258 lines
30 KiB
XML
258 lines
30 KiB
XML
<document id="0DE683CBF67496494511490FC2436FD2" ID-DOI="10.11646/phytotaxa.252.1.2" ID-ISSN="1179-3163" ID-Zenodo-Dep="13674915" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725405123220" checkinUser="felipe" docAuthor="Sang, Julia & Kiew, Ruth" docDate="2016" docId="03A8A444FF81C463FF08FB03FD5DFE5B" docLanguage="en" docName="phytotaxa.252.1.2.pdf" docOrigin="Phytotaxa 252 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.252.1.2" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Begonia tebiang S. Julia & Kiew 2016, spec. nov." docType="treatment" docVersion="2" lastPageNumber="30" masterDocId="FF91DC3CFF8AC46EFF80FFDBFFE9FFA6" masterDocTitle="Begonia (Begoniaceae) from Batang Ai National Park and vicinity, Sarawak, Borneo, including six new species" masterLastPageNumber="30" masterPageNumber="17" pageNumber="28" updateTime="1725426159086" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="0162D868BDA9400FD62F2067FBE1C8D1">
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<mods:title id="A5DA33D34B88C3907088BD8CE2EE500F">Begonia (Begoniaceae) from Batang Ai National Park and vicinity, Sarawak, Borneo, including six new species</mods:title>
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</mods:titleInfo>
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<mods:name id="A89B7E7FB1558CC175DC818A0E277A27" type="personal">
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<mods:roleTerm id="C4B80BD7B9B2FC6FAE0C84254A58E340">Author</mods:roleTerm>
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<mods:namePart id="09988865A9BFF98F76A578F18E04BD85">Sang, Julia</mods:namePart>
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<mods:nameIdentifier id="8BFB3E293D3F87C39B61EEA1DB8C725F" type="email">juliasang@sarawakforestry.com</mods:nameIdentifier>
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<mods:name id="7104555053FDD9F53A65882EE555A147" type="personal">
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<mods:roleTerm id="162D2CED63EB2CB233FA46B02B15F23B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="0E2EE7B05D6197B72EFCA0C27BF5B646">Kiew, Ruth</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="86CBBFEDC6E820DD0974DD750A1C0D83">text</mods:typeOfResource>
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<mods:title id="CA89CC19606C72EC5C7C12968659E80B">Phytotaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="DD52473CE50233D7D4C8F0A0252F2B0B">
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<mods:date id="8DB89BDCD09DC6DFF84E2F8825F1D5C2">2016</mods:date>
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<mods:detail id="12B61FA7297BEE4C76E3C36BB441A810" type="pubDate">
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<mods:number id="A4CD179C0A9F4B3BB7876FB610523488">2016-03-08</mods:number>
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<mods:detail id="A6B2FD8589701A59E7F180D64A912918" type="volume">
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<mods:number id="FAB5C76E96EBBB790ED12686C527E62F">252</mods:number>
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<mods:number id="956815C2F0E1C0B749FEC06968568690">1</mods:number>
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<mods:start id="E84D70AB10D25BE16A59FFC21BE84B20">17</mods:start>
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<mods:url id="4CBB17927685A6CB6F4D5AF88882C139">http://dx.doi.org/10.11646/phytotaxa.252.1.2</mods:url>
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<mods:classification id="0FA436B1111DBAEABEC3A4725839C9F3">journal article</mods:classification>
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<mods:identifier id="CAA83C25F45116415D01EA4D4899FE7D" type="DOI">10.11646/phytotaxa.252.1.2</mods:identifier>
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<mods:identifier id="4B5BA03B11BCA58D5116F659BCB04B0F" type="ISSN">1179-3163</mods:identifier>
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<treatment id="03A8A444FF81C463FF08FB03FD5DFE5B" LSID="urn:lsid:plazi:treatment:03A8A444FF81C463FF08FB03FD5DFE5B" httpUri="http://treatment.plazi.org/id/03A8A444FF81C463FF08FB03FD5DFE5B" lastPageId="13" lastPageNumber="30" pageId="11" pageNumber="28">
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<subSubSection id="C31B46D9FF81C465FF08FB03FCECFB54" box="[136,773,1239,1266]" pageId="11" pageNumber="28" type="nomenclature">
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<paragraph id="8BBE1552FF81C465FF08FB03FCECFB54" blockId="11.[136,773,1239,1266]" box="[136,773,1239,1266]" pageId="11" pageNumber="28">
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<heading id="D0F6A23EFF81C465FF08FB03FCECFB54" box="[136,773,1239,1266]" fontSize="11" level="2" pageId="11" pageNumber="28" reason="3">
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<emphasis id="B975C940FF81C465FF08FB03FF74FB54" bold="true" box="[136,157,1240,1266]" italics="true" pageId="11" pageNumber="28">9.</emphasis>
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<taxonomicName id="4C016ED1FF81C465FF24FB03FDFCFB54" ID-CoL="7SHRG" authority="S.Julia & Kiew" authorityName="S. Julia & Kiew" authorityYear="2016" box="[164,533,1239,1266]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="11" pageNumber="28" phylum="Tracheophyta" rank="species" species="tebiang" status="sp. nov.">
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<emphasis id="B975C940FF81C465FF24FB03FE8BFB54" bold="true" box="[164,354,1239,1266]" italics="true" pageId="11" pageNumber="28">Begonia tebiang</emphasis>
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S.Julia & Kiew
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</taxonomicName>
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,
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<emphasis id="B975C940FF81C465FDA0FB02FD67FB54" box="[544,654,1241,1266]" italics="true" pageId="11" pageNumber="28">
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<taxonomicNameLabel id="A246743BFF81C465FDA0FB02FD67FB54" box="[544,654,1241,1266]" pageId="11" pageNumber="28" rank="species">spec. nov.</taxonomicNameLabel>
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</emphasis>
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(
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<figureCitation id="133A09D7FF81C465FD1CFB02FD14FB54" box="[668,765,1241,1266]" captionStart="FIGURE 7" captionStartId="12.[136,229,1942,1963]" captionTargetBox="[357,1234,574,1917]" captionTargetId="figure-218@12.[353,1234,569,1917]" captionTargetPageId="12" captionText="FIGURE 7. Begonia tebiang. A & B. Habit; C. Male inflorescence with numerous pairs of bracteoles; D. Male flower; E & F. Female flowers; G. Old fruit; H. Cross section of ovary. All photos from SFC 2763 by Julia Sang (A–G) and Ling Chea Yiing (H)." figureDoi="http://doi.org/10.5281/zenodo.13674937" httpUri="https://zenodo.org/record/13674937/files/figure.png" pageId="11" pageNumber="28">Figure 7</figureCitation>
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||
)
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C31B46D9FF81C465FF08FB26FE9EFAB0" box="[136,375,1277,1302]" pageId="11" pageNumber="28" type="description">
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<paragraph id="8BBE1552FF81C465FF08FB26FE9EFAB0" blockId="11.[136,375,1277,1302]" box="[136,375,1277,1302]" pageId="11" pageNumber="28">
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<emphasis id="B975C940FF81C465FF08FB26FF36FAB0" bold="true" box="[136,223,1277,1302]" pageId="11" pageNumber="28">Section</emphasis>
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<taxonomicName id="4C016ED1FF81C465FF66FB26FE9EFAB0" box="[230,375,1277,1302]" class="Magnoliopsida" family="Begoniaceae" genus="Petermannia" kingdom="Plantae" order="Cucurbitales" pageId="11" pageNumber="28" phylum="Tracheophyta" rank="genus">
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<emphasis id="B975C940FF81C465FF66FB26FE9EFAB0" box="[230,375,1277,1302]" italics="true" pageId="11" pageNumber="28">Petermannia</emphasis>
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</taxonomicName>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C31B46D9FF81C465FF08FA93FBDCFA4B" pageId="11" pageNumber="28" type="diagnosis">
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<paragraph id="8BBE1552FF81C465FF08FA93FBDCFA4B" blockId="11.[136,1452,1352,1589]" pageId="11" pageNumber="28">
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<emphasis id="B975C940FF81C465FF08FA93FF1AFAFB" bold="true" box="[136,243,1352,1373]" pageId="11" pageNumber="28">Diagnosis:</emphasis>
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—Similar to
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<taxonomicName id="4C016ED1FF81C465FEEBFA93FCACFAFB" authority="S. Julia & C. Y. Ling (2015: 110)" authorityName="S. Julia & C. Y. Ling" authorityPageNumber="110" authorityYear="2015" box="[363,837,1352,1373]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="11" pageNumber="28" phylum="Tracheophyta" rank="species" species="johariana">
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<emphasis id="B975C940FF81C465FEEBFA93FDF7FAFB" box="[363,542,1352,1373]" italics="true" pageId="11" pageNumber="28">Begonia johariana</emphasis>
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S.Julia & C.Y.Ling (2015: 110)
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</taxonomicName>
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in its slender creeping stem, small leaves, racemose inflorescence with large bracteoles and its small fruits wider than long but it is different in its succulent stem
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<quantity id="4CF9B8B7FF81C465FBEEFAB7FB56FA27" box="[1134,1215,1388,1409]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="11" pageNumber="28" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 mm</quantity>
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thick (vs. wiry and
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||
<quantity id="4CF9B8B7FF81C465FA07FAB7FF0BFA03" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="11" pageNumber="28" unit="mm" value="2.0" valueMax="3.0" valueMin="1.0">1–3 mm</quantity>
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thick in
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<taxonomicName id="4C016ED1FF81C465FEB7FA4BFE58FA03" authorityName="S. Julia & C. Y. Ling" authorityYear="2015" box="[311,433,1424,1445]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="11" pageNumber="28" phylum="Tracheophyta" rank="species" species="johariana">
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<emphasis id="B975C940FF81C465FEB7FA4BFE58FA03" box="[311,433,1424,1445]" italics="true" pageId="11" pageNumber="28">B. johariana</emphasis>
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</taxonomicName>
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||
), its larger stipules
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<quantity id="4CF9B8B7FF81C465FDEEFA4BFD3CFA03" box="[622,725,1424,1445]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="11" pageNumber="28" unit="mm" value="12.5" valueMax="15.0" valueMin="10.0">10–15 mm</quantity>
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long with an entire margin (vs.
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<quantity id="4CF9B8B7FF81C465FB89FA4BFBB1FA03" box="[1033,1112,1424,1445]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="11" pageNumber="28" unit="mm" value="6.5" valueMax="8.0" valueMin="5.0">5–8 mm</quantity>
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long with a serrate margin), longer inflorescences
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<quantity id="4CF9B8B7FF81C465FECEFA6FFE41FA6F" box="[334,424,1460,1481]" metricMagnitude="-2" metricUnit="m" metricValue="6.25" metricValueMax="8.5" metricValueMin="4.0" pageId="11" pageNumber="28" unit="cm" value="6.25" valueMax="8.5" valueMin="4.0">4–8.5 cm</quantity>
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long with larger bracts 7–17 ×
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<quantity id="4CF9B8B7FF81C465FD5EFA6FFCD0FA6F" box="[734,825,1460,1481]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="11.0" metricValueMin="4.0" pageId="11" pageNumber="28" unit="mm" value="7.5" valueMax="11.0" valueMin="4.0">4–11 mm</quantity>
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||
(vs. inflorescences to
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||
<quantity id="4CF9B8B7FF81C465FB96FA6FFBAFFA6F" box="[1046,1094,1460,1481]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="28" unit="cm" value="5.0">5 cm</quantity>
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||
long and bracts 5–7 ×
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||
<quantity id="4CF9B8B7FF81C465FAA9FA6FFA92FA6F" box="[1321,1403,1460,1481]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="11" pageNumber="28" unit="mm" value="3.0" valueMax="4.0" valueMin="2.0">2–4 mm</quantity>
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||
) and outer tepals of female flower less than twice as long as wide (vs. 2–3 times as long as wide).
|
||
</paragraph>
|
||
</subSubSection>
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||
<subSubSection id="C31B46D9FF81C462FF08FA27FB96FD87" lastPageId="12" lastPageNumber="29" pageId="11" pageNumber="28" type="materials_examined">
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<paragraph id="8BBE1552FF81C465FF08FA27FA7BF993" blockId="11.[136,1452,1352,1589]" pageId="11" pageNumber="28">
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<emphasis id="B975C940FF81C465FF08FA27FF2BF9B7" bold="true" box="[136,194,1532,1553]" pageId="11" pageNumber="28">Type:</emphasis>
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—
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||
<materialsCitation id="3B691F0FFF81C465FF5BFA27FA67F993" collectingDate="2015-08-02" collectionCode="SAR, KEP, SNP" country="Malaysia" county="Lubok Antu District" elevation="200" latitude="1.3" location="Lubang Baya" longLatPrecision="21" longitude="112.06944" municipality="Batang Ai National Park" pageId="11" pageNumber="28" specimenCode="SFC 2763" specimenCount="2" stateProvince="Sarawak" typeStatus="holotype">
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<collectingCountry id="F31655C2FF81C465FF5BFA27FEB3F9B7" box="[219,346,1532,1553]" name="Malaysia" pageId="11" pageNumber="28">MALAYSIA</collectingCountry>
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. Borneo.
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<collectingRegion id="49C5DBB0FF81C465FE35FA27FDE2F9B7" box="[437,523,1532,1553]" country="Malaysia" name="Sarawak" pageId="11" pageNumber="28">Sarawak</collectingRegion>
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.
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<collectingCounty id="62DF6DDEFF81C465FD93FA27FD31F9B7" box="[531,728,1532,1553]" pageId="11" pageNumber="28">Lubok Antu District</collectingCounty>
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:
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<collectingMunicipality id="6BDA8F28FF81C465FD60FA27FC27F9B7" box="[736,974,1532,1553]" pageId="11" pageNumber="28">Batang Ai National Park</collectingMunicipality>
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,
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<location id="8EDE4389FF81C465FC56FA27FBB0F9B7" LSID="urn:lsid:plazi:treatment:03A8A444FF81C463FF08FB03FD5DFE5B:8EDE4389FF81C465FC56FA27FBB0F9B7" box="[982,1113,1532,1553]" country="Malaysia" county="Lubok Antu District" latitude="1.3" longLatPrecision="21" longitude="112.06944" municipality="Batang Ai National Park" name="Lubang Baya" pageId="11" pageNumber="28" stateProvince="Sarawak">Lubang Baya</location>
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,
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<location id="8EDE4389FF81C465FBE2FA27FB01F9B7" LSID="urn:lsid:plazi:treatment:03A8A444FF81C463FF08FB03FD5DFE5B:8EDE4389FF81C465FBE2FA27FB01F9B7" box="[1122,1256,1532,1553]" country="Malaysia" county="Lubok Antu District" latitude="1.3" longLatPrecision="21" longitude="112.06944" municipality="Batang Ai National Park" name="Bebiyong Mit" pageId="11" pageNumber="28" stateProvince="Sarawak">Bebiyong Mit</location>
|
||
trail, adjacent to the
|
||
<location id="8EDE4389FF81C465FF3DF9FBFEDCF993" LSID="urn:lsid:plazi:treatment:03A8A444FF81C463FF08FB03FD5DFE5B:8EDE4389FF81C465FF3DF9FBFEDCF993" box="[189,309,1568,1589]" country="Malaysia" county="Lubok Antu District" latitude="1.3" longLatPrecision="21" longitude="112.06944" municipality="Batang Ai National Park" name="Ranger Post" pageId="11" pageNumber="28" stateProvince="Sarawak">Ranger Post</location>
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,
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||
<geoCoordinate id="EE357395FF81C465FEBEF9FBFE41F993" box="[318,424,1568,1589]" degrees="1" direction="north" minutes="18" orientation="latitude" pageId="11" pageNumber="28" precision="15" seconds="00" value="1.3">1°18’00”N</geoCoordinate>
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and
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<geoCoordinate id="EE357395FF81C465FE56F9FBFDBEF993" box="[470,599,1568,1589]" degrees="112" direction="east" minutes="04" orientation="longitude" pageId="11" pageNumber="28" precision="15" seconds="10" value="112.06944">112°04’10”E</geoCoordinate>
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,
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<quantity id="4CF9B8B7FF81C465FDE0F9FBFD76F993" box="[608,671,1568,1589]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="28" unit="m" value="200.0">
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<elevation id="002CF261FF81C465FDE0F9FBFD76F993" box="[608,671,1568,1589]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="28" unit="m" value="200.0">200 m</elevation>
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</quantity>
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,
|
||
<date id="FFBF3392FF81C465FD29F9FBFCDEF993" box="[681,823,1568,1589]" pageId="11" pageNumber="28" value="2015-08-02">
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<collectingDate id="EFFBCA7AFF81C465FD29F9FBFCDEF993" box="[681,823,1568,1589]" pageId="11" pageNumber="28" value="2015-08-02">2 August 2015</collectingDate>
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</date>
|
||
,
|
||
<emphasis id="B975C940FF81C465FCC1F9FBFBE7F993" box="[833,1038,1568,1589]" italics="true" pageId="11" pageNumber="28">
|
||
Julia et al.
|
||
<specimenCode id="DBA7BD29FF81C465FC2DF9FBFBE7F993" box="[941,1038,1568,1589]" collectionCode="SFC" country="China" httpUri="http://grbio.org/cool/myie-bfyd" name="Laboratory of Fishes" pageId="11" pageNumber="28">SFC 2763</specimenCode>
|
||
</emphasis>
|
||
(
|
||
<typeStatus id="54BAABF0FF81C465FB9AF9FBFB99F993" box="[1050,1136,1568,1589]" pageId="11" pageNumber="28" type="holotype">holotype</typeStatus>
|
||
<collectionCode id="ED108D97FF81C465FBF6F9FBFB45F993" box="[1142,1196,1568,1589]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12931" lsid="urn:lsid:biocol.org:col:12931" name="Department of Forestry" pageId="11" pageNumber="28" type="Herbarium">SAR</collectionCode>
|
||
!;
|
||
<typeStatus id="54BAABF0FF81C465FB39F9FBFAE1F993" box="[1209,1288,1568,1589]" pageId="11" pageNumber="28" type="isotype">isotypes</typeStatus>
|
||
<collectionCode id="ED108D97FF81C465FA8EF9FBFAABF993" box="[1294,1346,1568,1589]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15093" lsid="urn:lsid:biocol.org:col:15093" name="Forest Research Institute Malaysia" pageId="11" pageNumber="28" type="Herbarium">KEP</collectionCode>
|
||
!,
|
||
<collectionCode id="ED108D97FF81C465FACFF9FBFA6AF993" box="[1359,1411,1568,1589]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15856" lsid="urn:lsid:biocol.org:col:15856" name="Sabah Parks" pageId="11" pageNumber="28" type="Herbarium">SNP</collectionCode>
|
||
!)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BBE1552FF81C462FF08F985FB96FD87" blockId="11.[136,1452,1630,2087]" lastBlockId="12.[136,1452,160,545]" lastPageId="12" lastPageNumber="29" pageId="11" pageNumber="28">
|
||
Creeping herb. Indumentum on stems and petioles pubescent. Stems dull brown, slender,
|
||
<quantity id="4CF9B8B7FF81C465FB0EF985FB04F9D1" box="[1166,1261,1630,1655]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="11" pageNumber="28" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 mm</quantity>
|
||
thick, branched, succulent, internodes
|
||
<quantity id="4CF9B8B7FF81C465FEFAF959FE1FF93D" box="[378,502,1666,1691]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" metricValueMax="8.5" metricValueMin="2.5" pageId="11" pageNumber="28" unit="cm" value="5.5" valueMax="8.5" valueMin="2.5">2.5–8.5 cm</quantity>
|
||
long, slightly thicker at nodes. Stipules pale green, broadly ovate, 10–15 ×
|
||
<quantity id="4CF9B8B7FF81C465FABBF959FA4EF93D" box="[1339,1447,1666,1691]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="10.0" metricValueMin="7.0" pageId="11" pageNumber="28" unit="mm" value="8.5" valueMax="10.0" valueMin="7.0">7–10 mm</quantity>
|
||
, margin entire and ciliate, apex acuminate, acumen ca.
|
||
<quantity id="4CF9B8B7FF81C465FD62F97DFCCBF919" box="[738,802,1702,1727]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="28" unit="mm" value="1.0">1 mm</quantity>
|
||
long, caducous. Leaves alternate, distant, slightly oblique, held more or less horizontally; petioles dull brown, 1–3.5(–5) cm long, slightly grooved; lamina plain green above, paler beneath, in life slightly fleshy, glossy, with scattered short hairs, asymmetric, ovate, 5.5–8(–10.5) × 4.5–6(–8.5) cm, broad side 2.5–4(–5.5) cm wide, base slightly cordate, sometimes overlapping, basal lobes
|
||
<quantity id="4CF9B8B7FF81C465FB66F8C9FA8EF88D" box="[1254,1383,1810,1835]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.5" metricValueMin="0.7" pageId="11" pageNumber="28" unit="cm" value="1.1" valueMax="1.5" valueMin="0.7">0.7–1.5 cm</quantity>
|
||
long, margin minutely serrate, teeth more prominent at the vein endings, apex acuminate, acumen to
|
||
<quantity id="4CF9B8B7FF81C465FB48F8EDFAE9F8E9" box="[1224,1280,1846,1871]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="28" unit="cm" value="1.0">1 cm</quantity>
|
||
long; venation palmate-pinnate, veins greenish above, dark brown towards the base above and beneath, minutely pubescent above and beneath, 2 pairs at the base and 3–4 veins on either side of the midrib, 2–3 veins in basal lobe, slightly impressed above, prominent beneath. Inflorescences protogynous, axillary in upper leaf axils, racemose, slender,
|
||
<quantity id="4CF9B8B7FF81C465FA83F879FA82F81D" box="[1283,1387,1954,1979]" metricMagnitude="-2" metricUnit="m" metricValue="6.25" metricValueMax="8.5" metricValueMin="4.0" pageId="11" pageNumber="28" unit="cm" value="6.25" valueMax="8.5" valueMin="4.0">4–8.5 cm</quantity>
|
||
long, sometimes with a single lateral branch
|
||
<quantity id="4CF9B8B7FF81C465FDBDF81DFD4CF879" box="[573,677,1990,2015]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="11" pageNumber="28" unit="cm" value="1.75" valueMax="2.0" valueMin="1.5">1.5–2 cm</quantity>
|
||
long with a female flower. Bracts pale green, broadly lanceolate, (7–)10–12(–17) × (4–)6–7(–11) mm, margin ciliate, persistent; bracteoles 3–4 pairs, lower bracteoles subtending female flowers oval, pale green, ca. 8 ×
|
||
<quantity id="4CF9B8B7FF81C465FD83F7D5FDB4F781" box="[515,605,2062,2087]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="11" pageNumber="28" unit="mm" value="4.5">4.5 mm</quantity>
|
||
, similar to bracts; upper bracteoles subtending male flowers broadly oval, 6–7 ×
|
||
<quantity id="4CF9B8B7FF86C462FF53FF7BFEDDFF1F" box="[211,308,160,185]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="12" pageNumber="29" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">5–6 mm</quantity>
|
||
, sometimes tinged reddish at the tip, keeled, margin ciliate, persistent. Male flowers with pinkish or white pedicels,
|
||
<quantity id="4CF9B8B7FF86C462FEBCFF1FFE75FF7B" box="[316,412,196,221]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="12" pageNumber="29" unit="mm" value="3.5" valueMax="5.0" valueMin="2.0">2–5 mm</quantity>
|
||
long, pubescent; tepals 4, white sometimes tinged red or reddish, glabrous, margin entire, apex rounded, outer 2 tepals broadly elliptic, 6–8 ×
|
||
<quantity id="4CF9B8B7FF86C462FD47FF33FCCFFEA7" box="[711,806,232,257]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="12" pageNumber="29" unit="mm" value="5.0" valueMax="6.0" valueMin="4.0">4–6 mm</quantity>
|
||
, inner 2 tepals lanceolate, 4–5 ×
|
||
<quantity id="4CF9B8B7FF86C462FB13FF33FB1AFEA7" box="[1171,1267,232,257]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="12" pageNumber="29" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
|
||
; stamens 42–46, cluster globose, subsessile; filaments pale yellow,
|
||
<quantity id="4CF9B8B7FF86C462FD33FED7FCD0FE83" box="[691,825,268,293]" metricMagnitude="-4" metricUnit="m" metricValue="9.5" metricValueMax="11.0" metricValueMin="8.0" pageId="12" pageNumber="29" unit="mm" value="0.95" valueMax="1.1" valueMin="0.8">0.8–1.1 mm</quantity>
|
||
long; anthers pale yellow, obovate, 0.6–1 ×
|
||
<quantity id="4CF9B8B7FF86C462FA9FFED7FA4EFE83" box="[1311,1447,268,293]" metricMagnitude="-4" metricUnit="m" metricValue="3" metricValueMax="4.0" metricValueMin="2.0" pageId="12" pageNumber="29" unit="mm" value="0.3" valueMax="0.4" valueMin="0.2">0.2–0.4 mm</quantity>
|
||
, apex emarginate. Female flowers with pale green pedicels, ca.
|
||
<quantity id="4CF9B8B7FF86C462FCD8FEEBFC72FEEF" box="[856,923,304,329]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="29" unit="mm" value="1.0">1 mm</quantity>
|
||
long, glabrous; ovary pale green with reddish wings, broadly ovate, 5–6 ×
|
||
<quantity id="4CF9B8B7FF86C462FE45FE8FFDA9FECB" box="[453,576,340,365]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="12" pageNumber="29" unit="mm" value="14.5" valueMax="15.0" valueMin="14.0">14–15 mm</quantity>
|
||
, glabrous, wings 3, unequal, locules 3, placentas 2 per locule; tepals 5, reddish or greenish outside, white tinged reddish inside, glabrous, outer 3 tepals elliptic, 5–6 ×
|
||
<quantity id="4CF9B8B7FF86C462FBE5FEA3FB2CFE37" box="[1125,1221,376,401]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="12" pageNumber="29" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">3–5 mm</quantity>
|
||
, margin entire, apex acute, inner 2 tepals slightly smaller, ca. 4 ×
|
||
<quantity id="4CF9B8B7FF86C462FD03FE47FD21FE13" box="[643,712,412,437]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="29" unit="mm" value="2.0">2 mm</quantity>
|
||
; styles 3, pale yellow, ca.
|
||
<quantity id="4CF9B8B7FF86C462FC70FE47FBD8FE13" box="[1008,1073,412,437]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="29" unit="mm" value="1.0">1 mm</quantity>
|
||
long, divided to base, shallow Yshaped; stigma pale yellow, papillose forming a continuous twisted band. Capsules broadly obovate, 6–7 ×
|
||
<quantity id="4CF9B8B7FF86C462FABAFE1BFA4EFE7F" box="[1338,1447,448,473]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.4" metricValueMin="0.9" pageId="12" pageNumber="29" unit="mm" value="11.5" valueMax="14.0" valueMin="9.0">9–14 mm</quantity>
|
||
, glabrous, locules 3, wings 3, equal, broadly rounded,acute proximally, rounded or truncate distally,
|
||
<quantity id="4CF9B8B7FF86C462FA85FE3FFA8CFE5B" box="[1285,1381,484,509]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="12" pageNumber="29" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 mm</quantity>
|
||
wide, thinly fibrous, dehiscing between locule and wing; pedicel slender, pendent,
|
||
<quantity id="4CF9B8B7FF86C462FC62FDD3FBD6FD87" box="[994,1087,520,545]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="12" pageNumber="29" unit="mm" value="6.5" valueMax="8.0" valueMin="5.0">5–8 mm</quantity>
|
||
long.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF7E45DAFF86C462FF08F84DFAF6F869" ID-DOI="http://doi.org/10.5281/zenodo.13674937" ID-Zenodo-Dep="13674937" httpUri="https://zenodo.org/record/13674937/files/figure.png" pageId="12" pageNumber="29" startId="12.[136,229,1942,1963]" targetBox="[357,1234,574,1917]" targetPageId="12" targetType="figure">
|
||
<paragraph id="8BBE1552FF86C462FF08F84DFAF6F869" blockId="12.[136,1452,1942,1999]" pageId="12" pageNumber="29">
|
||
<emphasis id="B975C940FF86C462FF08F84DFF16F80D" bold="true" box="[136,255,1942,1963]" pageId="12" pageNumber="29">FIGURE 7.</emphasis>
|
||
<emphasis id="B975C940FF86C462FE86F84DFE45F80D" box="[262,428,1942,1963]" italics="true" pageId="12" pageNumber="29">
|
||
<taxonomicName id="4C016ED1FF86C462FE86F84DFE41F80D" authorityName="S. Julia & Kiew" authorityYear="2016" box="[262,424,1942,1963]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="12" pageNumber="29" phylum="Tracheophyta" rank="species" species="tebiang">Begonia tebiang</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
A & B. Habit; C. Male inflorescence with numerous pairs of bracteoles; D. Male flower; E & F. Female flowers; G. Old fruit; H. Cross section of ovary. All photos from SFC 2763 by Julia Sang (A–G) and Ling Chea Yiing (H).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C31B46D9FF87C463FF3DFF7BFEE3FF7B" pageId="13" pageNumber="30" type="etymology">
|
||
<paragraph id="8BBE1552FF87C463FF3DFF7BFEE3FF7B" blockId="13.[136,1452,160,509]" pageId="13" pageNumber="30">
|
||
<emphasis id="B975C940FF87C463FF3DFF7BFEA1FF1F" bold="true" box="[189,328,160,185]" pageId="13" pageNumber="30">Etymology:</emphasis>
|
||
—From local Iban word
|
||
<emphasis id="B975C940FF87C463FD02FF7BFD10FF1F" box="[642,761,160,185]" italics="true" pageId="13" pageNumber="30">
|
||
<taxonomicName id="4C016ED1FF87C463FD02FF7BFD3FFF1F" authorityName="S. Julia & Kiew" authorityYear="2016" box="[642,726,160,185]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="13" pageNumber="30" phylum="Tracheophyta" rank="species" species="tebiang">tebiang</taxonomicName>
|
||
=
|
||
</emphasis>
|
||
slope, referring to the habitat where the species grows abundantly.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31B46D9FF87C463FF3DFF33FD5DFE5B" pageId="13" pageNumber="30" type="distribution">
|
||
<paragraph id="8BBE1552FF87C463FF3DFF33FB7FFEA7" blockId="13.[136,1452,160,509]" box="[189,1174,232,257]" pageId="13" pageNumber="30">
|
||
<emphasis id="B975C940FF87C463FF3DFF33FEB0FEA7" bold="true" box="[189,345,232,257]" pageId="13" pageNumber="30">Distribution:</emphasis>
|
||
—Endemic in
|
||
<collectingCountry id="F31655C2FF87C463FE7CFF33FDB6FEA7" box="[508,607,232,257]" name="Malaysia" pageId="13" pageNumber="30">Sarawak</collectingCountry>
|
||
, as yet known only from Batang Ai National Park.
|
||
</paragraph>
|
||
<paragraph id="8BBE1552FF87C463FF3DFED7FEA3FEEF" blockId="13.[136,1452,160,509]" pageId="13" pageNumber="30">
|
||
<emphasis id="B975C940FF87C463FF3DFED7FECAFE83" bold="true" box="[189,291,268,293]" pageId="13" pageNumber="30">Habitat:</emphasis>
|
||
—Lowland mixed dipterocarp forest at elevations below
|
||
<quantity id="4CF9B8B7FF87C463FC1DFED7FC0DFE83" box="[925,996,268,293]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="30" unit="m" value="200.0">200 m</quantity>
|
||
on steep or gentle earth slopes in deep to light shade areas.
|
||
</paragraph>
|
||
<paragraph id="8BBE1552FF87C463FF3DFE8FFD7FFE13" blockId="13.[136,1452,160,509]" pageId="13" pageNumber="30">
|
||
<emphasis id="B975C940FF87C463FF3DFE8FFDAFFECB" bold="true" box="[189,582,340,365]" pageId="13" pageNumber="30">Additional specimens examined:</emphasis>
|
||
—
|
||
<collectingCountry id="F31655C2FF87C463FDE2FE8FFD1FFECB" box="[610,758,340,365]" name="Malaysia" pageId="13" pageNumber="30">MALAYSIA</collectingCountry>
|
||
. Borneo.
|
||
<collectingRegion id="49C5DBB0FF87C463FCE6FE8FFC23FECB" box="[870,970,340,365]" country="Malaysia" name="Sarawak" pageId="13" pageNumber="30">Sarawak</collectingRegion>
|
||
. Lubok Antu District: Batang Ai National Park, Nanga Lubang Baya, Bebiyong Mit Trail,
|
||
<emphasis id="B975C940FF87C463FD1BFEA3FC6CFE37" box="[667,901,376,401]" italics="true" pageId="13" pageNumber="30">Julia et al. SFC 2764</emphasis>
|
||
(KEP, SAR),
|
||
<emphasis id="B975C940FF87C463FBA0FEA3FAEEFE37" box="[1056,1287,376,401]" italics="true" pageId="13" pageNumber="30">Rantai et al. S 88048</emphasis>
|
||
(SAR), Nanga Sungai Lelayang,
|
||
<emphasis id="B975C940FF87C463FED3FE47FDD6FE13" box="[339,575,412,437]" italics="true" pageId="13" pageNumber="30">Rantai et al. S 82793</emphasis>
|
||
(SAR).
|
||
</paragraph>
|
||
<paragraph id="8BBE1552FF87C463FF3DFE1BFD5DFE5B" blockId="13.[136,1452,160,509]" pageId="13" pageNumber="30">
|
||
<emphasis id="B975C940FF87C463FF3DFE1BFDCDFE7F" bold="true" box="[189,548,448,473]" pageId="13" pageNumber="30">Proposed conservation status:</emphasis>
|
||
—Least Concern as the species occurs within a Totally Protected Area and their habitat is not threatened by any local disturbance.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |