237 lines
32 KiB
XML
237 lines
32 KiB
XML
<document id="15E0984A537E5D4EA4E152C96577F4FB" ID-DOI="10.26107/RBZ-2024-0008" ID-ISSN="2345-7600" ID-Zenodo-Dep="10941987" ID-ZooBank="F653A024-6EB9-43E3-9025-713FC5DE9805" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1712577800390" checkinUser="felipe" docAuthor="Kottelat, Maurice & Hui, Tan Heok" docDate="2024" docId="03AB4F27FFFF7B0AFB81F97E46DAFF68" docLanguage="en" docName="RafflesBZool.72.105-109.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Kapuasia Kottelat & Hui 2024, new genus" docType="treatment" docVersion="2" lastPageNumber="108" masterDocId="FF92375FFFFF7B09FF8AFFBE4507FFE2" masterDocTitle="Kapuasia, a genus name for ‘ Nemacheilus’ maculiceps (Teleostei: Nemacheilidae)" masterLastPageNumber="109" masterPageNumber="105" pageNumber="105" updateTime="1712578978962" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
|
||
<mods:mods id="BF73D41698AC47BF823E448BE89238C5" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="603A0168187FD3FE178903E2E0AEBA39">
|
||
<mods:title id="A633DDF3AE05CD4168F25996C400AE5D">Kapuasia, a genus name for ‘ Nemacheilus’ maculiceps (Teleostei: Nemacheilidae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="429F41C6D99C8D98799AD09CB484BF68" type="personal">
|
||
<mods:role id="DF7D6C6D990B548D79205C3CAE203BE6">
|
||
<mods:roleTerm id="BFA79341429A334960D8A11688F95A0C">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="2C49C98638C7A9684D9DA9242CD7F30C">Kottelat, Maurice</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="A8AF141DAC6188F5DD5CD85476600975" type="personal">
|
||
<mods:role id="43C41C09CB1B5B1ED9B56C5CBB7704E1">
|
||
<mods:roleTerm id="44A17BF8A8F34A9B8359AFCC902FCC82">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="DE111689EAF48BFAE4064D9A69EFD157">Hui, Tan Heok</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="8AEF3C18BA27571CA741DF9402D6A810">text</mods:typeOfResource>
|
||
<mods:relatedItem id="33B2A833CDCDC8B11C33E78BEB7F09EF" type="host">
|
||
<mods:titleInfo id="43A588096C266D822D4AE0168D93F32A">
|
||
<mods:title id="628BCC791969CCBF4C7FF73AFDF41C9E">Raffles Bulletin of Zoology</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="9DAB038F4CAE01B98D605268614E3DDE">
|
||
<mods:date id="6998D68FEAE058B5684C708C52665BDE">2024</mods:date>
|
||
<mods:detail id="0C2112ED7818EDB7EC725C2CAD7AE7D1" type="pubDate">
|
||
<mods:number id="AC3BD5BCE7DD7F807881E4CA607898FB">2024-03-15</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="7BE20A81B527041B6C29040663F6BB84" type="volume">
|
||
<mods:number id="2E31EF7B73360FA6A6CCE842CD1D2594">72</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="3233BAB2588FC9471202DBACFCF92349" unit="page">
|
||
<mods:start id="F931CC7A5DB1FD6EE8D364044DE47147">105</mods:start>
|
||
<mods:end id="E5851533694FD7DDE6C9848243F9CCF2">109</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="5D237564AC24F3040C3D00A331C4B89D">journal article</mods:classification>
|
||
<mods:identifier id="E16EE898BEC0B5C22F7CDBD0769ED131" type="DOI">10.26107/RBZ-2024-0008</mods:identifier>
|
||
<mods:identifier id="FC0A3C7E4BC46B8D24AB171AF86C7A22" type="ISSN">2345-7600</mods:identifier>
|
||
<mods:identifier id="46B9EF4411791E8BE08EDB8A3EB1B3AC" type="Zenodo-Dep">10941987</mods:identifier>
|
||
<mods:identifier id="D2829124CDC7FE329C8AD4B9A11CA6B3" type="ZooBank">F653A024-6EB9-43E3-9025-713FC5DE9805</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03AB4F27FFFF7B0AFB81F97E46DAFF68" LSID="urn:lsid:plazi:treatment:03AB4F27FFFF7B0AFB81F97E46DAFF68" httpUri="http://treatment.plazi.org/id/03AB4F27FFFF7B0AFB81F97E46DAFF68" lastPageId="3" lastPageNumber="108" pageId="0" pageNumber="105">
|
||
<subSubSection id="C318ADBAFFFF7B09FB81F97E41FDF93A" box="[1035,1274,1728,1752]" pageId="0" pageNumber="105" type="nomenclature">
|
||
<paragraph id="8BBDFE31FFFF7B09FB81F97E41FDF93A" blockId="0.[1035,1274,1728,1752]" box="[1035,1274,1728,1752]" pageId="0" pageNumber="105">
|
||
<heading id="D0F5495DFFFF7B09FB81F97E41FDF93A" bold="true" box="[1035,1274,1728,1752]" centered="true" fontSize="10" level="2" pageId="0" pageNumber="105" reason="2">
|
||
<emphasis id="B9762223FFFF7B09FB81F97E41FDF93A" bold="true" box="[1035,1274,1728,1752]" pageId="0" pageNumber="105">
|
||
<taxonomicName id="4C0285B2FFFF7B09FB81F97E4174F93A" authority="Kottelat & Hui, 2024" authorityName="Kottelat & Hui" authorityYear="2024" box="[1035,1139,1728,1752]" family="Nemacheilidae" genus="Kapuasia" kingdom="Animalia" order="Cypriniformes" pageId="0" pageNumber="105" phylum="Chordata" rank="genus" status="gen. nov.">
|
||
<emphasis id="B9762223FFFF7B09FB81F97E4174F93A" bold="true" box="[1035,1139,1728,1752]" italics="true" pageId="0" pageNumber="105">Kapuasia</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A2459F58FFFF7B09FB08F97E41FDF93A" box="[1154,1274,1728,1752]" pageId="0" pageNumber="105" rank="genus">new genus</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C318ADBAFFFF7B09FCC9F8BE4091F8FA" box="[835,1430,1792,1816]" pageId="0" pageNumber="105" type="type_taxon">
|
||
<paragraph id="8BBDFE31FFFF7B09FCC9F8BE4091F8FA" blockId="0.[835,1430,1792,1816]" box="[835,1430,1792,1816]" pageId="0" pageNumber="105">
|
||
<emphasis id="B9762223FFFF7B09FCC9F8BE46DCF8FA" bold="true" box="[835,987,1792,1816]" pageId="0" pageNumber="105">
|
||
<typeStatus id="54B94093FFFF7B09FCC9F8BE467AF8FA" box="[835,893,1792,1816]" pageId="0" pageNumber="105">Type</typeStatus>
|
||
species.
|
||
</emphasis>
|
||
<taxonomicName id="4C0285B2FFFF7B09FC6EF8BE4095F8FA" ID-CoL="465VF" authority="Roberts, 1989" authorityName="Roberts" authorityYear="1989" box="[996,1426,1792,1816]" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="0" pageNumber="105" phylum="Chordata" rank="species" species="maculiceps">
|
||
<emphasis id="B9762223FFFF7B09FC6EF8BE41E9F8FA" box="[996,1262,1792,1816]" italics="true" pageId="0" pageNumber="105">Nemacheilus maculiceps</emphasis>
|
||
<bibRefCitation id="EF9383C0FFFF7B09FB7DF8BE4095F8FA" author="Roberts TR" box="[1271,1426,1792,1816]" pageId="0" pageNumber="105" pagination="1 - 210" refId="ref2887" refString="Roberts TR (1989) The freshwater fishes of western Borneo (Kalimantan Barat, Indonesia). Memoirs of the California Academy of Sciences, 14: 1 - 210." type="journal article" year="1989">Roberts, 1989</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C318ADBAFFFF7B09FCC9F8FE4106F87A" pageId="0" pageNumber="105" type="etymology">
|
||
<paragraph id="8BBDFE31FFFF7B09FCC9F8FE4106F87A" blockId="0.[835,1474,1856,1944]" pageId="0" pageNumber="105">
|
||
<emphasis id="B9762223FFFF7B09FCC9F8FE46C2F8BA" bold="true" box="[835,965,1856,1880]" pageId="0" pageNumber="105">Etymology.</emphasis>
|
||
Name derived from the Kapuas River in
|
||
<collectingRegion id="49C630D3FFFF7B09FA01F8FF46CFF89A" country="Indonesia" name="Kalimantan Barat" pageId="0" pageNumber="105">West Kalimantan</collectingRegion>
|
||
, where the
|
||
<typeStatus id="54B94093FFFF7B09FBD8F8DF4185F89B" box="[1106,1154,1889,1913]" pageId="0" pageNumber="105">type</typeStatus>
|
||
species was first collected. Gender feminine.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C318ADBAFFFF7B0BFCC9F87E4744F8D9" lastPageId="2" lastPageNumber="107" pageId="0" pageNumber="105" type="diagnosis">
|
||
<paragraph id="8BBDFE31FFFF7B08FCC9F87E473FF933" blockId="0.[835,1475,1984,2104]" lastBlockId="1.[113,753,1337,1745]" lastPageId="1" lastPageNumber="106" pageId="0" pageNumber="105">
|
||
<emphasis id="B9762223FFFF7B09FCC9F87E46BFF83A" bold="true" box="[835,952,1984,2008]" pageId="0" pageNumber="105">Diagnosis.</emphasis>
|
||
<taxonomicName id="4C0285B2FFFF7B09FC4AF87F4121F83A" authorityName="Kottelat & Hui" authorityYear="2024" box="[960,1062,1985,2008]" family="Nemacheilidae" genus="Kapuasia" kingdom="Animalia" order="Cypriniformes" pageId="0" pageNumber="105" phylum="Chordata" rank="genus">
|
||
<emphasis id="B9762223FFFF7B09FC4AF87F4121F83A" box="[960,1062,1985,2008]" italics="true" pageId="0" pageNumber="105">Kapuasia</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from all other genera of
|
||
<taxonomicName id="4C0285B2FFFF7B09FCECF85E410CF81A" box="[870,1035,2016,2040]" family="Nemacheilidae" kingdom="Animalia" order="Cypriniformes" pageId="0" pageNumber="105" phylum="Chordata" rank="family">Nemacheilidae</taxonomicName>
|
||
in the morphology of the mouth. The mouth is strongly arched, U-shaped (
|
||
<figureCitation id="1339E2B4FFFF7B09FB5AF7BE4019F7FA" box="[1232,1310,2048,2072]" captionStart="Fig" captionStartId="3.[113,148,2035,2057]" captionTargetBox="[147,719,1258,2020]" captionTargetId="figure-10@3.[147,719,1258,2020]" captionTargetPageId="3" captionText="Fig. 4. Kapuasia maculiceps; a, ZRC 61464, 64.3 mm SL; mouth; b, ZRC 65285, 86.6 mm SL; close up of anterior part of lower lip. Arrow: mental groove." figureDoi="http://doi.org/10.5281/zenodo.10941997" httpUri="https://zenodo.org/record/10941997/files/figure.png" pageId="0" pageNumber="105">Fig. 4a</figureCitation>
|
||
). The upper lip is swollen and smooth, the posterior edge is entire, without median incision, with a deep groove between the lip and the rostral cap. The edge of the rostral cap between the anterior rostral barbels is slightly crenulated. The processus dentiformis is present. The lower lip is swollen, with a deep groove (postlabial) along the lateral parts; its anterior edge is continuous, with a narrow shallow median longitudinal groove (mental) extending from the mental area to the anterior edge of the lip. The median part of the lower lip has 8–10 ridges on each side of the mental groove (
|
||
<figureCitation id="1339E2B4FFFE7B08FD4EF9874588F993" captionStart="Fig" captionStartId="3.[113,148,2035,2057]" captionTargetBox="[147,719,1258,2020]" captionTargetId="figure-10@3.[147,719,1258,2020]" captionTargetPageId="3" captionText="Fig. 4. Kapuasia maculiceps; a, ZRC 61464, 64.3 mm SL; mouth; b, ZRC 65285, 86.6 mm SL; close up of anterior part of lower lip. Arrow: mental groove." figureDoi="http://doi.org/10.5281/zenodo.10941997" httpUri="https://zenodo.org/record/10941997/files/figure.png" pageId="1" pageNumber="106">Fig. 4b</figureCitation>
|
||
), radiating from the anterior extremity of the postlabial groove, across the whole lip, resulting in a crenulated inner edge of the lip; in some specimens, there is a longitudinal row of black pigments along most ridges.
|
||
</paragraph>
|
||
<caption id="DF7DAEB9FFFE7B08FFFBFB524699FAE3" ID-DOI="http://doi.org/10.5281/zenodo.10941989" ID-Zenodo-Dep="10941989" box="[113,926,1260,1282]" httpUri="https://zenodo.org/record/10941989/files/figure.png" pageId="1" pageNumber="106" startId="1.[113,148,1260,1282]" targetBox="[113,1451,189,1245]" targetPageId="1" targetType="figure">
|
||
<paragraph id="8BBDFE31FFFE7B08FFFBFB524699FAE3" blockId="1.[113,926,1260,1282]" box="[113,926,1260,1282]" pageId="1" pageNumber="106">
|
||
Fig. 1.
|
||
<taxonomicName id="4C0285B2FFFE7B08FF30FB5344E7FAE0" authority=", Borneo" authorityName="Borneo" box="[186,480,1260,1282]" family="Nemacheilidae" genus="Kapuasia" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="106" phylum="Chordata" rank="species" species="maculiceps">
|
||
<emphasis id="B9762223FFFE7B08FF30FB53448FFAE3" box="[186,392,1260,1282]" italics="true" pageId="1" pageNumber="106">Kapuasia maculiceps</emphasis>
|
||
, Borneo
|
||
</taxonomicName>
|
||
: Kapuas drainage. ZRC 61464, 56.2 mm SL.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BBDFE31FFFE7B08FFFBF94441B4F8D3" blockId="1.[113,753,1785,2098]" lastBlockId="1.[813,1452,1337,1841]" pageId="1" pageNumber="106">
|
||
<taxonomicName id="4C0285B2FFFE7B08FFFBF94445D0F8F3" authorityName="Kottelat & Hui" authorityYear="2024" box="[113,215,1786,1809]" family="Nemacheilidae" genus="Kapuasia" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="106" phylum="Chordata" rank="genus">
|
||
<emphasis id="B9762223FFFE7B08FFFBF94445D0F8F3" box="[113,215,1786,1809]" italics="true" pageId="1" pageNumber="106">Kapuasia</emphasis>
|
||
</taxonomicName>
|
||
is also distinguished from all nemacheilid loaches known to us by the presence of a conspicuous suprapectoral flap above the pectoral fin (
|
||
<figureCitation id="1339E2B4FFFE7B08FE22F88744EAF8B3" box="[424,493,1849,1873]" captionStart="Fig" captionStartId="3.[812,848,1596,1618]" captionTargetBox="[812,1452,1258,1581]" captionTargetId="figure-41@3.[812,1452,1258,1581]" captionTargetPageId="3" captionText="Fig. 5. Kapuasia maculiceps, ZRC 61464, 64.3 mm SL; head, pectoral fin and suprapectoral flap (arrow)." figureDoi="http://doi.org/10.5281/zenodo.10941995" httpUri="https://zenodo.org/record/10941995/files/figure.png" pageId="1" pageNumber="106">Fig. 5</figureCitation>
|
||
). Additional characters useful to distinguish the genus (but none unique to it) are: a well-developed pelvic axillary lobe, extending along the complete base of the fin; a small ridge along both dorsal and ventral midlines on the posterior part of the caudal peduncle; the anus about midway between the base of the pelvic fin and the origin of the anal fin; the pelvic fin with 1 unbranched and 7 branched rays; the pectoral fin with 1 unbranched and 9 or 10 branched rays [Roberts, 1990: 108 reported 10–12 branched rays]; the anal fin with 3 unbranched and 5½ branched rays; the dorsal fin with 4 unbranched and 8½ branched rays, its distal margin straight to slightly concave; the caudal fin deeply forked, the tip of the lobes slightly rounded, with 9+8 branched rays; the body entirely covered by scales, including on the predorsal area and the throat; the scales are very small, embedded; the lateral line is complete, with the pored scales difficult to count, estimated around 120–130; the cephalic lateral line canals have 5 (or 3?) supratemporal, 6 supraorbital, 11+3 infraorbital, and 8–10 preoperculo-mandibular pores; the anterior nostril is on the anterior side of a flap-like tube; there are no enlarged scales along the caudal peduncle; the eye protrudes slightly above the dorsal profile of the head; and the interorbital area is concave.
|
||
</paragraph>
|
||
<paragraph id="8BBDFE31FFFE7B08FCA7F8E74072F833" blockId="1.[811,1452,1881,2001]" pageId="1" pageNumber="106">There is no black spot at the base of the anterior dorsal-fin rays or on their lower part, no ocellus at the upper extremity of the base of the caudal fin. There is a large, roughly triangular, black mark at the base of each lobe of the caudal fin.</paragraph>
|
||
<paragraph id="8BBDFE31FFFE7B0BFCA7F8474744F8D9" blockId="1.[813,1452,2041,2097]" lastBlockId="2.[135,777,1347,1851]" lastPageId="2" lastPageNumber="107" pageId="1" pageNumber="106">
|
||
The examined specimens have features for which sexual dimorphism has been documented in various nemacheilid genera. The pectoral-fin rays are hard and rigid, not curled. The unbranched and the three anterior branched rays are thicker than the remaining rays. The first branched pectoral-fin ray is branched only once, the branches are thick and the space between them narrow. In the second and third branched rays, the anterior branch is unbranched and thicker than the posterior one, with a membrane between the two branches; the posterior branch is branched and these sub-branches are adjacent, without or with only very narrow membrane in between. In the remaining branched rays, both branches are branched and there are membranes between the sub-branches. In the 64.3 mm SL specimen, there is a short row of short, small tubercles on the dorsal side of each branch of the first branched ray (
|
||
<figureCitation id="1339E2B4FFFD7B0BFEA8F95D4465F919" box="[290,354,1763,1787]" captionStart="Fig" captionStartId="3.[812,846,2064,2086]" captionTargetBox="[812,1452,1717,2049]" captionTargetId="figure-60@3.[812,1452,1717,2049]" captionTargetPageId="3" captionText="Fig. 6.Kapuasia maculiceps, ZRC 61464, 56.2. mm SL; pectoral fin, dorsal view; note tubercles near tip of first and second branched rays." figureDoi="http://doi.org/10.5281/zenodo.10941999" httpUri="https://zenodo.org/record/10941999/files/figure.png" pageId="2" pageNumber="107">Fig. 6</figureCitation>
|
||
) and much less developed tubercles on the second branched ray. There are no tubercles in the other specimens. No suborbital flap is present.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF7DAEB9FFFD7B0BFF0DFB4846B3FAE9" ID-DOI="http://doi.org/10.5281/zenodo.10941991" ID-Zenodo-Dep="10941991" box="[135,948,1270,1292]" httpUri="https://zenodo.org/record/10941991/files/figure.png" pageId="2" pageNumber="107" startId="2.[135,170,1270,1292]" targetBox="[135,1473,189,1255]" targetPageId="2" targetType="figure">
|
||
<paragraph id="8BBDFE31FFFD7B0BFF0DFB4846B3FAE9" blockId="2.[135,948,1270,1292]" box="[135,948,1270,1292]" pageId="2" pageNumber="107">
|
||
Fig. 2.
|
||
<taxonomicName id="4C0285B2FFFD7B0BFF5BFB4944F1FAEE" authority=", Borneo" authorityName="Borneo" box="[209,502,1270,1292]" family="Nemacheilidae" genus="Kapuasia" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="107" phylum="Chordata" rank="species" species="maculiceps">
|
||
<emphasis id="B9762223FFFD7B0BFF5BFB494499FAE9" box="[209,414,1270,1292]" italics="true" pageId="2" pageNumber="107">Kapuasia maculiceps</emphasis>
|
||
, Borneo
|
||
</taxonomicName>
|
||
: Kapuas drainage. ZRC 61464, 64.3 mm SL.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C318ADBAFFFD7B0BFF02F8DA4154F999" pageId="2" pageNumber="107" type="discussion">
|
||
<paragraph id="8BBDFE31FFFD7B0BFF02F8DA4154F999" blockId="2.[136,775,1891,2075]" lastBlockId="2.[835,1476,1347,1659]" pageId="2" pageNumber="107">
|
||
<emphasis id="B9762223FFFD7B0BFF02F8DA45F0F89E" bold="true" box="[136,247,1892,1916]" pageId="2" pageNumber="107">Remarks.</emphasis>
|
||
In a number of nemacheilid genera, the presence of tubercles on the dorsal side of the anterior pectoral-fin rays of the males, the pattern of their distribution, their size etc. are diagnostic features (e.g., see
|
||
<bibRefCitation id="EF9383C0FFFD7B0BFDE8F87D4604F83E" author="Kottelat M" box="[610,771,1987,2012]" pageId="2" pageNumber="107" pagination="289 - 314" refId="ref2662" refString="Kottelat M (2018) Mustura celata, a new genus and species of loaches from northern Myanmar, and an overview of Physoschistura and related taxa (Teleostei: Nemacheilidae). Ichthyological Exploration of Freshwaters, 28 (4): 289 - 314." type="journal article" year="2018">Kottelat, 2018</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF9383C0FFFD7B0BFF02F85A45C5F81E" author="Kottelat M" box="[136,194,2020,2044]" pageId="2" pageNumber="107" pagination="151 - 170" refId="ref2709" refString="Kottelat M (2019) Rhyacoschistura larreci, a new genus and species of loach from Laos and redescription of R. suber (Teleostei: Nemacheilidae). Zootaxa, 4612 (2): 151 - 170." type="journal article" year="2019">2019</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9383C0FFFD7B0BFF5BF85A44FCF81E" author="Conway KW & Kottelat M" box="[209,507,2019,2044]" pageId="2" pageNumber="107" pagination="681 - 701" refId="ref2145" refString="Conway KW & Kottelat M (2023) Physoschistura mango, a new miniature species of loach from Myanmar (Teleostei: Nemacheilidae). Raffles Bulletin of Zoology, 71: 681 - 701" type="journal article" year="2023">Conway & Kottelat, 2023</bibRefCitation>
|
||
). However, in males of some species their presence and development apparently vary with age, season, and sexual activity (MK, pers. obs.). Therefore, at this stage, we do not want to use the presence and shape of tubercles in a single specimen of
|
||
<taxonomicName id="4C0285B2FFFD7B0BFAB5FA3A40A2FA79" authorityName="Kottelat & Hui" authorityYear="2024" box="[1343,1445,1412,1435]" family="Nemacheilidae" genus="Kapuasia" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="107" phylum="Chordata" rank="genus">
|
||
<emphasis id="B9762223FFFD7B0BFAB5FA3A40A2FA79" box="[1343,1445,1412,1435]" italics="true" pageId="2" pageNumber="107">Kapuasia</emphasis>
|
||
</taxonomicName>
|
||
as a character usable to diagnose the genus. Also, sex has not been confirmed by dissection. In all nemacheilid genera in which the morphology of the pectoral fin has been observed or reported with enough details, in the female, all branched rays are similarly slender and the branches of branched rays are all separated by membranes. This was not observed in the examined specimens.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C318ADBAFFFD7B0AFCCFF91D46DAFF68" lastPageId="3" lastPageNumber="108" pageId="2" pageNumber="107" type="materials_examined">
|
||
<paragraph id="8BBDFE31FFFD7B0BFCCFF91D40C4F7F9" blockId="2.[834,1477,1699,2076]" pageId="2" pageNumber="107">
|
||
<emphasis id="B9762223FFFD7B0BFCCFF91D413BF959" bold="true" box="[837,1084,1699,1723]" pageId="2" pageNumber="107">Material examined.</emphasis>
|
||
<taxonomicName id="4C0285B2FFFD7B0BFBCCF91A4043F959" box="[1094,1348,1699,1723]" family="Nemacheilidae" genus="Kapuasia" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="107" phylum="Chordata" rank="species" species="maculiceps">
|
||
<emphasis id="B9762223FFFD7B0BFBCCF91A4043F959" box="[1094,1348,1699,1723]" italics="true" pageId="2" pageNumber="107">Kapuasia maculiceps</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<materialsCitation id="3B6AF46CFFFD7B0BFAD0F91D40B9F7F9" collectingDate="1976-07-24" collectionCode="ZRC, MZB, BMNH, RMNH" collectorName="Mempawah R. & Sungai Pinoh & T. R. Roberts" country="Indonesia" county="All" latitude="-0.65833336" location="Kapuas" longLatPrecision="930" longitude="111.666664" municipality="Borneo" pageId="2" pageNumber="107" specimenCode="ZRC 56404, 1, 69.7, ZRC 61464, 2, 56.2, 64.3, ZRC 65285, 86.6, MZB 3543, BMNH 1982.3, RMNH 28878" specimenCount="1" stateProvince="Kalimantan Barat" typeStatus="holotype">
|
||
<collectingCounty id="62DC86BDFFFD7B0BFAD0F91D4078F959" box="[1370,1407,1699,1723]" pageId="2" pageNumber="107">All</collectingCounty>
|
||
from
|
||
<collectingCountry id="F315BEA1FFFD7B0BFCC9F97D46AAF939" box="[835,941,1731,1755]" name="Indonesia" pageId="2" pageNumber="107">Indonesia</collectingCountry>
|
||
:
|
||
<collectingMunicipality id="6BD9644BFFFD7B0BFC3CF97A410FF939" box="[950,1032,1732,1755]" pageId="2" pageNumber="107">Borneo</collectingMunicipality>
|
||
:
|
||
<collectingRegion id="49C630D3FFFD7B0BFB85F97D41CBF939" box="[1039,1228,1731,1755]" country="Indonesia" name="Kalimantan Barat" pageId="2" pageNumber="107">Kalimantan Barat</collectingRegion>
|
||
:
|
||
<location id="8EDDA8EAFFFD7B0BFB5EF97A4024F939" LSID="urn:lsid:plazi:treatment:03AB4F27FFFF7B0AFB81F97E46DAFF68:8EDDA8EAFFFD7B0BFB5EF97A4024F939" box="[1236,1315,1732,1755]" country="Indonesia" county="All" latitude="-0.65833336" longLatPrecision="930" longitude="111.666664" municipality="Borneo" name="Kapuas" pageId="2" pageNumber="107" stateProvince="Kalimantan Barat">Kapuas</location>
|
||
drainage:
|
||
<specimenCode id="DBA4564AFFFD7B0BFA05F97A4680F919" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="107">ZRC 56404</specimenCode>
|
||
,
|
||
<specimenCode id="DBA4564AFFFD7B0BFC1AF95A469AF91E" box="[912,925,1764,1788]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="107">1</specimenCode>
|
||
,
|
||
<specimenCode id="DBA4564AFFFD7B0BFC23F95D46DFF919" box="[937,984,1763,1787]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="107">69.7</specimenCode>
|
||
mm SL;
|
||
<location id="8EDDA8EAFFFD7B0BFBB2F95D41E9F919" LSID="urn:lsid:plazi:treatment:03AB4F27FFFF7B0AFB81F97E46DAFF68:8EDDA8EAFFFD7B0BFBB2F95D41E9F919" box="[1080,1262,1763,1787]" country="Indonesia" county="All" latitude="-0.65833336" longLatPrecision="930" longitude="111.666664" municipality="Borneo" name="Bengkayang area" pageId="2" pageNumber="107" stateProvince="Kalimantan Barat">Bengkayang area</location>
|
||
(
|
||
<geoCoordinate id="EE3698F6FFFD7B0BFB71F95C404FF919" box="[1275,1352,1762,1787]" degrees="0" direction="north" minutes="49" orientation="latitude" pageId="2" pageNumber="107" precision="925" value="0.81666666">0°49′N</geoCoordinate>
|
||
<geoCoordinate id="EE3698F6FFFD7B0BFAC7F95C40B2F919" box="[1357,1461,1762,1787]" degrees="109" direction="east" minutes="29" orientation="longitude" pageId="2" pageNumber="107" precision="925" value="109.48333">109°29′E</geoCoordinate>
|
||
); aquarium-fish trade. —
|
||
<specimenCode id="DBA4564AFFFD7B0BFBC3F8BA41CCF8F9" box="[1097,1227,1795,1820]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="107">ZRC 61464</specimenCode>
|
||
,
|
||
<specimenCode id="DBA4564AFFFD7B0BFB52F8BA41E0F8FE" box="[1240,1255,1796,1820]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="107">2</specimenCode>
|
||
,
|
||
<specimenCode id="DBA4564AFFFD7B0BFB7FF8BD4023F8F9" box="[1269,1316,1795,1819]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="107">56.2</specimenCode>
|
||
–
|
||
<specimenCode id="DBA4564AFFFD7B0BFAB8F8BD4067F8F9" box="[1330,1376,1795,1819]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="107">64.3</specimenCode>
|
||
mm SL; Kec. Banyuke,
|
||
<location id="8EDDA8EAFFFD7B0BFC61F89D4168F8D9" LSID="urn:lsid:plazi:treatment:03AB4F27FFFF7B0AFB81F97E46DAFF68:8EDDA8EAFFFD7B0BFC61F89D4168F8D9" box="[1003,1135,1827,1851]" country="Indonesia" county="All" latitude="-0.65833336" longLatPrecision="930" longitude="111.666664" municipality="Borneo" name="Ulu Landak" pageId="2" pageNumber="107" stateProvince="Kalimantan Barat">Ulu Landak</location>
|
||
,
|
||
<location id="8EDDA8EAFFFD7B0BFBF1F89D4066F8D9" LSID="urn:lsid:plazi:treatment:03AB4F27FFFF7B0AFB81F97E46DAFF68:8EDDA8EAFFFD7B0BFBF1F89D4066F8D9" box="[1147,1377,1827,1851]" country="Indonesia" county="All" latitude="-0.65833336" longLatPrecision="930" longitude="111.666664" municipality="Borneo" name="Lokasi Desa Kampet" pageId="2" pageNumber="107" stateProvince="Kalimantan Barat">Lokasi Desa Kampet</location>
|
||
,
|
||
<location id="8EDDA8EAFFFD7B0BFAE4F89A4687F8B9" LSID="urn:lsid:plazi:treatment:03AB4F27FFFF7B0AFB81F97E46DAFF68:8EDDA8EAFFFD7B0BFAE4F89A4687F8B9" country="Indonesia" county="All" latitude="-0.65833336" longLatPrecision="930" longitude="111.666664" municipality="Borneo" name="Kampet River" pageId="2" pageNumber="107" stateProvince="Kalimantan Barat">Kampet River</location>
|
||
(flowing into
|
||
<collectorName id="26F79BE7FFFD7B0BFB90F8FD41BEF8BE" box="[1050,1209,1859,1884]" pageId="2" pageNumber="107">Mempawah R.</collectorName>
|
||
), near
|
||
<location id="8EDDA8EAFFFD7B0BFA8EF8FA40C2F8B9" LSID="urn:lsid:plazi:treatment:03AB4F27FFFF7B0AFB81F97E46DAFF68:8EDDA8EAFFFD7B0BFA8EF8FA40C2F8B9" box="[1284,1477,1859,1883]" country="Indonesia" county="All" latitude="-0.65833336" longLatPrecision="930" longitude="111.666664" municipality="Borneo" name="Mt. Marabukatan." pageId="2" pageNumber="107" stateProvince="Kalimantan Barat">Mt. Marabukatan.</location>
|
||
—
|
||
<specimenCode id="DBA4564AFFFD7B0BFCE0F8DA46F1F899" box="[874,1014,1891,1916]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="107">ZRC 65285</specimenCode>
|
||
,
|
||
<specimenCode id="DBA4564AFFFD7B0BFB8CF8DA4113F89E" box="[1030,1044,1892,1916]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="107">1</specimenCode>
|
||
,
|
||
<specimenCode id="DBA4564AFFFD7B0BFBADF8DD415EF899" box="[1063,1113,1891,1915]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="107">86.6</specimenCode>
|
||
mm SL; Putussibau. —
|
||
<specimenCode id="DBA4564AFFFD7B0BFA0CF8DA467AF879" collectionCode="MZB" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="2" pageNumber="107" type="Museum">MZB 3543</specimenCode>
|
||
,
|
||
<typeStatus id="54B94093FFFD7B0BFC06F83D46E8F879" box="[908,1007,1923,1947]" pageId="2" pageNumber="107" type="holotype">holotype</typeStatus>
|
||
, 78.9 mm SL (photograph only);
|
||
<specimenCode id="DBA4564AFFFD7B0BFAF8F83A4688F859" collectionCode="BMNH" pageId="2" pageNumber="107">BMNH 1982.3</specimenCode>
|
||
.29.135,
|
||
<specimenCount id="9D0435B8FFFD7B0BFC61F81A4158F85E" box="[1003,1119,1956,1980]" count="1" pageId="2" pageNumber="107" type="generic" typeStatus="paratype">1 paratype</specimenCount>
|
||
, 72.0 mm SL (photograph only);
|
||
<specimenCode id="DBA4564AFFFD7B0BFCC9F87A46E3F83E" box="[835,996,1988,2012]" collectionCode="RMNH" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="2" pageNumber="107" type="Museum">RMNH 28878</specimenCode>
|
||
,
|
||
<specimenCode id="DBA4564AFFFD7B0BFC79F87A4106F83E" box="[1011,1025,1988,2012]" collectionCode="RMNH" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="2" pageNumber="107" type="Museum">1</specimenCode>
|
||
, 79.0 mm SL (photograph only); rocky channel in mainstream of
|
||
<collectorName id="26F79BE7FFFD7B0BFBE7F85D4002F819" box="[1133,1285,2019,2043]" pageId="2" pageNumber="107">Sungai Pinoh</collectorName>
|
||
,
|
||
<quantity id="4CFA53D4FFFD7B0BFA9EF85A405BF819" box="[1300,1372,2019,2044]" metricMagnitude="4" metricUnit="m" metricValue="3.7" pageId="2" pageNumber="107" unit="km" value="37.0">37 km</quantity>
|
||
south of
|
||
<location id="8EDDA8EAFFFD7B0BFCCEF7BD46CFF7F9" LSID="urn:lsid:plazi:treatment:03AB4F27FFFF7B0AFB81F97E46DAFF68:8EDDA8EAFFFD7B0BFCCEF7BD46CFF7F9" box="[836,968,2051,2075]" country="Indonesia" county="All" latitude="-0.65833336" longLatPrecision="930" longitude="111.666664" municipality="Borneo" name="Nangapinoh" pageId="2" pageNumber="107" stateProvince="Kalimantan Barat">Nangapinoh</location>
|
||
,
|
||
<geoCoordinate id="EE3698F6FFFD7B0BFC5AF7BC412DF7F9" box="[976,1066,2050,2075]" degrees="0" direction="south" minutes="39.5" orientation="latitude" pageId="2" pageNumber="107" precision="92" value="-0.65833336">0°39.5′S</geoCoordinate>
|
||
<geoCoordinate id="EE3698F6FFFD7B0BFBBAF7BC4192F7F9" box="[1072,1173,2050,2075]" degrees="111" direction="east" minutes="40" orientation="longitude" pageId="2" pageNumber="107" precision="925" value="111.666664">111°40′E</geoCoordinate>
|
||
;
|
||
<collectorName id="26F79BE7FFFD7B0BFB15F7BA402AF7F9" box="[1183,1325,2051,2075]" pageId="2" pageNumber="107">T. R. Roberts</collectorName>
|
||
,
|
||
<date id="FFBCD8F1FFFD7B0BFABCF7BA40B9F7F9" box="[1334,1470,2051,2076]" pageId="2" pageNumber="107" value="1976-07-24">
|
||
<collectingDate id="EFF82119FFFD7B0BFABCF7BA40B9F7F9" box="[1334,1470,2051,2076]" pageId="2" pageNumber="107" value="1976-07-24">24 July 1976</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BBDFE31FFFC7B0AFDB5FFCC46DAFF68" blockId="3.[575,989,114,138]" box="[575,989,114,138]" pageId="3" pageNumber="108">
|
||
Kottelat & Tan:
|
||
<taxonomicName id="4C0285B2FFFC7B0AFD7DFFCC465CFF6B" authority="Kottelat & Hui, 2024" authorityName="Kottelat & Hui" authorityYear="2024" box="[759,859,114,137]" family="Nemacheilidae" genus="Kapuasia" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="108" phylum="Chordata" rank="genus" status="gen. nov.">
|
||
<emphasis id="B9762223FFFC7B0AFD7DFFCC465CFF6B" box="[759,859,114,137]" italics="true" pageId="3" pageNumber="108">Kapuasia</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A2459F58FFFC7B0AFCE0FFCC46DAFF68" box="[874,989,114,138]" pageId="3" pageNumber="108" rank="genus">new genus</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |