi del gruppo Calamaridae. Archivio per la Zoologia, l'Anatomia e la Fisiologia, 2, 1 - 76." type="journal article" year="1862">Jan 1862</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
by having eight supralabials (six in
|
||
<taxonomicName id="4C4BCAB8FF85FFD2973BA67076A1AA54" box="[772,909,604,629]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="22" phylum="Chordata" rank="species" species="poeppigi">
|
||
<emphasis id="B93F6D29FF85FFD2973BA67076A1AA54" box="[772,909,604,629]" italics="true" pageId="3" pageNumber="22">A. poeppigi</emphasis>
|
||
</taxonomicName>
|
||
), 1+2 temporals (0+1 or 0+2), the loreal much longer than high (slightly higher than long), and a yellowish cream venter heavily mottled with brownish black blotches (reddish cream or red and black); and from
|
||
<taxonomicName id="4C4BCAB8FF85FFD29537A6F87769AACF" authority="Wagler 1828" authorityName="Wagler" authorityYear="1828" box="[264,581,724,750]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="22" phylum="Chordata" rank="species" species="trilineatus">
|
||
<emphasis id="B93F6D29FF85FFD29537A6F8748FAACC" box="[264,419,724,749]" italics="true" pageId="3" pageNumber="22">A. trilineatus</emphasis>
|
||
<bibRefCitation id="EFDACCCAFF85FFD29593A6F87769AACF" author="Wagler" box="[428,581,724,750]" pageId="3" pageNumber="34" refString="Wagler, J. G. (1828) Auzuge aus seinem Systema Aphibiorem. Isis von Oken, 21, 740 - 744." type="journal article" year="1828">Wagler 1828</bibRefCitation>
|
||
</taxonomicName>
|
||
by having 28–31 subcaudal scales (
|
||
<quantity id="4CB31CDEFF85FFD297C9A6F87173AACF" box="[1014,1119,724,750]" metricMagnitude="-1" metricUnit="m" metricValue="3.81" metricValueMax="4.826" metricValueMin="2.794" pageId="3" pageNumber="22" unit="in" value="15.0" valueMax="19.0" valueMin="11.0">11–19 in</quantity>
|
||
<taxonomicName id="4C4BCAB8FF85FFD29057A6F8702FAACC" box="[1128,1283,724,749]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="22" phylum="Chordata" rank="species" species="trilineatus">
|
||
<emphasis id="B93F6D29FF85FFD29057A6F8702FAACC" box="[1128,1283,724,749]" italics="true" pageId="3" pageNumber="22">A. trilineatus</emphasis>
|
||
</taxonomicName>
|
||
), a yellowish cream venter heavily mottled with brownish black blotches (cream with light brown spots on posteriormost ventrals and anal), and by lacking well-defined longitudinal stripes (3–4 well-defined stripes present).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C351E2B0FF85FFD79507A758773FAECA" lastPageId="6" lastPageNumber="25" pageId="3" pageNumber="22" type="description">
|
||
<paragraph id="8BF4B13BFF85FFD29507A7587695AEA7" blockId="3.[264,1323,284,1870]" pageId="3" pageNumber="22">
|
||
<emphasis id="B93F6D29FF85FFD29507A75877BEABAF" bold="true" box="[312,658,884,910]" pageId="3" pageNumber="22">
|
||
Description of the
|
||
<typeStatus id="54F00F99FF85FFD29620A75877A1ABAF" box="[543,653,884,910]" pageId="3" pageNumber="22" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
An adult male, TTL
|
||
<quantity id="4CB31CDEFF85FFD297B2A75876DDABAF" box="[909,1009,884,910]" metricMagnitude="-1" metricUnit="m" metricValue="3.97" pageId="3" pageNumber="22" unit="mm" value="397.0">397 mm</quantity>
|
||
, SVL
|
||
<quantity id="4CB31CDEFF85FFD29003A758718DABAF" box="[1084,1185,884,910]" metricMagnitude="-1" metricUnit="m" metricValue="3.46" pageId="3" pageNumber="22" unit="mm" value="346.0">346 mm</quantity>
|
||
, TL
|
||
<quantity id="4CB31CDEFF85FFD290E7A7587007ABAF" box="[1240,1323,884,910]" metricMagnitude="-2" metricUnit="m" metricValue="5.1" pageId="3" pageNumber="22" unit="mm" value="51.0">51 mm</quantity>
|
||
(12.8% of TTL); head slightly distinct from neck, length 1.4 times width (HW: 7.6 mm; HL: 10.8 mm); snout length (from eye to tip of snout) 4.0 mm, snout rounded in dorsal and lateral view; eye small (1.6 mm); pupil semi-elliptical; eye-nostril distance 29% of HL; rostral small, visible in dorsal view, width about 1.8 times height; internasals small, as long as wide, laterally in contact with anterior and posterior nasals; prefrontals large, length 1.4 times width, much larger than the internasals, irregularly hexagonal, touching eye, their medial suture asymmetrical, noticeably dextral to the medial internasal suture; frontal pentagonal, slightly wider than long, 1.1 times length of medial prefrontal suture; parietals large, length 1.5 times width, medial suture of parietals 1.2 times length of frontal. Nasal divided, posterior nasal as long as anterior nasal; preocular absent; loreal long, length 2.8 times height; postoculars 2/2, small, upper slightly larger than lower; temporals 1+2, lower posterior temporal slightly longer than upper; supralabials 8/8, eighth largest, first contacting nasals, second contacting posterior nasal and loreal, third and fourth contacting loreal, fourth and fifth touching eye, sixth contacting postocular and anterior temporal, seventh contacting anterior and posterior temporals, eighth contacting posterior temporal; infralabials 8/8, first four in contact with chinshields, first pair meeting at ventral midline, separating mental from chinshields; one pair of chinshields, length 2.5 times width (
|
||
<figureCitation id="1370ADBEFF85FFD29598A26874C3AE7F" box="[423,495,1604,1630]" captionStart="FIGURE 1" captionStartId="4.[264,367,1841,1865]" captionTargetBox="[340,1223,325,1778]" captionTargetId="figure@4.[327,1250,279,1805]" captionTargetPageId="4" captionText="FIGURE 1. Atractus tamessari sp. nov. (holotype, IRSNB 2640): (A) dorsal, (B) lateral, and (C) ventral views of the head." httpUri="https://zenodo.org/record/175025/files/figure.png" pageId="3" pageNumber="22">Fig. 1</figureCitation>
|
||
). Maxillary teeth 6; dorsal scales in 15-15-15 rows, smooth without apical pits; ventrals 153; anal single; paired subcaudals 31.
|
||
</paragraph>
|
||
<paragraph id="8BF4B13BFF85FFD79507A2B8779DAA37" blockId="3.[264,1323,284,1870]" lastBlockId="6.[264,1323,284,1891]" lastPageId="6" lastPageNumber="25" pageId="3" pageNumber="22">
|
||
<emphasis id="B93F6D29FF85FFD29507A2B87742AE8F" bold="true" box="[312,622,1684,1710]" pageId="3" pageNumber="22">
|
||
Color of
|
||
<typeStatus id="54F00F99FF85FFD29595A2B87739AE8F" box="[426,533,1684,1710]" pageId="3" pageNumber="22" type="holotype">holotype</typeStatus>
|
||
in life.
|
||
</emphasis>
|
||
The dorsal ground color is brownish black with numerous irregular bright red and paler red markings. The tail is brownish black with a few bright red lateral markings. The top of the head is brownish black. The snout is lighter, with a slight yellow wash. The supralabials are mostly yellow with the upper third dark brown, except for the eighth, which is almost completely dark brown with a small yellow spot anteriorly, and the first and the second, which are speckled with dark brown. The infralabials are mostly yellow with the lower third and posterior portion of each scale dark brown. The chinshields are mostly yellow with dark brown anteriorly and laterally. The mental groove area has a few dark brown speckles. The venter is yellowish cream with many irregular brownish black blotches, fewer blotches anteriorly, heavily mottled posteriorly. Underside of tail is brownish black with a few small irregular yellowish cream markings. The iris is brown (
|
||
<figureCitation id="1370ADBEFF80FFD79664A5D07788AA37" box="[603,676,508,534]" captionStart="FIGURE 2" captionStartId="5.[264,367,1805,1829]" captionTargetBox="[439,1155,315,1772]" captionTargetId="figure@5.[439,1157,302,1784]" captionTargetPageId="5" captionText="FIGURE 2. Atractus tamessari sp. nov.: (A) the male holotype (IRSNB 2640) in life, (B) ventral view of the male holotype (IRSNB 2640) in life, and (C) a subadult female paratopotype (IRSNB 2641) in life." httpUri="https://zenodo.org/record/175026/files/figure.png" pageId="6" pageNumber="25">Fig. 2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF34E1B3FF82FFD59537A31D7737AF4A" httpUri="https://zenodo.org/record/175025/files/figure.png" pageId="4" pageNumber="23" targetBox="[340,1223,325,1778]" targetPageId="4">
|
||
<paragraph id="8BF4B13BFF82FFD59537A31D7737AF4A" blockId="4.[264,1322,1841,1899]" pageId="4" pageNumber="23">
|
||
<emphasis id="B93F6D29FF82FFD59537A31D74A7AF68" bold="true" box="[264,395,1841,1865]" pageId="4" pageNumber="23">FIGURE 1.</emphasis>
|
||
<taxonomicName id="4C4BCAB8FF82FFD595ABA31D7773AF69" box="[404,607,1841,1864]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="23" phylum="Chordata" rank="species" species="tamessari" status="sp. nov.">
|
||
<emphasis id="B93F6D29FF82FFD595ABA31D7773AF69" box="[404,607,1841,1864]" italics="true" pageId="4" pageNumber="23">Atractus tamessari</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20CD052FF82FFD59659A31D7791AF68" box="[614,701,1841,1865]" pageId="4" pageNumber="23" rank="species">
|
||
<emphasis id="B93F6D29FF82FFD59659A31D7791AF68" bold="true" box="[614,701,1841,1865]" pageId="4" pageNumber="23">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(holotype, IRSNB 2640): (A) dorsal, (B) lateral, and (C) ventral views of the head.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF34E1B3FF83FFD49537A32174BFAF4B" httpUri="https://zenodo.org/record/175026/files/figure.png" pageId="5" pageNumber="24" targetBox="[439,1155,315,1772]" targetPageId="5">
|
||
<paragraph id="8BF4B13BFF83FFD49537A32174BFAF4B" blockId="5.[264,1322,1805,1898]" pageId="5" pageNumber="24">
|
||
<emphasis id="B93F6D29FF83FFD49537A32174A7AF04" bold="true" box="[264,395,1805,1829]" pageId="5" pageNumber="24">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C4BCAB8FF83FFD495ABA3227772AF04" box="[404,606,1806,1829]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="24" phylum="Chordata" rank="species" species="tamessari" status="sp. nov.">
|
||
<emphasis id="B93F6D29FF83FFD495ABA3227772AF04" box="[404,606,1806,1829]" italics="true" pageId="5" pageNumber="24">Atractus tamessari</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20CD052FF83FFD4965AA3217790AF04" box="[613,700,1805,1829]" pageId="5" pageNumber="24" rank="species">
|
||
<emphasis id="B93F6D29FF83FFD4965AA3217790AF04" bold="true" box="[613,700,1805,1829]" pageId="5" pageNumber="24">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
: (A) the male holotype (IRSNB 2640) in life, (B) ventral view of the male holotype (IRSNB 2640) in life, and (C) a subadult female paratopotype (IRSNB 2641) in life.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF4B13BFF80FFD79507A60D76E1AAAA" blockId="6.[264,1323,284,1891]" pageId="6" pageNumber="25">
|
||
<emphasis id="B93F6D29FF80FFD79507A60D77C7AA1A" bold="true" box="[312,747,545,571]" pageId="6" pageNumber="25">
|
||
Color of
|
||
<typeStatus id="54F00F99FF80FFD79590A60D7736AA1A" box="[431,538,545,571]" pageId="6" pageNumber="25" type="holotype">holotype</typeStatus>
|
||
in preservative.
