202 lines
24 KiB
XML
202 lines
24 KiB
XML
<document id="B9DEAD934F33D6AB95FCC943062AC2D8" ID-DOI="10.5281/zenodo.273398" ID-GBIF-Dataset="ae452b89-2798-4a2a-8bd9-963dead78ebc" ID-ISSN="11755326" ID-Zenodo-Dep="273398" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459761708870" checkinUser="plazi" docAuthor="Howden, Henry F." docDate="2006" docId="0380EE1A1954FFE0FE94F8E0391C07B9" docLanguage="en" docName="zt01137p052.pdf" docOrigin="Zootaxa 1137" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Neoathyreus (Mesoathyreus) guyanensis Howden, 2006, new species" docType="treatment" docVersion="7" lastPageNumber="51" masterDocId="FFB996621959FFEEFF9CFB0C3B2E025F" masterDocTitle="New species and a new subgenus of South American Neoathyreus Howden and Martínez (Coleoptera: Geotrupidae: Athyreini)" masterLastPageNumber="52" masterPageNumber="37" pageNumber="50" updateTime="1698215286212" updateUser="plazi">
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<mods:mods id="76E84480020174EAF4FAD65E01DF78E9" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="0C72EA26CAC5E054AE9180BDCFEF43EA">
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<mods:title id="3D2C947FD5AFB15CCEBD33AE2121301F">New species and a new subgenus of South American Neoathyreus Howden and Martínez (Coleoptera: Geotrupidae: Athyreini)</mods:title>
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</mods:titleInfo>
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<mods:name id="6EEA44C3BA7E88C797219FA43EC2AAD5" type="personal">
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<mods:role id="14E21FB2FAD19021953C18B3C5EAD918">
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<mods:roleTerm id="3FFD3C3EB5E3BC42AE5D52B434D9D423">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="29D10A36C8E10ACB6574A3C18A337091">Howden, Henry F.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="C81712B18E2B6DBB2C5C99FBA524138C">text</mods:typeOfResource>
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<mods:titleInfo id="D8272DD7D8787C03260E8E0A83F3701A">
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<mods:title id="5F6FA120D7C69A0DDD6CBFE4B75E5373">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="0BD17A23BD8D614B9543902660446CEB">
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<mods:date id="9510BC418A91DB3BD900C06C30AD33F0">2006</mods:date>
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<mods:detail id="2BEB09A353E90E0AB2462E803ACECF9D" type="volume">
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<mods:number id="E60EB0291FBFEF8CDC6710960DA42629">1137</mods:number>
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</mods:detail>
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<mods:extent id="6E4D679E0966904E2C0EC5726FC1155E" unit="page">
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<mods:start id="D24D8D6CBC5FE63B3BDF738CD8A90FAF">37</mods:start>
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<mods:end id="8FFA59FCE14068690C0426BDC82F879A">52</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="365B046ECAA57FF647DDD813C2B3495D">journal article</mods:classification>
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<mods:identifier id="069D09728B94AC0C5255CFCF08372153" type="DOI">10.5281/zenodo.273398</mods:identifier>
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<mods:identifier id="037CD24DA40119860E8F1DAE13A8F35A" type="GBIF-Dataset">ae452b89-2798-4a2a-8bd9-963dead78ebc</mods:identifier>
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<mods:identifier id="C654C96E9B5558B997E478058ED8B61E" type="ISSN">11755326</mods:identifier>
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<mods:identifier id="427E8211342FCE5CC176AE48AEF29B47" type="Zenodo-Dep">273398</mods:identifier>
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</mods:mods>
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<treatment id="0380EE1A1954FFE0FE94F8E0391C07B9" ID-DOI="http://doi.org/10.5281/zenodo.6260975" ID-GBIF-Taxon="119345039" ID-Zenodo-Dep="6260975" LSID="urn:lsid:plazi:treatment:0380EE1A1954FFE0FE94F8E0391C07B9" httpUri="http://treatment.plazi.org/id/0380EE1A1954FFE0FE94F8E0391C07B9" lastPageId="14" lastPageNumber="51" pageId="13" pageNumber="50">
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<subSubSection id="C3330C871954FFE3FE94F8E03F170659" box="[264,1081,1004,1030]" pageId="13" pageNumber="50" type="nomenclature">
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<paragraph id="8B965F0C1954FFE3FE94F8E03F170659" blockId="13.[264,1081,1004,1030]" box="[264,1081,1004,1030]" pageId="13" pageNumber="50">
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<heading id="D0DEE8601954FFE3FE94F8E03F170659" bold="true" box="[264,1081,1004,1030]" fontSize="11" level="1" pageId="13" pageNumber="50" reason="1">
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||
<emphasis id="B95D831E1954FFE3FE94F8E03F170659" bold="true" box="[264,1081,1004,1030]" pageId="13" pageNumber="50">
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<taxonomicName id="4C29248F1954FFE3FE94F8E039DD0659" box="[264,755,1004,1030]" class="Insecta" family="Bolboceratidae" genus="Neoathyreus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="50" phylum="Arthropoda" rank="species" species="guyanensis" status="sp. nov." subGenus="Mesoathyreus">
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<emphasis id="B95D831E1954FFE3FE94F8E03A8F0659" bold="true" box="[264,417,1004,1030]" italics="true" pageId="13" pageNumber="50">Neoathyreus</emphasis>
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(
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<emphasis id="B95D831E1954FFE3FE2EF8E039750659" bold="true" box="[434,603,1004,1030]" italics="true" pageId="13" pageNumber="50">Mesoathyreus</emphasis>
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)
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<emphasis id="B95D831E1954FFE3FDF0F8E039DD0659" bold="true" box="[620,755,1004,1030]" italics="true" pageId="13" pageNumber="50">guyanensis</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A26E3E651954FFE3FC9EF8E038BC0659" box="[770,914,1004,1030]" pageId="13" pageNumber="50" rank="species">new species</taxonomicNameLabel>
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(
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<figureCitation id="131243891954FFE3FC3EF8E03F1F0659" box="[930,1073,1004,1030]" captionStart="FIGURES 16 – 21" captionStartId="12.[264,382,1784,1808]" captionTargetBox="[274,1313,592,1772]" captionTargetId="figure@12.[274,1314,592,1772]" captionTargetPageId="12" captionText="FIGURES 16 – 21. Neoathyreus (Mesoathyreus) guyanensis, n. sp.: 16, Dorsum of male major; 17, Male major, head and pronotum, lateral view; 18, Dorsum of female; 19, Head of female, note right mandible; 20, Male genitalia, dorsal view; 21, Male genitalia, lateral view. Scale lines for genitalia = 0.2 mm." httpUri="https://zenodo.org/record/273402/files/figure.png" pageId="13" pageNumber="50">Figs. 16–21</figureCitation>
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)
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3330C871954FFE0FE94FF3A3FAF00B1" lastPageId="14" lastPageNumber="51" pageId="13" pageNumber="50" type="materials_examined">
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<paragraph id="8B965F0C1954FFE3FE94FF3A3F3804F6" blockId="13.[264,1323,1078,1905]" pageId="13" pageNumber="50">
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<emphasis id="B95D831E1954FFE3FE94FF3A3AAF060F" bold="true" box="[264,385,1078,1104]" pageId="13" pageNumber="50">
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||
<typeStatus id="5492E1AE1954FFE3FE94FF3A3A52060F" box="[264,380,1078,1104]" pageId="13" pageNumber="50" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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Male, length
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<quantity id="4CD1F2E91954FFE3FDB9FF3B39BF060E" box="[549,657,1079,1105]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" pageId="13" pageNumber="50" unit="mm" value="14.5">14.5 mm</quantity>
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, greatest width 9.0 mm. Dorsally shiny reddish brown. Clypeus (
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<figureCitation id="131243891954FFE3FEE4FF533AE20626" box="[376,460,1119,1145]" captionStart="FIGURES 16 – 21" captionStartId="12.[264,382,1784,1808]" captionTargetBox="[274,1313,592,1772]" captionTargetId="figure@12.[274,1314,592,1772]" captionTargetPageId="12" captionText="FIGURES 16 – 21. Neoathyreus (Mesoathyreus) guyanensis, n. sp.: 16, Dorsum of male major; 17, Male major, head and pronotum, lateral view; 18, Dorsum of female; 19, Head of female, note right mandible; 20, Male genitalia, dorsal view; 21, Male genitalia, lateral view. Scale lines for genitalia = 0.2 mm." httpUri="https://zenodo.org/record/273402/files/figure.png" pageId="13" pageNumber="50">Fig. 17</figureCitation>
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) with pronounced, erect horn,
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<quantity id="4CD1F2E91954FFE3FCA8FF5338A30626" box="[820,909,1119,1145]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="13" pageNumber="50" unit="mm" value="2.1">2.1 mm</quantity>
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long from anterior edge of clypeus to apex; anterior clypeal carina on each side extending medially up anterior face of horn, joining
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<quantity id="4CD1F2E91954FFE3FEFEFFA33A880696" box="[354,422,1199,1225]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="50" unit="mm" value="1.0">1 mm</quantity>
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from apex of horn; posterior carina on each side extending up posterior side of horn to apex. Lateral clypeal tubercle on each side above antennal insertion small; clypeal surface between carinae with scattered small punctures. Vertex slightly convex medially anterior to shallow, transverse depression between eyes; surface with widely scattered, small punctures. Pronotum (
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<figureCitation id="131243891954FFE3FD4BFE43381E0736" box="[727,816,1359,1385]" captionStart="FIGURES 16 – 21" captionStartId="12.[264,382,1784,1808]" captionTargetBox="[274,1313,592,1772]" captionTargetId="figure@12.[274,1314,592,1772]" captionTargetPageId="12" captionText="FIGURES 16 – 21. Neoathyreus (Mesoathyreus) guyanensis, n. sp.: 16, Dorsum of male major; 17, Male major, head and pronotum, lateral view; 18, Dorsum of female; 19, Head of female, note right mandible; 20, Male genitalia, dorsal view; 21, Male genitalia, lateral view. Scale lines for genitalia = 0.2 mm." httpUri="https://zenodo.org/record/273402/files/figure.png" pageId="13" pageNumber="50">Fig. 16</figureCitation>
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) at anterior margin behind each eye with small fovea, diameter slightly more than width of adjacent marginal bead; anterior half of pronotum transversely concave for distance equal to width of head; surface of concavity anterior to ends of inner carinae with scattered punctures, surface of anterior half of concavity between carinae alutaceous, impunctate, posterior to that surface with scattered small punctures; most pronotal punctures each with short, fine, erect seta. Outer pronotal carina on each side indicated by irregular, impunctate, very slightly elevated area; lateral pronotal margin not indented below lateral fovea. Scutellum, elytra, and ventral characters as described in subgeneric description. Genitalia as in
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<figureCitation id="131243891954FFE3FC19FD833F3C04F6" box="[901,1042,1679,1705]" captionStart="FIGURES 16 – 21" captionStartId="12.[264,382,1784,1808]" captionTargetBox="[274,1313,592,1772]" captionTargetId="figure@12.[274,1314,592,1772]" captionTargetPageId="12" captionText="FIGURES 16 – 21. Neoathyreus (Mesoathyreus) guyanensis, n. sp.: 16, Dorsum of male major; 17, Male major, head and pronotum, lateral view; 18, Dorsum of female; 19, Head of female, note right mandible; 20, Male genitalia, dorsal view; 21, Male genitalia, lateral view. Scale lines for genitalia = 0.2 mm." httpUri="https://zenodo.org/record/273402/files/figure.png" pageId="13" pageNumber="50">Figs. 20, 21</figureCitation>
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.
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</paragraph>
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<paragraph id="8B965F0C1954FFE0FEA4FDBA3F8E03A1" blockId="13.[264,1323,1078,1905]" lastBlockId="14.[264,1324,284,1510]" lastPageId="14" lastPageNumber="51" pageId="13" pageNumber="50">
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||
<emphasis id="B95D831E1954FFE3FEA4FDBA3A86048F" bold="true" box="[312,424,1718,1744]" pageId="13" pageNumber="50">Allotype.</emphasis>
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Female, length
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<quantity id="4CD1F2E91954FFE3FDEDFDBB39CE048E" box="[625,736,1719,1745]" metricMagnitude="-2" metricUnit="m" metricValue="1.46" pageId="13" pageNumber="50" unit="mm" value="14.6">14.6 mm</quantity>
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, greatest width 9.0 mm. Differing from male as follows: clypeus (
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<figureCitation id="131243891954FFE3FE46FDD3394904A6" box="[474,615,1759,1785]" captionStart="FIGURES 16 – 21" captionStartId="12.[264,382,1784,1808]" captionTargetBox="[274,1313,592,1772]" captionTargetId="figure@12.[274,1314,592,1772]" captionTargetPageId="12" captionText="FIGURES 16 – 21. Neoathyreus (Mesoathyreus) guyanensis, n. sp.: 16, Dorsum of male major; 17, Male major, head and pronotum, lateral view; 18, Dorsum of female; 19, Head of female, note right mandible; 20, Male genitalia, dorsal view; 21, Male genitalia, lateral view. Scale lines for genitalia = 0.2 mm." httpUri="https://zenodo.org/record/273402/files/figure.png" pageId="13" pageNumber="50">Figs. 18, 19</figureCitation>
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||
) lacking horn, median tubercle about twice height of lateral tubercle on each side, anterior carina on each side joining medially just anterior to apex of median tubercle; surface of clypeus and vertex more closely punctate than in male, punctures small, usually separated by two or more diameters, most punctures each with small anterior tubercle and short, fine central seta; pronotum (
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<figureCitation id="131243891957FFE0FC77FA103F6F0369" box="[1003,1089,284,310]" captionStart="FIGURES 16 – 21" captionStartId="12.[264,382,1784,1808]" captionTargetBox="[274,1313,592,1772]" captionTargetId="figure@12.[274,1314,592,1772]" captionTargetPageId="12" captionText="FIGURES 16 – 21. Neoathyreus (Mesoathyreus) guyanensis, n. sp.: 16, Dorsum of male major; 17, Male major, head and pronotum, lateral view; 18, Dorsum of female; 19, Head of female, note right mandible; 20, Male genitalia, dorsal view; 21, Male genitalia, lateral view. Scale lines for genitalia = 0.2 mm." httpUri="https://zenodo.org/record/273402/files/figure.png" pageId="14" pageNumber="51">Fig. 18</figureCitation>
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||
) with low, rounded inner and outer carinae, inner carinae slightly more elevated in anterior half of length; pronotal concavity shallow; except for carinae, entire pronotum punctatetuberculate as on vertex, spacing closer, particularly on pronotal sides; pronotal margin below lateral fovea slightly sinuous; pygidium apically more widely rounded than in male; foretibia with seven teeth on outer margin, minute eighth basal tooth indicated on right tibia.
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</paragraph>
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||
<paragraph id="8B965F0C1957FFE0FEA4F9073FAF00B1" blockId="14.[264,1324,284,1510]" pageId="14" pageNumber="51">
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||
<emphasis id="B95D831E1957FFE0FEA4F9073AFF007A" bold="true" box="[312,465,523,549]" pageId="14" pageNumber="51">
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||
<typeStatus id="5492E1AE1957FFE0FEA4F9073A5B007A" box="[312,373,523,549]" pageId="14" pageNumber="51">Type</typeStatus>
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Series.
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</emphasis>
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||
<typeStatus id="5492E1AE1957FFE0FE47F90039650079" box="[475,587,524,550]" pageId="14" pageNumber="51" type="holotype">Holotype</typeStatus>
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, male, [
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<collectingCountry id="F33E1F9C1957FFE0FD37F90038270079" box="[683,777,524,550]" name="French Guiana" pageId="14" pageNumber="51">Guyane</collectingCountry>
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=
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||
<collectingCountry id="F33E1F9C1957FFE0FCBEF90038F70079" box="[802,985,524,550]" name="French Guiana" pageId="14" pageNumber="51">French Guiana</collectingCountry>
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], Dd. Saramaca Pk. Rte des Compagnons,
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<collectingCountry id="F33E1F9C1957FFE0FE2EF93839230011" box="[434,525,564,590]" name="French Guiana" pageId="14" pageNumber="51">Guyane</collectingCountry>
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||
Fse,
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<date id="FF9779CC1957FFE0FDD3F93839F80011" box="[591,726,564,590]" pageId="14" pageNumber="51" value="1984-09-11">11.IX.1984</date>
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||
, M. Duranton Recolt. (
|
||
<collectionCode id="ED38C7C91957FFE0FC76F9383F690011" box="[1002,1095,564,590]" pageId="14" pageNumber="51">CMNC</collectionCode>
|
||
). Allotype, female, Dd. Saramaca, Mgne des Singes,
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<collectingCountry id="F33E1F9C1957FFE0FD02F95039D70029" box="[670,761,604,630]" name="French Guiana" pageId="14" pageNumber="51">Guyane</collectingCountry>
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||
Fse,
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||
<date id="FF9779CC1957FFE0FCA0F95038980029" box="[828,950,604,630]" pageId="14" pageNumber="51" value="1984-09-08">8.IX.1984</date>
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||
, M. Duranton Recol. (
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||
<collectionCode id="ED38C7C91957FFE0FB59F9503E310029" box="[1221,1311,604,630]" pageId="14" pageNumber="51">HAHC</collectionCode>
|
||
).
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||
<typeStatus id="5492E1AE1957FFE0FE94F9883A5200C1" box="[264,380,644,670]" pageId="14" pageNumber="51" type="paratype">Paratypes</typeStatus>
|
||
,
|
||
<specimenCount id="9D2F94851957FFE0FE16F9883AF000C1" box="[394,478,644,670]" pageId="14" pageNumber="51" type="male">1 male</specimenCount>
|
||
,
|
||
<specimenCount id="9D2F94851957FFE0FE70F988394A00C1" box="[492,612,644,670]" pageId="14" pageNumber="51" type="female">2 females</specimenCount>
|
||
:
|
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<specimenCount id="9D2F94851957FFE0FDEDF98839F200C1" box="[625,732,644,670]" pageId="14" pageNumber="51" type="female">1 female</specimenCount>
|
||
, same data as allotype except
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<date id="FF9779CC1957FFE0FBD6F9883FEF00C1" box="[1098,1217,644,670]" pageId="14" pageNumber="51" value="1985-08">VIII.1985</date>
|
||
;
|
||
<specimenCount id="9D2F94851957FFE0FB53F9883E0A00C1" box="[1231,1316,644,670]" pageId="14" pageNumber="51" type="male">1 male</specimenCount>
|
||
,
|
||
<collectingCountry id="F33E1F9C1957FFE0FE94F9A03A4D0099" box="[264,355,684,710]" name="French Guiana" pageId="14" pageNumber="51">Guyane</collectingCountry>
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Francaise, Kourou RN 1 Pk84, cryldé,
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<date id="FF9779CC1957FFE0FCD9F9A038EA0099" box="[837,964,684,710]" pageId="14" pageNumber="51" value="2001-10-30">30.X.2001</date>
|
||
, M. Duranton coll.;
|
||
<specimenCount id="9D2F94851957FFE0FB27F9A03E090099" box="[1211,1319,684,710]" pageId="14" pageNumber="51" type="female">1 female</specimenCount>
|
||
, same data as male
|
||
<typeStatus id="5492E1AE1957FFE0FE78F9D8396700B1" box="[484,585,724,750]" pageId="14" pageNumber="51" type="paratype">paratype</typeStatus>
|
||
except Pk 71,
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||
<date id="FF9779CC1957FFE0FD6AF9D8387E00B1" box="[758,848,724,750]" pageId="14" pageNumber="51" value="2001-10">X.2001</date>
|
||
.
|
||
<typeStatus id="5492E1AE1957FFE0FCC0F9D838E000B1" box="[860,974,724,750]" pageId="14" pageNumber="51" type="paratype">Paratypes</typeStatus>
|
||
are in:
|
||
<collectionCode id="ED38C7C91957FFE0FBBAF9D83F5300B1" box="[1062,1149,724,750]" pageId="14" pageNumber="51">HAHC</collectionCode>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3330C871957FFE0FEA4F9F0381D07C9" pageId="14" pageNumber="51" type="discussion">
|
||
<paragraph id="8B965F0C1957FFE0FEA4F9F03E3606F9" blockId="14.[264,1324,284,1510]" pageId="14" pageNumber="51">
|
||
<emphasis id="B95D831E1957FFE0FEA4F9F03A9F0149" bold="true" box="[312,433,764,790]" pageId="14" pageNumber="51">Remarks.</emphasis>
|
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Variation between the two female
|
||
<typeStatus id="5492E1AE1957FFE0FCC9F9F038EB0149" box="[853,965,764,790]" pageId="14" pageNumber="51" type="paratype">paratypes</typeStatus>
|
||
and the allotype is slight, the two
|
||
<typeStatus id="5492E1AE1957FFE0FEA0F8283A820161" box="[316,428,804,830]" pageId="14" pageNumber="51" type="paratype">paratypes</typeStatus>
|
||
measure 13.5 and
|
||
<quantity id="4CD1F2E91957FFE0FD17F82839DA0161" box="[651,756,804,830]" metricMagnitude="-2" metricUnit="m" metricValue="1.43" pageId="14" pageNumber="51" unit="mm" value="14.3">14.3 mm</quantity>
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in length and 8.0 and 9.0 in greatest width. The smaller female has six teeth on the foretibia, the larger one seven. The larger female has a slightly deeper pronotal concavity, but other variation is not obvious. The one male
|
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<typeStatus id="5492E1AE1957FFE0FE94F8903A4301E9" box="[264,365,924,950]" pageId="14" pageNumber="51" type="paratype">paratype</typeStatus>
|
||
measures
|
||
<quantity id="4CD1F2E91957FFE0FE75F890397C01E9" box="[489,594,924,950]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="14" pageNumber="51" unit="mm" value="12.5">12.5 mm</quantity>
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in length and
|
||
<quantity id="4CD1F2E91957FFE0FD65F890387D01E9" box="[761,851,924,950]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="14" pageNumber="51" unit="mm" value="7.1">7.1 mm</quantity>
|
||
in greatest width. Its small size, making it a minor male, causes it to resemble a female rather than the major male
|
||
<typeStatus id="5492E1AE1957FFE0FB19F8C83FC10181" box="[1157,1263,964,990]" pageId="14" pageNumber="51" type="holotype">holotype</typeStatus>
|
||
. It is similar to the females except that the median clypeal tubercle is distinctly higher, three or more times the height of the lateral tubercles. The pronotal concavity resembles those of the larger females, and there is a small, almost impunctate, alutaceous area near the middle of the concavity. Other dorsal characters resemble those of a female. The outer margin of the foretibia has seven teeth. The male genital characters resemble those of the
|
||
<typeStatus id="5492E1AE1957FFE0FB35FF803E3A06F9" box="[1193,1300,1164,1190]" pageId="14" pageNumber="51" type="holotype">holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B965F0C1957FFE0FEA4FFB8381D07C9" blockId="14.[264,1324,284,1510]" pageId="14" pageNumber="51">
|
||
The minor male and the females will key to couplet
|
||
<quantity id="4CD1F2E91957FFE0FC47FFB83F330691" box="[987,1053,1204,1230]" metricMagnitude="-1" metricUnit="m" metricValue="6.604" pageId="14" pageNumber="51" unit="in" value="26.0">26 in</quantity>
|
||
my 1985 revision of
|
||
<taxonomicName id="4C29248F1957FFE0FE94FFD03AB506AA" box="[264,411,1244,1269]" class="Insecta" family="Bolboceratidae" genus="Neoathyreus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="51" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95D831E1957FFE0FE94FFD03AB506AA" box="[264,411,1244,1269]" italics="true" pageId="14" pageNumber="51">Neoathyreus</emphasis>
|
||
</taxonomicName>
|
||
. The male major, lacking a distinct outer carina, will not key past couplet 10, partly because it does not fit the generic description of either
|
||
<taxonomicName id="4C29248F1957FFE0FC53FE083F4C0742" box="[975,1122,1284,1309]" class="Insecta" family="Bolboceratidae" genus="Neoathyreus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="51" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95D831E1957FFE0FC53FE083F4C0742" box="[975,1122,1284,1309]" italics="true" pageId="14" pageNumber="51">Neoathyreus</emphasis>
|
||
</taxonomicName>
|
||
or
|
||
<emphasis id="B95D831E1957FFE0FB14FE083FD90742" box="[1160,1271,1284,1309]" italics="true" pageId="14" pageNumber="51">
|
||
<taxonomicName id="4C29248F1957FFE0FB14FE083FDD0742" box="[1160,1267,1284,1309]" class="Insecta" family="Bolboceratidae" genus="Athyreus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="51" phylum="Arthropoda" rank="genus">Athyreus</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
The sexual differences, a character of
|
||
<emphasis id="B95D831E1957FFE0FD2CFE203830071A" box="[688,798,1324,1349]" italics="true" pageId="14" pageNumber="51">
|
||
<taxonomicName id="4C29248F1957FFE0FD2CFE203834071A" box="[688,794,1324,1349]" class="Insecta" family="Bolboceratidae" genus="Athyreus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="51" phylum="Arthropoda" rank="genus">Athyreus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and the unique blunt apical lobe of each mandible (
|
||
<figureCitation id="131243891957FFE0FE1AFE583AF20731" box="[390,476,1364,1390]" captionStart="FIGURES 16 – 21" captionStartId="12.[264,382,1784,1808]" captionTargetBox="[274,1313,592,1772]" captionTargetId="figure@12.[274,1314,592,1772]" captionTargetPageId="12" captionText="FIGURES 16 – 21. Neoathyreus (Mesoathyreus) guyanensis, n. sp.: 16, Dorsum of male major; 17, Male major, head and pronotum, lateral view; 18, Dorsum of female; 19, Head of female, note right mandible; 20, Male genitalia, dorsal view; 21, Male genitalia, lateral view. Scale lines for genitalia = 0.2 mm." httpUri="https://zenodo.org/record/273402/files/figure.png" pageId="14" pageNumber="51">Fig. 19</figureCitation>
|
||
) distinguish this species from any other
|
||
<taxonomicName id="4C29248F1957FFE0FC22FE583F7F0732" box="[958,1105,1364,1389]" class="Insecta" family="Bolboceratidae" genus="Neoathyreus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="51" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95D831E1957FFE0FC22FE583F7F0732" box="[958,1105,1364,1389]" italics="true" pageId="14" pageNumber="51">Neoathyreus</emphasis>
|
||
</taxonomicName>
|
||
and it is therefore placed in the separate subgenus,
|
||
<taxonomicName id="4C29248F1957FFE0FD14FE70380507CA" box="[648,811,1404,1429]" class="Insecta" family="Bolboceratidae" genus="Mesoathyreus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="51" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95D831E1957FFE0FD14FE70380507CA" box="[648,811,1404,1429]" italics="true" pageId="14" pageNumber="51">Mesoathyreus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3330C871957FFE0FEA4FEA8391C07B9" pageId="14" pageNumber="51" type="etymology">
|
||
<paragraph id="8B965F0C1957FFE0FEA4FEA8391C07B9" blockId="14.[264,1324,284,1510]" pageId="14" pageNumber="51">
|
||
<emphasis id="B95D831E1957FFE0FEA4FEA83AEA07E1" bold="true" box="[312,452,1444,1470]" pageId="14" pageNumber="51">Etymology.</emphasis>
|
||
The species name “
|
||
<taxonomicName id="4C29248F1957FFE0FD5BFEA8386107E1" box="[711,847,1444,1470]" class="Insecta" family="Bolboceratidae" genus="Neoathyreus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="51" phylum="Arthropoda" rank="species" species="guyanensis">guyanensis</taxonomicName>
|
||
” refers to
|
||
<collectingCountry id="F33E1F9C1957FFE0FC47FEA83FBC07E1" box="[987,1170,1444,1470]" name="French Guiana" pageId="14" pageNumber="51">French Guiana</collectingCountry>
|
||
, the area in which the species occurs.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |