263 lines
30 KiB
XML
263 lines
30 KiB
XML
<document id="7BF1EFF3CBAC832AFA9CF854605EF7FA" ID-DOI="10.11646/phytotaxa.252.1.2" ID-ISSN="1179-3163" ID-Zenodo-Dep="13674915" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725405123220" checkinUser="felipe" docAuthor="Sang, Julia & Kiew, Ruth" docDate="2016" docId="03A8A444FF83C465FF08F8E9FD5DFB0F" docLanguage="en" docName="phytotaxa.252.1.2.pdf" docOrigin="Phytotaxa 252 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.252.1.2" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Begonia jenginensis S. Julia & Kiew 2016, spec. nov." docType="treatment" docVersion="2" lastPageNumber="28" masterDocId="FF91DC3CFF8AC46EFF80FFDBFFE9FFA6" masterDocTitle="Begonia (Begoniaceae) from Batang Ai National Park and vicinity, Sarawak, Borneo, including six new species" masterLastPageNumber="30" masterPageNumber="17" pageNumber="26" updateTime="1725426159086" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:mods id="B7A7D597E6BBC25B0270B30229AD75B6" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="7691F6A65B22F867AC6EE65B76CC2068">
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<mods:title id="2F86D1114607EA7EE8D2CAD4F98B2078">Begonia (Begoniaceae) from Batang Ai National Park and vicinity, Sarawak, Borneo, including six new species</mods:title>
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</mods:titleInfo>
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<mods:name id="DD2D3FBD9CB1888070F9F1F0A3621184" type="personal">
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<mods:role id="A04D32A516D9324EB935F2594AD02179">
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<mods:roleTerm id="F94F5B31ECB0274EF4271617CCF2FEC3">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="3A29C1D620BAABC15B02F793928EF1E5">Sang, Julia</mods:namePart>
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<mods:nameIdentifier id="F84803603DB97AD43862176F53EEFDF8" type="email">juliasang@sarawakforestry.com</mods:nameIdentifier>
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</mods:name>
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<mods:name id="F41EA57D69534FA68582FA5B335B0AE6" type="personal">
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<mods:role id="22778B646C32AA9744104B70CA0DEB60">
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<mods:roleTerm id="41BB14ADD8BA7B9D20E7D4C68FD687E3">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="ABAFD9C2C9A105795667DD3719F19753">Kiew, Ruth</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="E2754A6BCA02210537379F9A09AC14EE">text</mods:typeOfResource>
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<mods:titleInfo id="7BA4193E210DC9DA361B7FD6162E9E6F">
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<mods:title id="1E352325732A64F3E0D4A680343E2F0F">Phytotaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="EE27BB8ABEAAD88E82922A50D15A76F9">
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<mods:date id="D00DCD10D2537ECE8A8B6DA889A78772">2016</mods:date>
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<mods:detail id="98AABDCB6BEBA7D39F88CA69B875E860" type="pubDate">
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<mods:number id="D2D1B6C93385E71F4B99FA853E40A6F1">2016-03-08</mods:number>
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</mods:detail>
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<mods:detail id="F78C1F9B95CE6F53685238D2752464F3" type="volume">
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<mods:number id="BF9B35AD113918C120B2658E12A721E2">252</mods:number>
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</mods:detail>
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<mods:detail id="5373088E72171268200CAF63E20CCDF9" type="issue">
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<mods:number id="C5626D8029EAC85F4FA6DEC7B6A9B6D1">1</mods:number>
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</mods:detail>
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<mods:extent id="6F7AADE566823DBB56DB670EB1EC8457" unit="page">
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<mods:start id="8558CD6BCDC8F375EBF5CD98FD02CC21">17</mods:start>
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<mods:end id="68074610CB2C78E3E616CCDB624C3870">30</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location id="1074014A2C24EDC3EACED63EC727DF65">
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<mods:url id="33482709613F10EE84513B04629FAE37">http://dx.doi.org/10.11646/phytotaxa.252.1.2</mods:url>
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</mods:location>
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<mods:classification id="971C144C19A0F232676151C3B214BB6E">journal article</mods:classification>
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<mods:identifier id="744E6C8116F100E5B24C631898F6EA76" type="DOI">10.11646/phytotaxa.252.1.2</mods:identifier>
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<mods:identifier id="9BFAC1EF0829FC33F8AC0599CD158A0A" type="ISSN">1179-3163</mods:identifier>
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<mods:identifier id="91D3C2E0F1B13D3F01C3D63FAD2D7E47" type="Zenodo-Dep">13674915</mods:identifier>
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</mods:mods>
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<treatment id="03A8A444FF83C465FF08F8E9FD5DFB0F" LSID="urn:lsid:plazi:treatment:03A8A444FF83C465FF08F8E9FD5DFB0F" httpUri="http://treatment.plazi.org/id/03A8A444FF83C465FF08F8E9FD5DFB0F" lastPageId="11" lastPageNumber="28" pageId="9" pageNumber="26">
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<subSubSection id="C31B46D9FF83C467FF08F8E9FCDBF8EB" box="[136,818,1842,1869]" pageId="9" pageNumber="26" type="nomenclature">
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<paragraph id="8BBE1552FF83C467FF08F8E9FCDBF8EB" blockId="9.[136,818,1842,1869]" box="[136,818,1842,1869]" pageId="9" pageNumber="26">
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<heading id="D0F6A23EFF83C467FF08F8E9FCDBF8EB" box="[136,818,1842,1869]" fontSize="11" level="2" pageId="9" pageNumber="26" reason="3">
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<emphasis id="B975C940FF83C467FF08F8E9FF74F8EA" bold="true" box="[136,157,1842,1868]" italics="true" pageId="9" pageNumber="26">8.</emphasis>
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<taxonomicName id="4C016ED1FF83C467FF24F8E9FDABF8EB" ID-CoL="7SG2W" authority="S.Julia & Kiew" authorityName="S. Julia & Kiew" authorityYear="2016" box="[164,578,1842,1869]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="9" pageNumber="26" phylum="Tracheophyta" rank="species" species="jenginensis" status="sp. nov.">
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<emphasis id="B975C940FF83C467FF24F8E9FE66F8EA" bold="true" box="[164,399,1842,1868]" italics="true" pageId="9" pageNumber="26">Begonia jenginensis</emphasis>
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S.Julia & Kiew
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</taxonomicName>
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,
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<emphasis id="B975C940FF83C467FDCDF8EFFD52F8EB" box="[589,699,1844,1869]" italics="true" pageId="9" pageNumber="26">
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<taxonomicNameLabel id="A246743BFF83C467FDCDF8EFFD52F8EB" box="[589,699,1844,1869]" pageId="9" pageNumber="26" rank="species">spec. nov.</taxonomicNameLabel>
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||
</emphasis>
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||
(
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<figureCitation id="133A09D7FF83C467FD49F8EFFCC3F8EB" box="[713,810,1844,1869]" captionStart="FIGURE 6" captionStartId="10.[136,229,1868,1889]" captionTargetBox="[317,1270,320,1832]" captionTargetId="figure-77@10.[310,1278,314,1842]" captionTargetPageId="10" captionText="FIGURE 6. Begonia jenginensis. A. Habit; B. Stipule; C. Fruits and male inflorescence; D. Male inflorescence; E. Male flower with one inner tepal missing; F. Female flower; G. Female flower showing pistil; H. Fruit; I. Cross section of ovary.All photos from SFC 2750 by Julia Sang." figureDoi="http://doi.org/10.5281/zenodo.13674933" httpUri="https://zenodo.org/record/13674933/files/figure.png" pageId="9" pageNumber="26">Figure 6</figureCitation>
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)
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</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C31B46D9FF83C467FF08F883FE9EF8D7" box="[136,375,1880,1905]" pageId="9" pageNumber="26" type="description">
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||
<paragraph id="8BBE1552FF83C467FF08F883FE9EF8D7" blockId="9.[136,375,1880,1905]" box="[136,375,1880,1905]" pageId="9" pageNumber="26">
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||
<emphasis id="B975C940FF83C467FF08F883FF36F8D7" bold="true" box="[136,223,1880,1905]" pageId="9" pageNumber="26">Section</emphasis>
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<taxonomicName id="4C016ED1FF83C467FF66F883FE9EF8D7" box="[230,375,1880,1905]" class="Magnoliopsida" family="Begoniaceae" genus="Petermannia" kingdom="Plantae" order="Cucurbitales" pageId="9" pageNumber="26" phylum="Tracheophyta" rank="genus">
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<emphasis id="B975C940FF83C467FF66F883FE9EF8D7" box="[230,375,1880,1905]" italics="true" pageId="9" pageNumber="26">Petermannia</emphasis>
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</taxonomicName>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C31B46D9FF83C464FF08F878FE04FF7F" lastPageId="10" lastPageNumber="27" pageId="9" pageNumber="26" type="diagnosis">
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<paragraph id="8BBE1552FF83C464FF08F878FE04FF7F" blockId="9.[136,1452,1955,2084]" lastBlockId="10.[136,1451,160,289]" lastPageId="10" lastPageNumber="27" pageId="9" pageNumber="26">
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<emphasis id="B975C940FF83C467FF08F878FF1AF81E" bold="true" box="[136,243,1955,1976]" pageId="9" pageNumber="26">Diagnosis:</emphasis>
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—Similar to
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<taxonomicName id="4C016ED1FF83C467FEF6F878FD32F81E" authority="Sands (1997: 432)" authorityName="Sands" authorityPageNumber="432" authorityYear="1997" box="[374,731,1955,1976]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="9" pageNumber="26" phylum="Tracheophyta" rank="species" species="awongii">
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<emphasis id="B975C940FF83C467FEF6F878FDF5F81E" box="[374,540,1955,1976]" italics="true" pageId="9" pageNumber="26">Begonia awongii</emphasis>
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<bibRefCitation id="EF9068A3FF83C467FDA5F878FD32F81E" author="Sands, M. S." box="[549,731,1955,1976]" pageId="9" pageNumber="26" refId="ref9572" refString="Sands, M. S. (1997 ' 1996 ') In: Coode, M. J. E., Dransfield, J., Forman, L. L., Kirkup, D. W. & Idris, M. S. (Eds.) A Checklist of Flowering Plants and Gymnosperms of Brunei Darussalam. Brunei Forest Department and The Royal Botanic Gardens, Kew, p. 432." type="journal volume" year="1997">Sands (1997: 432)</bibRefCitation>
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</taxonomicName>
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in its habit, hispid inflorescence, leaf shape and racemose inflorescence but differs in the shallowly cordate base with a basal lobe
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<quantity id="4CF9B8B7FF83C467FD72F81CFCD0F87A" box="[754,825,1991,2012]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="9" pageNumber="26" unit="cm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 cm</quantity>
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long (vs. deeply cordate with a basal lobe
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<quantity id="4CF9B8B7FF83C467FB5AF81CFAACF87A" box="[1242,1349,1991,2012]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" metricValueMax="8.5" metricValueMin="5.5" pageId="9" pageNumber="26" unit="cm" value="7.0" valueMax="8.5" valueMin="5.5">5.5–8.5 cm</quantity>
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long in
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<taxonomicName id="4C016ED1FF83C467FA17F81CFEE3F7A6" authorityName="Sands" authorityYear="1997" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="9" pageNumber="26" phylum="Tracheophyta" rank="species" species="awongii">
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<emphasis id="B975C940FF83C467FA17F81CFEE3F7A6" italics="true" pageId="9" pageNumber="26">B. awongii</emphasis>
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</taxonomicName>
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||
), male flowers with pale pink male tepals and fewer (28–30) but larger stamens with filaments ca.
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<quantity id="4CF9B8B7FF83C467FB33F830FB00F7A6" box="[1203,1257,2027,2048]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="26" unit="mm" value="2.0">2 mm</quantity>
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||
long and anthers ca.
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||
<quantity id="4CF9B8B7FF83C467FF3DF7D4FEEEF782" box="[189,263,2063,2084]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="26" unit="mm" value="1.5">1.5 mm</quantity>
|
||
long (vs. white tepals and ca. 46 subsessile stamens with filaments ca.
|
||
<quantity id="4CF9B8B7FF83C467FC35F7D4FC17F782" box="[949,1022,2063,2084]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="26" unit="mm" value="0.5">0.5 mm</quantity>
|
||
long and anthers ca.
|
||
<quantity id="4CF9B8B7FF83C467FB4BF7D4FAC9F782" box="[1227,1312,2063,2084]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="9" pageNumber="26" unit="mm" value="0.25">0.25 mm</quantity>
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||
long), smaller tepals in the female flower, 6–7 ×
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||
<quantity id="4CF9B8B7FF80C464FD90FF7BFD88FF13" box="[528,609,160,181]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="10" pageNumber="27" unit="mm" value="5.0" valueMax="6.0" valueMin="4.0">4–6 mm</quantity>
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||
(vs. 12–14 ×
|
||
<quantity id="4CF9B8B7FF80C464FD6CFF7BFCD6FF13" box="[748,831,160,181]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="10" pageNumber="27" unit="mm" value="8.0" valueMax="9.0" valueMin="7.0">7–9 mm</quantity>
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||
) and larger glabrous capsules, 12–14 ×
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||
<quantity id="4CF9B8B7FF80C464FB47FF7BFAD9FF13" box="[1223,1328,160,181]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" metricValueMax="2.4" metricValueMin="1.4" pageId="10" pageNumber="27" unit="mm" value="19.0" valueMax="24.0" valueMin="14.0">14–24 mm</quantity>
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||
(vs. densely hispid and 7–12 ×
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||
<quantity id="4CF9B8B7FF80C464FEF6FF1FFE08FF7F" box="[374,481,196,217]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" metricValueMax="1.8" metricValueMin="1.6" pageId="10" pageNumber="27" unit="mm" value="17.0" valueMax="18.0" valueMin="16.0">16–18 mm</quantity>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C31B46D9FF80C465FF08FF33FEA3FCE7" lastPageId="11" lastPageNumber="28" pageId="10" pageNumber="27" type="materials_examined">
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||
<paragraph id="8BBE1552FF80C464FF08FF33FC48FE87" blockId="10.[136,1451,160,289]" pageId="10" pageNumber="27">
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||
<emphasis id="B975C940FF80C464FF08FF33FF2BFF5B" bold="true" box="[136,194,232,253]" pageId="10" pageNumber="27">Type:</emphasis>
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||
—
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||
<materialsCitation id="3B691F0FFF80C464FF5BFF33FC74FE87" collectingDate="2015-08-01" collectionCode="SAR, BRUN, KEP, SNP" country="Malaysia" county="Lubok Antu District" elevation="194" latitude="1.25" location="Sungai Jengin" longLatPrecision="21" longitude="112.1" municipality="Batang Ai" pageId="10" pageNumber="27" specimenCode="SFC 2750" specimenCount="2" stateProvince="Sarawak" typeStatus="holotype">
|
||
<collectingCountry id="F31655C2FF80C464FF5BFF33FEB3FF5B" box="[219,346,232,253]" name="Malaysia" pageId="10" pageNumber="27">MALAYSIA</collectingCountry>
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||
. Borneo.
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||
<collectingRegion id="49C5DBB0FF80C464FE36FF33FDE5FF5B" box="[438,524,232,253]" country="Malaysia" name="Sarawak" pageId="10" pageNumber="27">Sarawak</collectingRegion>
|
||
.
|
||
<collectingCounty id="62DF6DDEFF80C464FD96FF33FD35FF5B" box="[534,732,232,253]" pageId="10" pageNumber="27">Lubok Antu District</collectingCounty>
|
||
:
|
||
<collectingMunicipality id="6BDA8F28FF80C464FD65FF33FCA1FF5B" box="[741,840,232,253]" pageId="10" pageNumber="27">Batang Ai</collectingMunicipality>
|
||
,
|
||
<location id="8EDE4389FF80C464FCD3FF33FC34FF5B" LSID="urn:lsid:plazi:treatment:03A8A444FF83C465FF08F8E9FD5DFB0F:8EDE4389FF80C464FCD3FF33FC34FF5B" box="[851,989,232,253]" country="Malaysia" county="Lubok Antu District" latitude="1.25" longLatPrecision="21" longitude="112.1" municipality="Batang Ai" name="Sungai Jengin" pageId="10" pageNumber="27" stateProvince="Sarawak">Sungai Jengin</location>
|
||
,
|
||
<geoCoordinate id="EE357395FF80C464FC67FF33FBB8FF5B" box="[999,1105,232,253]" degrees="1" direction="north" minutes="15" orientation="latitude" pageId="10" pageNumber="27" precision="15" seconds="00" value="1.25">1°15’00”N</geoCoordinate>
|
||
and
|
||
<geoCoordinate id="EE357395FF80C464FB01FF33FAE9FF5B" box="[1153,1280,232,253]" degrees="112" direction="east" minutes="06" orientation="longitude" pageId="10" pageNumber="27" precision="15" seconds="00" value="112.1">112°06’00”E</geoCoordinate>
|
||
,
|
||
<quantity id="4CF9B8B7FF80C464FA8BFF33FAA2FF5B" box="[1291,1355,232,253]" metricMagnitude="2" metricUnit="m" metricValue="1.94" pageId="10" pageNumber="27" unit="m" value="194.0">
|
||
<elevation id="002CF261FF80C464FA8BFF33FAA2FF5B" box="[1291,1355,232,253]" metricMagnitude="2" metricUnit="m" metricValue="1.94" pageId="10" pageNumber="27" unit="m" value="194.0">194 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFBF3392FF80C464FAD5FF33FF06FE87" pageId="10" pageNumber="27" value="2015-08-01">
|
||
<collectingDate id="EFFBCA7AFF80C464FAD5FF33FF06FE87" pageId="10" pageNumber="27" value="2015-08-01">1 August 2015</collectingDate>
|
||
</date>
|
||
,
|
||
<emphasis id="B975C940FF80C464FF79FED7FE2FFE87" box="[249,454,268,289]" italics="true" pageId="10" pageNumber="27">
|
||
Julia et al.
|
||
<specimenCode id="DBA7BD29FF80C464FEE5FED7FE2FFE87" box="[357,454,268,289]" collectionCode="SFC" country="China" httpUri="http://grbio.org/cool/myie-bfyd" name="Laboratory of Fishes" pageId="10" pageNumber="27">SFC 2750</specimenCode>
|
||
</emphasis>
|
||
(
|
||
<typeStatus id="54BAABF0FF80C464FE52FED7FDC1FE87" box="[466,552,268,289]" pageId="10" pageNumber="27" type="holotype">holotype</typeStatus>
|
||
<collectionCode id="ED108D97FF80C464FDAEFED7FD8DFE87" box="[558,612,268,289]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12931" lsid="urn:lsid:biocol.org:col:12931" name="Department of Forestry" pageId="10" pageNumber="27" type="Herbarium">SAR</collectionCode>
|
||
!;
|
||
<typeStatus id="54BAABF0FF80C464FDF1FED7FD29FE87" box="[625,704,268,289]" pageId="10" pageNumber="27" type="isotype">isotypes</typeStatus>
|
||
<collectionCode id="ED108D97FF80C464FD46FED7FCF9FE87" box="[710,784,268,289]" country="Brunei" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12726" lsid="urn:lsid:biocol.org:col:12726" name="Brunei Forestry Centre" pageId="10" pageNumber="27" type="Herbarium">BRUN</collectionCode>
|
||
!,
|
||
<collectionCode id="ED108D97FF80C464FC9DFED7FCB8FE87" box="[797,849,268,289]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15093" lsid="urn:lsid:biocol.org:col:15093" name="Forest Research Institute Malaysia" pageId="10" pageNumber="27" type="Herbarium">KEP</collectionCode>
|
||
!,
|
||
<collectionCode id="ED108D97FF80C464FCDEFED7FC7BFE87" box="[862,914,268,289]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15856" lsid="urn:lsid:biocol.org:col:15856" name="Sabah Parks" pageId="10" pageNumber="27" type="Herbarium">SNP</collectionCode>
|
||
!)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF7E45DAFF80C464FF08F897FF1BF80F" ID-DOI="http://doi.org/10.5281/zenodo.13674933" ID-Zenodo-Dep="13674933" httpUri="https://zenodo.org/record/13674933/files/figure.png" pageId="10" pageNumber="27" startId="10.[136,229,1868,1889]" targetBox="[317,1270,320,1832]" targetPageId="10" targetType="figure">
|
||
<paragraph id="8BBE1552FF80C464FF08F897FF1BF80F" blockId="10.[136,1451,1868,1961]" pageId="10" pageNumber="27">
|
||
<emphasis id="B975C940FF80C464FF08F897FF14F8C7" bold="true" box="[136,253,1868,1889]" pageId="10" pageNumber="27">FIGURE 6.</emphasis>
|
||
<emphasis id="B975C940FF80C464FE83F897FE25F8C7" box="[259,460,1868,1889]" italics="true" pageId="10" pageNumber="27">
|
||
<taxonomicName id="4C016ED1FF80C464FE83F897FE21F8C7" authorityName="S. Julia & Kiew" authorityYear="2016" box="[259,456,1868,1889]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="10" pageNumber="27" phylum="Tracheophyta" rank="species" species="jenginensis">Begonia jenginensis</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
A. Habit; B. Stipule; C. Fruits and male inflorescence; D. Male inflorescence; E. Male flower with one inner tepal missing; F. Female flower; G. Female flower showing pistil; H. Fruit; I. Cross section of ovary. All photos from SFC 2750 by Julia Sang.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BBE1552FF80C465FF08F802FEA3FCE7" blockId="10.[136,1452,2009,2070]" lastBlockId="11.[136,1452,160,1193]" lastPageId="11" lastPageNumber="28" pageId="10" pageNumber="27">
|
||
Cane-like begonia to
|
||
<quantity id="4CF9B8B7FF80C464FEF7F802FE2CF854" box="[375,453,2009,2034]" metricMagnitude="0" metricUnit="m" metricValue="1.35" pageId="10" pageNumber="27" unit="m" value="1.35">1.35 m</quantity>
|
||
tall. Stems reddish brown, pilose, hairs brownish, terete and finely ridged,
|
||
<quantity id="4CF9B8B7FF80C464FA8AF802FA8FF854" box="[1290,1382,2009,2034]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="10" pageNumber="27" unit="mm" value="7.0" valueMax="8.0" valueMin="6.0">6–8 mm</quantity>
|
||
thick, internodes 4–6(–10) cm, swollen at the nodes. Stipules narrowly or broadly lanceolate, 12–18 ×
|
||
<quantity id="4CF9B8B7FF80C464FB66F826FABDF7B0" box="[1254,1364,2045,2070]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="11.0" metricValueMin="4.0" pageId="10" pageNumber="27" unit="mm" value="7.5" valueMax="11.0" valueMin="4.0">4–11 mm</quantity>
|
||
, keeled terminating in a seta
|
||
<quantity id="4CF9B8B7FF81C465FEFFFF7BFE36FF1F" box="[383,479,160,185]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="11" pageNumber="28" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 mm</quantity>
|
||
long, hispid outside. Leaves alternate, very oblique, held horizontally or sometimes slightly vertical; petioles reddish brown, (3–)5–10(–13.5) cm long, grooved, hispid; lamina plain green above, paler beneath, above sparsely hispid between veins, veins densely hispid, glabrous beneath except on veins, in life succulent, membranous when dry, matt, broadly ovate, 13.5–25 ×
|
||
<quantity id="4CF9B8B7FF81C465FC8CFED7FC43FE83" box="[780,938,268,293]" metricMagnitude="-1" metricUnit="m" metricValue="1.6" metricValueMax="2.05" metricValueMin="1.15" pageId="11" pageNumber="28" unit="cm" value="16.0" valueMax="20.5" valueMin="11.5">11.5–20.5 cm</quantity>
|
||
, broad side
|
||
<quantity id="4CF9B8B7FF81C465FBB8FED7FB58FE83" box="[1080,1201,268,293]" metricMagnitude="-1" metricUnit="m" metricValue="1.175" metricValueMax="1.45" metricValueMin="0.9" pageId="11" pageNumber="28" unit="cm" value="11.75" valueMax="14.5" valueMin="9.0">9–14.5 cm</quantity>
|
||
wide, base shallowly cordate, basal lobes
|
||
<quantity id="4CF9B8B7FF81C465FEEFFEEBFE2FFEEF" box="[367,454,304,329]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="11" pageNumber="28" unit="cm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 cm</quantity>
|
||
, margin minutely dentate, apex acuminate,
|
||
<quantity id="4CF9B8B7FF81C465FC35FEEBFBF7FEEF" box="[949,1054,304,329]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="11" pageNumber="28" unit="cm" value="1.75" valueMax="2.0" valueMin="1.5">1.5–2 cm</quantity>
|
||
long; venation palmate, prominent on both surfaces, 4–6 veins radiating from the base, in life veins pale green on both surfaces, reddish brown towards the base, drying reddish, 3–4 veins along the midrib on the broad side, 1–2 veins in the basal lobe. Inflorescences protogynous, axillary in the upper leaf axils, racemose, rachis unbranched, (2.5–)
|
||
<quantity id="4CF9B8B7FF81C465FBB5FE47FB4EFE13" box="[1077,1191,412,437]" metricMagnitude="-1" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="11" pageNumber="28" unit="cm" value="12.5" valueMax="15.0" valueMin="10.0">10–15 cm</quantity>
|
||
long. Bracts greenish, lanceolate, 12–17 ×
|
||
<quantity id="4CF9B8B7FF81C465FEF5FE1BFE3EFE7F" box="[373,471,448,473]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="11" pageNumber="28" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4–5 mm</quantity>
|
||
, margin entire, apex setose to
|
||
<quantity id="4CF9B8B7FF81C465FCBDFE1BFC68FE7F" box="[829,897,448,473]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="11" pageNumber="28" unit="mm" value="4.0">4 mm</quantity>
|
||
long, caducous; bracteoles similar to bracts but smaller, 4–10 ×
|
||
<quantity id="4CF9B8B7FF81C465FEC2FE3FFE6EFE5B" box="[322,391,484,509]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="11" pageNumber="28" unit="mm" value="4.0">4 mm</quantity>
|
||
, caducous. Male flowers with pale pink pedicels,
|
||
<quantity id="4CF9B8B7FF81C465FC42FE3FFBC7FE5B" box="[962,1070,484,509]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="11" pageNumber="28" unit="mm" value="7.5" valueMax="10.0" valueMin="5.0">5–10 mm</quantity>
|
||
long; tepals 4, pale pink, reddish towards base, oval, outer 2 tepals 8–10 ×
|
||
<quantity id="4CF9B8B7FF81C465FDD5FDD3FD5AFD87" box="[597,691,520,545]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="11" pageNumber="28" unit="mm" value="6.5" valueMax="8.0" valueMin="5.0">5–8 mm</quantity>
|
||
, inner 2 tepals 5–8 ×
|
||
<quantity id="4CF9B8B7FF81C465FC20FDD3FC16FD87" box="[928,1023,520,545]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="11" pageNumber="28" unit="mm" value="3.0" valueMax="4.0" valueMin="2.0">2–4 mm</quantity>
|
||
, margin entire, apex rounded; stamens 28–30, cluster globose, subsessile; filaments pale yellow, ca.
|
||
<quantity id="4CF9B8B7FF81C465FCB0FDF7FC99FDE3" box="[816,880,556,581]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="28" unit="mm" value="2.0">2 mm</quantity>
|
||
long; anthers pale yellow, obovate, ca. 1.5 ×
|
||
<quantity id="4CF9B8B7FF81C465FAE5FDF7FA4EFDE3" box="[1381,1447,556,581]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="28" unit="mm" value="1.0">1 mm</quantity>
|
||
, apex emarginate. Female flowers with greenish pedicels,
|
||
<quantity id="4CF9B8B7FF81C465FC9CFD8BFC95FDCF" box="[796,892,592,617]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="8.0" metricValueMin="4.0" pageId="11" pageNumber="28" unit="mm" value="6.0" valueMax="8.0" valueMin="4.0">4–8 mm</quantity>
|
||
long, glabrous; ovary pale green, wing tips pale green, ovate, 7–8 ×
|
||
<quantity id="4CF9B8B7FF81C465FEE5FDAFFE38FD2B" box="[357,465,628,653]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="11.0" metricValueMin="8.0" pageId="11" pageNumber="28" unit="mm" value="9.5" valueMax="11.0" valueMin="8.0">8–11 mm</quantity>
|
||
, wings 3, equal, locules 3, placentas 1 per locule; tepals 5, ovate, 6–7 ×
|
||
<quantity id="4CF9B8B7FF81C465FB73FDAFFABBFD2B" box="[1267,1362,628,653]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="11" pageNumber="28" unit="mm" value="5.0" valueMax="6.0" valueMin="4.0">4–6 mm</quantity>
|
||
, margin entire, apex rounded; styles 3, pale yellow,
|
||
<quantity id="4CF9B8B7FF81C465FDEEFD43FD25FD17" box="[622,716,664,689]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="11" pageNumber="28" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
|
||
long, divided to base, widely Y-shaped; stigma reddish, papillose cells large and pale yellow forming a continuous twisted band. Capsules 2–4 pairs along the rachis, greenish yellow, glabrous, 12–14 ×
|
||
<quantity id="4CF9B8B7FF81C465FED8FD3BFE3AFD5F" box="[344,467,736,761]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" metricValueMax="2.4" metricValueMin="1.4" pageId="11" pageNumber="28" unit="mm" value="19.0" valueMax="24.0" valueMin="14.0">14–24 mm</quantity>
|
||
, glabrous, locules 3, wings 3, equal, rounded proximally and distally or truncate distally,
|
||
<quantity id="4CF9B8B7FF81C465FF08FCDFFF0FFCBB" box="[136,230,772,797]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="11" pageNumber="28" unit="mm" value="4.5" valueMax="6.0" valueMin="3.0">3–6 mm</quantity>
|
||
wide, thinly fibrous, dehiscing between locule and wing; pedicel pendent,
|
||
<quantity id="4CF9B8B7FF81C465FBB6FCDFFB4BFCBB" box="[1078,1186,772,797]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.5" metricValueMin="0.6" pageId="11" pageNumber="28" unit="mm" value="10.5" valueMax="15.0" valueMin="6.0">6–15 mm</quantity>
|
||
long, glabrous, reddish towards the base.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31B46D9FF81C465FF3DFC97FA79FCC3" box="[189,1424,844,869]" pageId="11" pageNumber="28" type="etymology">
|
||
<paragraph id="8BBE1552FF81C465FF3DFC97FA79FCC3" blockId="11.[136,1452,160,1193]" box="[189,1424,844,869]" pageId="11" pageNumber="28">
|
||
<emphasis id="B975C940FF81C465FF3DFC97FEA1FCC3" bold="true" box="[189,328,844,869]" pageId="11" pageNumber="28">Etymology:</emphasis>
|
||
—Named after the
|
||
<typeStatus id="54BAABF0FF81C465FD9FFC97FDA6FCC3" box="[543,591,844,869]" pageId="11" pageNumber="28">type</typeStatus>
|
||
locality, Jengin River (Sungai = River) where the species was discovered.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31B46D9FF81C465FF3DFCABFD5DFB0F" pageId="11" pageNumber="28" type="distribution">
|
||
<paragraph id="8BBE1552FF81C465FF3DFCABFBDFFC0B" blockId="11.[136,1452,160,1193]" pageId="11" pageNumber="28">
|
||
<emphasis id="B975C940FF81C465FF3DFCABFEB0FC2F" bold="true" box="[189,345,880,905]" pageId="11" pageNumber="28">Distribution:</emphasis>
|
||
—
|
||
<collectingCountry id="F31655C2FF81C465FEF6FCABFDE3FC2F" box="[374,522,880,905]" name="Malaysia" pageId="11" pageNumber="28">MALAYSIA</collectingCountry>
|
||
. Borneo.
|
||
<collectingRegion id="49C5DBB0FF81C465FDFDFCABFD08FC2F" box="[637,737,880,905]" country="Malaysia" name="Sarawak" pageId="11" pageNumber="28">Sarawak</collectingRegion>
|
||
. Endemic in
|
||
<collectingRegion id="49C5DBB0FF81C465FCFFFCABFC36FC2F" box="[895,991,880,905]" country="Malaysia" name="Sarawak" pageId="11" pageNumber="28">Sarawak</collectingRegion>
|
||
where the species is known only from Sungai Bebiyong Besai in Batang Ai National Park and Sungai Jengin in Batang Ai.
|
||
</paragraph>
|
||
<paragraph id="8BBE1552FF81C465FF3DFC63FDF6FC53" blockId="11.[136,1452,160,1193]" pageId="11" pageNumber="28">
|
||
<emphasis id="B975C940FF81C465FF3DFC63FECAFC77" bold="true" box="[189,291,952,977]" pageId="11" pageNumber="28">Habitat:</emphasis>
|
||
—Riparian forest at
|
||
<quantity id="4CF9B8B7FF81C465FD88FC63FDA6FC77" box="[520,591,952,977]" metricMagnitude="2" metricUnit="m" metricValue="1.94" pageId="11" pageNumber="28" unit="m" value="194.0">194 m</quantity>
|
||
elevation, growing on steep earth slopes above the river bank or on boulders with thin layer of soil in light shade.
|
||
</paragraph>
|
||
<paragraph id="8BBE1552FF81C465FF3DFBDBFCCCFB9B" blockId="11.[136,1452,160,1193]" pageId="11" pageNumber="28">
|
||
<emphasis id="B975C940FF81C465FF3DFBDBFDD4FBBF" bold="true" box="[189,573,1024,1049]" pageId="11" pageNumber="28">Additional specimen examined:</emphasis>
|
||
—
|
||
<collectingCountry id="F31655C2FF81C465FDD9FBDBFD04FBBF" box="[601,749,1024,1049]" name="Malaysia" pageId="11" pageNumber="28">MALAYSIA</collectingCountry>
|
||
. Borneo.
|
||
<collectingRegion id="49C5DBB0FF81C465FCDFFBDBFC2AFBBF" box="[863,963,1024,1049]" country="Malaysia" name="Sarawak" pageId="11" pageNumber="28">Sarawak</collectingRegion>
|
||
. Lubok Antu District: Batang Ai National Park, Sungai Bebiyong Besai,
|
||
<emphasis id="B975C940FF81C465FE5EFBFFFD24FB9B" box="[478,717,1060,1085]" italics="true" pageId="11" pageNumber="28">Julia et al. SFC 2755</emphasis>
|
||
(SAR).
|
||
</paragraph>
|
||
<paragraph id="8BBE1552FF81C465FF3DFB93FA84FBC7" blockId="11.[136,1452,160,1193]" box="[189,1389,1096,1121]" pageId="11" pageNumber="28">
|
||
<emphasis id="B975C940FF81C465FF3DFB93FEE0FBC7" bold="true" box="[189,265,1096,1121]" pageId="11" pageNumber="28">Notes:</emphasis>
|
||
—The species is more common along the bank of Sungai Jengin than along Sungai Bebiyong Besai.
|
||
</paragraph>
|
||
<paragraph id="8BBE1552FF81C465FF3DFBB7FD5DFB0F" blockId="11.[136,1452,160,1193]" pageId="11" pageNumber="28">
|
||
<emphasis id="B975C940FF81C465FF3DFBB7FDCDFB23" bold="true" box="[189,548,1132,1157]" pageId="11" pageNumber="28">Proposed conservation status:</emphasis>
|
||
—Least Concern as the species occurs within a Totally Protected Area and their habitat is not threatened by any local disturbance.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |