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<document id="91FE1F6A614DAC87F4D21E3869522A0F" ID-DOI="10.5281/zenodo.177584" ID-GBIF-Dataset="c64bb094-8a87-4f11-b28d-282dea53fa07" ID-ISSN="1175-5326" ID-Zenodo-Dep="177584" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459868439420" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2007" docId="03820808FFF7FFA9FF365845559DFC1C" docLanguage="en" docName="zt01527p029.pdf" docOrigin="Zootaxa 1527" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Hughleechia Perkins 1981" docType="treatment" docVersion="10" lastPageNumber="20" masterDocId="FFBB7070FFF5FFAAFFA15C475754FFE6" masterDocTitle="A revision of the Australian intertidal water beetle genus Hughleechia Perkins (Coleoptera: Hydraenidae)" masterLastPageNumber="29" masterPageNumber="17" pageNumber="19" updateTime="1698226390393" updateUser="plazi">
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<mods:title id="3405E5D860988E96930AEBCCFFD9C4DD">A revision of the Australian intertidal water beetle genus Hughleechia Perkins (Coleoptera: Hydraenidae)</mods:title>
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<treatment id="03820808FFF7FFA9FF365845559DFC1C" ID-DOI="http://doi.org/10.5281/zenodo.6243245" ID-GBIF-Taxon="119369034" ID-Zenodo-Dep="6243245" LSID="urn:lsid:plazi:treatment:03820808FFF7FFA9FF365845559DFC1C" httpUri="http://treatment.plazi.org/id/03820808FFF7FFA9FF365845559DFC1C" lastPageId="3" lastPageNumber="20" pageId="2" pageNumber="19">
<subSubSection id="C331EA95FFF7FFA8FF36584556B0FBFA" box="[151,484,1026,1052]" pageId="2" pageNumber="19" type="nomenclature">
<paragraph id="8B94B91EFFF7FFA8FF36584556B0FBFA" blockId="2.[151,484,1026,1052]" box="[151,484,1026,1052]" pageId="2" pageNumber="19">
<heading id="D0DC0E72FFF7FFA8FF36584556B0FBFA" bold="true" box="[151,484,1026,1052]" fontSize="11" level="1" pageId="2" pageNumber="19" reason="1">
<taxonomicName id="4C2BC29DFFF7FFA8FF36584556B0FBFA" authority="Perkins, 1981" authorityName="Perkins" authorityYear="1981" box="[151,484,1026,1052]" class="Insecta" family="Hydraenidae" genus="Hughleechia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B95F650CFFF7FFA8FF36584556B0FBFA" bold="true" box="[151,484,1026,1052]" pageId="2" pageNumber="19">
<emphasis id="B95F650CFFF7FFA8FF3658455667FBFA" bold="true" box="[151,307,1026,1052]" italics="true" pageId="2" pageNumber="19">Hughleechia</emphasis>
<bibRefCitation id="EFBAC4EFFFF7FFA8FE9B584556B0FBFA" author="Perkins" box="[314,484,1026,1052]" pageId="2" pageNumber="29" refString="Perkins, P. D. (1981). Hughleechia giulianii, a new genus and new species of intertidal beetle from Australia (Coleoptera: Hydraenidae). Pan-Pacific Entomologist, 57 (1), 295 - 302." type="journal article" year="1981">Perkins, 1981</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C331EA95FFF7FFA8FF36580A527DFB82" box="[151,1321,1101,1124]" pageId="2" pageNumber="19" type="reference_group">
<paragraph id="8B94B91EFFF7FFA8FF36580A527DFB82" blockId="2.[151,1321,1101,1124]" box="[151,1321,1101,1124]" pageId="2" pageNumber="19">
<treatmentCitationGroup id="AB3B9E30FFF7FFA8FF36580A527DFB82" box="[151,1321,1101,1124]" pageId="2" pageNumber="19">
<taxonomicName id="4C2BC29DFFF7FFA8FF36580A56AFFB82" authority="Perkins, 1981: 299" authorityName="Perkins" authorityPageNumber="299" authorityYear="1981" box="[151,507,1101,1124]" class="Insecta" family="Hydraenidae" genus="Hughleechia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B95F650CFFF7FFA8FF36580A564BFB82" box="[151,287,1101,1124]" italics="true" pageId="2" pageNumber="19">Hughleechia</emphasis>
<treatmentCitation id="0A8A9F0FFFF7FFA8FE8D580A56AFFB82" author="Perkins" box="[300,507,1101,1124]" page="299" pageId="2" pageNumber="19" year="1981">
<bibRefCitation id="EFBAC4EFFFF7FFA8FE8D580A5696FB82" author="Perkins" box="[300,450,1101,1124]" pageId="2" pageNumber="29" refString="Perkins, P. D. (1981). Hughleechia giulianii, a new genus and new species of intertidal beetle from Australia (Coleoptera: Hydraenidae). Pan-Pacific Entomologist, 57 (1), 295 - 302." type="journal article" year="1981">Perkins, 1981</bibRefCitation>
: 299
</treatmentCitation>
</taxonomicName>
(
<typeStatus id="549007BCFFF7FFA8FDA8580A556CFB82" box="[521,568,1101,1124]" pageId="2" pageNumber="19">type</typeStatus>
species
<taxonomicName id="4C2BC29DFFF7FFA8FD32580A534CFB82" ID-CoL="9PSQ2" authority="Perkins, 1981" authorityName="Perkins" authorityYear="1981" box="[659,1048,1101,1124]" class="Insecta" family="Hydraenidae" genus="Hughleechia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="giulianii">
<emphasis id="B95F650CFFF7FFA8FD32580A5428FB82" box="[659,892,1101,1124]" italics="true" pageId="2" pageNumber="19">Hughleechia giulianii</emphasis>
<bibRefCitation id="EFBAC4EFFFF7FFA8FC22580A534CFB82" author="Perkins" box="[899,1048,1101,1124]" pageId="2" pageNumber="29" refString="Perkins, P. D. (1981). Hughleechia giulianii, a new genus and new species of intertidal beetle from Australia (Coleoptera: Hydraenidae). Pan-Pacific Entomologist, 57 (1), 295 - 302." type="journal article" year="1981">Perkins, 1981</bibRefCitation>
</taxonomicName>
, by original designation).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C331EA95FFF7FFA8FF3658D2546FFAC1" pageId="2" pageNumber="19" type="diagnosis">
<paragraph id="8B94B91EFFF7FFA8FF3658D2546FFAC1" blockId="2.[151,1437,1173,1999]" pageId="2" pageNumber="19">
<emphasis id="B95F650CFFF7FFA8FF3658D25644FB49" bold="true" box="[151,272,1173,1199]" pageId="2" pageNumber="19">Diagnosis</emphasis>
. Members of
<taxonomicName id="4C2BC29DFFF7FFA8FE1F58D25500FB48" box="[446,596,1173,1198]" class="Insecta" family="Hydraenidae" genus="Hughleechia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B95F650CFFF7FFA8FE1F58D25500FB48" box="[446,596,1173,1198]" italics="true" pageId="2" pageNumber="19">Hughleechia</emphasis>
</taxonomicName>
can be recognized by the combination of the short maxillary palpi, the absence of a lateral hyaline border on the pronotum, the arcuate posterior border of the pronotum which lacks distinct posterior angles and has a hyaline border, and the ESDS components of the antennal pocket, including a transgenal ridge (see
<bibRefCitation id="EFBAC4EFFFF7FFA8FE04594A556BFAC1" author="Perkins" box="[421,575,1293,1319]" pageId="2" pageNumber="29" refString="Perkins, P. D. (1997). Life on the effective bubble: exocrine secretion delivery systems (ESDS) and the evolution and classification of beetles in the family Hydraenidae (Insecta: Coleoptera). Annals of Carnegie Museum, 66 (2), 89 - 207." type="journal article" year="1997">Perkins 1997</bibRefCitation>
for other characters).
</paragraph>
</subSubSection>
<subSubSection id="C331EA95FFF7FFA9FF675972559DFC1C" lastPageId="3" lastPageNumber="20" pageId="2" pageNumber="19" type="description">
<paragraph id="8B94B91EFFF7FFA8FF675972544FFAA9" blockId="2.[151,1437,1173,1999]" box="[198,795,1333,1359]" pageId="2" pageNumber="19">
<emphasis id="B95F650CFFF7FFA8FF675972560BFAA9" bold="true" box="[198,351,1333,1359]" pageId="2" pageNumber="19">Description.</emphasis>
Body form elongate oval, depressed.
</paragraph>
<paragraph id="8B94B91EFFF7FFA8FF67591A55BAFA09" blockId="2.[151,1437,1173,1999]" pageId="2" pageNumber="19">Head with distinct ocelli next to protuberant eyes; dorsum of head flat, frons and clypeus in same plane. Maxillary palpus short, palpomere 1 very small, 2 shorter than or subequal in length to 3, 3 broader than other palpomeres, 4 (last) slightly shorter than 3. Nine antennomeres: 2 basal, 1 small intermediate, 1 small cupule, and 5 pubescent segments forming a compact club.</paragraph>
<paragraph id="8B94B91EFFF7FFA8FF6759BA5310F951" blockId="2.[151,1437,1173,1999]" pageId="2" pageNumber="19">
Pronotum with anterior margin weakly rounded in middle, posterior margin much more strongly rounded; anterior and posterior margins with narrow (
<taxonomicName id="4C2BC29DFFF7FFA8FD155A62546BF9D8" box="[692,831,1573,1598]" class="Insecta" family="Hydraenidae" genus="Hughleechia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="giulianii">
<emphasis id="B95F650CFFF7FFA8FD155A62546BF9D8" box="[692,831,1573,1598]" italics="true" pageId="2" pageNumber="19">H. giulianii</emphasis>
</taxonomicName>
) or moderately wide (
<taxonomicName id="4C2BC29DFFF7FFA8FBF25A625380F9D8" box="[1107,1236,1573,1598]" class="Insecta" family="Hydraenidae" genus="Hughleechia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B95F650CFFF7FFA8FBF25A625380F9D8" box="[1107,1236,1573,1598]" italics="true" pageId="2" pageNumber="19">H. gracilis</emphasis>
</taxonomicName>
) hyaline border, sides lacking hyaline border; disc with shallow median longitudinal depression which is slightly constricted at midlength; lateral fossulae well developed; pronotal surface with moderately dense and moderately long to very long hairs, except hairs absent in median depression and in lateral fossulae.
</paragraph>
<paragraph id="8B94B91EFFF7FFA8FF675A8257BEF8B1" blockId="2.[151,1437,1173,1999]" pageId="2" pageNumber="19">
Elytra ovoid or elongate, fused together at weakly raised suture; each elytron with ca. eight rows of punctures, except rows 68 much less distinct than discal rows, sometimes random; each puncture with a short, stiff, recumbent seta (
<taxonomicName id="4C2BC29DFFF7FFA8FE385B525575F8C8" box="[409,545,1813,1838]" class="Insecta" family="Hydraenidae" genus="Hughleechia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="giulianii">
<emphasis id="B95F650CFFF7FFA8FE385B525575F8C8" box="[409,545,1813,1838]" italics="true" pageId="2" pageNumber="19">H. giulianii</emphasis>
</taxonomicName>
) or a long, slender and flexible seta (
<taxonomicName id="4C2BC29DFFF7FFA8FC7C5B52530FF8C8" box="[989,1115,1813,1838]" class="Insecta" family="Hydraenidae" genus="Hughleechia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B95F650CFFF7FFA8FC7C5B52530FF8C8" box="[989,1115,1813,1838]" italics="true" pageId="2" pageNumber="19">H. gracilis</emphasis>
</taxonomicName>
) (
<figureCitation id="1310A59BFFF7FFA8FBD75B5253BCF8C9" box="[1142,1256,1813,1839]" captionStart="FIGURES 5 6" captionStartId="7.[151,269,1753,1777]" captionTargetBox="[240,1336,194,1753]" captionTargetId="figure@7.[240,1346,165,1753]" captionTargetPageId="7" captionText="FIGURES 5 6. Hughleechia, elytron punctation pattern and microsculpture detail. — 5. H. giulianii. — 6. H. gracilis." httpUri="https://zenodo.org/record/177588/files/figure.png" pageId="2" pageNumber="19">Figs. 5, 6</figureCitation>
). Flight wings absent.
</paragraph>
<paragraph id="8B94B91EFFF7FFA9FF675B225383FF3C" blockId="2.[151,1437,1173,1999]" lastBlockId="3.[151,1437,152,1018]" lastPageId="3" lastPageNumber="20" pageId="2" pageNumber="19">
Prosternum slightly and evenly elevated in midline, not carinate; procoxae contiguous. Mesoventrite narrow to broad between mesocoxae, with or without carinae; hydrofuge pubescent. Metaventrite reduced, length in midline approximately equal to that of mesoventrite; intercoxal process moderately to very broad; metacoxae narrowly (
<taxonomicName id="4C2BC29DFFF6FFA9FE395CDF5542FF57" box="[408,534,152,177]" class="Insecta" family="Hydraenidae" genus="Hughleechia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B95F650CFFF6FFA9FE395CDF5542FF57" box="[408,534,152,177]" italics="true" pageId="3" pageNumber="20">H. gracilis</emphasis>
</taxonomicName>
) to widely (
<taxonomicName id="4C2BC29DFFF6FFA9FD0B5CDF5466FF57" box="[682,818,152,177]" class="Insecta" family="Hydraenidae" genus="Hughleechia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="giulianii">
<emphasis id="B95F650CFFF6FFA9FD0B5CDF5466FF57" box="[682,818,152,177]" italics="true" pageId="3" pageNumber="20">H. giulianii</emphasis>
</taxonomicName>
) separated; hydrofuge pubescence entirely covering metaventrite, or restricted to lateral areas and narrow band along edge of mesocoxal cavities.
</paragraph>
<paragraph id="8B94B91EFFF6FFA9FF675CAF55E9FEB4" blockId="3.[151,1437,152,1018]" pageId="3" pageNumber="20">Abdomen with seven sternites; intercoxal sternite narrow, or broader than long; sternites 14 entirely hydrofuge pubescent, or with hydrofuge pubescence restricted to sides; segment 5 totally hydrofuge pubescent; segments 6 and 7 with a few sparse hairs.</paragraph>
<paragraph id="8B94B91EFFF6FFA9FF675D2755B3FE2C" blockId="3.[151,1437,152,1018]" pageId="3" pageNumber="20">Legs rather long. Tarsi long, slender for most of length, but distinctly broadened apically; tarsal formula 5-5-5; tarsomeres 13 very small, combined length slightly longer than, or subequal to length of tarsomere 4; tarsomere 5 longer than other segments combined.</paragraph>
<paragraph id="8B94B91EFFF6FFA9FF675D9F559DFC1C" blockId="3.[151,1437,152,1018]" pageId="3" pageNumber="20">
Structures examined in microslide mounts (terminology as in
<bibRefCitation id="EFBAC4EFFFF6FFA9FC365D9F536AFE14" author="Perkins" box="[919,1086,472,498]" pageId="3" pageNumber="29" refString="Perkins, P. D. (1997). Life on the effective bubble: exocrine secretion delivery systems (ESDS) and the evolution and classification of beetles in the family Hydraenidae (Insecta: Coleoptera). Annals of Carnegie Museum, 66 (2), 89 - 207." type="journal article" year="1997">Perkins, 1997</bibRefCitation>
): Head with distinctive transgenal ridge and more-or-less developed secretion sulcus or pores (
<figureCitation id="1310A59BFFF6FFA9FC395E4754BEFDFC" box="[920,1002,512,538]" captionStart="FIGURES 7 9" captionStartId="9.[151,269,1811,1835]" captionTargetBox="[296,1289,219,1783]" captionTargetId="figure@9.[291,1296,197,1784]" captionTargetPageId="9" captionText="FIGURES 7 9. Hughleechia giulianii. — 7. Ventral view of right eye, antennal pocket, and associated ESDS structures. — 8. Ventral view of antenna, head and prothorax. — 9. Detail of transgenal ridge and postocular secretion delivery setae (a, secretion delivery setae; i, hypomeral antennal pocket; n, prosternum; o, hypomeral hyaline border; t, postocular asperite hydrofuge; u, first antennomere; z, transgenal ridge, with exocrine secretion sulcus)." httpUri="https://zenodo.org/record/177589/files/figure.png" pageId="3" pageNumber="20">Figs. 9</figureCitation>
,
<figureCitation id="1310A59BFFF6FFA9FC565E475343FDFC" box="[1015,1047,512,538]" captionStart="FIGURES 10 13" captionStartId="10.[151,269,1616,1640]" captionTargetBox="[161,1427,203,1575]" captionTargetId="figure@10.[151,1436,183,1590]" captionTargetPageId="10" captionText="FIGURES 10 13. Hughleechia gracilis. — 10. Ventral view of head. — 11. Ventral view of right eye, antennal pocket of prothorax, and associated ESDS structures. — 12. Detail of transgenal secretion ridge and forked postocular secretion delivery setae on left side. — 13. Same, on right side (a, secretion delivery setae; o, hypomeral hyaline border; z, transgenal ridge, with exocrine secretion pores)." httpUri="https://zenodo.org/record/177590/files/figure.png" pageId="3" pageNumber="20">12</figureCitation>
,
<emphasis id="B95F650CFFF6FFA9FB835E475379FDFF" box="[1058,1069,512,537]" italics="true" pageId="3" pageNumber="20">z</emphasis>
). Three to four elongate secretion delivery setae in
<taxonomicName id="4C2BC29DFFF6FFA9FE075E6F5566FDA7" box="[422,562,552,577]" class="Insecta" family="Hydraenidae" genus="Hughleechia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="giulianii">
<emphasis id="B95F650CFFF6FFA9FE075E6F5566FDA7" box="[422,562,552,577]" italics="true" pageId="3" pageNumber="20">H. giulianii</emphasis>
</taxonomicName>
(
<figureCitation id="1310A59BFFF6FFA9FDE65E6F55C0FDA4" box="[583,660,552,578]" captionStart="FIGURES 7 9" captionStartId="9.[151,269,1811,1835]" captionTargetBox="[296,1289,219,1783]" captionTargetId="figure@9.[291,1296,197,1784]" captionTargetPageId="9" captionText="FIGURES 7 9. Hughleechia giulianii. — 7. Ventral view of right eye, antennal pocket, and associated ESDS structures. — 8. Ventral view of antenna, head and prothorax. — 9. Detail of transgenal ridge and postocular secretion delivery setae (a, secretion delivery setae; i, hypomeral antennal pocket; n, prosternum; o, hypomeral hyaline border; t, postocular asperite hydrofuge; u, first antennomere; z, transgenal ridge, with exocrine secretion sulcus)." httpUri="https://zenodo.org/record/177589/files/figure.png" pageId="3" pageNumber="20">Fig. 7</figureCitation>
,
<emphasis id="B95F650CFFF6FFA9FD045E6F55E7FDA7" box="[677,691,552,577]" italics="true" pageId="3" pageNumber="20">a</emphasis>
); two or three setae in
<taxonomicName id="4C2BC29DFFF6FFA9FC7E5E6F5335FDA7" box="[991,1121,552,577]" class="Insecta" family="Hydraenidae" genus="Hughleechia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B95F650CFFF6FFA9FC7E5E6F5335FDA7" box="[991,1121,552,577]" italics="true" pageId="3" pageNumber="20">H. gracilis</emphasis>
</taxonomicName>
, shorter than those of
<taxonomicName id="4C2BC29DFFF6FFA9FADE5E6F57AEFD8F" class="Insecta" family="Hydraenidae" genus="Hughleechia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="giulianii">
<emphasis id="B95F650CFFF6FFA9FADE5E6F57AEFD8F" italics="true" pageId="3" pageNumber="20">H. giulianii</emphasis>
</taxonomicName>
, which sometimes become stuck together in dry specimens (
<figureCitation id="1310A59BFFF6FFA9FC6B5E175303FD8C" box="[970,1111,592,618]" captionStart="FIGURES 10 13" captionStartId="10.[151,269,1616,1640]" captionTargetBox="[161,1427,203,1575]" captionTargetId="figure@10.[151,1436,183,1590]" captionTargetPageId="10" captionText="FIGURES 10 13. Hughleechia gracilis. — 10. Ventral view of head. — 11. Ventral view of right eye, antennal pocket of prothorax, and associated ESDS structures. — 12. Detail of transgenal secretion ridge and forked postocular secretion delivery setae on left side. — 13. Same, on right side (a, secretion delivery setae; o, hypomeral hyaline border; z, transgenal ridge, with exocrine secretion pores)." httpUri="https://zenodo.org/record/177590/files/figure.png" pageId="3" pageNumber="20">Figs. 1113</figureCitation>
). Postocular hydrofuge (
<emphasis id="B95F650CFFF6FFA9FA2B5E1752C6FD8F" box="[1418,1426,592,617]" italics="true" pageId="3" pageNumber="20">t</emphasis>
) asperite and cuticle with exocrine pores (
<emphasis id="B95F650CFFF6FFA9FDD85E3F55D3FD77" box="[633,647,632,657]" italics="true" pageId="3" pageNumber="20">p</emphasis>
) near base of secretion delivery setae. Hypomeral antennal pocket (
<figureCitation id="1310A59BFFF6FFA9FF3E5EE757BCFD5C" box="[159,232,672,698]" captionStart="FIGURES 7 9" captionStartId="9.[151,269,1811,1835]" captionTargetBox="[296,1289,219,1783]" captionTargetId="figure@9.[291,1296,197,1784]" captionTargetPageId="9" captionText="FIGURES 7 9. Hughleechia giulianii. — 7. Ventral view of right eye, antennal pocket, and associated ESDS structures. — 8. Ventral view of antenna, head and prothorax. — 9. Detail of transgenal ridge and postocular secretion delivery setae (a, secretion delivery setae; i, hypomeral antennal pocket; n, prosternum; o, hypomeral hyaline border; t, postocular asperite hydrofuge; u, first antennomere; z, transgenal ridge, with exocrine secretion sulcus)." httpUri="https://zenodo.org/record/177589/files/figure.png" pageId="3" pageNumber="20">Fig. 7</figureCitation>
,
<figureCitation id="1310A59BFFF6FFA9FF575EE75647FD5C" box="[246,275,672,698]" captionStart="FIGURES 10 13" captionStartId="10.[151,269,1616,1640]" captionTargetBox="[161,1427,203,1575]" captionTargetId="figure@10.[151,1436,183,1590]" captionTargetPageId="10" captionText="FIGURES 10 13. Hughleechia gracilis. — 10. Ventral view of head. — 11. Ventral view of right eye, antennal pocket of prothorax, and associated ESDS structures. — 12. Detail of transgenal secretion ridge and forked postocular secretion delivery setae on left side. — 13. Same, on right side (a, secretion delivery setae; o, hypomeral hyaline border; z, transgenal ridge, with exocrine secretion pores)." httpUri="https://zenodo.org/record/177590/files/figure.png" pageId="3" pageNumber="20">11</figureCitation>
) well developed, cuticle smooth except microreticulate medially and hydrofuge pubescent anteromedially. Exocrine pores moderately dense in antennal pocket. Hypomeral hyaline border moderately large (
<figureCitation id="1310A59BFFF6FFA9FF3E5EB757A3FCEC" box="[159,247,752,778]" captionStart="FIGURES 10 13" captionStartId="10.[151,269,1616,1640]" captionTargetBox="[161,1427,203,1575]" captionTargetId="figure@10.[151,1436,183,1590]" captionTargetPageId="10" captionText="FIGURES 10 13. Hughleechia gracilis. — 10. Ventral view of head. — 11. Ventral view of right eye, antennal pocket of prothorax, and associated ESDS structures. — 12. Detail of transgenal secretion ridge and forked postocular secretion delivery setae on left side. — 13. Same, on right side (a, secretion delivery setae; o, hypomeral hyaline border; z, transgenal ridge, with exocrine secretion pores)." httpUri="https://zenodo.org/record/177590/files/figure.png" pageId="3" pageNumber="20">Fig. 11</figureCitation>
,
<emphasis id="B95F650CFFF6FFA9FEA65EB75642FCEF" box="[263,278,752,777]" italics="true" pageId="3" pageNumber="20">o</emphasis>
) to quite large anteriorly (
<figureCitation id="1310A59BFFF6FFA9FDF85EB755F0FCEC" box="[601,676,752,778]" captionStart="FIGURES 7 9" captionStartId="9.[151,269,1811,1835]" captionTargetBox="[296,1289,219,1783]" captionTargetId="figure@9.[291,1296,197,1784]" captionTargetPageId="9" captionText="FIGURES 7 9. Hughleechia giulianii. — 7. Ventral view of right eye, antennal pocket, and associated ESDS structures. — 8. Ventral view of antenna, head and prothorax. — 9. Detail of transgenal ridge and postocular secretion delivery setae (a, secretion delivery setae; i, hypomeral antennal pocket; n, prosternum; o, hypomeral hyaline border; t, postocular asperite hydrofuge; u, first antennomere; z, transgenal ridge, with exocrine secretion sulcus)." httpUri="https://zenodo.org/record/177589/files/figure.png" pageId="3" pageNumber="20">Fig. 7</figureCitation>
,
<emphasis id="B95F650CFFF6FFA9FD155EB75597FCEF" box="[692,707,752,777]" italics="true" pageId="3" pageNumber="20">o</emphasis>
), in latter condition it contacts postocular secretion delivery setae. Hypomeral hyaline border posteriorly quite narrow, becoming contiguous with posterior hyaline bor- der. Posterior border of pronotum arcuate such that there are no posterior angles. Hypomeral hyaline border and anterior hyaline border contiguous (
<taxonomicName id="4C2BC29DFFF6FFA9FDD05F2F55BBFC67" box="[625,751,872,897]" class="Insecta" family="Hydraenidae" genus="Hughleechia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B95F650CFFF6FFA9FDD05F2F55BBFC67" box="[625,751,872,897]" italics="true" pageId="3" pageNumber="20">H. gracilis</emphasis>
</taxonomicName>
), or separated (
<taxonomicName id="4C2BC29DFFF6FFA9FC085F2F5365FC67" box="[937,1073,872,897]" class="Insecta" family="Hydraenidae" genus="Hughleechia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="giulianii">
<emphasis id="B95F650CFFF6FFA9FC085F2F5365FC67" box="[937,1073,872,897]" italics="true" pageId="3" pageNumber="20">H. giulianii</emphasis>
</taxonomicName>
). Wet-hypomeron well developed. Lateral hyaline border absent. Hypomeral antennal pocket setae slightly (
<figureCitation id="1310A59BFFF6FFA9FBF95FD753F5FC4C" box="[1112,1185,912,938]" captionStart="FIGURES 7 9" captionStartId="9.[151,269,1811,1835]" captionTargetBox="[296,1289,219,1783]" captionTargetId="figure@9.[291,1296,197,1784]" captionTargetPageId="9" captionText="FIGURES 7 9. Hughleechia giulianii. — 7. Ventral view of right eye, antennal pocket, and associated ESDS structures. — 8. Ventral view of antenna, head and prothorax. — 9. Detail of transgenal ridge and postocular secretion delivery setae (a, secretion delivery setae; i, hypomeral antennal pocket; n, prosternum; o, hypomeral hyaline border; t, postocular asperite hydrofuge; u, first antennomere; z, transgenal ridge, with exocrine secretion sulcus)." httpUri="https://zenodo.org/record/177589/files/figure.png" pageId="3" pageNumber="20">Fig. 7</figureCitation>
,
<taxonomicName id="4C2BC29DFFF6FFA9FB0E5FD75263FC4F" box="[1199,1335,912,937]" class="Insecta" family="Hydraenidae" genus="Hughleechia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="giulianii">
<emphasis id="B95F650CFFF6FFA9FB0E5FD75263FC4F" box="[1199,1335,912,937]" italics="true" pageId="3" pageNumber="20">H. giulianii</emphasis>
</taxonomicName>
) to well extended (
<figureCitation id="1310A59BFFF6FFA9FEB15FFF5630FC34" box="[272,356,952,978]" captionStart="FIGURES 10 13" captionStartId="10.[151,269,1616,1640]" captionTargetBox="[161,1427,203,1575]" captionTargetId="figure@10.[151,1436,183,1590]" captionTargetPageId="10" captionText="FIGURES 10 13. Hughleechia gracilis. — 10. Ventral view of head. — 11. Ventral view of right eye, antennal pocket of prothorax, and associated ESDS structures. — 12. Detail of transgenal secretion ridge and forked postocular secretion delivery setae on left side. — 13. Same, on right side (a, secretion delivery setae; o, hypomeral hyaline border; z, transgenal ridge, with exocrine secretion pores)." httpUri="https://zenodo.org/record/177590/files/figure.png" pageId="3" pageNumber="20">Fig. 11</figureCitation>
,
<taxonomicName id="4C2BC29DFFF6FFA9FED15FFF56B9FC37" box="[368,493,952,977]" class="Insecta" family="Hydraenidae" genus="Hughleechia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B95F650CFFF6FFA9FED15FFF56B9FC37" box="[368,493,952,977]" italics="true" pageId="3" pageNumber="20">H. gracilis</emphasis>
</taxonomicName>
) beyond hypomeral hyaline border. Profemur lacking distal spine cluster, spines in this area small, as are others on the profemur.
</paragraph>
</subSubSection>
</treatment>
</document>