|
||
</emphasis>
|
||
After one month in preservative (70% ethanol after formalin), the red markings have faded slightly but are still well visible. The yellow coloration became white. The dark markings are unchanged.
|
||
</paragraph>
|
||
<paragraph id="8BF4B13BFF80FFD79507A6B57494AB82" blockId="6.[264,1323,284,1891]" pageId="6" pageNumber="25">
|
||
<emphasis id="B93F6D29FF80FFD79507A6B574E8AA92" bold="true" box="[312,452,665,691]" pageId="6" pageNumber="25">Hemipenis.</emphasis>
|
||
Almost fully everted right hemipenis about 10.5 mm long, extending to level of seventh subcaudal when adpressed to the tail; hemipenis differentiated (
|
||
<emphasis id="B93F6D29FF80FFD790D6A6ED7006AAFB" box="[1257,1322,705,730]" italics="true" pageId="6" pageNumber="25">sensu</emphasis>
|
||
Savage 1960), capitate, slightly bilobed, shallowly forked; lobes distinct from the base, each lobe nearly flattened on apical portion, displaying series of spinulate calyces; capitation stronger on asulcate side; sulcus spermaticus centrifugal, bifurcate, dividing about half of organ, at the base of apex; basal portion of hemipenial body covered by large spines (
|
||
<figureCitation id="1370ADBEFF80FFD7955EA7A57484AB82" box="[353,424,905,931]" captionStart="FIGURE 3" captionStartId="7.[264,367,1007,1031]" captionTargetBox="[264,1323,326,983]" captionTargetId="figure@7.[264,1323,326,983]" captionTargetPageId="7" captionText="FIGURE 3. Almost fully everted right hemipenis of Atractus tamessari sp. nov. (holotype, IRSNB 2640): (left) sulcate, and (right) asulcate sides; scale = 5 mm." httpUri="https://zenodo.org/record/175027/files/figure.png" pageId="6" pageNumber="25">Fig. 3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF4B13BFF80FFD79507A79D773FAECA" blockId="6.[264,1323,284,1891]" pageId="6" pageNumber="25">
|
||
<emphasis id="B93F6D29FF80FFD79507A79D749DABEA" bold="true" box="[312,433,945,971]" pageId="6" pageNumber="25">Variation.</emphasis>
|
||
The
|
||
<typeStatus id="54F00F99FF80FFD79631A79D776DABEA" box="[526,577,945,971]" pageId="6" pageNumber="25">type</typeStatus>
|
||
series consists of the male
|
||
<typeStatus id="54F00F99FF80FFD797E4A79D716EABEA" box="[987,1090,945,971]" pageId="6" pageNumber="25" type="holotype">holotype</typeStatus>
|
||
and two female paratopotypes. The paratopotypes strongly resemble the
|
||
<typeStatus id="54F00F99FF80FFD797F2A7F57118ABD2" box="[973,1076,985,1011]" pageId="6" pageNumber="25" type="holotype">holotype</typeStatus>
|
||
with the following exceptions (those of
|
||
<typeStatus id="54F00F99FF80FFD79629A02D7750AC3A" box="[534,636,1025,1051]" pageId="6" pageNumber="25" type="holotype">holotype</typeStatus>
|
||
in parentheses): TTL from
|
||
<quantity id="4CB31CDEFF80FFD797D2A02D71B9AC3A" box="[1005,1173,1025,1051]" metricMagnitude="-1" metricUnit="m" metricValue="3.9899999999999998" metricValueMax="4.3" metricValueMin="3.6799999999999997" pageId="6" pageNumber="25" unit="mm" value="399.0" valueMax="430.0" valueMin="368.0">368–430 mm</quantity>
|
||
; SVL from
|
||
<quantity id="4CB31CDEFF80FFD79537A0057484AC62" box="[264,424,1065,1091]" metricMagnitude="-1" metricUnit="m" metricValue="3.59" metricValueMax="3.86" metricValueMin="3.32" pageId="6" pageNumber="25" unit="mm" value="359.0" valueMax="386.0" valueMin="332.0">332–386 mm</quantity>
|
||
; TL from
|
||
<quantity id="4CB31CDEFF80FFD7961DA0057789AC62" box="[546,677,1065,1091]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" metricValueMax="4.4" metricValueMin="3.6" pageId="6" pageNumber="25" unit="mm" value="40.0" valueMax="44.0" valueMin="36.0">36–44 mm</quantity>
|
||
, 9.8–10.2% of TTL (12.8%); HL from 10.6–11.5 mm; HW from 7.5–8.3 mm; 28–29 subcaudals (31); 157–163 ventrals (153); upper postocular 1.8 times the lower one in IRSNB 2641 (only slightly larger in the
|
||
<typeStatus id="54F00F99FF80FFD79029A0557150ACB2" box="[1046,1148,1145,1171]" pageId="6" pageNumber="25" type="holotype">holotype</typeStatus>
|
||
and in IRSNB 2642); seven infralabials in IRSNB 2642 (eight in the
|
||
<typeStatus id="54F00F99FF80FFD7978FA08D713AAC9A" box="[944,1046,1185,1211]" pageId="6" pageNumber="25" type="holotype">holotype</typeStatus>
|
||
and in IRSNB 2641), apparently due to the fusion of infralabials 5 and 6. The sexual dimorphism in the number of ventrals and subcaudals is typical for this genus (Savage 1960). Prefrontal and internasal medial sutures are asymmetrical in both sexes (see
|
||
<bibRefCitation id="EFDACCCAFF80FFD797E7A1357007AD12" author="Hoogmoed" box="[984,1323,1305,1331]" pageId="6" pageNumber="32" refString="Hoogmoed, M. S. & Prudente A. L. C. (2003). A new species of Atractus (Reptilia: Ophidia: Colubridae: Dipsadinae) from the Amazon forest region in Brazil. Zoologische Mededelingen, 77, 425 - 440." type="journal article" year="2003">Hoogmoed & Prudente 2003</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFDACCCAFF80FFD7957CA16D74F6AD7A" author="Myers" box="[323,474,1345,1371]" pageId="6" pageNumber="33" refString="Myers, C. W. (2003) Rare snakes - Five new species from eastern Panama: reviews of northern Atractus and southern Geophis (Colubridae: Dipsadinae). American Museum Novitates, 3391, 1 - 47." type="journal article" year="2003">Myers 2003</bibRefCitation>
|
||
for further discussion). The most striking difference between the paratopotypes and the
|
||
<typeStatus id="54F00F99FF80FFD79622A14577A8ADA2" box="[541,644,1385,1411]" pageId="6" pageNumber="25" type="holotype">holotype</typeStatus>
|
||
occurs in dorsal pattern and coloration, which is fairly variable. IRSNB 2641 has a medium brown dorsum with irregular pale reddish brown markings and two ventrolateral black stripes (instead of irregular bright red markings on a brownish black ground without ventrolateral stripes in the
|
||
<typeStatus id="54F00F99FF80FFD79781A1CD7108ADDA" box="[958,1060,1505,1531]" pageId="6" pageNumber="25" type="holotype">holotype</typeStatus>
|
||
and IRSNB 2642). In IRSNB 2642 the bright red markings form an incomplete broken dorsolateral stripe. The head of each
|
||
<typeStatus id="54F00F99FF80FFD7959BA21D7725AE6A" box="[420,521,1585,1611]" pageId="6" pageNumber="25" type="paratype">paratype</typeStatus>
|
||
is medium brown and, as in the
|
||
<typeStatus id="54F00F99FF80FFD797B5A21D76D8AE6A" box="[906,1012,1585,1611]" pageId="6" pageNumber="25" type="holotype">holotype</typeStatus>
|
||
, the snout is lighter with a yellow wash. The pale reddish brown markings on the dorsum of IRSNB 2641 disappeared in preservative, resulting in a uniform medium brown dorsum. Ventral patterns are similar, those of the male
|
||
<typeStatus id="54F00F99FF80FFD796D8A2857662AEE2" box="[743,846,1705,1731]" pageId="6" pageNumber="25" type="holotype">holotype</typeStatus>
|
||
slightly more mottled than the female paratopotypes (
|
||
<figureCitation id="1370ADBEFF80FFD79583A2FD7729AECA" box="[444,517,1745,1771]" captionStart="FIGURE 4" captionStartId="8.[264,367,1756,1780]" captionTargetBox="[264,1322,846,1732]" captionTargetId="figure@8.[264,1323,846,1732]" captionTargetPageId="8" captionText="FIGURE 4. Atractus tamessari sp. nov.: (A) dorsal view of the preserved male holotype (IRSNB 2640), (B) dorsal view of a preserved subadult female paratopotype (IRSNB 2641), (C) ventral view of the preserved male holotype (IRSNB 2640), and (D) ventral view of a preserved subadult female paratopotype (IRSNB 2641)." httpUri="https://zenodo.org/record/175028/files/figure.png" pageId="6" pageNumber="25">Fig. 4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C351E2B0FF80FFD69507A2D574E1AE5E" lastPageId="7" lastPageNumber="26" pageId="6" pageNumber="25" type="distribution">
|
||
<paragraph id="8BF4B13BFF80FFD69507A2D574E1AE5E" blockId="6.[264,1323,284,1891]" lastBlockId="7.[264,1323,1117,1663]" lastPageId="7" lastPageNumber="26" pageId="6" pageNumber="25">
|
||
<emphasis id="B93F6D29FF80FFD79507A2D57754AF32" bold="true" box="[312,632,1785,1811]" pageId="6" pageNumber="25">Distribution and ecology.</emphasis>
|
||
<taxonomicName id="4C4BCAB8FF80FFD796BDA2D5764EAF33" box="[642,866,1785,1810]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="25" phylum="Chordata" rank="species" species="tamessari">
|
||
<emphasis id="B93F6D29FF80FFD796BDA2D5764EAF33" box="[642,866,1785,1810]" italics="true" pageId="6" pageNumber="25">Atractus tamessari</emphasis>
|
||
</taxonomicName>
|
||
is known only from the
|
||
<typeStatus id="54F00F99FF80FFD790A9A2D571E5AF32" box="[1174,1225,1785,1811]" pageId="6" pageNumber="25">type</typeStatus>
|
||
locality (
|
||
<figureCitation id="1370ADBEFF80FFD7952EA30D7474AF1A" box="[273,344,1825,1851]" captionStart="FIGURE 5" captionStartId="9.[264,367,1365,1389]" captionTargetBox="[312,1274,326,1347]" captionTargetId="figure@9.[312,1275,326,1347]" captionTargetPageId="9" captionText="FIGURE 5. Map of Guyana showing the distribution of Atractus tamessari sp. nov. (dot = type locality)." httpUri="https://zenodo.org/record/175029/files/figure.png" pageId="6" pageNumber="25">Fig. 5</figureCitation>
|
||
), which is located in the southeastern part of Kaieteur National Park, at the eastern edge of the
|
||
<taxonomicName id="4C4BCAB8FF80FFD795AEA3657721AF42" box="[401,525,1865,1891]" class="Reptilia" family="Colubridae" genus="Dipsas" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="26" phylum="Chordata" rank="species" species="pakaraima">Pakaraima</taxonomicName>
|
||
Mountains (
|
||
<figureCitation id="1370ADBEFF80FFD79697A36577DCAF42" box="[680,752,1865,1891]" captionStart="FIGURE 6" captionStartId="11.[264,367,1085,1109]" captionTargetBox="[264,1322,326,1061]" captionTargetId="figure@11.[264,1323,326,1061]" captionTargetPageId="11" captionText="FIGURE 6. Area map of the eastern part of Kaieteur National Park showing the collection site of Atractus tamessari sp. nov. (square). Map after “ Kaieteur Sheet 43 SW ” published by the Survey Department of Guyana, 1972." httpUri="https://zenodo.org/record/175030/files/figure.png" pageId="6" pageNumber="25">Fig. 6</figureCitation>
|
||
). The
|
||
<typeStatus id="54F00F99FF80FFD79704A365768EAF42" box="[827,930,1865,1891]" pageId="6" pageNumber="25" type="holotype">holotype</typeStatus>
|
||
and IRSNB 2641 were collected at night (
|
||
<emphasis id="B93F6D29FF81FFD6954BA07274BAAC56" box="[372,406,1118,1143]" italics="true" pageId="7" pageNumber="26">ca.</emphasis>
|
||
22:30 h), crawling on the ground a few meters from a stream, almost exactly at the same place and the same time, at a 24h interval. IRSNB 2642 was collected by day, apparently sleeping, on a stump in the river. The habitat can be characterized as a submontane forest of the
|
||
<taxonomicName id="4C4BCAB8FF81FFD6960AA0F9779DACCE" box="[565,689,1237,1263]" class="Reptilia" family="Colubridae" genus="Dipsas" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="26" phylum="Chordata" rank="species" species="pakaraima">Pakaraima</taxonomicName>
|
||
uplands on white sands dominated by clump wallaba (
|
||
<taxonomicName id="4C4BCAB8FF81FFD6952DA0D27456AD36" box="[274,378,1278,1303]" class="Magnoliopsida" family="Fabaceae" genus="Dicymbe" kingdom="Plantae" order="Fabales" pageId="7" pageNumber="26" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B93F6D29FF81FFD6952DA0D27456AD36" box="[274,378,1278,1303]" italics="true" pageId="7" pageNumber="26">Dicymbe</emphasis>
|
||
</taxonomicName>
|
||
spp) (ter
|
||
<bibRefCitation id="EFDACCCAFF81FFD6963BA0D1778CAD36" author="Steege" box="[516,672,1277,1303]" pageId="7" pageNumber="34" refString="ter Steege, H. (2001) National Vegetation map of Guyana, Guyana Forestry Commission: Georgetown." type="book" year="2001">Steege 2001</bibRefCitation>
|
||
). Examination of stomachs revealed no identifiable remains of prey, although X-rays of the male body showed the presence of sand in the digestive tract, which may suggest the recent ingestion of an earthworm, a very common food of
|
||
<taxonomicName id="4C4BCAB8FF81FFD69553A15A74FCADAE" box="[364,464,1398,1423]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="26" phylum="Chordata" rank="genus">
|
||
<emphasis id="B93F6D29FF81FFD69553A15A74FCADAE" box="[364,464,1398,1423]" italics="true" pageId="7" pageNumber="26">Atractus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFDACCCAFF81FFD695DEA159763EADAE" author="Martins" box="[481,786,1397,1423]" pageId="7" pageNumber="33" refString="Martins, M. & Oliveira, M. E. (1993) The snakes of the genus Atractus Wagler (Reptilia: Squamata: Colubridae) from the Manaus region, central Amazonia, Brazil. Zoologische Mededelingen, 67, 21 - 40." type="journal article" year="1993">Martins & Oliveira 1993</bibRefCitation>
|
||
, “
|
||
<bibRefCitation id="EFDACCCAFF81FFD69712A1597640ADAE" author="Oliveira" box="[813,876,1397,1423]" pageId="7" pageNumber="33" refString="Martins, M. & Oliveira, M. E. (" 1998 ") [1999] Natural history of snakes in forests in the Manaus region, Central Amazonia, Brazil. Herpetological Natural History, 6, 78 - 150." type="journal article" year="1998">1998</bibRefCitation>
|
||
” [1999]). No enlarged follicles were found in IRSNB 2641 ovaries; one egg was found in the abdominal cavity of IRSNB 2642, but as the specimen was cut in two by the collector, it is highly probable that other eggs were lost. All specimens have abdominal fat deposits. Other snake species collected in the immediate vicinity are the colubrids
|
||
<taxonomicName id="4C4BCAB8FF81FFD69685A23A7653AE0E" box="[698,895,1558,1583]" class="Reptilia" family="Colubridae" genus="Chironius" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="26" phylum="Chordata" rank="species" species="fuscus">
|
||
<emphasis id="B93F6D29FF81FFD69685A23A7653AE0E" box="[698,895,1558,1583]" italics="true" pageId="7" pageNumber="26">Chironius fuscus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4BCAB8FF81FFD697B0A23A71E5AE0E" box="[911,1225,1558,1583]" class="Reptilia" family="Colubridae" genus="Erythrolamprus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="26" phylum="Chordata" rank="species" species="aesculapii">
|
||
<emphasis id="B93F6D29FF81FFD697B0A23A71E5AE0E" box="[911,1225,1558,1583]" italics="true" pageId="7" pageNumber="26">Erythrolamprus aesculapii</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4BCAB8FF81FFD690E6A23A7444AE76" class="Reptilia" family="Colubridae" genus="Dipsas" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="26" phylum="Chordata" rank="species" species="catesbyi">
|
||
<emphasis id="B93F6D29FF81FFD690E6A23A7444AE76" italics="true" pageId="7" pageNumber="26">Dipsas catesbyi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4BCAB8FF81FFD69548A212775AAE76" box="[375,630,1597,1623]" class="Reptilia" family="Colubridae" genus="Dipsas" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="26" phylum="Chordata" rank="species" species="pakaraima">
|
||
<emphasis id="B93F6D29FF81FFD69548A21274E4AE76" box="[375,456,1598,1623]" italics="true" pageId="7" pageNumber="26">Dipsas</emphasis>
|
||
cf.
|
||
<emphasis id="B93F6D29FF81FFD695C8A212775AAE76" box="[503,630,1598,1623]" italics="true" pageId="7" pageNumber="26">pakaraima</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4BCAB8FF81FFD696BAA2127644AE76" box="[645,872,1598,1623]" class="Reptilia" family="Colubridae" genus="Helicops" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="26" phylum="Chordata" rank="species" species="angulatus">
|
||
<emphasis id="B93F6D29FF81FFD696BAA2127644AE76" box="[645,872,1598,1623]" italics="true" pageId="7" pageNumber="26">Helicops angulatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4BCAB8FF81FFD69747A2127147AE76" box="[888,1131,1598,1623]" class="Reptilia" family="Colubridae" genus="Pseudoboa" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="26" phylum="Chordata" rank="species" species="coronata">
|
||
<emphasis id="B93F6D29FF81FFD69747A2127147AE76" box="[888,1131,1598,1623]" italics="true" pageId="7" pageNumber="26">Pseudoboa coronata</emphasis>
|
||
</taxonomicName>
|
||
, and the viperid
|
||
<taxonomicName id="4C4BCAB8FF81FFD69537A24A74EAAE5E" box="[264,454,1638,1663]" class="Reptilia" family="Viperidae" genus="Bothrops" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="26" phylum="Chordata" rank="species" species="brazili">
|
||
<emphasis id="B93F6D29FF81FFD69537A24A74EAAE5E" box="[264,454,1638,1663]" italics="true" pageId="7" pageNumber="26">Bothrops brazili</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |