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<document id="A2153F565E5AB079CBF82749C8F290FC" ID-CLB-Dataset="4901" ID-DOI="10.11646/zootaxa.412.1.1" ID-GBIF-Dataset="57fb6e48-c07a-46ed-91a7-150614ef477a" ID-ISSN="1175­5334" ID-Zenodo-Dep="5027840" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624552792396" checkinUser="felipe" docAuthor="Ibáñez-Bernal, Sergio" docDate="2004" docId="03AF1D19FFDFFF999B04822DFB7E6526" docLanguage="en" docName="zt00412.pdf" docOrigin="Zootaxa 412 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.412.1.1" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Feuerborniella veracruzana Ibanez-Bernal 2004, sp. nov." docType="treatment" docVersion="8" lastPageNumber="7" masterDocId="FF966561FFDEFF9F9A0C8331FFE46275" masterDocTitle="A new species of the genus Feuerborniella Vaillant, from Mexico (Diptera: Psychodidae)" masterLastPageNumber="8" masterPageNumber="1" pageNumber="2" updateTime="1699020402584" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="711BF2FCEC26D7F2284F75507BDB62F1">A new species of the genus Feuerborniella Vaillant, from Mexico (Diptera: Psychodidae)</mods:title>
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<mods:namePart id="F5B610DA1C82596ACA84EDA6C8652129">Ibáñez-Bernal, Sergio</mods:namePart>
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<mods:date id="81ACB4CA2083F96E954B643C04550D82">2004</mods:date>
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<mods:identifier id="F959FD6D93FE96B073552BAAE68577C5" type="DOI">10.11646/zootaxa.412.1.1</mods:identifier>
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<treatment id="03AF1D19FFDFFF999B04822DFB7E6526" ID-DOI="http://doi.org/10.5281/zenodo.5228236" ID-GBIF-Taxon="184643076" ID-Zenodo-Dep="5228236" LSID="urn:lsid:plazi:treatment:03AF1D19FFDFFF999B04822DFB7E6526" httpUri="http://treatment.plazi.org/id/03AF1D19FFDFFF999B04822DFB7E6526" lastPageId="6" lastPageNumber="7" pageId="1" pageNumber="2">
<subSubSection id="C31CFF84FFDFFF9E9B04822DFC666340" box="[264,898,283,310]" pageId="1" pageNumber="2" type="nomenclature">
<paragraph id="8BB9AC0FFFDFFF9E9B04822DFC666340" blockId="1.[264,898,283,345]" box="[264,898,283,310]" pageId="1" pageNumber="2">
<heading id="D0F11B63FFDFFF9E9B04822DFC666340" bold="true" box="[264,898,283,310]" fontSize="11" level="1" pageId="1" pageNumber="2" reason="1">
<emphasis id="B972701DFFDFFF9E9B04822DFC666340" bold="true" box="[264,898,283,310]" pageId="1" pageNumber="2">
<taxonomicName id="4C06D78CFFDFFF9E9B04822DFCFC6340" ID-CoL="6HTJQ" authority="Ibanez-Bernal" authorityName="Ibanez-Bernal" authorityYear="2004" box="[264,792,283,310]" class="Insecta" family="Psychodidae" genus="Feuerborniella" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="veracruzana" status="sp. nov.">
<emphasis id="B972701DFFDFFF9E9B04822DFDB86343" bold="true" box="[264,604,284,310]" italics="true" pageId="1" pageNumber="2">Feuerborniella veracruzana</emphasis>
Ibáñez­Bernal
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,
<taxonomicNameLabel id="A241CD66FFDFFF9E9929822AFC666340" box="[805,898,283,309]" pageId="1" pageNumber="2" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C31CFF84FFDFFF999B04820EFB7E6526" lastPageId="6" lastPageNumber="7" pageId="1" pageNumber="2" type="description">
<paragraph id="8BB9AC0FFFDFFF9E9B04820EFE73632C" blockId="1.[264,898,283,345]" box="[264,407,319,345]" pageId="1" pageNumber="2">
(
<figureCitation id="133DB08AFFDFFF9E9B1D820EFE6A632C" box="[273,398,319,345]" captionStart-0="FIGURE" captionStart-1="FIGURE 8–15" captionStartId-0="2.[264,370,1776,1800]" captionStartId-1="3.[264,367,1419,1443]" captionTargetBox-0="[394,1196,652,1732]" captionTargetBox-1="[408,1179,321,1409]" captionTargetId-0="figure-159@2.[386,1203,639,1760]" captionTargetId-1="figure-141@3.[399,1187,312,1415]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE. 1–7. Feuerborniella veracruzana sp. nov. Male. 1. Head, frontal view; 2. Head, posterior view; 3. base of antenna (scape, pedicel and first flagellomere, showing one ascoid); 4. Apical four flagellomeres; 5. Ventral view of aedeagus and one gonopod; 6. Dorsal view of epandrium, tergite 10 and one cercopod; 7. Male genitalia, lateral view." captionText-1="FIGURE 8–15. Feuerborniella veracruzana sp. nov. Female. 8. Lateral view of hind leg tarsomere 5; 9. Frontal view of labellum; 10. Wing; 11. Base of antenna (scape, pedicel, and first flagellomere, showing one ascoid); 12. Apical four flagellomeres; 13. Spermatheca complex; 14. Subgenital plate, ventral view; 15. Cerci, ventral view." figureDoi-0="http://doi.org/10.5281/zenodo.5027842" figureDoi-1="http://doi.org/10.5281/zenodo.5027844" httpUri-0="https://zenodo.org/record/5027842/files/figure.png" httpUri-1="https://zenodo.org/record/5027844/files/figure.png" pageId="1" pageNumber="2">Figs. 1–15</figureCitation>
)
</paragraph>
<paragraph id="8BB9AC0FFFDFFF9D9B0482BEFE216053" blockId="1.[264,1323,399,1870]" lastBlockId="2.[264,1323,284,550]" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2">
Male description (
<figureCitation id="133DB08AFFDFFF9E9BD282BEFDB463DC" box="[478,592,399,425]" captionStart="FIGURE" captionStartId="2.[264,370,1776,1800]" captionTargetBox="[394,1196,652,1732]" captionTargetId="figure-159@2.[386,1203,639,1760]" captionTargetPageId="2" captionText="FIGURE. 1–7. Feuerborniella veracruzana sp. nov. Male. 1. Head, frontal view; 2. Head, posterior view; 3. base of antenna (scape, pedicel and first flagellomere, showing one ascoid); 4. Apical four flagellomeres; 5. Ventral view of aedeagus and one gonopod; 6. Dorsal view of epandrium, tergite 10 and one cercopod; 7. Male genitalia, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5027842" httpUri="https://zenodo.org/record/5027842/files/figure.png" pageId="1" pageNumber="2">Figs. 1–7</figureCitation>
). Body vestiture apparently uniform pale gray, the integument yellowish brown. Head (
<figureCitation id="133DB08AFFDFFF9E98228286FD7363A4" box="[558,663,439,465]" captionStart="FIGURE" captionStartId="2.[264,370,1776,1800]" captionTargetBox="[394,1196,652,1732]" captionTargetId="figure-159@2.[386,1203,639,1760]" captionTargetPageId="2" captionText="FIGURE. 1–7. Feuerborniella veracruzana sp. nov. Male. 1. Head, frontal view; 2. Head, posterior view; 3. base of antenna (scape, pedicel and first flagellomere, showing one ascoid); 4. Apical four flagellomeres; 5. Ventral view of aedeagus and one gonopod; 6. Dorsal view of epandrium, tergite 10 and one cercopod; 7. Male genitalia, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5027842" httpUri="https://zenodo.org/record/5027842/files/figure.png" pageId="1" pageNumber="2">Fig. 1–2</figureCitation>
): Eyes separated by one or one and a half facet diameters; eye­bridge with four rows of facets; interocular suture somewhat arched and with a poorly visible dorsal middlemost spur; a row of five setae along the upper margin of each eye; the vertex rounded in outline, with a single median patch of scale alveoli leaving a small naked area in the upper central area which extends downward without reaching the level of the interocular suture; frontal patch of setae alveoli undivided, extending upward between the eyes to reach the level of the third facet (
<figureCitation id="133DB08AFFDFFF9E99908196FC0060B4" box="[924,996,679,705]" captionStart="FIGURE" captionStartId="2.[264,370,1776,1800]" captionTargetBox="[394,1196,652,1732]" captionTargetId="figure-159@2.[386,1203,639,1760]" captionTargetPageId="2" captionText="FIGURE. 1–7. Feuerborniella veracruzana sp. nov. Male. 1. Head, frontal view; 2. Head, posterior view; 3. base of antenna (scape, pedicel and first flagellomere, showing one ascoid); 4. Apical four flagellomeres; 5. Ventral view of aedeagus and one gonopod; 6. Dorsal view of epandrium, tergite 10 and one cercopod; 7. Male genitalia, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5027842" httpUri="https://zenodo.org/record/5027842/files/figure.png" pageId="1" pageNumber="2">Fig. 1</figureCitation>
). Ascendant tentorial arms straight, narrow, as long as 8x its width; posterior tentorial arms composed of a pair of laterally sclerotized, nearly parallel, concave flanges in posterior view; maxillary apodeme nearly straight; posterior margin of the eyes sigmoidal in outline, and a postoccipital plate well separated by a suture (
<figureCitation id="133DB08AFFDFFF9E98588076FD7A6114" box="[596,670,839,865]" captionStart="FIGURE" captionStartId="2.[264,370,1776,1800]" captionTargetBox="[394,1196,652,1732]" captionTargetId="figure-159@2.[386,1203,639,1760]" captionTargetPageId="2" captionText="FIGURE. 1–7. Feuerborniella veracruzana sp. nov. Male. 1. Head, frontal view; 2. Head, posterior view; 3. base of antenna (scape, pedicel and first flagellomere, showing one ascoid); 4. Apical four flagellomeres; 5. Ventral view of aedeagus and one gonopod; 6. Dorsal view of epandrium, tergite 10 and one cercopod; 7. Male genitalia, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5027842" httpUri="https://zenodo.org/record/5027842/files/figure.png" pageId="1" pageNumber="2">Fig. 2</figureCitation>
). Labellum bulbous with three or four long anteapical setae and two to four short spiniform setae in the internal margin near the apex, but without blunt teeth (as in
<figureCitation id="133DB08AFFDFFF9E980180A6FDB361C4" box="[525,599,919,945]" captionStart="FIGURE 8–15" captionStartId="3.[264,367,1419,1443]" captionTargetBox="[408,1179,321,1409]" captionTargetId="figure-141@3.[399,1187,312,1415]" captionTargetPageId="3" captionText="FIGURE 8–15. Feuerborniella veracruzana sp. nov. Female. 8. Lateral view of hind leg tarsomere 5; 9. Frontal view of labellum; 10. Wing; 11. Base of antenna (scape, pedicel, and first flagellomere, showing one ascoid); 12. Apical four flagellomeres; 13. Spermatheca complex; 14. Subgenital plate, ventral view; 15. Cerci, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5027844" httpUri="https://zenodo.org/record/5027844/files/figure.png" pageId="1" pageNumber="2">Fig. 9</figureCitation>
). Palpus as long as 0.36x antenna length, and about 1.1x as long as head height; proportion of palpomeres: 1.00: 1.42: 1.36: 1.46 (
<figureCitation id="133DB08AFFDFFF9E9E31808EFB6661AC" box="[1085,1154,959,985]" captionStart="FIGURE" captionStartId="2.[264,370,1776,1800]" captionTargetBox="[394,1196,652,1732]" captionTargetId="figure-159@2.[386,1203,639,1760]" captionTargetPageId="2" captionText="FIGURE. 1–7. Feuerborniella veracruzana sp. nov. Male. 1. Head, frontal view; 2. Head, posterior view; 3. base of antenna (scape, pedicel and first flagellomere, showing one ascoid); 4. Apical four flagellomeres; 5. Ventral view of aedeagus and one gonopod; 6. Dorsal view of epandrium, tergite 10 and one cercopod; 7. Male genitalia, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5027842" httpUri="https://zenodo.org/record/5027842/files/figure.png" pageId="1" pageNumber="2">Fig. 1</figureCitation>
). Antennae as long as 0.5x wing length, with 14 flagellomeres; scape barrel­shaped, longer than the spherical pedicel; flagellomere 1 with node 2x as long as its own internode; subsequent flagellomeres similar to 1, but nodes progressively decreasing in size and internodes increasing in length to such an extent that flagellomere 10 has the node equal in length to the internode; each internode parallel sided, uniformly sclerotized but with a median transverse sulcus (
<figureCitation id="133DB08AFFDFFF9E9BA0879EFE1166BC" box="[428,501,1199,1225]" captionStart="FIGURE" captionStartId="2.[264,370,1776,1800]" captionTargetBox="[394,1196,652,1732]" captionTargetId="figure-159@2.[386,1203,639,1760]" captionTargetPageId="2" captionText="FIGURE. 1–7. Feuerborniella veracruzana sp. nov. Male. 1. Head, frontal view; 2. Head, posterior view; 3. base of antenna (scape, pedicel and first flagellomere, showing one ascoid); 4. Apical four flagellomeres; 5. Ventral view of aedeagus and one gonopod; 6. Dorsal view of epandrium, tergite 10 and one cercopod; 7. Male genitalia, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5027842" httpUri="https://zenodo.org/record/5027842/files/figure.png" pageId="1" pageNumber="2">Fig. 3</figureCitation>
); flagellomere 11 without internode; flagellomeres 12–14 small and globular, not fused, 12–13 nearly sessile but in a short broad pedunculus, 14 with a long pedunculus; a pair of “Y”­shaped ascoids present on flagellomeres 1–11, each with two large anterior branches and one shorter posterior branch; the last three flagellomeres only with the short spiniform sensilla (
<figureCitation id="133DB08AFFDFFF9E989C867EFD3D671C" box="[656,729,1359,1385]" captionStart="FIGURE" captionStartId="2.[264,370,1776,1800]" captionTargetBox="[394,1196,652,1732]" captionTargetId="figure-159@2.[386,1203,639,1760]" captionTargetPageId="2" captionText="FIGURE. 1–7. Feuerborniella veracruzana sp. nov. Male. 1. Head, frontal view; 2. Head, posterior view; 3. base of antenna (scape, pedicel and first flagellomere, showing one ascoid); 4. Apical four flagellomeres; 5. Ventral view of aedeagus and one gonopod; 6. Dorsal view of epandrium, tergite 10 and one cercopod; 7. Male genitalia, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5027842" httpUri="https://zenodo.org/record/5027842/files/figure.png" pageId="1" pageNumber="2">Fig. 4</figureCitation>
). Thorax: Wing length about 2.5x breadth; Sc ending at the level of the base of R
<subScript id="1782AE4AFFDFFF9E988886B2FD6A67E1" attach="right" box="[644,654,1411,1428]" fontSize="7" pageId="1" pageNumber="2">1</subScript>
; R
<subScript id="1782AE4AFFDFFF9E98BE86B2FD5867E1" attach="left" box="[690,700,1411,1428]" fontSize="7" pageId="1" pageNumber="2">5</subScript>
ending at wing tip; base of M
<subScript id="1782AE4AFFDFFF9E9E2B86B2FBD567E1" attach="left" box="[1063,1073,1411,1428]" fontSize="7" pageId="1" pageNumber="2">2</subScript>
not connected; wing membrane without hairs; proportion of R
<subScript id="1782AE4AFFDFFF9E98E7869FFCED67CA" attach="left" box="[747,777,1454,1471]" fontSize="7" pageId="1" pageNumber="2">2+3</subScript>
:R
<subScript id="1782AE4AFFDFFF9E9929869FFCCB67CA" attach="left" box="[805,815,1454,1471]" fontSize="7" pageId="1" pageNumber="2">3</subScript>
= 2.9:2.8; M
<subScript id="1782AE4AFFDFFF9E99CE869FFC0467CA" attach="left" box="[962,992,1454,1471]" fontSize="7" pageId="1" pageNumber="2">1+2</subScript>
:M
<subScript id="1782AE4AFFDFFF9E9E0E869FFBE867CA" attach="left" box="[1026,1036,1454,1471]" fontSize="7" pageId="1" pageNumber="2">2</subScript>
= 1.5:4.0. Genitalia with elaborate and symmetric aedeagus (
<figureCitation id="133DB08AFFDFFF9E98BA86FDFCCF6793" box="[694,811,1484,1510]" captionStart="FIGURE" captionStartId="2.[264,370,1776,1800]" captionTargetBox="[394,1196,652,1732]" captionTargetId="figure-159@2.[386,1203,639,1760]" captionTargetPageId="2" captionText="FIGURE. 1–7. Feuerborniella veracruzana sp. nov. Male. 1. Head, frontal view; 2. Head, posterior view; 3. base of antenna (scape, pedicel and first flagellomere, showing one ascoid); 4. Apical four flagellomeres; 5. Ventral view of aedeagus and one gonopod; 6. Dorsal view of epandrium, tergite 10 and one cercopod; 7. Male genitalia, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5027842" httpUri="https://zenodo.org/record/5027842/files/figure.png" pageId="1" pageNumber="2">Figs. 5–7</figureCitation>
). Cercopod with setae alveoli covering its surface, as long as 1.7x the lateral length of epandrium, bearing a clavate retinaculum which is as long as 0.3x the cercopod length; epandrium strongly bilobed apically, a little longer than wide, with two well separated lateral patches that do not reach the proximal portion and with a small pseudospiracular opening (
<figureCitation id="133DB08AFFDFFF9E998E855DFC2A64F3" box="[898,974,1644,1670]" captionStart="FIGURE" captionStartId="2.[264,370,1776,1800]" captionTargetBox="[394,1196,652,1732]" captionTargetId="figure-159@2.[386,1203,639,1760]" captionTargetPageId="2" captionText="FIGURE. 1–7. Feuerborniella veracruzana sp. nov. Male. 1. Head, frontal view; 2. Head, posterior view; 3. base of antenna (scape, pedicel and first flagellomere, showing one ascoid); 4. Apical four flagellomeres; 5. Ventral view of aedeagus and one gonopod; 6. Dorsal view of epandrium, tergite 10 and one cercopod; 7. Male genitalia, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5027842" httpUri="https://zenodo.org/record/5027842/files/figure.png" pageId="1" pageNumber="2">Fig. 6</figureCitation>
). Gonocoxite subcylindrical, with an external lateral patch of setae alveoli, 0.64x as long as the gonostylus; gonocoxal apodemes broad and plate­like meeting beneath the aedeagal apodeme, only slightly expanded, with the internal margin nearly straight and convergent posteriorly. Gonostylus hirsute, straight, but ending in a sharp, entally curved apex, and with an external basal patch of setae alveoli and an irregular row of small setae in the internal apical margin (
<figureCitation id="133DB08AFFDFFF9E9EF78405FACF653B" box="[1275,1323,1844,1870]" captionStart="FIGURE" captionStartId="2.[264,370,1776,1800]" captionTargetBox="[394,1196,652,1732]" captionTargetId="figure-159@2.[386,1203,639,1760]" captionTargetPageId="2" captionText="FIGURE. 1–7. Feuerborniella veracruzana sp. nov. Male. 1. Head, frontal view; 2. Head, posterior view; 3. base of antenna (scape, pedicel and first flagellomere, showing one ascoid); 4. Apical four flagellomeres; 5. Ventral view of aedeagus and one gonopod; 6. Dorsal view of epandrium, tergite 10 and one cercopod; 7. Male genitalia, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5027842" httpUri="https://zenodo.org/record/5027842/files/figure.png" pageId="1" pageNumber="2">Fig.</figureCitation>
5). Aedeagus fusiform, with the apex acutely rounded in dorso­ventral view (
<figureCitation id="133DB08AFFDCFF9D9EC3822DFAFF6343" box="[1231,1307,284,310]" captionStart="FIGURE" captionStartId="2.[264,370,1776,1800]" captionTargetBox="[394,1196,652,1732]" captionTargetId="figure-159@2.[386,1203,639,1760]" captionTargetPageId="2" captionText="FIGURE. 1–7. Feuerborniella veracruzana sp. nov. Male. 1. Head, frontal view; 2. Head, posterior view; 3. base of antenna (scape, pedicel and first flagellomere, showing one ascoid); 4. Apical four flagellomeres; 5. Ventral view of aedeagus and one gonopod; 6. Dorsal view of epandrium, tergite 10 and one cercopod; 7. Male genitalia, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5027842" httpUri="https://zenodo.org/record/5027842/files/figure.png" pageId="2" pageNumber="3">Fig. 5</figureCitation>
), canoe­like in profile (
<figureCitation id="133DB08AFFDCFF9D980A8275FDA9632B" box="[518,589,324,350]" captionStart="FIGURE" captionStartId="2.[264,370,1776,1800]" captionTargetBox="[394,1196,652,1732]" captionTargetId="figure-159@2.[386,1203,639,1760]" captionTargetPageId="2" captionText="FIGURE. 1–7. Feuerborniella veracruzana sp. nov. Male. 1. Head, frontal view; 2. Head, posterior view; 3. base of antenna (scape, pedicel and first flagellomere, showing one ascoid); 4. Apical four flagellomeres; 5. Ventral view of aedeagus and one gonopod; 6. Dorsal view of epandrium, tergite 10 and one cercopod; 7. Male genitalia, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5027842" httpUri="https://zenodo.org/record/5027842/files/figure.png" pageId="2" pageNumber="3">Fig. 7</figureCitation>
), flanked by one pair of relatively long and nearly straight lateral shafts, except for a slightly ectally directed curvature of the apical portion; apex of lateral shafts reaching the apical fourth of aedeagus (
<figureCitation id="133DB08AFFDCFF9D996782A5FC5163DB" box="[875,949,404,430]" captionStart="FIGURE" captionStartId="2.[264,370,1776,1800]" captionTargetBox="[394,1196,652,1732]" captionTargetId="figure-159@2.[386,1203,639,1760]" captionTargetPageId="2" captionText="FIGURE. 1–7. Feuerborniella veracruzana sp. nov. Male. 1. Head, frontal view; 2. Head, posterior view; 3. base of antenna (scape, pedicel and first flagellomere, showing one ascoid); 4. Apical four flagellomeres; 5. Ventral view of aedeagus and one gonopod; 6. Dorsal view of epandrium, tergite 10 and one cercopod; 7. Male genitalia, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5027842" httpUri="https://zenodo.org/record/5027842/files/figure.png" pageId="2" pageNumber="3">Fig. 5</figureCitation>
); aedeagal apodeme broad and truncated; aedeagal sheath trapezoidal in dorso­ventral view. The hypandrium is slender and diadem­like (
<figureCitation id="133DB08AFFDCFF9D9BDB82D5FDC4638B" box="[471,544,484,510]" captionStart="FIGURE" captionStartId="2.[264,370,1776,1800]" captionTargetBox="[394,1196,652,1732]" captionTargetId="figure-159@2.[386,1203,639,1760]" captionTargetPageId="2" captionText="FIGURE. 1–7. Feuerborniella veracruzana sp. nov. Male. 1. Head, frontal view; 2. Head, posterior view; 3. base of antenna (scape, pedicel and first flagellomere, showing one ascoid); 4. Apical four flagellomeres; 5. Ventral view of aedeagus and one gonopod; 6. Dorsal view of epandrium, tergite 10 and one cercopod; 7. Male genitalia, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5027842" httpUri="https://zenodo.org/record/5027842/files/figure.png" pageId="2" pageNumber="3">Fig. 5</figureCitation>
). Tergite 10 is globular with a long conical median fleshy projection (
<figureCitation id="133DB08AFFDCFF9D9B4A813DFE5C6053" box="[326,440,524,550]" captionStart="FIGURE" captionStartId="2.[264,370,1776,1800]" captionTargetBox="[394,1196,652,1732]" captionTargetId="figure-159@2.[386,1203,639,1760]" captionTargetPageId="2" captionText="FIGURE. 1–7. Feuerborniella veracruzana sp. nov. Male. 1. Head, frontal view; 2. Head, posterior view; 3. base of antenna (scape, pedicel and first flagellomere, showing one ascoid); 4. Apical four flagellomeres; 5. Ventral view of aedeagus and one gonopod; 6. Dorsal view of epandrium, tergite 10 and one cercopod; 7. Male genitalia, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5027842" httpUri="https://zenodo.org/record/5027842/files/figure.png" pageId="2" pageNumber="3">Figs. 6–7</figureCitation>
).
</paragraph>
<caption id="DF79FC87FFDCFF9D9B0485C1FCD3651A" ID-DOI="http://doi.org/10.5281/zenodo.5027842" ID-Zenodo-Dep="5027842" httpUri="https://zenodo.org/record/5027842/files/figure.png" pageId="2" pageNumber="3" startId="2.[264,370,1776,1800]" targetBox="[394,1196,652,1732]" targetPageId="2">
<paragraph id="8BB9AC0FFFDCFF9D9B0485C1FCD3651A" blockId="2.[264,1323,1776,1903]" pageId="2" pageNumber="3">
<emphasis id="B972701DFFDCFF9D9B0485C1FE4F657D" bold="true" box="[264,427,1776,1800]" pageId="2" pageNumber="3">FIGURE. 1–7.</emphasis>
<taxonomicName id="4C06D78CFFDCFF9D9BBD85C0FD3A657D" authority="Ibáñez-Bernal, 2004" authorityName="Ibáñez-Bernal" authorityYear="2004" box="[433,734,1777,1800]" class="Insecta" family="Psychodidae" genus="Feuerborniella" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="veracruzana" status="sp. nov.">
<emphasis id="B972701DFFDCFF9D9BBD85C0FD3A657D" box="[433,734,1777,1800]" italics="true" pageId="2" pageNumber="3">Feuerborniella veracruzana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A241CD66FFDCFF9D98E885C1FCD26572" box="[740,822,1776,1799]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
Male. 1. Head, frontal view; 2. Head, posterior view; 3. base of antenna (scape, pedicel and first flagellomere, showing one ascoid); 4. Apical four flagellomeres; 5. Ventral view of aedeagus and one gonopod; 6. Dorsal view of epandrium, tergite 10 and one cercopod; 7. Male genitalia, lateral view.
</paragraph>
</caption>
<caption id="DF79FC87FFDDFF9C9B0486BAFD31647E" ID-DOI="http://doi.org/10.5281/zenodo.5027844" ID-Zenodo-Dep="5027844" httpUri="https://zenodo.org/record/5027844/files/figure.png" pageId="3" pageNumber="4" startId="3.[264,367,1419,1443]" targetBox="[408,1179,321,1409]" targetPageId="3">
<paragraph id="8BB9AC0FFFDDFF9C9B0486BAFD31647E" blockId="3.[264,1325,1419,1547]" pageId="3" pageNumber="4">
<emphasis id="B972701DFFDDFF9C9B0486BAFE5667D6" bold="true" box="[264,434,1419,1443]" pageId="3" pageNumber="4">FIGURE 8–15.</emphasis>
<taxonomicName id="4C06D78CFFDDFF9C9BB586BDFD0267D6" authority="Ibáñez-Bernal, 2004" authorityName="Ibáñez-Bernal" authorityYear="2004" box="[441,742,1420,1443]" class="Insecta" family="Psychodidae" genus="Feuerborniella" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="veracruzana" status="sp. nov.">
<emphasis id="B972701DFFDDFF9C9BB586BDFD0267D6" box="[441,742,1420,1443]" italics="true" pageId="3" pageNumber="4">Feuerborniella veracruzana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A241CD66FFDDFF9C98E086BDFCDB67D6" box="[748,831,1420,1443]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
Female. 8. Lateral view of hind leg tarsomere 5; 9. Frontal view of labellum; 10. Wing; 11. Base of antenna (scape, pedicel, and first flagellomere, showing one ascoid); 12. Apical four flagellomeres; 13. Spermatheca complex; 14. Subgenital plate, ventral view; 15. Cerci, ventral view.
</paragraph>
</caption>
<paragraph id="8BB9AC0FFFDDFF9C9B348556FC7C6554" blockId="3.[264,1322,1639,1905]" pageId="3" pageNumber="4">Measurements. Head height: 0.315 (0.31–0.32) n=2; proboscis length: 0.08; palpus length: 0.35 (0.35–0.37) n=4; antenna length: 0.98 (0.94–1.03) n=5; wing length: 1.91 (1.71–2.25) n=5; wing width: 0.77 (0.68–0.76) n=5; cercopod length: 0.28 (0.27–0.30) n=5; gonostylus length: 0.13 (0.13–0.14) n=5; gonocoxite length: 0.08 (0.08–0.09) n=5; aedeagus length (from apodeme): 0.19 (0.17–0.21) n=5.</paragraph>
<paragraph id="8BB9AC0FFFDDFF9C9B34841EFB916504" blockId="3.[264,1322,1639,1905]" pageId="3" pageNumber="4">
Female description (
<figureCitation id="133DB08AFFDDFF9C982B841EFD43653C" box="[551,679,1839,1865]" captionStart="FIGURE 8–15" captionStartId="3.[264,367,1419,1443]" captionTargetBox="[408,1179,321,1409]" captionTargetId="figure-141@3.[399,1187,312,1415]" captionTargetPageId="3" captionText="FIGURE 8–15. Feuerborniella veracruzana sp. nov. Female. 8. Lateral view of hind leg tarsomere 5; 9. Frontal view of labellum; 10. Wing; 11. Base of antenna (scape, pedicel, and first flagellomere, showing one ascoid); 12. Apical four flagellomeres; 13. Spermatheca complex; 14. Subgenital plate, ventral view; 15. Cerci, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5027844" httpUri="https://zenodo.org/record/5027844/files/figure.png" pageId="3" pageNumber="4">Figs. 8–15</figureCitation>
). Similar to the male except for the following features: palpus 0.36x as long as antenna; proportion of palpomeres: 1.0:1.2:1.2:1.3.
</paragraph>
<paragraph id="8BB9AC0FFFDAFF9B9B34822DFB486053" blockId="4.[264,1325,284,1873]" pageId="4" pageNumber="5">
Antennae (
<figureCitation id="133DB08AFFDAFF9B9BBB822DFDA66343" box="[439,578,284,310]" captionStart="FIGURE 8–15" captionStartId="3.[264,367,1419,1443]" captionTargetBox="[408,1179,321,1409]" captionTargetId="figure-141@3.[399,1187,312,1415]" captionTargetPageId="3" captionText="FIGURE 8–15. Feuerborniella veracruzana sp. nov. Female. 8. Lateral view of hind leg tarsomere 5; 9. Frontal view of labellum; 10. Wing; 11. Base of antenna (scape, pedicel, and first flagellomere, showing one ascoid); 12. Apical four flagellomeres; 13. Spermatheca complex; 14. Subgenital plate, ventral view; 15. Cerci, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5027844" httpUri="https://zenodo.org/record/5027844/files/figure.png" pageId="4" pageNumber="5">Figs. 11–12</figureCitation>
) 0.46x as long as wing; flagellomeres 12–14 each with a longer petiole (
<figureCitation id="133DB08AFFDAFF9B9B798275FE3E632B" box="[373,474,324,350]" captionStart="FIGURE 8–15" captionStartId="3.[264,367,1419,1443]" captionTargetBox="[408,1179,321,1409]" captionTargetId="figure-141@3.[399,1187,312,1415]" captionTargetPageId="3" captionText="FIGURE 8–15. Feuerborniella veracruzana sp. nov. Female. 8. Lateral view of hind leg tarsomere 5; 9. Frontal view of labellum; 10. Wing; 11. Base of antenna (scape, pedicel, and first flagellomere, showing one ascoid); 12. Apical four flagellomeres; 13. Spermatheca complex; 14. Subgenital plate, ventral view; 15. Cerci, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5027844" httpUri="https://zenodo.org/record/5027844/files/figure.png" pageId="4" pageNumber="5">Fig. 12</figureCitation>
). Wing 2.7x as long as width; proportion of R2+3:R3=1.0:1.0; M1+2:M2=1.0:2.2 (
<figureCitation id="133DB08AFFDAFF9B9BFB825DFDB763F3" box="[503,595,364,390]" captionStart="FIGURE 8–15" captionStartId="3.[264,367,1419,1443]" captionTargetBox="[408,1179,321,1409]" captionTargetId="figure-141@3.[399,1187,312,1415]" captionTargetPageId="3" captionText="FIGURE 8–15. Feuerborniella veracruzana sp. nov. Female. 8. Lateral view of hind leg tarsomere 5; 9. Frontal view of labellum; 10. Wing; 11. Base of antenna (scape, pedicel, and first flagellomere, showing one ascoid); 12. Apical four flagellomeres; 13. Spermatheca complex; 14. Subgenital plate, ventral view; 15. Cerci, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5027844" httpUri="https://zenodo.org/record/5027844/files/figure.png" pageId="4" pageNumber="5">Fig. 10</figureCitation>
). Genitalia (
<figureCitation id="133DB08AFFDAFF9B98E0825DFC6563F3" box="[748,897,364,390]" captionStart="FIGURE 8–15" captionStartId="3.[264,367,1419,1443]" captionTargetBox="[408,1179,321,1409]" captionTargetId="figure-141@3.[399,1187,312,1415]" captionTargetPageId="3" captionText="FIGURE 8–15. Feuerborniella veracruzana sp. nov. Female. 8. Lateral view of hind leg tarsomere 5; 9. Frontal view of labellum; 10. Wing; 11. Base of antenna (scape, pedicel, and first flagellomere, showing one ascoid); 12. Apical four flagellomeres; 13. Spermatheca complex; 14. Subgenital plate, ventral view; 15. Cerci, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5027844" httpUri="https://zenodo.org/record/5027844/files/figure.png" pageId="4" pageNumber="5">Figs. 13–15</figureCitation>
): subgenital plate 0.67x as long as broad; distal lobes “X”­shaped, each with one sensory setae originating near the basal margin of the lobes at the level of the triangular genital sac (
<figureCitation id="133DB08AFFDAFF9B99A6828DFBE663A3" box="[938,1026,444,470]" captionStart="FIGURE 8–15" captionStartId="3.[264,367,1419,1443]" captionTargetBox="[408,1179,321,1409]" captionTargetId="figure-141@3.[399,1187,312,1415]" captionTargetPageId="3" captionText="FIGURE 8–15. Feuerborniella veracruzana sp. nov. Female. 8. Lateral view of hind leg tarsomere 5; 9. Frontal view of labellum; 10. Wing; 11. Base of antenna (scape, pedicel, and first flagellomere, showing one ascoid); 12. Apical four flagellomeres; 13. Spermatheca complex; 14. Subgenital plate, ventral view; 15. Cerci, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5027844" httpUri="https://zenodo.org/record/5027844/files/figure.png" pageId="4" pageNumber="5">Fig. 14</figureCitation>
). Spermathecae complex as figured (
<figureCitation id="133DB08AFFDAFF9B9B9F82D5FE09638B" box="[403,493,484,510]" captionStart="FIGURE 8–15" captionStartId="3.[264,367,1419,1443]" captionTargetBox="[408,1179,321,1409]" captionTargetId="figure-141@3.[399,1187,312,1415]" captionTargetPageId="3" captionText="FIGURE 8–15. Feuerborniella veracruzana sp. nov. Female. 8. Lateral view of hind leg tarsomere 5; 9. Frontal view of labellum; 10. Wing; 11. Base of antenna (scape, pedicel, and first flagellomere, showing one ascoid); 12. Apical four flagellomeres; 13. Spermatheca complex; 14. Subgenital plate, ventral view; 15. Cerci, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5027844" httpUri="https://zenodo.org/record/5027844/files/figure.png" pageId="4" pageNumber="5">Fig. 13</figureCitation>
); corpus spermathecae sculptured; median arms blade­like; posterior arms slightly divergent posteriorly; cercus long and gradually tapered (
<figureCitation id="133DB08AFFDAFF9B9E4B813DFB7A6053" box="[1095,1182,524,550]" captionStart="FIGURE 8–15" captionStartId="3.[264,367,1419,1443]" captionTargetBox="[408,1179,321,1409]" captionTargetId="figure-141@3.[399,1187,312,1415]" captionTargetPageId="3" captionText="FIGURE 8–15. Feuerborniella veracruzana sp. nov. Female. 8. Lateral view of hind leg tarsomere 5; 9. Frontal view of labellum; 10. Wing; 11. Base of antenna (scape, pedicel, and first flagellomere, showing one ascoid); 12. Apical four flagellomeres; 13. Spermatheca complex; 14. Subgenital plate, ventral view; 15. Cerci, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5027844" httpUri="https://zenodo.org/record/5027844/files/figure.png" pageId="4" pageNumber="5">Fig. 15</figureCitation>
).
</paragraph>
<paragraph id="8BB9AC0FFFDAFF9B9B348105FBBC60EB" blockId="4.[264,1325,284,1873]" pageId="4" pageNumber="5">Measurements. Head height: 0.315 (0.31–0.32) n=2; proboscis length: 0.08, n=2; palpus length: 0.36 (0.35–0.37) n=2; antenna length: 1.0; wing length: 1.99 (1.89–2.09) n=2; wing width: 0745 (0.73–0.76) n=2; cercus length: 0.24 (0.21–0.27) n=2.</paragraph>
<paragraph id="8BB9AC0FFFDAFF9B9B34819DFD9A60B3" blockId="4.[264,1325,284,1873]" box="[312,638,684,710]" pageId="4" pageNumber="5">Immature stages: unknown.</paragraph>
<paragraph id="8BB9AC0FFFDAFF9B9B3481E5FBC4609B" blockId="4.[264,1325,284,1873]" box="[312,1056,724,750]" pageId="4" pageNumber="5">
Etymology:
<taxonomicName id="4C06D78CFFDAFF9B9BC581E5FDBF6098" authorityName="Ibáñez-Bernal" authorityYear="2004" box="[457,603,724,749]" class="Insecta" family="Psychodidae" genus="Feuerborniella" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="veracruzana">
<emphasis id="B972701DFFDAFF9B9BC581E5FDBF6098" box="[457,603,724,749]" italics="true" pageId="4" pageNumber="5">veracruzana</emphasis>
</taxonomicName>
, pertaining to
<collectingRegion id="49C262EDFFDAFF9B990B81E5FC57609B" box="[775,947,724,750]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="4" pageNumber="5">Veracruz State</collectingRegion>
,
<collectingCountry id="F311EC9FFFDAFF9B99B381E5FBF8609B" box="[959,1052,724,750]" name="Mexico" pageId="4" pageNumber="5">Mexico</collectingCountry>
.
</paragraph>
<paragraph id="8BB9AC0FFFDAFF9B9B3481CDFE976113" blockId="4.[264,1325,284,1873]" pageId="4" pageNumber="5">
Type locality.
<collectingCountry id="F311EC9FFFDAFF9B9BE081CDFDAC6163" box="[492,584,764,790]" name="Mexico" pageId="4" pageNumber="5">Mexico</collectingCountry>
,
<collectingRegion id="49C262EDFFDAFF9B985781CDFD226163" box="[603,710,764,790]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="4" pageNumber="5">Veracruz</collectingRegion>
, Tlilapan, Cueva Matzinga. The specimens were obtained by aspirating resting adults from the cave walls, somewhat beyond the cave entrance.
</paragraph>
<paragraph id="8BB9AC0FFFDAFF9B9B348045FB066623" blockId="4.[264,1325,284,1873]" pageId="4" pageNumber="5">
<materialsCitation id="3B6EA652FFDAFF9B9B348045FC7061AB" ID-GBIF-Occurrence="3336545302" collectingDate="1989-10-25" collectorName="F. Lledias &amp; All" country="Mexico" location="Cueva Matzinga" municipality="Tlilapan" pageId="4" pageNumber="5" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="holotype">
<typeStatus id="54BD12ADFFDAFF9B9B348045FE4161FB" box="[312,421,884,910]" pageId="4" pageNumber="5" type="holotype">Holotype</typeStatus>
male,
<typeStatus id="54BD12ADFFDAFF9B98098045FD8761FB" box="[517,611,884,910]" pageId="4" pageNumber="5" type="allotype">allotype</typeStatus>
female, and
<specimenCount id="9D006786FFDAFF9B99198045FC4761FB" box="[789,931,884,910]" pageId="4" pageNumber="5" type="male">three males</specimenCount>
and
<specimenCount id="9D006786FFDAFF9B99E28045FB9C61FB" box="[1006,1144,884,910]" pageId="4" pageNumber="5" type="female">one female</specimenCount>
<typeStatus id="54BD12ADFFDAFF9B9E848045FB1C61FB" box="[1160,1272,884,910]" pageId="4" pageNumber="5" type="paratype">paratypes</typeStatus>
(as labeled):
<collectingCountry id="F311EC9FFFDAFF9B9B7780ADFE3C61C3" box="[379,472,924,950]" name="Mexico" pageId="4" pageNumber="5">Mexico</collectingCountry>
,
<collectingRegion id="49C262EDFFDAFF9B9BE480ADFDB761C3" box="[488,595,924,950]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="4" pageNumber="5">Veracruz</collectingRegion>
,
<collectingMunicipality id="6BDD3675FFDAFF9B986880ADFD2D61C3" box="[612,713,924,950]" pageId="4" pageNumber="5">Tlilapan</collectingMunicipality>
,
<location id="8ED9FAD4FFDAFF9B98D680ADFC4761C3" LSID="urn:lsid:plazi:treatment:03AF1D19FFDFFF999B04822DFB7E6526:8ED9FAD4FFDAFF9B98D680ADFC4761C3" box="[730,931,924,950]" country="Mexico" municipality="Tlilapan" name="Cueva Matzinga" pageId="4" pageNumber="5" stateProvince="Veracruz de Ignacio de la Llave">Cueva Matzinga</location>
,
<date id="FFB88ACFFFDAFF9B99B880ADFBD461C3" box="[948,1072,924,950]" pageId="4" pageNumber="5" value="1989-10-25">
<collectingDate id="EFFC7327FFDAFF9B99B880ADFBD461C3" box="[948,1072,924,950]" pageId="4" pageNumber="5" value="1989-10-25">25­x­1989</collectingDate>
</date>
,
<collectorName id="26F3C9D9FFDAFF9B9E4E80ADFB5961C3" box="[1090,1213,924,950]" pageId="4" pageNumber="5">F. Lledías</collectorName>
, col.
<collectorName id="26F3C9D9FFDAFF9B9F0A80ADFACF61C3" box="[1286,1323,924,950]" pageId="4" pageNumber="5">All</collectorName>
specimens are mounted on slides with
<collectingCountry id="F311EC9FFFDAFF9B98D080F5FCD061AB" box="[732,820,964,990]" name="Canada" pageId="4" pageNumber="5">Canada</collectingCountry>
balsam
</materialsCitation>
.
<materialsCitation id="3B6EA652FFDAFF9B99AD80F5FD4B665B" ID-GBIF-Occurrence="3336545301" collectionCode="IEXA" collectorName="Instituto de Ecologia, A. C." country="Mexico" location="Instituto de Ecologia" municipality="Xalapa" pageId="4" pageNumber="5" specimenCount="3" specimenCount-male="1" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="holotype">
<typeStatus id="54BD12ADFFDAFF9B99AD80F5FBF561AB" box="[929,1041,964,990]" pageId="4" pageNumber="5" type="holotype">Holotype</typeStatus>
,
<typeStatus id="54BD12ADFFDAFF9B9E2C80F5FB9A61AB" box="[1056,1150,964,990]" pageId="4" pageNumber="5" type="allotype">allotype</typeStatus>
and
<specimenCount id="9D006786FFDAFF9B9EB080F5FACF61AB" box="[1212,1323,964,990]" pageId="4" pageNumber="5" type="male">two male</specimenCount>
<typeStatus id="54BD12ADFFDAFF9B9B0480DDFE9C6673" box="[264,376,1004,1030]" pageId="4" pageNumber="5" type="paratype">paratypes</typeStatus>
are deposited in the entomological collection at the
<collectorName id="26F3C9D9FFDAFF9B99E780DDFACF6673" box="[1003,1323,1004,1030]" pageId="4" pageNumber="5">
<location id="8ED9FAD4FFDAFF9B99E780DDFB026673" LSID="urn:lsid:plazi:treatment:03AF1D19FFDFFF999B04822DFB7E6526:8ED9FAD4FFDAFF9B99E780DDFB026673" box="[1003,1254,1004,1030]" country="Mexico" municipality="Xalapa" name="Instituto de Ecologia" pageId="4" pageNumber="5" stateProvince="Veracruz de Ignacio de la Llave">Instituto de Ecología</location>
, A.C.
</collectorName>
(
<collectionCode id="ED1734CAFFDAFF9B9B1C8725FEB8665B" box="[272,348,1044,1070]" pageId="4" pageNumber="5">IEXA</collectionCode>
),
<collectingMunicipality id="6BDD3675FFDAFF9B9B7F8725FE2D665B" box="[371,457,1044,1070]" pageId="4" pageNumber="5">Xalapa</collectingMunicipality>
,
<collectingRegion id="49C262EDFFDAFF9B9BD48725FDA0665B" box="[472,580,1044,1070]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="4" pageNumber="5">Veracruz</collectingRegion>
,
<collectingCountry id="F311EC9FFFDAFF9B985D8725FD4B665B" box="[593,687,1044,1070]" name="Mexico" pageId="4" pageNumber="5">Mexico</collectingCountry>
</materialsCitation>
.
<materialsCitation id="3B6EA652FFDAFF9B98B18725FB3A6623" ID-GBIF-Occurrence="3336545303" collectionCode="LACMNH" country="United States of America" location="Los Angeles County Museum of Natural History" pageId="4" pageNumber="5" specimenCount="1" specimenCount-female="1" stateProvince="California" typeStatus="paratype">
One male and
<specimenCount id="9D006786FFDAFF9B997C8725FC10665B" box="[880,1012,1044,1070]" pageId="4" pageNumber="5" type="female">one female</specimenCount>
<typeStatus id="54BD12ADFFDAFF9B99F28725FB87665B" box="[1022,1123,1044,1070]" pageId="4" pageNumber="5" type="paratype">paratype</typeStatus>
are deposited in the
<location id="8ED9FAD4FFDAFF9B9B3F870DFC956623" LSID="urn:lsid:plazi:treatment:03AF1D19FFDFFF999B04822DFB7E6526:8ED9FAD4FFDAFF9B9B3F870DFC956623" box="[307,881,1084,1110]" country="United States of America" name="Los Angeles County Museum of Natural History" pageId="4" pageNumber="5" stateProvince="California">Los Angeles County Museum of Natural History</location>
(
<collectionCode id="ED1734CAFFDAFF9B998D870DFBE36623" box="[897,1031,1084,1110]" pageId="4" pageNumber="5">LACMNH</collectionCode>
),
<collectingRegion id="49C262EDFFDAFF9B9E16870DFB706623" box="[1050,1172,1084,1110]" country="United States of America" name="California" pageId="4" pageNumber="5">California</collectingRegion>
,
<collectingCountry id="F311EC9FFFDAFF9B9EAC870DFB3A6623" box="[1184,1246,1084,1110]" name="United States of America" pageId="4" pageNumber="5">USA</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="8BB9AC0FFFDAFF9B9B348755FD62679C" blockId="4.[264,1325,284,1873]" pageId="4" pageNumber="5">
Comments.
<bibRefCitation id="EF97D1FEFFDAFF9B9BDD8755FD92660B" author="Eaton, A. E." box="[465,630,1124,1150]" pageId="4" pageNumber="5" pagination="55 - 59" refId="ref3816" refString="Eaton, A. E. (1904) New genera of European Psychodidae. The Entomologist's Monthly Magazine, 15 (2), 55 - 59." type="journal article" year="1904">Eaton (1904)</bibRefCitation>
erected the subgenus
<taxonomicName id="4C06D78CFFDAFF9B99A48755FBAC6608" authorityName="Eaton" authorityYear="1904" box="[936,1096,1124,1149]" class="Insecta" family="Psychodidae" genus="Philosepedon" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B972701DFFDAFF9B99A48755FBAC6608" box="[936,1096,1124,1149]" italics="true" pageId="4" pageNumber="5">Philosepedon</emphasis>
</taxonomicName>
within
<taxonomicName id="4C06D78CFFDAFF9B9EB58755FE8A66D3" authority="Latreille" authorityName="Latreille" authorityYear="1797" class="Insecta" family="Psychodidae" genus="Psychoda" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B972701DFFDAFF9B9EB58755FACF6608" box="[1209,1323,1124,1149]" italics="true" pageId="4" pageNumber="5">Psychoda</emphasis>
Latreille
</taxonomicName>
, based on the presence of two retinacula in the male cercopodium.
<bibRefCitation id="EF97D1FEFFDAFF9B9E8287BDFACE66D3" author="Quate, L. W." box="[1166,1322,1164,1190]" pageId="4" pageNumber="5" pagination="444 - 451" refId="ref3883" refString="Quate, L. W. (1959) Classification of the Psychodini (Psychodidae: Diptera). Annals of the Entomological Society of America, 52, 444 - 451." type="journal article" year="1959">Quate (1959)</bibRefCitation>
maintained that this taxon is a valid genus and gave the diagnostic characteristics for its recognition based on the presence of a bulbous labellum without teeth, 16 antennomeres (scape, pedicel and 14 flagellomeres) with the apical three reduced, ascoids “Y”­shaped, wing with radial fork distad of the median fork by a distance equal to more than 3x the width of cell R
<subScript id="1782AE4AFFDAFF9B9BB78651FE216704" attach="left" box="[443,453,1376,1393]" fontSize="7" pageId="4" pageNumber="5">2</subScript>
at the point of bifurcation, male with cercopodium not much longer than epandrium, with 2 or 3 retinacula (some species with only one), aedeagus symmetric, with one or two pairs of lateral shafts, female genitalia without digit and cercus short, equal or shorter than the subgenital plate.
</paragraph>
<paragraph id="8BB9AC0FFFDAFF9A9B3486C6FE1763AC" blockId="4.[264,1325,284,1873]" lastBlockId="5.[264,1323,284,1878]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
<bibRefCitation id="EF97D1FEFFDAFF9B9B3486C6FD5A6464" author="Botosaneanu, L. &amp; Vaillant F." box="[312,702,1527,1553]" pageId="4" pageNumber="5" refId="ref3491" refString="Botosaneanu, L. &amp; Vaillant F. (1970) Trois Dipteres Psychodidae nouveaux de Cuba. Travailles Laboratoire de Hydrobiologie, 61,173 - 183." type="book" year="1970">Botosaneanu and Vaillant (1970)</bibRefCitation>
erected the genus
<taxonomicName id="4C06D78CFFDAFF9B99AD86C6FBF46465" authorityName="Botosaneanu &amp; Vaillant" authorityYear="1970" box="[929,1040,1527,1552]" class="Insecta" family="Psychodidae" genus="Quatiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B972701DFFDAFF9B99AD86C6FBF46465" box="[929,1040,1527,1552]" italics="true" pageId="4" pageNumber="5">Quatiella</emphasis>
</taxonomicName>
based on the discovery of the species
<taxonomicName id="4C06D78CFFDAFF9B9BA3852EFC4C644C" authority="Botosaneanu and Vaillant. They" authorityName="Botosaneanu and Vaillant. They" box="[431,936,1567,1593]" class="Insecta" family="Psychodidae" genus="Quatiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="cubana">
<emphasis id="B972701DFFDAFF9B9BA3852EFDCD644D" box="[431,553,1567,1592]" italics="true" pageId="4" pageNumber="5">Q. cubana</emphasis>
Botosaneanu and Vaillant. They
</taxonomicName>
used the following generic characteristics: eyes approximate but not contiguous, bulbous labellum with setae and short spines (without blunt teeth), 16 antennomeres with the terminal three reduced, ascoids trifid, male genitalia with cercopodium bearing one retinaculum, aedeagus symmetric, female genitalia with a very short cercus. Additionally, they denoted in a short complementary key for that refered by them as published in 1970, actually published one year later (
<bibRefCitation id="EF97D1FEFFDAFF9B9B43843EFE18655C" author="Vaillant, F." box="[335,508,1807,1833]" pageId="4" pageNumber="5" pagination="1 - 48" refId="ref3952" refString="Vaillant, F. (1971) Psychodidae Psychodinae. In: Linder, E. (Ed): Die Fliegen der Palearktischen Region, 9 d, pp. 1 - 48. Sttutgart: E. Schweizerbartsche Verlangsbuchhandlung." type="book chapter" year="1971">Vaillant, 1971</bibRefCitation>
), that epadrium is much longer than wide and that the base of the gonopods are contiguous with each other, characteristics used to separate
<taxonomicName id="4C06D78CFFDAFF9B9E768406FB0D6525" authorityName="Botosaneanu &amp; Vaillant" authorityYear="1970" box="[1146,1257,1847,1872]" class="Insecta" family="Psychodidae" genus="Quatiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B972701DFFDAFF9B9E768406FB0D6525" box="[1146,1257,1847,1872]" italics="true" pageId="4" pageNumber="5">Quatiella</emphasis>
</taxonomicName>
from
<taxonomicName id="4C06D78CFFDBFF9A9B04822DFE5D6340" authorityName="Vaillant" authorityYear="1974" box="[264,441,284,309]" class="Insecta" family="Psychodidae" genus="Feuerborniella" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B972701DFFDBFF9A9B04822DFE5D6340" box="[264,441,284,309]" italics="true" pageId="5" pageNumber="6">Feuerborniella</emphasis>
</taxonomicName>
. Based on the figure of the wing presented by
<bibRefCitation id="EF97D1FEFFDBFF9A99F6822DFEAB632B" author="Botosaneanu, L. &amp; Vaillant F." pageId="5" pageNumber="6" refId="ref3491" refString="Botosaneanu, L. &amp; Vaillant F. (1970) Trois Dipteres Psychodidae nouveaux de Cuba. Travailles Laboratoire de Hydrobiologie, 61,173 - 183." type="book" year="1970">Botosaneanu and Vaillant (1970</bibRefCitation>
, p. 175) and the figure of the type­species
<taxonomicName id="4C06D78CFFDBFF9A999A8275FACE632B" authority="Vaillant" authorityName="Vaillant" authorityYear="1968" box="[918,1322,324,350]" class="Insecta" family="Psychodidae" genus="Philosepedon" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="leonhardti">
<emphasis id="B972701DFFDBFF9A999A8275FB5B6328" box="[918,1215,324,349]" italics="true" pageId="5" pageNumber="6">Philosepedon leonhardti</emphasis>
Vaillant
</taxonomicName>
(=
<taxonomicName id="4C06D78CFFDBFF9A9B2F825DFDB463F3" authority="Dyar" authorityName="Dyar" box="[291,592,364,390]" class="Insecta" family="Psychodidae" genus="Quatiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="interdicta">
<emphasis id="B972701DFFDBFF9A9B2F825DFDEF63F0" box="[291,523,364,389]" italics="true" pageId="5" pageNumber="6">Quatiella interdicta</emphasis>
Dyar
</taxonomicName>
) (
<bibRefCitation id="EF97D1FEFFDBFF9A9864825DFCE963F3" author="Vaillant, F." box="[616,781,364,390]" pageId="5" pageNumber="6" pagination="423 - 430" refId="ref3916" refString="Vaillant, F. (1968) Un diptere Psychodide nouveau d'Amerique du Nord et ses stades preimaginaux. Bulletin de la Societe Zoologique de France, 93 (3), 423 - 430." type="journal article" year="1968">Vaillant 1968</bibRefCitation>
), we can infer that at least these species have the radial fork distad of the median fork by a distance about the width of cell R
<subScript id="1782AE4AFFDBFF9A9ED88291FB3A63C4" attach="left" box="[1236,1246,416,433]" fontSize="7" pageId="5" pageNumber="6">2</subScript>
at the point of bifurcation.
</paragraph>
<paragraph id="8BB9AC0FFFDBFF9A9B3482D6FD656194" blockId="5.[264,1323,284,1878]" pageId="5" pageNumber="6">
<bibRefCitation id="EF97D1FEFFDBFF9A9B3482D6FE0E6074" author="Vaillant, F." box="[312,490,487,513]" pageId="5" pageNumber="6" pagination="345 - 379" refId="ref3994" refString="Vaillant, F. (1973) Some new Psychodidae Psychodinae from the United States. Annales de la Societe Entomologique de France (N. S.), 9 (1), 345 - 379." type="journal article" year="1973">Vaillant (1973)</bibRefCitation>
later described
<taxonomicName id="4C06D78CFFDBFF9A98A582D6FC7F6075" authorityName="Vaillant" authorityYear="1973" box="[681,923,487,512]" class="Insecta" family="Psychodidae" genus="Philosepedon" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="quatei">
<emphasis id="B972701DFFDBFF9A98A582D6FC7F6075" box="[681,923,487,512]" italics="true" pageId="5" pageNumber="6">Philosepedon quatei</emphasis>
</taxonomicName>
and presented a key to the North American genera of his
<taxonomicName id="4C06D78CFFDBFF9A9832813EFD48605D" authorityName="Eaton" authorityYear="1904" box="[574,684,527,552]" class="Insecta" family="Psychodidae" genus="Threticus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B972701DFFDBFF9A9832813EFD48605D" box="[574,684,527,552]" italics="true" pageId="5" pageNumber="6">Threticus</emphasis>
</taxonomicName>
group. In that work, the distinctive characteristics between
<taxonomicName id="4C06D78CFFDBFF9A9B7F8106FDF76025" authorityName="Eaton" authorityYear="1904" box="[371,531,567,592]" class="Insecta" family="Psychodidae" genus="Philosepedon" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B972701DFFDBFF9A9B7F8106FDF76025" box="[371,531,567,592]" italics="true" pageId="5" pageNumber="6">Philosepedon</emphasis>
</taxonomicName>
and
<taxonomicName id="4C06D78CFFDBFF9A98428106FD596025" authorityName="Botosaneanu &amp; Vaillant" authorityYear="1970" box="[590,701,567,592]" class="Insecta" family="Psychodidae" genus="Quatiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B972701DFFDBFF9A98428106FD596025" box="[590,701,567,592]" italics="true" pageId="5" pageNumber="6">Quatiella</emphasis>
</taxonomicName>
were: cercopodium with 2 or 2 groups of retinacula
<emphasis id="B972701DFFDBFF9A9B04816EFEB7600D" box="[264,339,607,632]" italics="true" pageId="5" pageNumber="6">versus</emphasis>
only one retinaculum, respectively. In the same year,
<bibRefCitation id="EF97D1FEFFDBFF9A99F9816EFB37600C" author="Duckhouse, D. A." box="[1013,1235,607,633]" pageId="5" pageNumber="6" pagination="1 - 29" refId="ref3566" refString="Duckhouse, D. A. (1973) Family Psychodidae, 6 A. In: Papavero, N. (Ed): A Catalogue of the Diptera of the Americas South of the United States. Departamento de Zoologia., Secretaria da Agricultura, Sao Paulo, Brazil, pp. 1 - 29." type="book chapter" year="1973">Duckhouse (1973)</bibRefCitation>
placed
<taxonomicName id="4C06D78CFFDBFF9A9B0481B6FE9360D5" authorityName="Botosaneanu &amp; Vaillant" authorityYear="1970" box="[264,375,647,672]" class="Insecta" family="Psychodidae" genus="Quatiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B972701DFFDBFF9A9B0481B6FE9360D5" box="[264,375,647,672]" italics="true" pageId="5" pageNumber="6">Quatiella</emphasis>
</taxonomicName>
and
<taxonomicName id="4C06D78CFFDBFF9A9BBA81B6FD6760D4" authority="Quate" authorityName="Quate" authorityYear="1959" box="[438,643,647,673]" class="Insecta" family="Psychodidae" genus="Eurygarka" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B972701DFFDBFF9A9BBA81B6FDD760D5" box="[438,563,647,672]" italics="true" pageId="5" pageNumber="6">Eurygarka</emphasis>
Quate
</taxonomicName>
as subgenera of
<taxonomicName id="4C06D78CFFDBFF9A995981B6FC1160D5" authorityName="Eaton" authorityYear="1904" box="[853,1013,647,672]" class="Insecta" family="Psychodidae" genus="Philosepedon" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B972701DFFDBFF9A995981B6FC1160D5" box="[853,1013,647,672]" italics="true" pageId="5" pageNumber="6">Philosepedon</emphasis>
</taxonomicName>
, considering
<taxonomicName id="4C06D78CFFDBFF9A9E9681B6FACF60D5" box="[1178,1323,647,672]" class="Insecta" family="Psychodidae" genus="Quatiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="interdicta">
<emphasis id="B972701DFFDBFF9A9E9681B6FACF60D5" box="[1178,1323,647,672]" italics="true" pageId="5" pageNumber="6">P. interdicta</emphasis>
</taxonomicName>
under the subgenus
<taxonomicName id="4C06D78CFFDBFF9A981F819EFD5660BD" authorityName="Eaton" authorityYear="1904" box="[531,690,687,712]" class="Insecta" family="Psychodidae" genus="Philosepedon" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B972701DFFDBFF9A981F819EFD5660BD" box="[531,690,687,712]" italics="true" pageId="5" pageNumber="6">Philosepedon</emphasis>
</taxonomicName>
. In addition, the Brazilian species he originally described as
<taxonomicName id="4C06D78CFFDBFF9A9BA581E6FD406085" box="[425,676,727,752]" class="Insecta" family="Psychodidae" genus="Psychoda" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="plaumanni">
<emphasis id="B972701DFFDBFF9A9BA581E6FD406085" box="[425,676,727,752]" italics="true" pageId="5" pageNumber="6">Psychoda plaumanni</emphasis>
</taxonomicName>
,
<taxonomicName id="4C06D78CFFDBFF9A98B481E6FB546084" authority="(Duckhouse, 1968)" baseAuthorityName="Duckhouse" baseAuthorityYear="1968" box="[696,1200,727,753]" class="Insecta" family="Psychodidae" genus="Psychoda" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="spathipenis">
<emphasis id="B972701DFFDBFF9A98B481E6FC5F6085" box="[696,955,727,752]" italics="true" pageId="5" pageNumber="6">Psychoda spathipenis</emphasis>
(
<bibRefCitation id="EF97D1FEFFDBFF9A99C381E6FB436084" author="Duckhouse, D. A." box="[975,1191,727,753]" pageId="5" pageNumber="6" pagination="29 - 40" refId="ref3518" refString="Duckhouse, D. A. (1968) Psychodidae (Diptera, Nematocera) collected by Mr. Plaumann in Southern Brazil. The Proceedings of the Royal Entomological Society of London (B), 37 (3 - 4), 29 - 40." type="journal article" year="1968">Duckhouse, 1968</bibRefCitation>
)
</taxonomicName>
, and
<taxonomicName id="4C06D78CFFDBFF9A9EFA81E6FE1C616C" authority="Knab" authorityName="Knab" authorityYear="1914" class="Insecta" family="Psychodidae" genus="Psychoda" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="fumata">
<emphasis id="B972701DFFDBFF9A9EFA81E6FE49616D" italics="true" pageId="5" pageNumber="6">Psychoda fumata</emphasis>
Knab
</taxonomicName>
(known from
<collectingCountry id="F311EC9FFFDBFF9A98BC81CEFCEF616C" box="[688,779,767,793]" name="Mexico" pageId="5" pageNumber="6">Mexico</collectingCountry>
) were also included. In another publication,
<bibRefCitation id="EF97D1FEFFDBFF9A9B048016FE006134" author="Duckhouse, D. A." box="[264,484,807,833]" pageId="5" pageNumber="6" pagination="141 - 152" refId="ref3625" refString="Duckhouse, D. A. (1974) Redescription of the Neotropical Psychodidae (Diptera, Nematocera) described by Knab, Dyar, and Coquillett. Journal of Entomology (B), 42 (2), 141 - 152." type="journal article" year="1974">Duckhouse (1974)</bibRefCitation>
expressed his opinion about the validity of
<taxonomicName id="4C06D78CFFDBFF9A9E0F8016FB966135" authorityName="Botosaneanu &amp; Vaillant" authorityYear="1970" box="[1027,1138,807,832]" class="Insecta" family="Psychodidae" genus="Quatiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B972701DFFDBFF9A9E0F8016FB966135" box="[1027,1138,807,832]" italics="true" pageId="5" pageNumber="6">Quatiella</emphasis>
</taxonomicName>
and concluded that there was no justification to retain it as a separate genus. The only conspicuous difference between
<taxonomicName id="4C06D78CFFDBFF9A9BA38046FDC861E5" authorityName="Quate" authorityYear="1959" box="[431,556,887,912]" class="Insecta" family="Psychodidae" genus="Eurygarka" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B972701DFFDBFF9A9BA38046FDC861E5" box="[431,556,887,912]" italics="true" pageId="5" pageNumber="6">Eurygarka</emphasis>
</taxonomicName>
and the others, as was first noted by
<bibRefCitation id="EF97D1FEFFDBFF9A99EA8046FB6561E4" author="Quate, L. W." box="[998,1153,887,913]" pageId="5" pageNumber="6" pagination="444 - 451" refId="ref3883" refString="Quate, L. W. (1959) Classification of the Psychodini (Psychodidae: Diptera). Annals of the Entomological Society of America, 52, 444 - 451." type="journal article" year="1959">Quate (1959)</bibRefCitation>
is the form of the antennal ascoids, whereas the other diagnostic characteristics overlap or are not sufficiently precise to separate them.
</paragraph>
<paragraph id="8BB9AC0FFFDBFF9A9B3480DEFCE46754" blockId="5.[264,1323,284,1878]" pageId="5" pageNumber="6">
<bibRefCitation id="EF97D1FEFFDBFF9A9B3480DEFE0D667C" author="Vaillant, F." box="[312,489,1007,1033]" pageId="5" pageNumber="6" pagination="109 - 142" refId="ref4033" refString="Vaillant, F. (1974) Psychodidae Psychodinae. In: Linder, E. (Ed): Die Fliegen der Palearktischen Region, 9 d. Sttutgart: E. Schweizerbartsche Verlangsbuchhandlung, pp. 109 - 142." type="book chapter" year="1974">Vaillant (1974)</bibRefCitation>
described the genus
<taxonomicName id="4C06D78CFFDBFF9A98EF80DEFC70667D" authorityName="Vaillant" authorityYear="1974" box="[739,916,1007,1032]" class="Insecta" family="Psychodidae" genus="Feuerborniella" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B972701DFFDBFF9A98EF80DEFC70667D" box="[739,916,1007,1032]" italics="true" pageId="5" pageNumber="6">Feuerborniella</emphasis>
</taxonomicName>
to harbor the Palearctic species
<taxonomicName id="4C06D78CFFDBFF9A9F1980DEFE006644" authority="(Tonnoir)" baseAuthorityName="Tonnoir" baseAuthorityYear="1919" class="Insecta" family="Psychodidae" genus="Feuerborniella" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="obscura">
<emphasis id="B972701DFFDBFF9A9F1980DEFE826645" italics="true" pageId="5" pageNumber="6">F. obscura</emphasis>
(Tonnoir)
</taxonomicName>
, and considered both Brazilian species of Duckhouse and
<taxonomicName id="4C06D78CFFDBFF9A9EA18726FDBC662C" authority="Satchell" authorityName="Satchell" authorityYear="1955" class="Insecta" family="Psychodidae" genus="Trichopsychoda" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="malayensis">
<emphasis id="B972701DFFDBFF9A9EA18726FE00662D" italics="true" pageId="5" pageNumber="6">Trichopsychoda malayensis</emphasis>
Satchell
</taxonomicName>
, under this genus. Vaillant considered
<taxonomicName id="4C06D78CFFDBFF9A9E53870EFAC2662C" authority="Eaton" authorityName="Eaton" authorityYear="1904" box="[1119,1318,1087,1113]" class="Insecta" family="Psychodidae" genus="Threticus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B972701DFFDBFF9A9E53870EFB29662D" box="[1119,1229,1087,1112]" italics="true" pageId="5" pageNumber="6">Threticus</emphasis>
Eaton
</taxonomicName>
,
<taxonomicName id="4C06D78CFFDBFF9A9B048756FE4366F5" authorityName="Eaton" authorityYear="1904" box="[264,423,1127,1152]" class="Insecta" family="Psychodidae" genus="Philosepedon" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B972701DFFDBFF9A9B048756FE4366F5" box="[264,423,1127,1152]" italics="true" pageId="5" pageNumber="6">Philosepedon</emphasis>
</taxonomicName>
,
<taxonomicName id="4C06D78CFFDBFF9A9BB58756FDCC66F5" authorityName="Botosaneanu &amp; Vaillant" authorityYear="1970" box="[441,552,1127,1152]" class="Insecta" family="Psychodidae" genus="Quatiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B972701DFFDBFF9A9BB58756FDCC66F5" box="[441,552,1127,1152]" italics="true" pageId="5" pageNumber="6">Quatiella</emphasis>
</taxonomicName>
,
<taxonomicName id="4C06D78CFFDBFF9A98348756FCC866F4" authority="Vaillant" authorityName="Vaillant" authorityYear="1972" box="[568,812,1127,1153]" class="Insecta" family="Psychodidae" genus="Nielseniella" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B972701DFFDBFF9A98348756FD2066F5" box="[568,708,1127,1152]" italics="true" pageId="5" pageNumber="6">Nielseniella</emphasis>
Vaillant
</taxonomicName>
,
<taxonomicName id="4C06D78CFFDBFF9A99368756FC0F66F5" authorityName="Vaillant" authorityYear="1974" box="[826,1003,1127,1152]" class="Insecta" family="Psychodidae" genus="Feuerborniella" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B972701DFFDBFF9A99368756FC0F66F5" box="[826,1003,1127,1152]" italics="true" pageId="5" pageNumber="6">Feuerborniella</emphasis>
</taxonomicName>
, and three no longer valid genera as the
<taxonomicName id="4C06D78CFFDBFF9A9BA287BEFDF866DD" authorityName="Eaton" authorityYear="1904" box="[430,540,1167,1192]" class="Insecta" family="Psychodidae" genus="Threticus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B972701DFFDBFF9A9BA287BEFDF866DD" box="[430,540,1167,1192]" italics="true" pageId="5" pageNumber="6">Threticus</emphasis>
</taxonomicName>
group of
<taxonomicName id="4C06D78CFFDBFF9A989987BEFCAD66DC" box="[661,841,1167,1193]" class="Insecta" family="Psychodidae" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="tribe" tribe="Telmatoscopini">Telmatoscopini</taxonomicName>
(
<bibRefCitation id="EF97D1FEFFDBFF9A995687BEFC1966DC" author="Vaillant, F." box="[858,1021,1167,1193]" pageId="5" pageNumber="6" pagination="345 - 379" refId="ref3994" refString="Vaillant, F. (1973) Some new Psychodidae Psychodinae from the United States. Annales de la Societe Entomologique de France (N. S.), 9 (1), 345 - 379." type="journal article" year="1973">Vaillant 1973</bibRefCitation>
,
<bibRefCitation id="EF97D1FEFFDBFF9A9E0087BEFBAF66DC" author="Vaillant, F." box="[1036,1099,1167,1193]" pageId="5" pageNumber="6" pagination="109 - 142" refId="ref4033" refString="Vaillant, F. (1974) Psychodidae Psychodinae. In: Linder, E. (Ed): Die Fliegen der Palearktischen Region, 9 d. Sttutgart: E. Schweizerbartsche Verlangsbuchhandlung, pp. 109 - 142." type="book chapter" year="1974">1974</bibRefCitation>
). It is important to note that the original description of
<taxonomicName id="4C06D78CFFDBFF9A98B18786FC8A66A5" authorityName="Vaillant" authorityYear="1974" box="[701,878,1207,1232]" class="Insecta" family="Psychodidae" genus="Feuerborniella" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B972701DFFDBFF9A98B18786FC8A66A5" box="[701,878,1207,1232]" italics="true" pageId="5" pageNumber="6">Feuerborniella</emphasis>
</taxonomicName>
was published 4 years after the first mention of the genus by
<bibRefCitation id="EF97D1FEFFDBFF9A983F87EEFC58668C" author="Botosaneanu, L. &amp; Vaillant F." box="[563,956,1247,1273]" pageId="5" pageNumber="6" refId="ref3491" refString="Botosaneanu, L. &amp; Vaillant F. (1970) Trois Dipteres Psychodidae nouveaux de Cuba. Travailles Laboratoire de Hydrobiologie, 61,173 - 183." type="book" year="1970">Botosaneanu and Vaillant (1970)</bibRefCitation>
, situation that could invalidate the name in accordance with ICNZ (2000).
</paragraph>
<paragraph id="8BB9AC0FFFDBFF9A9B34861EFDE56464" blockId="5.[264,1323,284,1878]" pageId="5" pageNumber="6">
<bibRefCitation id="EF97D1FEFFDBFF9A9B34861EFE0B673C" author="Vaillant, F." box="[312,495,1327,1353]" pageId="5" pageNumber="6" pagination="141 - 163" refId="ref4117" refString="Vaillant, F. (1990) Propositions pour une revision de la classification des Dipteres Psychodidae Psychodinae. Bulletin de la Societe Vaudoise de Sciences Naturelles, 80 (2), 141 - 163." type="journal article" year="1990">Vaillant (1990)</bibRefCitation>
retained
<taxonomicName id="4C06D78CFFDBFF9A9865861EFD32673D" authorityName="Eaton" authorityYear="1904" box="[617,726,1327,1352]" class="Insecta" family="Psychodidae" genus="Threticus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B972701DFFDBFF9A9865861EFD32673D" box="[617,726,1327,1352]" italics="true" pageId="5" pageNumber="6">Threticus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C06D78CFFDBFF9A98E0861EFC6F673D" authorityName="Eaton" authorityYear="1904" box="[748,907,1327,1352]" class="Insecta" family="Psychodidae" genus="Philosepedon" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B972701DFFDBFF9A98E0861EFC6F673D" box="[748,907,1327,1352]" italics="true" pageId="5" pageNumber="6">Philosepedon</emphasis>
</taxonomicName>
,
<taxonomicName id="4C06D78CFFDBFF9A99AD861EFB2C673C" authority="Tonnoir" authorityName="Tonnoir" authorityYear="1922" box="[929,1224,1327,1353]" class="Insecta" family="Psychodidae" genus="Trichopsychoda" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B972701DFFDBFF9A99AD861EFBB8673D" box="[929,1116,1327,1352]" italics="true" pageId="5" pageNumber="6">Trichopsychoda</emphasis>
Tonnoir
</taxonomicName>
,
<taxonomicName id="4C06D78CFFDBFF9A9ED7861EFE976705" authorityName="Vaillant" authorityYear="1974" class="Insecta" family="Psychodidae" genus="Feuerborniella" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B972701DFFDBFF9A9ED7861EFE976705" italics="true" pageId="5" pageNumber="6">Feuerborniella</emphasis>
</taxonomicName>
,
<taxonomicName id="4C06D78CFFDBFF9A9B888666FE176705" authorityName="Botosaneanu &amp; Vaillant" authorityYear="1970" box="[388,499,1367,1392]" class="Insecta" family="Psychodidae" genus="Quatiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B972701DFFDBFF9A9B888666FE176705" box="[388,499,1367,1392]" italics="true" pageId="5" pageNumber="6">Quatiella</emphasis>
</taxonomicName>
and
<taxonomicName id="4C06D78CFFDBFF9A98238666FD5F6705" authorityName="Vaillant" authorityYear="1972" box="[559,699,1367,1392]" class="Insecta" family="Psychodidae" genus="Nielseniella" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B972701DFFDBFF9A98238666FD5F6705" box="[559,699,1367,1392]" italics="true" pageId="5" pageNumber="6">Nielseniella</emphasis>
</taxonomicName>
as valid genera under the tribe
<taxonomicName id="4C06D78CFFDBFF9A9E348666FAC26704" authority="Enderlein" authorityName="Enderlein" authorityYear="1937" box="[1080,1318,1367,1393]" class="Insecta" family="Psychodidae" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="tribe" tribe="Mormiini">Mormiini Enderlein</taxonomicName>
, corresponding to the tribe
<taxonomicName id="4C06D78CFFDBFF9A9831864EFD1567EC" box="[573,753,1407,1433]" class="Insecta" family="Psychodidae" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="tribe" tribe="Telmatoscopini">Telmatoscopini</taxonomicName>
of
<bibRefCitation id="EF97D1FEFFDBFF9A9914864EFC2867EC" author="Vaillant, F." box="[792,972,1407,1433]" pageId="5" pageNumber="6" pagination="1 - 48" refId="ref3952" refString="Vaillant, F. (1971) Psychodidae Psychodinae. In: Linder, E. (Ed): Die Fliegen der Palearktischen Region, 9 d, pp. 1 - 48. Sttutgart: E. Schweizerbartsche Verlangsbuchhandlung." type="book chapter" year="1971">Vaillant (1971)</bibRefCitation>
, or to the tribes
<taxonomicName id="4C06D78CFFDBFF9A9E85864EFACE67EC" box="[1161,1322,1407,1433]" class="Insecta" family="Psychodidae" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="tribe" tribe="Paramormiini">Paramormiini</taxonomicName>
and part of
<taxonomicName id="4C06D78CFFDBFF9A9B948696FDFE67B4" authorityName="Newman" authorityYear="1834" box="[408,538,1447,1473]" class="Insecta" family="Psychodidae" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="tribe" tribe="Psychodini">Psychodini</taxonomicName>
of
<bibRefCitation id="EF97D1FEFFDBFF9A98448696FCC367B4" author="Duckhouse, D. A." box="[584,807,1447,1473]" pageId="5" pageNumber="6" pagination="231 - 282" refId="ref3717" refString="Duckhouse, D. A. (1987) A revision of Afrotropical Setomima, elucidation of their genealogical relationships and descriptions of other Afrotropical Psychodinae (Diptera: Psychodidae). Annals of the Natal Museum, 28 (2), 231 - 282." type="journal article" year="1987">Duckhouse (1987)</bibRefCitation>
. More recently,
<bibRefCitation id="EF97D1FEFFDBFF9A99E28696FB4567B4" author="Wagner, R." box="[1006,1185,1447,1473]" pageId="5" pageNumber="6" pagination="11 - 65" refId="ref4154" refString="Wagner, R. (1991) Family Psychodidae. In: Soos, A. &amp; Papp L. (Ed): Catalogue of Palearctic Diptera, Psychodidae-Chironomidae, Vol. 2. Elsevier, Netherlands, pp. 11 - 65." type="book chapter" year="1991">Wagner (1991)</bibRefCitation>
considered
<emphasis id="B972701DFFDBFF9A9B0486FEFC47679D" box="[264,931,1487,1512]" italics="true" pageId="5" pageNumber="6">
<taxonomicName id="4C06D78CFFDBFF9A9B0486FEFE58679D" authorityName="Vaillant" authorityYear="1974" box="[264,444,1487,1512]" class="Insecta" family="Psychodidae" genus="Feuerborniella" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">Feuerborniella</taxonomicName>
,
<taxonomicName id="4C06D78CFFDBFF9A9BC786FEFDBE679D" authorityName="Vaillant" authorityYear="1972" box="[459,602,1487,1512]" class="Insecta" family="Psychodidae" genus="Nielseniella" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">Nielseniella</taxonomicName>
,
<taxonomicName id="4C06D78CFFDBFF9A986486FEFCC2679D" authorityName="Tonnoir" authorityYear="1922" box="[616,806,1487,1512]" class="Insecta" family="Psychodidae" genus="Trichopsychoda" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">Trichopsychoda</taxonomicName>
,
<taxonomicName id="4C06D78CFFDBFF9A993986FEFC47679D" authorityName="Eaton" authorityYear="1904" box="[821,931,1487,1512]" class="Insecta" family="Psychodidae" genus="Threticus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">Threticus</taxonomicName>
</emphasis>
and
<taxonomicName id="4C06D78CFFDBFF9A99ED86FEFB65679D" authorityName="Eaton" authorityYear="1904" box="[993,1153,1487,1512]" class="Insecta" family="Psychodidae" genus="Philosepedon" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B972701DFFDBFF9A99ED86FEFB65679D" box="[993,1153,1487,1512]" italics="true" pageId="5" pageNumber="6">Philosepedon</emphasis>
</taxonomicName>
as part of the tribe
<taxonomicName id="4C06D78CFFDBFF9A9B4986C6FE196464" box="[325,509,1527,1553]" class="Insecta" family="Psychodidae" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="tribe" tribe="Telmatoscopini">Telmatoscopini</taxonomicName>
.
</paragraph>
<paragraph id="8BB9AC0FFFDBFF999B34852EFC70638B" blockId="5.[264,1323,284,1878]" lastBlockId="6.[264,1324,284,1875]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
<bibRefCitation id="EF97D1FEFFDBFF9A9B34852EFDF3644C" author="Duckhouse, D. A." box="[312,535,1567,1593]" pageId="5" pageNumber="6" pagination="601 - 620" refId="ref3669" refString="Duckhouse, D. A. (1985) A re-examination of Neomaruina (Diptera, Psychodidae), with observations on its life-history and affinities and redefinition of the tribe Psychodini. Annals of the Natal Museum, 26 (2), 601 - 620." type="journal article" year="1985">Duckhouse (1985)</bibRefCitation>
redefined the tribe
<taxonomicName id="4C06D78CFFDBFF9A991C852EFC76644C" authorityName="Newman" authorityYear="1834" box="[784,914,1567,1593]" class="Insecta" family="Psychodidae" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="tribe" tribe="Psychodini">Psychodini</taxonomicName>
based on the following apomorphies: antennal flagellomeres flask­shaped except the terminal three which are reduced and without ascoids; ascoids Y­shaped or derived from this
<typeStatus id="54BD12ADFFDBFF9A99B1855EFC0B64FC" box="[957,1007,1647,1673]" pageId="5" pageNumber="6">type</typeStatus>
by loss of one or two anterior branches; wing pointed with R
<subScript id="1782AE4AFFDBFF9A98B98592FD5B64C1" attach="left" box="[693,703,1699,1716]" fontSize="7" pageId="5" pageNumber="6">5</subScript>
ending at the apex; subcosta (Sc) short and thick; wings folding in tectiform position; erect hair present on R
<subScript id="1782AE4AFFDBFF9A99CF85FFFC2964AA" attach="right" box="[963,973,1742,1759]" fontSize="7" pageId="5" pageNumber="6">1</subScript>
, and epandrium of male with single pseudospiracular opening, as well as other characters observed in larvae. The simple characteristic of the flattened labellum is insufficient to separate the tribe, because the species of
<taxonomicName id="4C06D78CFFDBFF9A9B89840DFE176520" authorityName="Eaton" authorityYear="1904" box="[389,499,1852,1877]" class="Insecta" family="Psychodidae" genus="Threticus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B972701DFFDBFF9A9B89840DFE176520" box="[389,499,1852,1877]" italics="true" pageId="5" pageNumber="6">Threticus</emphasis>
</taxonomicName>
have both conditions of the labellum.
<bibRefCitation id="EF97D1FEFFDBFF9A99B5840DFB776523" author="Duckhouse, D. A." box="[953,1171,1852,1878]" pageId="5" pageNumber="6" pagination="601 - 620" refId="ref3669" refString="Duckhouse, D. A. (1985) A re-examination of Neomaruina (Diptera, Psychodidae), with observations on its life-history and affinities and redefinition of the tribe Psychodini. Annals of the Natal Museum, 26 (2), 601 - 620." type="journal article" year="1985">Duckhouse (1985)</bibRefCitation>
included the genera
<emphasis id="B972701DFFD8FF999B53822DFD7E6340" box="[351,666,284,309]" italics="true" pageId="6" pageNumber="7">
<taxonomicName id="4C06D78CFFD8FF999B53822DFE306340" authorityName="Latreille" authorityYear="1797" box="[351,468,284,309]" class="Insecta" family="Psychodidae" genus="Psychoda" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">Psychoda</taxonomicName>
,
<taxonomicName id="4C06D78CFFD8FF999BEE822DFD726340" authorityName="Vaillant" authorityYear="1974" box="[482,662,284,309]" class="Insecta" family="Psychodidae" genus="Feuerborniella" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">Feuerborniella</taxonomicName>
,
</emphasis>
<taxonomicName id="4C06D78CFFD8FF9998A8822DFC796343" authority="Vaillant" authorityName="Vaillant" authorityYear="1963" box="[676,925,284,310]" class="Insecta" family="Psychodidae" genus="Neomaruina" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B972701DFFD8FF9998A8822DFCD26340" box="[676,822,284,309]" italics="true" pageId="6" pageNumber="7">Neomaruina</emphasis>
Vaillant
</taxonomicName>
,
<taxonomicName id="4C06D78CFFD8FF9999A7822DFB926343" authority="Quate" authorityName="Quate" box="[939,1142,284,310]" class="Insecta" family="Psychodidae" genus="Epacreton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B972701DFFD8FF9999A7822DFBC06340" box="[939,1060,284,309]" italics="true" pageId="6" pageNumber="7">Epacreton</emphasis>
Quate
</taxonomicName>
,
<emphasis id="B972701DFFD8FF999E8F822DFE276328" italics="true" pageId="6" pageNumber="7">
<taxonomicName id="4C06D78CFFD8FF999E8F822DFAC16340" authorityName="Eaton" authorityYear="1904" box="[1155,1317,284,309]" class="Insecta" family="Psychodidae" genus="Philosepedon" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">Philosepedon</taxonomicName>
,
<taxonomicName id="4C06D78CFFD8FF999B048275FE276328" authorityName="Tonnoir" authorityYear="1922" box="[264,451,324,349]" class="Insecta" family="Psychodidae" genus="Trichopsychoda" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">Trichopsychoda</taxonomicName>
</emphasis>
and
<taxonomicName id="4C06D78CFFD8FF99980D8275FD8B6328" authorityName="Eaton" authorityYear="1904" box="[513,623,324,349]" class="Insecta" family="Psychodidae" genus="Threticus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B972701DFFD8FF99980D8275FD8B6328" box="[513,623,324,349]" italics="true" pageId="6" pageNumber="7">Threticus</emphasis>
</taxonomicName>
in tribe
<taxonomicName id="4C06D78CFFD8FF9998DB8275FCB9632B" authorityName="Newman" authorityYear="1834" box="[727,861,324,350]" class="Insecta" family="Psychodidae" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="tribe" tribe="Psychodini">Psychodini</taxonomicName>
. Other genera that have been included more recently in the
<taxonomicName id="4C06D78CFFD8FF99980B825DFD6D63F3" authorityName="Newman" authorityYear="1834" box="[519,649,364,390]" class="Insecta" family="Psychodidae" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="tribe" tribe="Psychodini">Psychodini</taxonomicName>
are:
<taxonomicName id="4C06D78CFFD8FF9998C7825DFC5D63F3" authority="Jezek" authorityName="Jezek" authorityYear="1984" box="[715,953,364,390]" class="Insecta" family="Psychodidae" genus="Chodopsycha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B972701DFFD8FF9998C7825DFC8E63F0" box="[715,874,364,389]" italics="true" pageId="6" pageNumber="7">Chodopsycha</emphasis>
Jezek
</taxonomicName>
,
<taxonomicName id="4C06D78CFFD8FF9999C4825DFB9763F3" authority="Eaton" authorityName="Eaton" authorityYear="1904" box="[968,1139,364,390]" class="Insecta" family="Psychodidae" genus="Logima" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B972701DFFD8FF9999C4825DFBC563F0" box="[968,1057,364,389]" italics="true" pageId="6" pageNumber="7">Logima</emphasis>
Eaton
</taxonomicName>
,
<taxonomicName id="4C06D78CFFD8FF999E8F825DFAC263F3" authority="Jezek" authorityName="Jezek" authorityYear="1984" box="[1155,1318,364,390]" class="Insecta" family="Psychodidae" genus="Psycha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B972701DFFD8FF999E8F825DFB3C63F0" box="[1155,1240,364,389]" italics="true" pageId="6" pageNumber="7">Psycha</emphasis>
Jezek
</taxonomicName>
,
<taxonomicName id="4C06D78CFFD8FF999B0482A5FE1663DB" authority="Jezek" authorityName="Jezek" authorityYear="1984" box="[264,498,404,430]" class="Insecta" family="Chironomidae" genus="Psychodocha" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B972701DFFD8FF999B0482A5FE4063D8" box="[264,420,404,429]" italics="true" pageId="6" pageNumber="7">Psychodocha</emphasis>
Jezek
</taxonomicName>
, and
<taxonomicName id="4C06D78CFFD8FF99983982A5FB6D63DB" authority="Jezek (Duckhouse and Lewis, 2000)" authorityName="Jezek (Duckhouse and Lewis" authorityYear="2000" box="[565,1161,404,430]" class="Insecta" family="Psychodidae" genus="Psychomora" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B972701DFFD8FF99983982A5FD2363D8" box="[565,711,404,429]" italics="true" pageId="6" pageNumber="7">Psychomora</emphasis>
Jezek (
<bibRefCitation id="EF97D1FEFFD8FF99992882A5FB6563DB" author="Duckhouse, D. A. &amp; Lewis, D. J." box="[804,1153,404,430]" pageId="6" pageNumber="7" refId="ref3764" refString="Duckhouse, D. A. &amp; Lewis, D. J. (2000) A / O Diptera Catalog. Psychodidae 15. Available from: http: // www. bishop. hawaii. org / bishop / ento / aocat / (September 12, 2003)." type="journal volume" year="2000">Duckhouse and Lewis, 2000</bibRefCitation>
)
</taxonomicName>
. These inclusions are well supported based on the available information and we follows the concept provided by
<bibRefCitation id="EF97D1FEFFD8FF999B9082D5FD92638B" author="Duckhouse, D. A." box="[412,630,484,510]" pageId="6" pageNumber="7" pagination="601 - 620" refId="ref3669" refString="Duckhouse, D. A. (1985) A re-examination of Neomaruina (Diptera, Psychodidae), with observations on its life-history and affinities and redefinition of the tribe Psychodini. Annals of the Natal Museum, 26 (2), 601 - 620." type="journal article" year="1985">Duckhouse (1985)</bibRefCitation>
on the tribe
<taxonomicName id="4C06D78CFFD8FF99990582D5FC6B638B" authorityName="Newman" authorityYear="1834" box="[777,911,484,510]" class="Insecta" family="Psychodidae" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="tribe" tribe="Psychodini">Psychodini</taxonomicName>
.
</paragraph>
<paragraph id="8BB9AC0FFFD8FF999B34813DFD22662E" blockId="6.[264,1324,284,1875]" pageId="6" pageNumber="7">
The new species described in this report correspond to the genus
<taxonomicName id="4C06D78CFFD8FF999E5E813DFAE76050" authorityName="Vaillant" authorityYear="1974" box="[1106,1283,524,549]" class="Insecta" family="Psychodidae" genus="Feuerborniella" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B972701DFFD8FF999E5E813DFAE76050" box="[1106,1283,524,549]" italics="true" pageId="6" pageNumber="7">Feuerborniella</emphasis>
</taxonomicName>
by the approximate eyes, ocular bridge with 4 rows of facets, 14 flagellomeres, “Y”­shaped ascoids, apical 3 flagellomeres reduced in size, labellum small and fleshy with setae and small spines but without blunt teeth, R
<subScript id="1782AE4AFFD8FF9998EF81BEFD0B60D1" attach="left" box="[739,751,655,676]" fontSize="9" pageId="6" pageNumber="7">5</subScript>
ending at the wing apex, males with cercopod longer than epandrium, bearing only one simple (but not considerably long) retinaculum, gonocoxites separated from each other, and female genitalia with long, gradually tapered cercus and without digit (
<bibRefCitation id="EF97D1FEFFD8FF9998238030FD36616E" author="Vaillant, F." box="[559,722,769,795]" pageId="6" pageNumber="7" pagination="109 - 142" refId="ref4033" refString="Vaillant, F. (1974) Psychodidae Psychodinae. In: Linder, E. (Ed): Die Fliegen der Palearktischen Region, 9 d. Sttutgart: E. Schweizerbartsche Verlangsbuchhandlung, pp. 109 - 142." type="book chapter" year="1974">Vaillant 1974</bibRefCitation>
). It is important to note that
<taxonomicName id="4C06D78CFFD8FF999E178030FB16616E" box="[1051,1266,769,795]" class="Insecta" family="Psychodidae" genus="Philosepedon" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B972701DFFD8FF999E178030FB5F616F" box="[1051,1211,769,794]" italics="true" pageId="6" pageNumber="7">Philosepedon</emphasis>
spp.
</taxonomicName>
may have two or two groups of cercopodial retinacula in the male and the cercus short in the female.
<taxonomicName id="4C06D78CFFD8FF999B6B8060FDF8611E" box="[359,540,849,875]" class="Insecta" family="Psychodidae" genus="Eurygarka" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B972701DFFD8FF999B6B8060FE00611F" box="[359,484,849,874]" italics="true" pageId="6" pageNumber="7">Eurygarka</emphasis>
spp.
</taxonomicName>
are characterized by the foliar sulcate branches of the “V”­shaped antennal ascoids.
<taxonomicName id="4C06D78CFFD8FF999BDB8048FD9961E6" box="[471,637,889,915]" class="Insecta" family="Psychodidae" genus="Quatiella" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B972701DFFD8FF999BDB8048FDA261E7" box="[471,582,889,914]" italics="true" pageId="6" pageNumber="7">Quatiella</emphasis>
spp.
</taxonomicName>
have the gonocoxites contiguous and epandrium considerably longer than wide in the male, and short cerci in the female.
<taxonomicName id="4C06D78CFFD8FF9999F78090FB4461CE" box="[1019,1184,929,955]" class="Insecta" family="Psychodidae" genus="Threticus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B972701DFFD8FF9999F78090FB8D61CF" box="[1019,1129,929,954]" italics="true" pageId="6" pageNumber="7">Threticus</emphasis>
spp.
</taxonomicName>
have 3 or 3 groups of cercopodial retinacula and the aedeagus asymmetric.
<taxonomicName id="4C06D78CFFD8FF999E0B80F8FB186196" box="[1031,1276,969,995]" class="Insecta" family="Psychodidae" genus="Trichopsychoda" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B972701DFFD8FF999E0B80F8FB266197" box="[1031,1218,969,994]" italics="true" pageId="6" pageNumber="7">Trichopsychoda</emphasis>
spp.
</taxonomicName>
are characterized by the long, thin retinacula finishing as a bud or bell­shape, with the wing membrane pilose.
<taxonomicName id="4C06D78CFFD8FF999BEE8728FD436646" box="[482,679,1049,1075]" class="Insecta" family="Psychodidae" genus="Nielseniella" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B972701DFFD8FF999BEE8728FD8A6647" box="[482,622,1049,1074]" italics="true" pageId="6" pageNumber="7">Nielseniella</emphasis>
spp.
</taxonomicName>
have antennal ascoids with only two branches basally directed and an asymmetric aedeagus.
</paragraph>
<paragraph id="8BB9AC0FFFD8FF999B348758FE246756" blockId="6.[264,1324,284,1875]" pageId="6" pageNumber="7">
Previous to this work, only 4 or 5 species had been considered to belong to
<emphasis id="B972701DFFD8FF999EA88758FEA266DF" italics="true" pageId="6" pageNumber="7">
<taxonomicName id="4C06D78CFFD8FF999EA88758FEDB66DF" authorityName="Vaillant" authorityYear="1974" class="Insecta" family="Psychodidae" genus="Feuerborniella" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">Feuerborniella</taxonomicName>
:
</emphasis>
<taxonomicName id="4C06D78CFFD8FF999B5E87A0FDDA66DE" authority="(Tonnoir)" baseAuthorityName="Tonnoir" box="[338,574,1169,1195]" class="Insecta" family="Psychodidae" genus="Feuerborniella" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="oscura">
<emphasis id="B972701DFFD8FF999B5E87A0FE2766DF" box="[338,451,1169,1194]" italics="true" pageId="6" pageNumber="7">F. oscura</emphasis>
(Tonnoir)
</taxonomicName>
from the Paleactic region,
<taxonomicName id="4C06D78CFFD8FF99999E87A0FB3766DE" authority="(Duckhouse)" baseAuthorityName="Duckhouse" box="[914,1235,1169,1195]" class="Insecta" family="Psychodidae" genus="Psychoda" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="plaumanni">
<emphasis id="B972701DFFD8FF99999E87A0FBD466DF" box="[914,1072,1169,1194]" italics="true" pageId="6" pageNumber="7">F. plaumanni</emphasis>
(Duckhouse)
</taxonomicName>
and
<taxonomicName id="4C06D78CFFD8FF999F1987A0FDCB66A6" authority="(Duckhouse)" baseAuthorityName="Duckhouse" baseAuthorityYear="1968" class="Insecta" family="Psychodidae" genus="Psychoda" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="spathipenis">
<emphasis id="B972701DFFD8FF999F1987A0FE6966A7" italics="true" pageId="6" pageNumber="7">F. spathipenis</emphasis>
(Duckhouse)
</taxonomicName>
both from
<collectingCountry id="F311EC9FFFD8FF9998B08788FCE266A6" box="[700,774,1209,1235]" name="Brazil" pageId="6" pageNumber="7">Brazil</collectingCountry>
,
<taxonomicName id="4C06D78CFFD8FF99991A8788FBD166A6" authority="(Satchell)" baseAuthorityName="Satchell" box="[790,1077,1209,1235]" class="Insecta" family="Psychodidae" genus="Trichopsychoda" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="malayensis">
<emphasis id="B972701DFFD8FF99991A8788FC5D66A7" box="[790,953,1209,1234]" italics="true" pageId="6" pageNumber="7">F. malayensis</emphasis>
(Satchell)
</taxonomicName>
of Malaya (
<bibRefCitation id="EF97D1FEFFD8FF999EC18788FEA2668E" author="Vaillant, F." pageId="6" pageNumber="7" pagination="109 - 142" refId="ref4033" refString="Vaillant, F. (1974) Psychodidae Psychodinae. In: Linder, E. (Ed): Die Fliegen der Palearktischen Region, 9 d. Sttutgart: E. Schweizerbartsche Verlangsbuchhandlung, pp. 109 - 142." type="book chapter" year="1974">Vaillant 1974</bibRefCitation>
), and apparently another species known only from a single larva from
<collectingCountry id="F311EC9FFFD8FF999E8187D0FACE668E" box="[1165,1322,1249,1275]" name="United States of America" pageId="6" pageNumber="7">United States</collectingCountry>
(
<bibRefCitation id="EF97D1FEFFD8FF999B1C8638FE576756" author="Vaillant, F." box="[272,435,1289,1315]" pageId="6" pageNumber="7" pagination="117 - 125" refId="ref4075" refString="Vaillant, F. (1983) Some Nearctic Psychodidae Psychodinae of the tribe Telmatoscopini (Diptera). Annales de la Societe Entomologique de France (N. S.), 19 (1), 117 - 125." type="journal article" year="1983">Vaillant 1983</bibRefCitation>
).
</paragraph>
<paragraph id="8BB9AC0FFFD8FF999B348600FCE16416" blockId="6.[264,1324,284,1875]" pageId="6" pageNumber="7">
The male of
<taxonomicName id="4C06D78CFFD8FF999BC78600FCF0673F" authority="Ibáñez-Bernal, 2004" authorityName="Ibáñez-Bernal" authorityYear="2004" box="[459,788,1329,1354]" class="Insecta" family="Psychodidae" genus="Feuerborniella" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="veracruzana" status="sp. nov.">
<emphasis id="B972701DFFD8FF999BC78600FCF0673F" box="[459,788,1329,1354]" italics="true" pageId="6" pageNumber="7">Feuerborniella veracruzana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A241CD66FFD8FF9999108600FC93673E" box="[796,887,1329,1355]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
is very similar to
<taxonomicName id="4C06D78CFFD8FF999E448600FB06673F" box="[1096,1250,1329,1354]" class="Insecta" family="Psychodidae" genus="Psychoda" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="plaumanni">
<emphasis id="B972701DFFD8FF999E448600FB06673F" box="[1096,1250,1329,1354]" italics="true" pageId="6" pageNumber="7">F. plaumanni</emphasis>
</taxonomicName>
, sharing the pattern of head setae patches, the length of the cercopods, general shape of aedeagus tapered to a narrow rounded tip, divergent parameres beneath aedeagus connected to a broad plate above the aedeagus, and the shape of tergite 10. Nevertheless,
<taxonomicName id="4C06D78CFFD8FF999E828698FACE67B7" box="[1166,1322,1449,1474]" class="Insecta" family="Psychodidae" genus="Psychoda" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="plaumanni">
<emphasis id="B972701DFFD8FF999E828698FACE67B7" box="[1166,1322,1449,1474]" italics="true" pageId="6" pageNumber="7">F. plaumanni</emphasis>
</taxonomicName>
can be separated because it has the antennal flagellomeres 11–13 partially fused to each other, the aedeagal apodeme comparatively longer and thinner, the aedeagus much longer than the lateral shafts, and the gonostylus with long apical setae in the male. The female of this species is unknown (
<bibRefCitation id="EF97D1FEFFD8FF9998208578FD1C6416" author="Duckhouse, D. A." box="[556,760,1609,1635]" pageId="6" pageNumber="7" pagination="29 - 40" refId="ref3518" refString="Duckhouse, D. A. (1968) Psychodidae (Diptera, Nematocera) collected by Mr. Plaumann in Southern Brazil. The Proceedings of the Royal Entomological Society of London (B), 37 (3 - 4), 29 - 40." type="journal article" year="1968">Duckhouse 1968</bibRefCitation>
).
</paragraph>
<paragraph id="8BB9AC0FFFD8FF999B348540FB7E6526" blockId="6.[264,1324,284,1875]" pageId="6" pageNumber="7">
<taxonomicName id="4C06D78CFFD8FF999B348540FD6064FF" authority="Ibáñez-Bernal, 2004" authorityName="Ibáñez-Bernal" authorityYear="2004" box="[312,644,1649,1674]" class="Insecta" family="Psychodidae" genus="Feuerborniella" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="veracruzana" status="sp. nov.">
<emphasis id="B972701DFFD8FF999B348540FD6064FF" box="[312,644,1649,1674]" italics="true" pageId="6" pageNumber="7">Feuerborniella veracruzana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A241CD66FFD8FF9998818540FD0F64FE" box="[653,747,1649,1675]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
, can be separated from the rest of the species in the genus:
<taxonomicName id="4C06D78CFFD8FF999B8A85A8FE1764C7" box="[390,499,1689,1714]" class="Insecta" family="Psychodidae" genus="Feuerborniella" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="oscura">
<emphasis id="B972701DFFD8FF999B8A85A8FE1764C7" box="[390,499,1689,1714]" italics="true" pageId="6" pageNumber="7">F. oscura</emphasis>
</taxonomicName>
has the cercopod as long as epandrium, the apical projection of tergite 10 is short and blunt, the aedeagus is not tapered and the lateral shafts are not curved ectally at the apex in the male, and there is an absence of two strong setae on the genital plate in the female;
<taxonomicName id="4C06D78CFFD8FF999BF88420FD7C655F" baseAuthorityName="Duckhouse" baseAuthorityYear="1968" box="[500,664,1809,1834]" class="Insecta" family="Psychodidae" genus="Psychoda" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="spathipenis">
<emphasis id="B972701DFFD8FF999BF88420FD7C655F" box="[500,664,1809,1834]" italics="true" pageId="6" pageNumber="7">F. spathipenis</emphasis>
</taxonomicName>
has two small projections at the vertex, the aedeagus is spoon­like, and the gonocoxite is nearly as long as the gonostylus in the male.
</paragraph>
</subSubSection>
</treatment>
</document>
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