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<mods:title id="4C702960ABEADC9FDE3BD478256FA1A5">A synthesis on the scolopendromorph centipedes (Chilopoda: Scolopendromorpha) of Martinique, with description of Cryptops (Trigonocryptops) amicitia n. sp. and new data on some Neotropical scolopendromorphs</mods:title>
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<mods:namePart id="6F1A3A97834B67BB843CA41BCBFE8945">Schileyko, Arkady</mods:namePart>
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<mods:namePart id="D397BF2312E9E55838A6D12C6542AC8B">Iorio, Etienne</mods:namePart>
<mods:affiliation id="F2AF0A3942D74B8BFA103316DC429F4A">Entomologie &amp; Myriapodologie, 36 impasse des Acacias, 84260 Sarrians, France</mods:affiliation>
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<mods:namePart id="E87DBC13014A3629F5149EAB0F88D66C">Coulis, Mathieu</mods:namePart>
<mods:affiliation id="D2AA54EBB56AE0DE23F17304AB454FBF">CIRAD, UPR GECO, F- 97285 Le Lamentin, Martinique, France &amp; GECO, University Montpellier, CIRAD, Montpellier, France</mods:affiliation>
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Subgenus
<taxonomicName id="4C6AC382FFB4FFB08739FF62DD9CFF38" ID-CoL="8FB7" authority="Verhoeff, 1906" authorityName="Verhoeff" authorityYear="1906" box="[279,665,151,177]" kingdom="Animalia" pageId="15" pageNumber="578" rank="subGenus" subGenus="Trigonocryptops">
<emphasis id="B91E6413FFB4FFB08739FF62DED8FF38" bold="true" box="[279,477,151,177]" italics="true" pageId="15" pageNumber="578">Trigonocryptops</emphasis>
<bibRefCitation id="EFFBC5F0FFB4FFB087CAFF62DD9CFF38" author="Verhoeff, K. W." box="[484,665,151,177]" pageId="15" pageNumber="578" pagination="353 - 500" refId="ref31109" refString="Verhoeff, K. W. (1906) Vergleichend-morphologische Studie uber die coxopleuralen Korperteile der Chilopoden, mit besonderer Berucksichtigung der Scolopendromorpha, ein Beitrag zur Anatomie und Systematik derselben, nebst physiologischen und phylogenetischen Mitteilungen und Aus. Nova Acta Leopoldina, 86, 353 - 500." type="journal article" year="1906">Verhoeff, 1906</bibRefCitation>
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Type-species.
<taxonomicName id="4C6AC382FFB4FFB08706FF29DD69FF7D" authority="Kraepelin, 1903" authorityName="Kraepelin" authorityYear="1903" box="[296,620,220,244]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="15" pageNumber="578" phylum="Arthropoda" rank="species" species="gigas">
<emphasis id="B91E6413FFB4FFB08706FF29DEC7FF7A" box="[296,450,220,243]" italics="true" pageId="15" pageNumber="578">Cryptops gigas</emphasis>
<bibRefCitation id="EFFBC5F0FFB4FFB087E6FF29DD69FF7D" author="Kraepelin, K." box="[456,620,220,244]" pageId="15" pageNumber="578" pagination="1 - 276" refId="ref28397" refString="Kraepelin, K. (1903) Revision der Scolopendriden. Mitteilungen aus dem Naturhistorischen Museum in Hamburg, 20, 1 - 276." type="journal article" year="1903">Kraepelin, 1903</bibRefCitation>
</taxonomicName>
(by subsequent designation of
<bibRefCitation id="EFFBC5F0FFB4FFB08588FF29DB39FF7D" author="Attems, C." box="[934,1084,220,244]" pageId="15" pageNumber="578" pagination="1 - 306" refId="ref26698" refString="Attems, C. (1930) Myriopoda. 2. Scolopendromorpha. Das Tierreich, Lfg. 54, 1 - 306. https: // doi. org / 10.1515 / 9783112373002" type="journal article" year="1930">Attems (1930)</bibRefCitation>
.
</paragraph>
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<subSubSection id="C370EB8AFFB4FFB086B9FED4DC4DFC99" pageId="15" pageNumber="578" type="diagnosis">
<paragraph id="8BD5B801FFB4FFB086B9FED4DB57FD41" blockId="15.[151,1437,289,1252]" pageId="15" pageNumber="578">
<emphasis id="B91E6413FFB4FFB086B9FED4DE0FFEB2" bold="true" box="[151,266,289,315]" pageId="15" pageNumber="578">Diagnosis</emphasis>
. Cephalic plate with complete or (more rarely) incomplete paramedian sutures. Clypeus with setose plate(s), which are delimited by sutures (
<figureCitation id="1351A484FFB4FFB0844FFEB3DDB1FEE9" box="[609,692,326,352]" captionStart="FIGURES 2025" captionStartId="17.[151,264,1352,1376]" captionTargetBox="[152,1435,182,1328]" captionTargetId="figure-17@17.[151,1435,181,1328]" captionTargetPageId="17" captionText="FIGURES 2025. Cryptops (Trigonocryptops) amicitia n. sp., holotype, 212-P8G (MNHN). 20. Head, ventrally. Scale bar: 0.2 mm. 21. LBS 34, ventrally. Scale bar: 0.3 mm. 22. LBS 1314, ventrally. Scale bar: 0.4 mm. 23. Head + LBS 1, dorsally. Scale bar: 0.4 mm. 24. LBS 2021, ventrally. Scale bar: 0.2 mm. 25. Right ultimate leg, medially. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210614" httpUri="https://zenodo.org/record/13210614/files/figure.png" pageId="15" pageNumber="578">Fig. 20</figureCitation>
, fig.
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<bibRefCitation id="EFFBC5F0FFB4FFB08502FEB3DB18FEE9" author="Schileyko, A. A. &amp; Iorio, E. &amp; Coulis, M." box="[812,1053,326,352]" pageId="15" pageNumber="578" pagination="559 - 574" refId="ref30390" refString="Schileyko, A. A., Iorio, E. &amp; Coulis, M. (2018) A contribution to the knowledge of scolopendromorph centipedes of Martinique Island, with descriptions of two new species (Chilopoda: Scolopendromorpha). Zootaxa, 4486 (4), 559 - 574. https: // doi. org / 10.11646 / zootaxa. 4486.4.9" type="journal article" year="2018">
Schileyko
<emphasis id="B91E6413FFB4FFB0858AFEB2DCD0FEE9" box="[932,981,326,352]" italics="true" pageId="15" pageNumber="578">et al</emphasis>
. 2018
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) and bear 210 setae. Pretarsus of maxillae 2 apically pointed, with a ventral projection or without it. Dorsal brush of maxilla 2 as long as or slightly longer than the corresponding pretarsus.Anterior margin of forcipular coxosternite bilobed (
<figureCitation id="1351A484FFB4FFB082BEFE7BDBE7FE21" box="[1168,1250,398,424]" captionStart="FIGURES 2025" captionStartId="17.[151,264,1352,1376]" captionTargetBox="[152,1435,182,1328]" captionTargetId="figure-17@17.[151,1435,181,1328]" captionTargetPageId="17" captionText="FIGURES 2025. Cryptops (Trigonocryptops) amicitia n. sp., holotype, 212-P8G (MNHN). 20. Head, ventrally. Scale bar: 0.2 mm. 21. LBS 34, ventrally. Scale bar: 0.3 mm. 22. LBS 1314, ventrally. Scale bar: 0.4 mm. 23. Head + LBS 1, dorsally. Scale bar: 0.4 mm. 24. LBS 2021, ventrally. Scale bar: 0.2 mm. 25. Right ultimate leg, medially. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210614" httpUri="https://zenodo.org/record/13210614/files/figure.png" pageId="15" pageNumber="578">Fig. 20</figureCitation>
), in most species with long enlarged marginal setae; tarsungula long, overlapping each other by at least 1/3 of their length when adducted. Tergite 1 mainly with anterior transverse suture, often with (complete or incomplete) paramedian sutures (fig.
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Schileyko
<emphasis id="B91E6413FFB4FFB087A8FE0EDEB2FD9D" box="[390,439,506,532]" italics="true" pageId="15" pageNumber="578">et al</emphasis>
. 2018
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). Sternites with well-developed transverse thickening between coxae of legs (
<figureCitation id="1351A484FFB4FFB08344FE0FDFE5FDB1" captionStart="FIGURES 2025" captionStartId="17.[151,264,1352,1376]" captionTargetBox="[152,1435,182,1328]" captionTargetId="figure-17@17.[151,1435,181,1328]" captionTargetPageId="17" captionText="FIGURES 2025. Cryptops (Trigonocryptops) amicitia n. sp., holotype, 212-P8G (MNHN). 20. Head, ventrally. Scale bar: 0.2 mm. 21. LBS 34, ventrally. Scale bar: 0.3 mm. 22. LBS 1314, ventrally. Scale bar: 0.4 mm. 23. Head + LBS 1, dorsally. Scale bar: 0.4 mm. 24. LBS 2021, ventrally. Scale bar: 0.2 mm. 25. Right ultimate leg, medially. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210614" httpUri="https://zenodo.org/record/13210614/files/figure.png" pageId="15" pageNumber="578">Figs 21, 22</figureCitation>
); at least some anterior sternites with endosternites bordered by trigonal sutures of various configurations (K-, C- and X-shaped,
<figureCitation id="1351A484FFB4FFB0874EFDB7DEB4FDD5" box="[352,433,578,604]" captionStart="FIGURES 2025" captionStartId="17.[151,264,1352,1376]" captionTargetBox="[152,1435,182,1328]" captionTargetId="figure-17@17.[151,1435,181,1328]" captionTargetPageId="17" captionText="FIGURES 2025. Cryptops (Trigonocryptops) amicitia n. sp., holotype, 212-P8G (MNHN). 20. Head, ventrally. Scale bar: 0.2 mm. 21. LBS 34, ventrally. Scale bar: 0.3 mm. 22. LBS 1314, ventrally. Scale bar: 0.4 mm. 23. Head + LBS 1, dorsally. Scale bar: 0.4 mm. 24. LBS 2021, ventrally. Scale bar: 0.2 mm. 25. Right ultimate leg, medially. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210614" httpUri="https://zenodo.org/record/13210614/files/figure.png" pageId="15" pageNumber="578">Fig. 21</figureCitation>
, figs
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Schileyko
<emphasis id="B91E6413FFB4FFB084E4FDB6DDFFFDD5" box="[714,762,578,604]" italics="true" pageId="15" pageNumber="578">et al</emphasis>
. 2018
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). Anterior corners of the endosternite of some anterior sternites with lateral projections (fig.
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Schileyko
<emphasis id="B91E6413FFB4FFB084DDFD92DC21FD09" box="[755,804,614,640]" italics="true" pageId="15" pageNumber="578">et al</emphasis>
. 2018
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). In the most species katopleure bipartite. In most species femur of the ultimate legs with 1 well-developed saw tooth (in
<taxonomicName id="4C6AC382FFB4FFB08581FD7FDE39FD41" authority="Schileyko, Iorio &amp; Coulis 2018" authorityName="Schileyko, Iorio &amp; Coulis" authorityYear="2018" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="15" pageNumber="578" phylum="Arthropoda" rank="species" species="martinicensis" subGenus="Trigonocryptops">
<emphasis id="B91E6413FFB4FFB08581FD7FDB13FD2D" box="[943,1046,650,676]" italics="true" pageId="15" pageNumber="578">Cryptops</emphasis>
(
<emphasis id="B91E6413FFB4FFB08208FD7EDB3EFD2D" box="[1062,1083,651,676]" italics="true" pageId="15" pageNumber="578">T.</emphasis>
)
<emphasis id="B91E6413FFB4FFB08264FD7EDBE4FD2D" box="[1098,1249,651,676]" italics="true" pageId="15" pageNumber="578">martinicensis</emphasis>
<bibRefCitation id="EFFBC5F0FFB4FFB082C6FD7FDE39FD41" author="Schileyko, A. A. &amp; Iorio, E. &amp; Coulis, M." pageId="15" pageNumber="578" pagination="559 - 574" refId="ref30390" refString="Schileyko, A. A., Iorio, E. &amp; Coulis, M. (2018) A contribution to the knowledge of scolopendromorph centipedes of Martinique Island, with descriptions of two new species (Chilopoda: Scolopendromorpha). Zootaxa, 4486 (4), 559 - 574. https: // doi. org / 10.11646 / zootaxa. 4486.4.9" type="journal article" year="2018">Schileyko, Iorio &amp; Coulis 2018</bibRefCitation>
</taxonomicName>
this character may be a subject of intraspecific variability, see below).
</paragraph>
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<emphasis id="B91E6413FFB4FFB086E9FD27DEABFD62" bold="true" box="[199,430,721,748]" pageId="15" pageNumber="578">Number of species.</emphasis>
30 (29 species according to
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Schileyko
<emphasis id="B91E6413FFB4FFB08556FD26DCA9FD65" box="[888,940,722,748]" italics="true" pageId="15" pageNumber="578">et al</emphasis>
. 2020
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plus
<taxonomicName id="4C6AC382FFB4FFB08218FD27DBEDFD65" authorityName="Schileyko &amp; Iorio &amp; Coulis" authorityYear="2024" box="[1078,1256,722,748]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="15" pageNumber="599" phylum="Arthropoda" rank="species" species="amicitia" status="sp. nov." subGenus="Trigonocryptops">
<emphasis id="B91E6413FFB4FFB08218FD27DB4CFD65" box="[1078,1097,722,748]" italics="true" pageId="15" pageNumber="578">C</emphasis>
. (
<emphasis id="B91E6413FFB4FFB0824CFD26DB77FD65" box="[1122,1138,723,748]" italics="true" pageId="15" pageNumber="578">T</emphasis>
.)
<emphasis id="B91E6413FFB4FFB082A2FD26DBEDFD65" box="[1164,1256,723,748]" italics="true" pageId="15" pageNumber="578">amicitia</emphasis>
</taxonomicName>
<emphasis id="B91E6413FFB4FFB082DFFD27DA2CFD65" bold="true" box="[1265,1321,722,748]" pageId="15" pageNumber="578">
<taxonomicNameLabel id="A22DD968FFB4FFB082DFFD27DA2CFD65" box="[1265,1321,722,748]" pageId="15" pageNumber="578" rank="species">n.sp.</taxonomicNameLabel>
</emphasis>
; thus, the corresponding list of
<bibRefCitation id="EFFBC5F0FFB4FFB087A8FD03DD50FC99" author="Bonato, L. &amp; Chagas-Junior, A. &amp; Edgecombe, G. D. &amp; Lewis, J. G. E. &amp; Minelli, A. &amp; Pereira, L. A. &amp; Shelley, R. M. &amp; Stoev, P. &amp; Zapparoli, M." box="[390,597,758,784]" pageId="15" pageNumber="578" refId="ref26901" refString="Bonato, L., Chagas-Junior, A., Edgecombe, G. D., Lewis, J. G. E., Minelli, A., Pereira, L. A., Shelley, R. M., Stoev, P. &amp; Zapparoli, M. (2016) ChiloBase 2.0 - A World Catalogue of Centipedes (Chilopoda). Available from: https: // chilobase. biologia. unipd. it (accessed 18 February 2021)" type="url" year="2016">
Bonato
<emphasis id="B91E6413FFB4FFB087F0FD02DD0AFC99" box="[478,527,758,784]" italics="true" pageId="15" pageNumber="578">et al</emphasis>
. 2016
</bibRefCitation>
should be corrected).
</paragraph>
</subSubSection>
<subSubSection id="C370EB8AFFB4FFB086E9FCEFDE41FC29" pageId="15" pageNumber="578" type="description">
<paragraph id="8BD5B801FFB4FFB086E9FCEFDE41FC29" blockId="15.[151,1437,289,1252]" pageId="15" pageNumber="578">
<emphasis id="B91E6413FFB4FFB086E9FCEFDE16FCBD" bold="true" box="[199,275,794,820]" pageId="15" pageNumber="578">Range</emphasis>
(after
<bibRefCitation id="EFFBC5F0FFB4FFB08773FCEFDD62FCBD" author="Schileyko &amp; Ark, A. &amp; Vahtera, V. &amp; Edgecombe, G. D." box="[349,615,794,820]" pageId="15" pageNumber="578" refId="ref30456" refString="Schileyko, Ark, A., Vahtera, V. &amp; Edgecombe, G. D. (2020) An overview of the extant genera and subgenera of the order Scolopendromorpha (Chilopoda): a new identification key and updated diagnoses. Zootaxa, 4825 (1), 001 - 064. https: // doi. org / 10.11646 / zootaxa. 4825.1.1" type="journal volume" year="2020">
Schileyko
<emphasis id="B91E6413FFB4FFB087FDFCEEDD06FCBD" box="[467,515,794,820]" italics="true" pageId="15" pageNumber="578">et al</emphasis>
. 2020: 7
</bibRefCitation>
, corrected). Europe:
<collectingCountry id="F37DF891FFB4FFB0857EFCEFDC96FCBD" box="[848,915,794,820]" name="Spain" pageId="15" pageNumber="578">Spain</collectingCountry>
; Northern and Central Africa; South
<collectingCountry id="F37DF891FFB4FFB08302FCEFDA9CFCBD" box="[1324,1433,794,820]" name="China" pageId="15" pageNumber="578">East Asia</collectingCountry>
:
<collectingCountry id="F37DF891FFB4FFB086B9FCCBDFD4FCD1" box="[151,209,830,856]" name="India" pageId="15" pageNumber="578">India</collectingCountry>
(Nagpur),
<collectingCountry id="F37DF891FFB4FFB08761FCCBDEB4FCD1" box="[335,433,830,856]" name="Vietnam" pageId="15" pageNumber="578">Vietnam</collectingCountry>
; Australasia: West and North East
<collectingCountry id="F37DF891FFB4FFB08515FCCBDCA0FCD1" box="[827,933,830,856]" name="Australia" pageId="15" pageNumber="578">Australia</collectingCountry>
, Sumba,
<collectingCountry id="F37DF891FFB4FFB08220FCCBDB88FCD1" box="[1038,1165,830,856]" name="Timor-Leste" pageId="15" pageNumber="578">East Timor</collectingCountry>
, New
<collectingCountry id="F37DF891FFB4FFB082F9FCCBDA2EFCD1" box="[1239,1323,830,856]" name="Guinea-Bissau" pageId="15" pageNumber="578">Guinea</collectingCountry>
,
<collectingCountry id="F37DF891FFB4FFB08319FCCBDFECFCF5" name="Solomon Islands" pageId="15" pageNumber="578">Solomon Islands</collectingCountry>
,
<collectingCountry id="F37DF891FFB4FFB086D9FC97DEAFFCF5" box="[247,426,866,892]" name="New Caledonia" pageId="15" pageNumber="578">New Caledonia</collectingCountry>
,
<collectingCountry id="F37DF891FFB4FFB08796FC97DD56FCF5" box="[440,595,866,892]" name="New Zealand" pageId="15" pageNumber="578">New Zealand</collectingCountry>
,
<collectingCountry id="F37DF891FFB4FFB0844FFC97DD89FCF5" box="[609,652,866,892]" name="Fiji" pageId="15" pageNumber="578">Fiji</collectingCountry>
; Antilles:
<collectingCountry id="F37DF891FFB4FFB0852CFC97DC45FCF5" box="[770,832,866,892]" name="Cuba" pageId="15" pageNumber="578">Cuba</collectingCountry>
, Martinique; South America:
<collectingCountry id="F37DF891FFB4FFB082B8FC97DBC9FCF5" box="[1174,1228,866,892]" name="Peru" pageId="15" pageNumber="578">Peru</collectingCountry>
, Southeast
<collectingCountry id="F37DF891FFB4FFB0837FFC97DA9DFCF5" box="[1361,1432,866,892]" name="Brazil" pageId="15" pageNumber="578">Brazil</collectingCountry>
, East
<collectingCountry id="F37DF891FFB4FFB086E3FC73DE45FC29" box="[205,320,902,928]" name="Argentina" pageId="15" pageNumber="578">Argentina</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C370EB8AFFB4FFB086E9FC5FDC6FFB6D" pageId="15" pageNumber="578" type="discussion">
<paragraph id="8BD5B801FFB4FFB086E9FC5FDC6FFB6D" blockId="15.[151,1437,289,1252]" pageId="15" pageNumber="578">
<emphasis id="B91E6413FFB4FFB086E9FC5FDE3EFC4D" bold="true" box="[199,315,938,964]" pageId="15" pageNumber="578">Remarks.</emphasis>
<bibRefCitation id="EFFBC5F0FFB4FFB0876FFC5FDD70FC4D" author="Schileyko, A. A. &amp; Stoev, P. E." box="[321,629,938,964]" pageId="15" pageNumber="578" pagination="247 - 280" refId="ref30268" refString="Schileyko, A. A. &amp; Stoev, P. E. (2016) Scolopendromorpha of New Guinea and adjacent islands (Myriapoda, Chilopoda). Zootaxa, 4147 (3), 247 - 280. https: // doi. org / 10.11646 / zootaxa. 4147.3.3" type="journal article" year="2016">Schileyko and Stoev (2016)</bibRefCitation>
regarded the subgeneric status of
<taxonomicName id="4C6AC382FFB4FFB085C3FC5EDBA3FC4D" authorityName="Verhoeff" authorityYear="1906" box="[1005,1190,939,964]" pageId="15" pageNumber="599" rank="subGenus" subGenus="Trigonocryptops">
<emphasis id="B91E6413FFB4FFB085C3FC5EDBA3FC4D" box="[1005,1190,939,964]" italics="true" pageId="15" pageNumber="578">Trigonocryptops</emphasis>
</taxonomicName>
as questionable. They revised the number of species in
<taxonomicName id="4C6AC382FFB4FFB08436FC3ADDD4FC61" authorityName="Verhoeff" authorityYear="1906" box="[536,721,975,1000]" pageId="15" pageNumber="599" rank="subGenus" subGenus="Trigonocryptops">
<emphasis id="B91E6413FFB4FFB08436FC3ADDD4FC61" box="[536,721,975,1000]" italics="true" pageId="15" pageNumber="578">Trigonocryptops</emphasis>
</taxonomicName>
in the former sense, mentioning (p. 267) that “at the moment
<taxonomicName id="4C6AC382FFB4FFB086B9FC06DE55FB85" authorityName="Verhoeff" authorityYear="1906" box="[151,336,1011,1036]" pageId="15" pageNumber="599" rank="subGenus" subGenus="Trigonocryptops">
<emphasis id="B91E6413FFB4FFB086B9FC06DE55FB85" box="[151,336,1011,1036]" italics="true" pageId="15" pageNumber="578">Trigonocryptops</emphasis>
</taxonomicName>
should include 28 species”, thus including in this subgenus all species described under this name. However, providing below the new diagnosis for this subgenus they wrote: ”Thus, we believe that the recent concept of
<taxonomicName id="4C6AC382FFB4FFB0873AFBCEDEC8FBDD" authorityName="Verhoeff" authorityYear="1906" box="[276,461,1083,1108]" pageId="15" pageNumber="599" rank="subGenus" subGenus="Trigonocryptops">
<emphasis id="B91E6413FFB4FFB0873AFBCEDEC8FBDD" box="[276,461,1083,1108]" italics="true" pageId="15" pageNumber="578">Trigonocryptops</emphasis>
</taxonomicName>
must be reconsidered and only species having anterior sternites with complete trigonal sutures and clypeus with setose plate(s) should be assigned to
<taxonomicName id="4C6AC382FFB4FFB08549FBAADB25FBF1" authorityName="Verhoeff" authorityYear="1906" box="[871,1056,1119,1144]" pageId="15" pageNumber="599" rank="subGenus" subGenus="Trigonocryptops">
<emphasis id="B91E6413FFB4FFB08549FBAADB25FBF1" box="[871,1056,1119,1144]" italics="true" pageId="15" pageNumber="578">Trigonocryptops</emphasis>
</taxonomicName>
”. However a few former species of
<taxonomicName id="4C6AC382FFB4FFB08699FB76DE75FB15" authorityName="Verhoeff" authorityYear="1906" box="[183,368,1155,1180]" pageId="15" pageNumber="599" rank="subGenus" subGenus="Trigonocryptops">
<emphasis id="B91E6413FFB4FFB08699FB76DE75FB15" box="[183,368,1155,1180]" italics="true" pageId="15" pageNumber="578">Trigonocryptops</emphasis>
</taxonomicName>
may not meet one or both of these diagnostic criteria, thus being not related to this subgenus. Summing up, this large and significant taxon has to be revised (preferably using both morphological and molecular methods) and the corresponding species list should be corrected.
</paragraph>
</subSubSection>
</treatment>
</document>