treatments-xml/data/03/DB/9B/03DB9B4DFFD7AF2FC98E9F78FA31947A.xml
2024-06-21 12:22:17 +02:00

267 lines
42 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="0E2B197E5C8FFCE3F99BCC2DABFE4ED7" ID-CLB-Dataset="56014" ID-DOI="10.11646/zootaxa.5125.5.6" ID-GBIF-Dataset="c9f37f27-8103-4768-8e05-7b4520854b1d" ID-ISSN="1175-5326" ID-Zenodo-Dep="6457396" ID-ZooBank="FC49517F-DD23-4281-A782-5BB6F9DC8560" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1649835338441" checkinUser="plazi" docAuthor="Li, Yan-Da, Hsiao, Yun, Huang, Di-Ying &amp; Cai, Chen-Yang" docDate="2022" docId="03DB9B4DFFD7AF2FC98E9F78FA31947A" docLanguage="en" docName="zootaxa.5125.5.6.pdf" docOrigin="Zootaxa 5125 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Vetuprostomis Engel &amp; Grimaldi 2008" docType="treatment" docVersion="5" lastPageNumber="566" masterDocId="FFE2E335FFD5AF2CC9199B5EFFAD957F" masterDocTitle="New species of Vetuprostomis from mid-Cretaceous amber of northern Myanmar (Coleoptera: Tenebrionoidea: Prostomidae)" masterLastPageNumber="574" masterPageNumber="563" pageNumber="565" updateTime="1699328905843" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="BBA94579852F7998B6DD801A06AE02B7" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="96E39964FD30C7B4A70857D9DF6F858F">
<mods:title id="79BD7525AD934BE09605E5EF7E01497D">New species of Vetuprostomis from mid-Cretaceous amber of northern Myanmar (Coleoptera: Tenebrionoidea: Prostomidae)</mods:title>
</mods:titleInfo>
<mods:name id="B32D39F434D4A10806937E267569257C" type="personal">
<mods:role id="2951EA9EF6C62543A6240BE1EFECDCBE">
<mods:roleTerm id="B9570BC6C16012DDBF269F9B88BE824C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="26FDDFC923207BA91598AECDC72207C7">Li, Yan-Da</mods:namePart>
<mods:nameIdentifier id="A3A63325CFC7580A740E6EAB52F5953F" type="ORCID">0000-0002-9439-202X</mods:nameIdentifier>
<mods:affiliation id="3344EF6C7891DE541185878D46C03F7E">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, and Centre for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, China &amp; ydli @ pku. edu. cn; https: // orcid. org / 0000 - 0002 - 9439 - 202 X</mods:affiliation>
<mods:nameIdentifier id="FBA594274D2E8279E2B19BDA97D95142" type="email">ydli@pku.edu.cn</mods:nameIdentifier>
</mods:name>
<mods:name id="D8D1864AF3105127DB98C2CF693AA336" type="personal">
<mods:role id="6790E00520E7D4C420AB248341FE6DB8">
<mods:roleTerm id="154F0C12345355EB28B659FA8CC496AF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AAE100DC0825145BDB4E11277953BCFD">Hsiao, Yun</mods:namePart>
<mods:nameIdentifier id="35EF611909C30882C997BA2F56492A5A" type="ORCID">0000-0003-2619-3296</mods:nameIdentifier>
<mods:affiliation id="2B0620AE0C75D90BB98DB057BCB5639B">Australian National Insect Collection, CSIRO, GPO Box 1700, Canberra ACT 2601, Australia &amp; Division of Ecology and Evolution, Research School of Biology, The Australian National University, Canberra, ACT 2601, Australia School of Earth Sciences, University of Bristol, Life Sciences Building, Tyndall Avenue, Bristol BS 8 1 TQ, UK &amp; yunhsiao @ outlook. com; https: // orcid. org / 0000 - 0003 - 2619 - 3296</mods:affiliation>
<mods:nameIdentifier id="18908DF6A1BEF315DE33665B73DC46BE" type="email">yunhsiao@outlook.com</mods:nameIdentifier>
</mods:name>
<mods:name id="493AC8CACDFF1F07413C9B30C789852E" type="personal">
<mods:role id="4284C1E6C9117F291523815B27B4D020">
<mods:roleTerm id="227FD6781A86A05DE0C4106B4C543C04">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7E01FD0AC9C328BAB7E06D39C4A93758">Huang, Di-Ying</mods:namePart>
<mods:nameIdentifier id="86C142048A30BAE0FE3172FFCD5DF06D" type="ORCID">0000-0002-5637-4867</mods:nameIdentifier>
<mods:affiliation id="EF33037BC419D83884E4EC7385362CDB">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, and Centre for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, China &amp; dyhuang @ nigpas. ac. cn; https: // orcid. org / 0000 - 0002 - 5637 - 4867</mods:affiliation>
<mods:nameIdentifier id="3815F5161B11E7892CA8F95C1C4E64BF" type="email">dyhuang@nigpas.ac.cn</mods:nameIdentifier>
</mods:name>
<mods:name id="D1F276299AE6088798B0726665FC6AF9" type="personal">
<mods:role id="C3272D62047A084CE941DA3149E00B0B">
<mods:roleTerm id="511C2E99CC179DE5E6BD8FBEECCFCC4E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="81C328AB3B091159804183D40E827991">Cai, Chen-Yang</mods:namePart>
<mods:affiliation id="28923A6ECE8E92D40342A8000542236D">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, and Centre for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, China</mods:affiliation>
</mods:name>
<mods:typeOfResource id="9883171B5E81D2E2FBBFAB2FF50A12D7">text</mods:typeOfResource>
<mods:relatedItem id="02DA13CECE34DFE83EF15D295BBA88B5" type="host">
<mods:titleInfo id="00A990EB0A67E83D3CA43F4BD06B3685">
<mods:title id="C368F391DF5A439355855E60E0001363">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="EF110C63C25B11949F1B8E6B456A8FEA">
<mods:date id="2E865A500257CEE4304486BD585D32D1">2022</mods:date>
<mods:detail id="2950112EF3417CF3CB5829BB71585736" type="pubDate">
<mods:number id="9FE02D3E906E5C377D59791E2E3A1CF0">2022-04-13</mods:number>
</mods:detail>
<mods:detail id="19F325FFC223C3D893CA5496F6D69A6F" type="volume">
<mods:number id="C0BB03D9EAF754D71D1B261CDD3B06EF">5125</mods:number>
</mods:detail>
<mods:detail id="B4E7B58DA1DCA23201B60C5DF996E313" type="issue">
<mods:number id="6EC5837285CB6A9EA631E906ADED1D1E">5</mods:number>
</mods:detail>
<mods:extent id="7CF967E3E889ABD2F126F5DFD221CD70" unit="page">
<mods:start id="4494E580A5F22A7C81A128FAA4E76EF8">563</mods:start>
<mods:end id="3B8AAD00DDB98555DD4EBD6182A666B1">574</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="74C405D26D9C8CFC5FB8AB10CF7D4472">journal article</mods:classification>
<mods:identifier id="3B5B75CDFA67657EBE4AB62935C9669E" type="CLB-Dataset">56014</mods:identifier>
<mods:identifier id="E8B4958F0E57E9BA06E7029756F39172" type="DOI">10.11646/zootaxa.5125.5.6</mods:identifier>
<mods:identifier id="1782586E53AEE4051946C6AB50729A10" type="GBIF-Dataset">c9f37f27-8103-4768-8e05-7b4520854b1d</mods:identifier>
<mods:identifier id="D3F150093A8BF2C6BD70CC62032B7C7B" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="6F819CDB69505ADBA400EE45DFAB131B" type="Zenodo-Dep">6457396</mods:identifier>
<mods:identifier id="2D404E662F3D9D0CCF07A23715E62D93" type="ZooBank">FC49517F-DD23-4281-A782-5BB6F9DC8560</mods:identifier>
</mods:mods>
<treatment id="03DB9B4DFFD7AF2FC98E9F78FA31947A" ID-DOI="http://doi.org/10.5281/zenodo.6457410" ID-GBIF-Taxon="194591677" ID-Zenodo-Dep="6457410" LSID="urn:lsid:plazi:treatment:03DB9B4DFFD7AF2FC98E9F78FA31947A" httpUri="http://treatment.plazi.org/id/03DB9B4DFFD7AF2FC98E9F78FA31947A" lastPageId="3" lastPageNumber="566" pageId="2" pageNumber="565">
<subSubSection id="C36879D0FFD7AF2EC98E9F78FD60913F" box="[151,717,1062,1089]" pageId="2" pageNumber="565" type="nomenclature">
<paragraph id="8BCD2A5BFFD7AF2EC98E9F78FD60913F" blockId="2.[151,717,1062,1089]" box="[151,717,1062,1089]" pageId="2" pageNumber="565">
<heading id="D0859D37FFD7AF2EC98E9F78FD60913F" bold="true" box="[151,717,1062,1089]" fontSize="11" level="1" pageId="2" pageNumber="565" reason="1">
<emphasis id="B906F649FFD7AF2EC98E9F78FD60913F" bold="true" box="[151,717,1062,1089]" italics="true" pageId="2" pageNumber="565">
Genus
<taxonomicName id="4C7251D8FFD7AF2EC9F79F79FD60913F" authority="Engel &amp; Grimaldi, 2008" authorityName="Engel &amp; Grimaldi" authorityYear="2008" box="[238,717,1062,1089]" class="Insecta" family="Prostomidae" genus="Vetuprostomis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="565" phylum="Arthropoda" rank="genus">
Vetuprostomis
<bibRefCitation id="EFE357AAFFD7AF2EC8B89F79FD60913F" author="Engel, M. S. &amp; Grimaldi, D. A." box="[417,717,1062,1089]" pageId="2" pageNumber="565" pagination="215 - 218" refId="ref5071" refString="Engel, M. S. &amp; Grimaldi, D. A. (2008) A jugular-horned beetle in Cretaceous amber from Myanmar (Coleoptera: Prostomidae). Alavesia, 2, 215 - 218." type="journal article" year="2008">Engel &amp; Grimaldi, 2008</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36879D0FFD7AF2EC98E9F31FAD09346" pageId="2" pageNumber="565" type="discussion">
<paragraph id="8BCD2A5BFFD7AF2EC98E9F31FE5390D6" blockId="2.[151,1437,1135,1595]" pageId="2" pageNumber="565">
<emphasis id="B906F649FFD7AF2EC98E9F31FEA691F6" bold="true" box="[151,267,1135,1161]" pageId="2" pageNumber="565">Remarks.</emphasis>
Although
<bibRefCitation id="EFE357AAFFD7AF2EC8989F31FD3091F6" author="Engel, M. S. &amp; Grimaldi, D. A." box="[385,669,1135,1161]" pageId="2" pageNumber="565" pagination="215 - 218" refId="ref5071" refString="Engel, M. S. &amp; Grimaldi, D. A. (2008) A jugular-horned beetle in Cretaceous amber from Myanmar (Coleoptera: Prostomidae). Alavesia, 2, 215 - 218." type="journal article" year="2008">Engel &amp; Grimaldi (2008)</bibRefCitation>
mentioned that
<taxonomicName id="4C7251D8FFD7AF2ECA489F31FC7091F6" authorityName="Thomson" authorityYear="1859" box="[849,989,1135,1161]" class="Insecta" family="Prostomidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="565" phylum="Arthropoda" rank="family">Prostomidae</taxonomicName>
consist of two extant genera,
<taxonomicName id="4C7251D8FFD7AF2ECC359F31FA3091F7" box="[1324,1437,1135,1160]" class="Insecta" family="Prostomidae" genus="Prostomis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="565" phylum="Arthropoda" rank="genus">
<emphasis id="B906F649FFD7AF2ECC359F31FA3091F7" box="[1324,1437,1135,1160]" italics="true" pageId="2" pageNumber="565">Prostomis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7251D8FFD7AF2EC9D19FCDFE9891D3" authorityName="Pascoe" authorityYear="1868" box="[200,309,1171,1196]" class="Insecta" family="Prostomidae" genus="Dryocora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="565" phylum="Arthropoda" rank="genus">
<emphasis id="B906F649FFD7AF2EC9D19FCDFE9891D3" box="[200,309,1171,1196]" italics="true" pageId="2" pageNumber="565">Dryocora</emphasis>
</taxonomicName>
, it seems that they did not compare their fossil with members of
<taxonomicName id="4C7251D8FFD7AF2ECD069FCDFB2191D3" authorityName="Pascoe" authorityYear="1868" box="[1055,1164,1171,1196]" class="Insecta" family="Prostomidae" genus="Dryocora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="565" phylum="Arthropoda" rank="genus">
<emphasis id="B906F649FFD7AF2ECD069FCDFB2191D3" box="[1055,1164,1171,1196]" italics="true" pageId="2" pageNumber="565">Dryocora</emphasis>
</taxonomicName>
at all. Their differential diagnosis was proposed based on comparison with
<taxonomicName id="4C7251D8FFD7AF2ECBC79FE9FCE291AF" box="[734,847,1207,1232]" class="Insecta" family="Prostomidae" genus="Prostomis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="565" phylum="Arthropoda" rank="genus">
<emphasis id="B906F649FFD7AF2ECBC79FE9FCE291AF" box="[734,847,1207,1232]" italics="true" pageId="2" pageNumber="565">Prostomis</emphasis>
</taxonomicName>
solely.
<bibRefCitation id="EFE357AAFFD7AF2ECABE9FE9FB6A91AE" author="Engel, M. S. &amp; Grimaldi, D. A." box="[935,1223,1207,1233]" pageId="2" pageNumber="565" pagination="215 - 218" refId="ref5071" refString="Engel, M. S. &amp; Grimaldi, D. A. (2008) A jugular-horned beetle in Cretaceous amber from Myanmar (Coleoptera: Prostomidae). Alavesia, 2, 215 - 218." type="journal article" year="2008">Engel &amp; Grimaldi (2008)</bibRefCitation>
claimed that
<taxonomicName id="4C7251D8FFD7AF2ECC7A9FE9FEA9918B" authorityName="Engel &amp; Grimaldi" authorityYear="2008" class="Insecta" family="Prostomidae" genus="Vetuprostomis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="565" phylum="Arthropoda" rank="genus">
<emphasis id="B906F649FFD7AF2ECC7A9FE9FEA9918B" italics="true" pageId="2" pageNumber="565">Vetuprostomis</emphasis>
</taxonomicName>
differs from modern prostomids in the shorter jugular processes, and the wider (about as wide as head) and the anteriorly constricted pronotum. However, extant
<taxonomicName id="4C7251D8FFD7AF2ECA3B9FA1FC229067" authorityName="Pascoe" authorityYear="1868" box="[802,911,1279,1304]" class="Insecta" family="Prostomidae" genus="Dryocora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="565" phylum="Arthropoda" rank="genus">
<emphasis id="B906F649FFD7AF2ECA3B9FA1FC229067" box="[802,911,1279,1304]" italics="true" pageId="2" pageNumber="565">Dryocora</emphasis>
</taxonomicName>
also has short jugular processes and anteriorly constricted pronotum. The pronotum can even be wider than head at least in some of
<taxonomicName id="4C7251D8FFD7AF2ECD6B9E7DFAED9042" box="[1138,1344,1315,1341]" class="Insecta" family="Prostomidae" genus="Dryocora" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="565" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B906F649FFD7AF2ECD6B9E7DFB729043" box="[1138,1247,1315,1340]" italics="true" pageId="2" pageNumber="565">Dryocora</emphasis>
species.
</taxonomicName>
Indeed,
<taxonomicName id="4C7251D8FFD7AF2EC98E9E19FE98901F" authorityName="Engel &amp; Grimaldi" authorityYear="2008" box="[151,309,1351,1376]" class="Insecta" family="Prostomidae" genus="Vetuprostomis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="565" phylum="Arthropoda" rank="genus">
<emphasis id="B906F649FFD7AF2EC98E9E19FE98901F" box="[151,309,1351,1376]" italics="true" pageId="2" pageNumber="565">Vetuprostomis</emphasis>
</taxonomicName>
is more similar to
<taxonomicName id="4C7251D8FFD7AF2ECB0D9E19FD2C901F" authorityName="Pascoe" authorityYear="1868" box="[532,641,1351,1376]" class="Insecta" family="Prostomidae" genus="Dryocora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="565" phylum="Arthropoda" rank="genus">
<emphasis id="B906F649FFD7AF2ECB0D9E19FD2C901F" box="[532,641,1351,1376]" italics="true" pageId="2" pageNumber="565">Dryocora</emphasis>
</taxonomicName>
than
<taxonomicName id="4C7251D8FFD7AF2ECBDA9E19FC99901F" box="[707,820,1351,1376]" class="Insecta" family="Prostomidae" genus="Prostomis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="565" phylum="Arthropoda" rank="genus">
<emphasis id="B906F649FFD7AF2ECBDA9E19FC99901F" box="[707,820,1351,1376]" italics="true" pageId="2" pageNumber="565">Prostomis</emphasis>
</taxonomicName>
. In addition to the characters mentioned above,
<taxonomicName id="4C7251D8FFD7AF2ECC7A9E19FEA990FB" authorityName="Engel &amp; Grimaldi" authorityYear="2008" class="Insecta" family="Prostomidae" genus="Vetuprostomis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="565" phylum="Arthropoda" rank="genus">
<emphasis id="B906F649FFD7AF2ECC7A9E19FEA990FB" italics="true" pageId="2" pageNumber="565">Vetuprostomis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7251D8FFD7AF2EC8269E35FE0190FB" authorityName="Pascoe" authorityYear="1868" box="[319,428,1387,1412]" class="Insecta" family="Prostomidae" genus="Dryocora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="565" phylum="Arthropoda" rank="genus">
<emphasis id="B906F649FFD7AF2EC8269E35FE0190FB" box="[319,428,1387,1412]" italics="true" pageId="2" pageNumber="565">Dryocora</emphasis>
</taxonomicName>
share the shorter and more simple mandibles and the posterolaterally projected temples compared to the nominal genus.
</paragraph>
<paragraph id="8BCD2A5BFFD7AF2EC9DE9EEDFAD09346" blockId="2.[151,1437,1135,1595]" pageId="2" pageNumber="565">
Extant prostomids have mandibles with three apical teeth. In
<bibRefCitation id="EFE357AAFFD7AF2ECA8B9EEDFB1790B2" author="Engel, M. S. &amp; Grimaldi, D. A." box="[914,1210,1459,1485]" pageId="2" pageNumber="565" pagination="215 - 218" refId="ref5071" refString="Engel, M. S. &amp; Grimaldi, D. A. (2008) A jugular-horned beetle in Cretaceous amber from Myanmar (Coleoptera: Prostomidae). Alavesia, 2, 215 - 218." type="journal article" year="2008">Engel &amp; Grimaldi (2008)</bibRefCitation>
,
<taxonomicName id="4C7251D8FFD7AF2ECDD09EEDFACA90B3" authorityName="Engel &amp; Grimaldi" authorityYear="2008" box="[1225,1383,1459,1484]" class="Insecta" family="Prostomidae" genus="Vetuprostomis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="565" phylum="Arthropoda" rank="genus">
<emphasis id="B906F649FFD7AF2ECDD09EEDFACA90B3" box="[1225,1383,1459,1484]" italics="true" pageId="2" pageNumber="565">Vetuprostomis</emphasis>
</taxonomicName>
was described as having apically bidentate mandibles. This difference is merely due to a different terminology. The first apicomarginal tooth
<emphasis id="B906F649FFD7AF2EC8999EA5FE13936B" box="[384,446,1531,1556]" italics="true" pageId="2" pageNumber="565">sensu</emphasis>
<bibRefCitation id="EFE357AAFFD7AF2EC8DF9EA5FD48936A" author="Engel, M. S. &amp; Grimaldi, D. A." box="[454,741,1531,1557]" pageId="2" pageNumber="565" pagination="215 - 218" refId="ref5071" refString="Engel, M. S. &amp; Grimaldi, D. A. (2008) A jugular-horned beetle in Cretaceous amber from Myanmar (Coleoptera: Prostomidae). Alavesia, 2, 215 - 218." type="journal article" year="2008">Engel &amp; Grimaldi (2008)</bibRefCitation>
is identical to the dorsal apical tooth in extant prostomids. In the present paper we follow the terminology used for the extant
<taxonomicName id="4C7251D8FFD7AF2ECA7C9D41FC5C9346" authorityName="Thomson" authorityYear="1859" box="[869,1009,1567,1593]" class="Insecta" family="Prostomidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="565" phylum="Arthropoda" rank="family">Prostomidae</taxonomicName>
(e.g.,
<bibRefCitation id="EFE357AAFFD7AF2ECD219D41FADC9346" author="Lawrence, J. F. &amp; Slipinski, A." box="[1080,1393,1567,1595]" pageId="2" pageNumber="565" refId="ref5620" refString="Lawrence, J. F. &amp; Slipinski, A. (2013) Australian beetles. Vol. 1. Morphology, classification and keys. CSIRO Publishing, Clayton, 520 pp. https: // doi. org / 10.1071 / 9780643097292" type="book" year="2013">Lawrence &amp; Ślipiński 2013</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36879D0FFD7AF2EC98E9DDBFDB493E0" box="[151,537,1669,1695]" pageId="2" pageNumber="565" type="multiple">
<paragraph id="8BCD2A5BFFD7AF2EC98E9DDBFDB493E0" blockId="2.[151,537,1669,1695]" box="[151,537,1669,1695]" pageId="2" pageNumber="565">
<heading id="D0859D37FFD7AF2EC98E9DDBFDB493E0" bold="true" box="[151,537,1669,1695]" fontSize="11" level="1" pageId="2" pageNumber="565" reason="1">
<emphasis id="B906F649FFD7AF2EC98E9DDBFDB493E0" bold="true" box="[151,537,1669,1695]" pageId="2" pageNumber="565">
Key to species of
<taxonomicName id="4C7251D8FFD7AF2EC8779DDBFDB493E0" authorityName="Engel &amp; Grimaldi" authorityYear="2008" box="[366,537,1669,1695]" class="Insecta" family="Prostomidae" genus="Vetuprostomis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="565" phylum="Arthropoda" rank="genus">
<emphasis id="B906F649FFD7AF2EC8779DDBFDB493E0" bold="true" box="[366,537,1669,1695]" italics="true" pageId="2" pageNumber="565">Vetuprostomis</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36879D0FFD7AF2FC98E9D94FA31947A" lastPageId="3" lastPageNumber="566" pageId="2" pageNumber="565" type="key">
<keyStep id="308632FEFFD7AF2EC98E9D94FA3192D3" pageId="2" pageNumber="565">
<paragraph id="8BCD2A5BFFD7AF2EC98E9D94FA309247" blockId="2.[151,1437,1738,2024]" pageId="2" pageNumber="565">
<keyLead id="3083896EFFD7AF2EC98E9D94FA309247" pageId="2" pageNumber="565">
<emphasis id="B906F649FFD7AF2EC98E9D94FF0793A0" bold="true" box="[151,170,1738,1759]" pageId="2" pageNumber="565">1.</emphasis>
Lateral sides of clypeus anteriorly converging (fig. 2A in
<bibRefCitation id="EFE357AAFFD7AF2ECA189D94FC48939F" author="Engel, M. S. &amp; Grimaldi, D. A." box="[769,997,1738,1760]" pageId="2" pageNumber="565" pagination="215 - 218" refId="ref5071" refString="Engel, M. S. &amp; Grimaldi, D. A. (2008) A jugular-horned beetle in Cretaceous amber from Myanmar (Coleoptera: Prostomidae). Alavesia, 2, 215 - 218." type="journal article" year="2008">Engel &amp; Grimaldi 2008</bibRefCitation>
). Penultimate maxillary palpomere wider than long, shorter than half length of apical one (?) (fig. 2C in
<bibRefCitation id="EFE357AAFFD7AF2ECA1B9DB6FC4A9381" author="Engel, M. S. &amp; Grimaldi, D. A." box="[770,999,1768,1790]" pageId="2" pageNumber="565" pagination="215 - 218" refId="ref5071" refString="Engel, M. S. &amp; Grimaldi, D. A. (2008) A jugular-horned beetle in Cretaceous amber from Myanmar (Coleoptera: Prostomidae). Alavesia, 2, 215 - 218." type="journal article" year="2008">Engel &amp; Grimaldi 2008</bibRefCitation>
). Jugal processes symmetrical, without lateral teeth or knobs (fig. 2C in
<bibRefCitation id="EFE357AAFFD7AF2EC8C49C5BFD6A9264" author="Engel, M. S. &amp; Grimaldi, D. A." box="[477,711,1797,1819]" pageId="2" pageNumber="565" pagination="215 - 218" refId="ref5071" refString="Engel, M. S. &amp; Grimaldi, D. A. (2008) A jugular-horned beetle in Cretaceous amber from Myanmar (Coleoptera: Prostomidae). Alavesia, 2, 215 - 218." type="journal article" year="2008">Engel &amp; Grimaldi 2008</bibRefCitation>
). Median groove of pronotum well developed, (almost) as long as pronotal length (fig.
<quantity id="4C8A87BEFFD7AF2EC84A9C7CFED59247" box="[339,376,1826,1848]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="2" pageNumber="565" unit="in" value="1.0">1 in</quantity>
<bibRefCitation id="EFE357AAFFD7AF2EC8649C7CFDCF9247" author="Engel, M. S. &amp; Grimaldi, D. A." box="[381,610,1826,1848]" pageId="2" pageNumber="565" pagination="215 - 218" refId="ref5071" refString="Engel, M. S. &amp; Grimaldi, D. A. (2008) A jugular-horned beetle in Cretaceous amber from Myanmar (Coleoptera: Prostomidae). Alavesia, 2, 215 - 218." type="journal article" year="2008">Engel &amp; Grimaldi 2008</bibRefCitation>
)..........................................
<taxonomicName id="4C7251D8FFD7AF2ECD799C7DFA309247" authority="Engel &amp; Grimaldi" authorityName="Engel &amp; Grimaldi" authorityYear="2008" box="[1120,1437,1826,1848]" class="Insecta" family="Prostomidae" genus="Vetuprostomis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="565" phylum="Arthropoda" rank="species" species="consimilis">
<emphasis id="B906F649FFD7AF2ECD799C7DFA309247" bold="true" box="[1120,1437,1826,1848]" pageId="2" pageNumber="565">
<emphasis id="B906F649FFD7AF2ECD799C7DFB709247" bold="true" box="[1120,1245,1826,1848]" italics="true" pageId="2" pageNumber="565">V. consimilis</emphasis>
Engel &amp; Grimaldi
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BCD2A5BFFD7AF2EC98E9C1EFA3192D3" blockId="2.[151,1437,1738,2024]" pageId="2" pageNumber="565">
<keyLead id="3083896EFFD7AF2EC98E9C1EFA3192D3" pageId="2" pageNumber="565">
<emphasis id="B906F649FFD7AF2EC98E9C1EFF32922A" bold="true" box="[151,159,1856,1877]" pageId="2" pageNumber="565">-</emphasis>
Lateral sides of clypeus subparallel (or weakly diverging) in anterior portion (
<figureCitation id="134936DEFFD7AF2ECAC99C1EFB8D922A" box="[976,1056,1856,1878]" captionStart="FIGURE 4" captionStartId="5.[151,260,1841,1867]" captionTargetBox="[151,1436,954,1816]" captionTargetId="figure-323@5.[151,1436,954,1816]" captionTargetPageId="5" captionText="FIGURE 4. Vetuprostomis angularis Li &amp; Cai sp. nov., holotype, NIGP177907, under confocal microscopy. A, Mouthparts, ventral view. B, Prothorax, ventral view. C, Mesothorax, ventral view. D, Mouthparts, dorsal view. E, Prothorax, dorsal view. F, Fore leg. Abbreviations: an, antenna; cl, clypeus; ey, compound eye; jp, jugular process; lb, labrum; md, mandible; msv, mesoventrite; mxp, maxillary palp; pf, profemur; pn, pronotum; ps, prosternum; ptb, protibia; pts, protarsus. Scale bars: 300 μm." figureDoi="http://doi.org/10.5281/zenodo.6457412" httpUri="https://zenodo.org/record/6457412/files/figure.png" pageId="2" pageNumber="565">Figs 4D</figureCitation>
,
<figureCitation id="134936DEFFD7AF2ECD339C1EFBE6922A" box="[1066,1099,1856,1877]" captionStart="FIGURE 7" captionStartId="8.[151,260,1790,1816]" captionTargetBox="[161,1427,486,1765]" captionTargetId="figure-152@8.[161,1427,486,1765]" captionTargetPageId="8" captionText="FIGURE 7. Vetuprostomis gaoi Li &amp; Cai sp. nov., holotype, NIGP177908, under confocal microscopy. A, Mouthparts, ventral view. B, Temporal projection (arrowhead), ventral view. C, Prothorax, ventral view. D, Fore leg. E, Mesothorax, dorsal view. F, Hind leg. G, Abdomen, ventral view. E, Mouthparts, dorsal view. F, Elytral apex, dorsal view. Abbreviations: cl, clypeus; el, elytron; ey, compound eye; jp, jugular process; lb, labrum; lbp, labial palp; md, mandible; msv, mesoventrite; mt, mentum; mtf, metafemur; mttb, metatibia; mtts, metatarsus; mxp, maxillary palp; pc, procoxa; pf, profemur; ps, prosternum; ptb, protibia; ptc, protrochanter; pts, protarsus; v15, ventrites 15. Scale bars: 200 μm." figureDoi="http://doi.org/10.5281/zenodo.6457422" httpUri="https://zenodo.org/record/6457422/files/figure.png" pageId="2" pageNumber="565">7H</figureCitation>
). Penultimate maxillary palpomere longer than wide, distinctly longer than half length of apical one (
<figureCitation id="134936DEFFD7AF2ECA459C03FC69920D" box="[860,964,1885,1907]" captionStart="FIGURE 4" captionStartId="5.[151,260,1841,1867]" captionTargetBox="[151,1436,954,1816]" captionTargetId="figure-323@5.[151,1436,954,1816]" captionTargetPageId="5" captionText="FIGURE 4. Vetuprostomis angularis Li &amp; Cai sp. nov., holotype, NIGP177907, under confocal microscopy. A, Mouthparts, ventral view. B, Prothorax, ventral view. C, Mesothorax, ventral view. D, Mouthparts, dorsal view. E, Prothorax, dorsal view. F, Fore leg. Abbreviations: an, antenna; cl, clypeus; ey, compound eye; jp, jugular process; lb, labrum; md, mandible; msv, mesoventrite; mxp, maxillary palp; pf, profemur; pn, pronotum; ps, prosternum; ptb, protibia; pts, protarsus. Scale bars: 300 μm." figureDoi="http://doi.org/10.5281/zenodo.6457412" httpUri="https://zenodo.org/record/6457412/files/figure.png" pageId="2" pageNumber="565">Figs 4A,D</figureCitation>
,
<figureCitation id="134936DEFFD7AF2ECAD79C03FBAB920C" box="[974,1030,1885,1907]" captionStart="FIGURE 7" captionStartId="8.[151,260,1790,1816]" captionTargetBox="[161,1427,486,1765]" captionTargetId="figure-152@8.[161,1427,486,1765]" captionTargetPageId="8" captionText="FIGURE 7. Vetuprostomis gaoi Li &amp; Cai sp. nov., holotype, NIGP177908, under confocal microscopy. A, Mouthparts, ventral view. B, Temporal projection (arrowhead), ventral view. C, Prothorax, ventral view. D, Fore leg. E, Mesothorax, dorsal view. F, Hind leg. G, Abdomen, ventral view. E, Mouthparts, dorsal view. F, Elytral apex, dorsal view. Abbreviations: cl, clypeus; el, elytron; ey, compound eye; jp, jugular process; lb, labrum; lbp, labial palp; md, mandible; msv, mesoventrite; mt, mentum; mtf, metafemur; mttb, metatibia; mtts, metatarsus; mxp, maxillary palp; pc, procoxa; pf, profemur; ps, prosternum; ptb, protibia; ptc, protrochanter; pts, protarsus; v15, ventrites 15. Scale bars: 200 μm." figureDoi="http://doi.org/10.5281/zenodo.6457422" httpUri="https://zenodo.org/record/6457422/files/figure.png" pageId="2" pageNumber="565">7A,H</figureCitation>
). Jugal processes asymmetrical; right process longer; inner sides with a tooth (
<figureCitation id="134936DEFFD7AF2ECB5E9C24FD3A92F0" box="[583,663,1914,1936]" captionStart="FIGURE 4" captionStartId="5.[151,260,1841,1867]" captionTargetBox="[151,1436,954,1816]" captionTargetId="figure-323@5.[151,1436,954,1816]" captionTargetPageId="5" captionText="FIGURE 4. Vetuprostomis angularis Li &amp; Cai sp. nov., holotype, NIGP177907, under confocal microscopy. A, Mouthparts, ventral view. B, Prothorax, ventral view. C, Mesothorax, ventral view. D, Mouthparts, dorsal view. E, Prothorax, dorsal view. F, Fore leg. Abbreviations: an, antenna; cl, clypeus; ey, compound eye; jp, jugular process; lb, labrum; md, mandible; msv, mesoventrite; mxp, maxillary palp; pf, profemur; pn, pronotum; ps, prosternum; ptb, protibia; pts, protarsus. Scale bars: 300 μm." figureDoi="http://doi.org/10.5281/zenodo.6457412" httpUri="https://zenodo.org/record/6457412/files/figure.png" pageId="2" pageNumber="565">Figs 4A</figureCitation>
,
<figureCitation id="134936DEFFD7AF2ECBB89C24FD6C92EF" box="[673,705,1914,1936]" captionStart="FIGURE 7" captionStartId="8.[151,260,1790,1816]" captionTargetBox="[161,1427,486,1765]" captionTargetId="figure-152@8.[161,1427,486,1765]" captionTargetPageId="8" captionText="FIGURE 7. Vetuprostomis gaoi Li &amp; Cai sp. nov., holotype, NIGP177908, under confocal microscopy. A, Mouthparts, ventral view. B, Temporal projection (arrowhead), ventral view. C, Prothorax, ventral view. D, Fore leg. E, Mesothorax, dorsal view. F, Hind leg. G, Abdomen, ventral view. E, Mouthparts, dorsal view. F, Elytral apex, dorsal view. Abbreviations: cl, clypeus; el, elytron; ey, compound eye; jp, jugular process; lb, labrum; lbp, labial palp; md, mandible; msv, mesoventrite; mt, mentum; mtf, metafemur; mttb, metatibia; mtts, metatarsus; mxp, maxillary palp; pc, procoxa; pf, profemur; ps, prosternum; ptb, protibia; ptc, protrochanter; pts, protarsus; v15, ventrites 15. Scale bars: 200 μm." figureDoi="http://doi.org/10.5281/zenodo.6457422" httpUri="https://zenodo.org/record/6457422/files/figure.png" pageId="2" pageNumber="565">7A</figureCitation>
). Median groove of pronotum absent or weakly developed, shorter than half length of pronotum (
<figureCitation id="134936DEFFD7AF2EC8B19CC6FE5C92D1" box="[424,497,1944,1966]" captionStart="FIGURE 4" captionStartId="5.[151,260,1841,1867]" captionTargetBox="[151,1436,954,1816]" captionTargetId="figure-323@5.[151,1436,954,1816]" captionTargetPageId="5" captionText="FIGURE 4. Vetuprostomis angularis Li &amp; Cai sp. nov., holotype, NIGP177907, under confocal microscopy. A, Mouthparts, ventral view. B, Prothorax, ventral view. C, Mesothorax, ventral view. D, Mouthparts, dorsal view. E, Prothorax, dorsal view. F, Fore leg. Abbreviations: an, antenna; cl, clypeus; ey, compound eye; jp, jugular process; lb, labrum; md, mandible; msv, mesoventrite; mxp, maxillary palp; pf, profemur; pn, pronotum; ps, prosternum; ptb, protibia; pts, protarsus. Scale bars: 300 μm." figureDoi="http://doi.org/10.5281/zenodo.6457412" httpUri="https://zenodo.org/record/6457412/files/figure.png" pageId="2" pageNumber="565">Fig. 4E</figureCitation>
)............................................................................
<emphasis id="B906F649FFD7AF2ECC899CC9FA3192D3" bold="true" box="[1424,1436,1943,1964]" pageId="2" pageNumber="565">2</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="308632FEFFD7AF2FC98E9CEBFA31947A" lastPageId="3" lastPageNumber="566" pageId="2" pageNumber="565">
<paragraph id="8BCD2A5BFFD7AF2FC98E9CEBFA3195D2" blockId="2.[151,1437,1738,2024]" lastBlockId="3.[151,1436,151,261]" lastPageId="3" lastPageNumber="566" pageId="2" pageNumber="565">
<keyLead id="3083896EFFD7AF2FC98E9CEBFA3195D2" lastPageId="3" lastPageNumber="566" pageId="2" pageNumber="565">
<emphasis id="B906F649FFD7AF2EC98E9CEBFF0492B5" bold="true" box="[151,169,1973,1994]" pageId="2" pageNumber="565">2.</emphasis>
Jugal processes longer (e.g., right process: portion above tooth more than twice as long as portion below tooth) (
<figureCitation id="134936DEFFD7AF2ECC129CEBFAF592B4" box="[1291,1368,1973,1995]" captionStart="FIGURE 4" captionStartId="5.[151,260,1841,1867]" captionTargetBox="[151,1436,954,1816]" captionTargetId="figure-323@5.[151,1436,954,1816]" captionTargetPageId="5" captionText="FIGURE 4. Vetuprostomis angularis Li &amp; Cai sp. nov., holotype, NIGP177907, under confocal microscopy. A, Mouthparts, ventral view. B, Prothorax, ventral view. C, Mesothorax, ventral view. D, Mouthparts, dorsal view. E, Prothorax, dorsal view. F, Fore leg. Abbreviations: an, antenna; cl, clypeus; ey, compound eye; jp, jugular process; lb, labrum; md, mandible; msv, mesoventrite; mxp, maxillary palp; pf, profemur; pn, pronotum; ps, prosternum; ptb, protibia; pts, protarsus. Scale bars: 300 μm." figureDoi="http://doi.org/10.5281/zenodo.6457412" httpUri="https://zenodo.org/record/6457412/files/figure.png" pageId="2" pageNumber="565">Fig. 4A</figureCitation>
). Teeth on incisor edge of left mandible moderately separated (
<figureCitation id="134936DEFFD7AF2ECBEA9C8CFCEE9298" box="[755,835,2002,2024]" captionStart="FIGURE 3" captionStartId="4.[151,260,1723,1749]" captionTargetBox="[151,1436,306,1698]" captionTargetId="figure-61@4.[151,1436,306,1698]" captionTargetPageId="4" captionText="FIGURE 3. Vetuprostomis angularis Li &amp; Cai sp. nov., holotype, NIGP177907, under widefield fluorescence. A, Head and prothorax, dorsal view. B, Head and prothorax, ventral view. C, Elytral apex, dorsal view. D, Pterothorax, ventral view. E, Abdomen, ventral view. Abbreviations: an, antenna; el, elytron; jp, jugular process; md, mandible; msf, mesofemur; msts, mesotarsus; msv, mesoventrite; mtf, metafemur; mttb, metatibia; mtts, metatarsus; mtv, metaventrite; mxp, maxillary palp; pf, profemur; pn, pronotum; ps, prosternum; ptb, protibia; pts, protarsus. Scale bars: 500 μm." figureDoi="http://doi.org/10.5281/zenodo.6457408" httpUri="https://zenodo.org/record/6457408/files/figure.png" pageId="2" pageNumber="565">Figs 3A</figureCitation>
,
<figureCitation id="134936DEFFD7AF2ECA549C8CFCC09297" box="[845,877,2002,2024]" captionStart="FIGURE 4" captionStartId="5.[151,260,1841,1867]" captionTargetBox="[151,1436,954,1816]" captionTargetId="figure-323@5.[151,1436,954,1816]" captionTargetPageId="5" captionText="FIGURE 4. Vetuprostomis angularis Li &amp; Cai sp. nov., holotype, NIGP177907, under confocal microscopy. A, Mouthparts, ventral view. B, Prothorax, ventral view. C, Mesothorax, ventral view. D, Mouthparts, dorsal view. E, Prothorax, dorsal view. F, Fore leg. Abbreviations: an, antenna; cl, clypeus; ey, compound eye; jp, jugular process; lb, labrum; md, mandible; msv, mesoventrite; mxp, maxillary palp; pf, profemur; pn, pronotum; ps, prosternum; ptb, protibia; pts, protarsus. Scale bars: 300 μm." figureDoi="http://doi.org/10.5281/zenodo.6457412" httpUri="https://zenodo.org/record/6457412/files/figure.png" pageId="2" pageNumber="565">4D</figureCitation>
). Anterior angles of pronotum pointed (
<figureCitation id="134936DEFFD7AF2ECDF39C8CFA859298" box="[1258,1320,2002,2024]" captionStart="FIGURE 2" captionStartId="3.[151,260,1962,1988]" captionTargetBox="[208,1379,824,1939]" captionTargetId="figure-319@3.[208,1379,824,1939]" captionTargetPageId="3" captionText="FIGURE 2. General habitus of Vetuprostomis angularis Li &amp; Cai sp. nov., holotype, NIGP177907, under incident light. A, Dorsal view. B, Ventral view. Scale bars: 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6457404" httpUri="https://zenodo.org/record/6457404/files/figure.png" pageId="2" pageNumber="565">
Figs
<geoCoordinate id="EE464C9CFFD7AF2ECC039C8CFA859298" box="[1306,1320,2002,2023]" degrees="2, 4" direction="east" orientation="longitude" pageId="2" pageNumber="565" precision="5555" value="2.4">2</geoCoordinate>
</figureCitation>
,
<figureCitation id="134936DEFFD7AF2ECC2B9C8CFAE29297" box="[1330,1359,2002,2024]" captionStart="FIGURE 4" captionStartId="5.[151,260,1841,1867]" captionTargetBox="[151,1436,954,1816]" captionTargetId="figure-323@5.[151,1436,954,1816]" captionTargetPageId="5" captionText="FIGURE 4. Vetuprostomis angularis Li &amp; Cai sp. nov., holotype, NIGP177907, under confocal microscopy. A, Mouthparts, ventral view. B, Prothorax, ventral view. C, Mesothorax, ventral view. D, Mouthparts, dorsal view. E, Prothorax, dorsal view. F, Fore leg. Abbreviations: an, antenna; cl, clypeus; ey, compound eye; jp, jugular process; lb, labrum; md, mandible; msv, mesoventrite; mxp, maxillary palp; pf, profemur; pn, pronotum; ps, prosternum; ptb, protibia; pts, protarsus. Scale bars: 300 μm." figureDoi="http://doi.org/10.5281/zenodo.6457412" httpUri="https://zenodo.org/record/6457412/files/figure.png" pageId="2" pageNumber="565">4E</figureCitation>
)................................................................................
<emphasis id="B906F649FFD6AF2FCD689BC6FA3195D2" bold="true" box="[1137,1436,151,173]" pageId="3" pageNumber="566">
<taxonomicName id="4C7251D8FFD6AF2FCD689BC6FAE795D2" authority="Li &amp; Cai" authorityName="Li &amp; Cai" authorityYear="2022" box="[1137,1354,151,173]" class="Insecta" family="Prostomidae" genus="Vetuprostomis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="566" phylum="Arthropoda" rank="species" species="angularis" status="sp. nov.">
<emphasis id="B906F649FFD6AF2FCD689BC6FB4495D2" bold="true" box="[1137,1257,151,173]" italics="true" pageId="3" pageNumber="566">V. angularis</emphasis>
Li &amp; Cai
</taxonomicName>
<taxonomicNameLabel id="A2354B32FFD6AF2FCC499BC6FA3195D2" box="[1360,1436,152,173]" pageId="3" pageNumber="566" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BCD2A5BFFD6AF2FC98E9BEBFA31947A" blockId="3.[151,1436,151,261]" pageId="3" pageNumber="566">
<keyLead id="3083896EFFD6AF2FC98E9BEBFA31947A" pageId="3" pageNumber="566">
<emphasis id="B906F649FFD6AF2FC98E9BEBFF3295B5" bold="true" box="[151,159,181,202]" pageId="3" pageNumber="566">-</emphasis>
Jugal processes shorter (e.g., right process: portion above tooth less than twice as long as portion below tooth) (
<figureCitation id="134936DEFFD6AF2FCC139BEBFAFA95B4" box="[1290,1367,181,203]" captionStart="FIGURE 7" captionStartId="8.[151,260,1790,1816]" captionTargetBox="[161,1427,486,1765]" captionTargetId="figure-152@8.[161,1427,486,1765]" captionTargetPageId="8" captionText="FIGURE 7. Vetuprostomis gaoi Li &amp; Cai sp. nov., holotype, NIGP177908, under confocal microscopy. A, Mouthparts, ventral view. B, Temporal projection (arrowhead), ventral view. C, Prothorax, ventral view. D, Fore leg. E, Mesothorax, dorsal view. F, Hind leg. G, Abdomen, ventral view. E, Mouthparts, dorsal view. F, Elytral apex, dorsal view. Abbreviations: cl, clypeus; el, elytron; ey, compound eye; jp, jugular process; lb, labrum; lbp, labial palp; md, mandible; msv, mesoventrite; mt, mentum; mtf, metafemur; mttb, metatibia; mtts, metatarsus; mxp, maxillary palp; pc, procoxa; pf, profemur; ps, prosternum; ptb, protibia; ptc, protrochanter; pts, protarsus; v15, ventrites 15. Scale bars: 200 μm." figureDoi="http://doi.org/10.5281/zenodo.6457422" httpUri="https://zenodo.org/record/6457422/files/figure.png" pageId="3" pageNumber="566">Fig. 7A</figureCitation>
). Teeth on incisor edge of left mandible more narrowly separated and strongly jointed basally; interspace only very weakly notched (
<figureCitation id="134936DEFFD6AF2FC9F09BB1FE9B947A" box="[233,310,239,261]" captionStart="FIGURE 7" captionStartId="8.[151,260,1790,1816]" captionTargetBox="[161,1427,486,1765]" captionTargetId="figure-152@8.[161,1427,486,1765]" captionTargetPageId="8" captionText="FIGURE 7. Vetuprostomis gaoi Li &amp; Cai sp. nov., holotype, NIGP177908, under confocal microscopy. A, Mouthparts, ventral view. B, Temporal projection (arrowhead), ventral view. C, Prothorax, ventral view. D, Fore leg. E, Mesothorax, dorsal view. F, Hind leg. G, Abdomen, ventral view. E, Mouthparts, dorsal view. F, Elytral apex, dorsal view. Abbreviations: cl, clypeus; el, elytron; ey, compound eye; jp, jugular process; lb, labrum; lbp, labial palp; md, mandible; msv, mesoventrite; mt, mentum; mtf, metafemur; mttb, metatibia; mtts, metatarsus; mxp, maxillary palp; pc, procoxa; pf, profemur; ps, prosternum; ptb, protibia; ptc, protrochanter; pts, protarsus; v15, ventrites 15. Scale bars: 200 μm." figureDoi="http://doi.org/10.5281/zenodo.6457422" httpUri="https://zenodo.org/record/6457422/files/figure.png" pageId="3" pageNumber="566">Fig. 7H</figureCitation>
). Anterior angles of pronotum not pointed (
<figureCitation id="134936DEFFD6AF2FCBC19BB1FCBD947A" box="[728,784,239,261]" captionStart="FIGURE 5" captionStartId="6.[151,260,1907,1933]" captionTargetBox="[151,1436,595,1883]" captionTargetId="figure-158@6.[151,1436,595,1883]" captionTargetPageId="6" captionText="FIGURE 5. General habitus of Vetuprostomis gaoi Li &amp; Cai sp. nov., holotype, NIGP177908, under incident light. A, Dorsal view. B, Ventral view. Scale bars: 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6457416" httpUri="https://zenodo.org/record/6457416/files/figure.png" pageId="3" pageNumber="566">Fig. 5</figureCitation>
).................................
<emphasis id="B906F649FFD6AF2FCDBC9BAEFA31947A" bold="true" box="[1189,1436,239,261]" pageId="3" pageNumber="566">
<taxonomicName id="4C7251D8FFD6AF2FCDBC9BAEFAE7947A" authority="Li &amp; Cai" authorityName="Li &amp; Cai" authorityYear="2022" box="[1189,1354,239,261]" class="Insecta" family="Prostomidae" genus="Vetuprostomis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="566" phylum="Arthropoda" rank="species" species="gaoi" status="sp. nov.">
<emphasis id="B906F649FFD6AF2FCDBC9BAEFB44947B" bold="true" box="[1189,1257,239,261]" italics="true" pageId="3" pageNumber="566">V. gaoi</emphasis>
Li &amp; Cai
</taxonomicName>
<taxonomicNameLabel id="A2354B32FFD6AF2FCC499BAEFA31947A" box="[1360,1436,240,261]" pageId="3" pageNumber="566" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>