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<document id="5CC1409EBF98375AFB37DFA364E771D4" ID-CLB-Dataset="7750" ID-DOI="10.11646/zootaxa.4938.4.1" ID-GBIF-Dataset="e5d1920d-c769-4aea-a8cb-237a0e58ae4c" ID-ISSN="1175-5326" ID-Zenodo-Dep="4574809" ID-ZooBank="6FEECE37-B2D3-4AE3-9878-CF212420AF9C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614750655839" checkinUser="plazi" docAuthor="Babenko, Anatoly, Kuznetsova, Natalia, Nakamori, Taizo &amp; Shveenkova, Yulia" docDate="2021" docId="BF05878D2329FFE1FF498CBFFEB6CE16" docLanguage="en" docName="zootaxa.4938.4.1.pdf" docOrigin="Zootaxa 4938 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudachorutes variabilis Babenko &amp; Kuznetsova &amp; Nakamori &amp; Shveenkova 2021, sp. nov." docType="treatment" docVersion="6" lastPageNumber="363" masterDocId="433CFFF52323FFEDFFDE8D09FF9DCA57" masterDocTitle="A review of Pseudachorutes Tullberg, 1871 (Collembola, Neanuridae) from the East Asia, with description of six new species" masterLastPageNumber="391" masterPageNumber="351" pageNumber="361" updateTime="1698931878726" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="161CA0380232EC085651E02334483A3E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="055719A0B83E6CF51C4BD1402E29ABCB">
<mods:title id="7010F243F0EA611E958DE91DFAD2C899">A review of Pseudachorutes Tullberg, 1871 (Collembola, Neanuridae) from the East Asia, with description of six new species</mods:title>
</mods:titleInfo>
<mods:name id="7CCB19D071A89EAB0F96D756FA35A6D8" type="personal">
<mods:role id="BEF738FE3517C39D75CBAC2FF1B12FAF">
<mods:roleTerm id="ED10EB19797DD6EAA3A4B5936B84E0C5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="762DCEF69E3BC758F01F048462BB28C0">Babenko, Anatoly</mods:namePart>
</mods:name>
<mods:name id="4B8769FFE0C035BEC5A73FE47C04D387" type="personal">
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</mods:role>
<mods:namePart id="D15F1690AD6523B38697063D6ED1C3FC">Kuznetsova, Natalia</mods:namePart>
<mods:affiliation id="0446668AF2E715E4C220B9B4E1306739">Moscow State Pedagogical University, Moscow 129278, Russia</mods:affiliation>
</mods:name>
<mods:name id="69CBF5E263DDFACAB2E78A751C44E486" type="personal">
<mods:role id="74332A38AF2D6E077852E5E62BA1B74C">
<mods:roleTerm id="D90A685CC8984AC48D374AC70A144CF3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="13BBC9871DA667220B3B9465CF0765E9">Nakamori, Taizo</mods:namePart>
<mods:affiliation id="5B2ADCC58FC22072D1561C66BEFCBFE3">Department of Environment and Natural Sciences, Graduate School of Environment and Information Sciences, Yokohama National University, Yokohama 240 - 8501, Japan</mods:affiliation>
</mods:name>
<mods:name id="AF76B5CB48A710313937866E4BE0908C" type="personal">
<mods:role id="7C10B0C2476E721E125D535C2F1E4C0C">
<mods:roleTerm id="C9C400C535E2BC06CD21ABF14CC76F28">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="23805518AEEEA8101B0F2D02FC3288DB">Shveenkova, Yulia</mods:namePart>
<mods:affiliation id="1FCC9F54A0437A94CD45E19FA7D68BAC">State Natural Reserve Privolzhskaya lesostep, Penza 440031, Russia</mods:affiliation>
</mods:name>
<mods:typeOfResource id="F85E4B12DA879CE264AC01D3735B8320">text</mods:typeOfResource>
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<mods:title id="509ECF08AE95C7E62A41744D581FEDD3">Zootaxa</mods:title>
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<mods:part id="1F782C3A0C24B953D6396D81B747122D">
<mods:date id="3EA75EE9713CAD98AF18FD13968D7A2A">2021</mods:date>
<mods:detail id="A122494AD51C9530E4DF392E85917190" type="pubDate">
<mods:number id="19DBB1BD7B0E1D5371C921D95DD82979">2021-03-02</mods:number>
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<mods:detail id="A800969115BFEF655CB1D1197BA2DD08" type="volume">
<mods:number id="C330C918CF922C72A2E090947009517B">4938</mods:number>
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<mods:classification id="BCA75A9B0395A66BDAA650A77B68396B">journal article</mods:classification>
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<mods:identifier id="8F7071890E80699D4998BF32E57A946C" type="DOI">10.11646/zootaxa.4938.4.1</mods:identifier>
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<mods:identifier id="5F44C11385F88F9E84A7F02DF5C65845" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="359DE7F3975FEA5CEBE6689840D177E9" type="Zenodo-Dep">4574809</mods:identifier>
<mods:identifier id="392A739D836C15778404F5EF0963E3A1" type="ZooBank">6FEECE37-B2D3-4AE3-9878-CF212420AF9C</mods:identifier>
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<treatment id="BF05878D2329FFE1FF498CBFFEB6CE16" ID-DOI="http://doi.org/10.5281/zenodo.4574819" ID-GBIF-Taxon="180568727" ID-Zenodo-Dep="4574819" LSID="urn:lsid:plazi:treatment:BF05878D2329FFE1FF498CBFFEB6CE16" httpUri="http://treatment.plazi.org/id/BF05878D2329FFE1FF498CBFFEB6CE16" lastPageId="12" lastPageNumber="363" pageId="10" pageNumber="361">
<subSubSection id="7FB665102329FFE7FF498CBFFCFACB86" box="[151,871,438,465]" pageId="10" pageNumber="361" type="nomenclature">
<paragraph id="3713369B2329FFE7FF498CBFFCFACB86" blockId="10.[151,871,438,501]" box="[151,871,438,465]" pageId="10" pageNumber="361">
<heading id="6C5B81F72329FFE7FF498CBFFCFACB86" bold="true" box="[151,871,438,465]" fontSize="11" level="1" pageId="10" pageNumber="361" reason="1">
<emphasis id="05D8EA892329FFE7FF498CBFFCFACB86" bold="true" box="[151,871,438,465]" pageId="10" pageNumber="361">
<taxonomicName id="F0AC4D182329FFE7FF498CBFFC9ECB87" authority="Babenko &amp; Shveenkova" authorityName="Babenko &amp; Kuznetsova &amp; Nakamori &amp; Shveenkova" authorityYear="2021" box="[151,771,438,465]" class="Entognatha" family="Neanuridae" genus="Pseudachorutes" kingdom="Animalia" order="Collembola" pageId="10" pageNumber="361" phylum="Arthropoda" rank="species" species="variabilis" status="sp. nov.">
<emphasis id="05D8EA892329FFE7FF498CBFFE52CB86" bold="true" box="[151,463,438,465]" italics="true" pageId="10" pageNumber="361">Pseudachorutes variabilis</emphasis>
Babenko &amp; Shveenkova
</taxonomicName>
<taxonomicNameLabel id="1EEB57F22329FFE7FCD48CBEFCFACB86" box="[778,871,439,465]" pageId="10" pageNumber="361" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="7FB665102329FFE7FF498CD2FE54CBA2" box="[151,457,474,501]" pageId="10" pageNumber="361" type="description">
<paragraph id="3713369B2329FFE7FF498CD2FE54CBA2" blockId="10.[151,871,438,501]" box="[151,457,474,501]" pageId="10" pageNumber="361">
<figureCitation id="AF972A1E2329FFE7FF498CD2FEF6CBA2" box="[151,363,474,501]" captionStart="FIGURES 31–39" captionStartId="11.[151,264,1906,1930]" captionTargetBox="[241,1346,192,1874]" captionTargetId="figure-20@11.[232,1356,181,1882]" captionTargetPageId="11" captionText="FIGURES 31–39. Pseudachorutes variabilis sp. nov. (31–35, 37–39) and P. subcrassus (36): 31, 33, head chaetotaxy (two different specimen); 32, 34, chaetotaxy of Abd. V–VI (two different specimen); 34a, shape of setae (Japanese specimen); 35–36, labium; 37, tip of tibiotarsus with unguis; 38, right and left mandibles; 39, Ant. III–IV (ventral side). Scales: Figs 31–34 = 0.1 mm, 35–39 = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.4574825" httpUri="https://zenodo.org/record/4574825/files/figure.png" pageId="10" pageNumber="361">Figs 31–35, 37–39</figureCitation>
,
<tableCitation id="7A2E03202329FFE7FEA88CD2FE54CBA2" box="[374,457,475,501]" captionStart="TABLE 1" captionStartId="34.[151,239,587,613]" captionText="TABLE 1. Main morphological characters of the East Asian species of the genus Pseudachorutes" pageId="10" pageNumber="361">Table 1</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="7FB665102329FFE7FF498F2AFD72CE2A" pageId="10" pageNumber="361" type="materials_examined">
<paragraph id="3713369B2329FFE7FF498F2AFB83CE46" blockId="10.[151,1437,546,2013]" pageId="10" pageNumber="361">
<emphasis id="05D8EA892329FFE7FF498F2AFED9C86B" bold="true" box="[151,324,546,573]" pageId="10" pageNumber="361">Type material.</emphasis>
<materialsCitation id="87C43CC62329FFE7FE958F2AFD53C8D2" ID-GBIF-Occurrence="3053021328" collectingDate="2017-08-08" collectionCode="MSPU" collectorName="N. Kuznetsova &amp; A. Kuprin &amp; A. Geraskina" country="Russia" county="Far East" elevation="930" latitude="45.138332" location="Sikhote-Alin State Nature Reserve" longLatPrecision="11" longitude="135.887" municipality="Southern Primor'e" pageId="10" pageNumber="361" specimenCount="1" typeStatus="holotype">
<collectingCountry id="4FBB760B2329FFE7FE958F2AFE07C86A" box="[331,410,547,573]" name="Russia" pageId="10" pageNumber="361">Russia</collectingCountry>
:
<typeStatus id="E81788392329FFE7FE7A8F2AFD94C86A" box="[420,521,547,573]" pageId="10" pageNumber="361" type="holotype">holotype</typeStatus>
, female,
<collectingCounty id="DE724E172329FFE7FDB38F2AFD51C86A" box="[621,716,547,573]" pageId="10" pageNumber="361">Far East</collectingCounty>
,
<collectingMunicipality id="D777ACE12329FFE7FD088F2AFC34C86A" box="[726,937,547,573]" pageId="10" pageNumber="361">Southern Primor’e</collectingMunicipality>
,
<location id="327360402329FFE7FC6A8F2AFAA7C86A" LSID="urn:lsid:plazi:treatment:BF05878D2329FFE1FF498CBFFEB6CE16:327360402329FFE7FC6A8F2AFAA7C86A" box="[948,1338,547,573]" country="Russia" county="Far East" latitude="45.138332" longLatPrecision="11" longitude="135.887" municipality="Southern Primor'e" name="Sikhote-Alin State Nature Reserve" pageId="10" pageNumber="361">Sikhote-Alin State Nature Reserve</location>
, coniferous forest with
<taxonomicName id="F0AC4D182329FFE7FEE18F4EFD10C836" box="[319,653,583,609]" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" order="Ericales" pageId="10" pageNumber="361" phylum="Tracheophyta" rank="species" species="brachycarpum">
<emphasis id="05D8EA892329FFE7FEE18F4EFD10C836" box="[319,653,583,609]" italics="true" pageId="10" pageNumber="361">Rhododendron brachycarpum</emphasis>
</taxonomicName>
,
<geoCoordinate id="5298505C2329FFE7FD478F4FFC92C836" box="[665,783,582,609]" degrees="45" direction="north" minutes="8.30" orientation="latitude" pageId="10" pageNumber="361" precision="9" value="45.138332">45°8.30’N</geoCoordinate>
<geoCoordinate id="5298505C2329FFE7FCCA8F4FFC38C836" box="[788,933,582,609]" degrees="135" direction="east" minutes="53.22" orientation="longitude" pageId="10" pageNumber="361" precision="9" value="135.887">135°53.22’E</geoCoordinate>
, ~
<quantity id="F0549B7E2329FFE7FC628F4EFB9EC836" box="[956,1027,583,609]" metricMagnitude="2" metricUnit="m" metricValue="9.3" pageId="10" pageNumber="361" unit="m" value="930.0">
<elevation id="BC81D1A82329FFE7FC628F4EFB9EC836" box="[956,1027,583,609]" metricMagnitude="2" metricUnit="m" metricValue="9.3" pageId="10" pageNumber="361" unit="m" value="930.0">930 m</elevation>
</quantity>
alt.,
<date id="4312105B2329FFE7FBE98F4EFB74C836" box="[1079,1257,583,609]" pageId="10" pageNumber="361" value="2017-08-08">
<collectingDate id="5356E9B32329FFE7FBE98F4EFB74C836" box="[1079,1257,583,609]" pageId="10" pageNumber="361" value="2017-08-08">08August 2017</collectingDate>
</date>
.
<collectorName id="9A59534D2329FFE7FB2F8F4EFA05C836" box="[1265,1432,583,609]" pageId="10" pageNumber="361">N. Kuznetsova</collectorName>
,
<collectorName id="9A59534D2329FFE7FF498F62FE95C8D2" box="[151,264,619,645]" pageId="10" pageNumber="361">A. Kuprin</collectorName>
&amp;
<collectorName id="9A59534D2329FFE7FEF58F62FE22C8D2" box="[299,447,619,645]" pageId="10" pageNumber="361">A. Geraskina</collectorName>
leg. [
<collectionCode id="51BDAE5E2329FFE7FE228F62FDD6C8D2" box="[508,587,619,645]" pageId="10" pageNumber="361">MSPU</collectionCode>
collection]
</materialsCitation>
.
<materialsCitation id="87C43CC62329FFE7FD068F62FCC1C8FE" ID-GBIF-Occurrence="3053021308" country="Russia" elevation="200" latitude="43.648" location="Ussuri State Nature Reserve" longLatPrecision="68" longitude="132.35167" pageId="10" pageNumber="361" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<typeStatus id="E81788392329FFE7FD068F62FCD5C8D2" box="[728,840,619,645]" pageId="10" pageNumber="361" type="paratype">Paratypes</typeStatus>
,
<specimenCount id="21AAFD122329FFE7FC8D8F63FC2AC8D2" box="[851,951,618,645]" pageId="10" pageNumber="361" type="female">
<emphasis id="05D8EA892329FFE7FC8D8F63FCFCC8D3" bold="true" box="[851,865,618,644]" pageId="10" pageNumber="361">1</emphasis>
female
</specimenCount>
, same region,
<location id="327360402329FFE7FB878F62FA05C8D2" LSID="urn:lsid:plazi:treatment:BF05878D2329FFE1FF498CBFFEB6CE16:327360402329FFE7FB878F62FA05C8D2" box="[1113,1432,619,645]" country="Russia" latitude="43.648" longLatPrecision="68" longitude="132.35167" name="Ussuri State Nature Reserve" pageId="10" pageNumber="361">Ussuri State Nature Reserve</location>
, forest with
<taxonomicName id="F0AC4D182329FFE7FEC28F86FE5BC8FE" authorityName="Rusek" authorityYear="1991" box="[284,454,655,681]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="10" pageNumber="361" phylum="Tracheophyta" rank="species" species="sibiricus">
<emphasis id="05D8EA892329FFE7FEC28F86FE5BC8FE" box="[284,454,655,681]" italics="true" pageId="10" pageNumber="361">Pinus sibiricus</emphasis>
</taxonomicName>
,
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<geoCoordinate id="5298505C2329FFE7FDBB8F86FD75C8FE" box="[613,744,655,681]" degrees="132" direction="east" minutes="21.1" orientation="longitude" pageId="10" pageNumber="361" precision="92" value="132.35167">132°21.1’E</geoCoordinate>
, ~
<quantity id="F0549B7E2329FFE7FCD18F86FCC1C8FE" box="[783,860,655,681]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="361" unit="m" value="200.0">
<specimenCount id="21AAFD122329FFE7FCD18F86FCC1C8FE" box="[783,860,655,681]" pageId="10" pageNumber="361" type="generic">
<elevation id="BC81D1A82329FFE7FCD18F86FCC1C8FE" box="[783,860,655,681]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="361" unit="m" value="200.0">200 m</elevation>
</specimenCount>
</quantity>
</materialsCitation>
. alt.,
<date id="4312105B2329FFE7FC438F86FBAAC8FE" box="[925,1079,655,681]" pageId="10" pageNumber="361" value="2016-07-22">22 July 2016</date>
. N. Kuznetsova &amp; M. Potapov leg.;
<materialsCitation id="87C43CC62329FFE7FF0F8FBBFE61C8A6" ID-GBIF-Occurrence="3053021355" collectingDate="2016-07-24" collectorName="N. Kuznetsova &amp; M. Potapov" country="Russia" elevation="380" latitude="43.636665" location="Nature Reserve" longLatPrecision="92" longitude="132.34967" pageId="10" pageNumber="361" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="21AAFD122329FFE7FF0F8FBBFEAAC89A" box="[209,311,690,717]" pageId="10" pageNumber="361" type="female">
<emphasis id="05D8EA892329FFE7FF0F8FBBFF42C89B" bold="true" box="[209,223,690,716]" pageId="10" pageNumber="361">1</emphasis>
female
</specimenCount>
, same
<location id="327360402329FFE7FE598FBAFDA7C89A" LSID="urn:lsid:plazi:treatment:BF05878D2329FFE1FF498CBFFEB6CE16:327360402329FFE7FE598FBAFDA7C89A" box="[391,570,691,717]" country="Russia" latitude="43.636665" longLatPrecision="92" longitude="132.34967" name="Nature Reserve" pageId="10" pageNumber="361">Nature Reserve</location>
, mixed forest,
<geoCoordinate id="5298505C2329FFE7FD388FBAFCC1C89A" box="[742,860,691,717]" degrees="43" direction="north" minutes="38.2" orientation="latitude" pageId="10" pageNumber="361" precision="92" value="43.636665">43°38.2’N</geoCoordinate>
<geoCoordinate id="5298505C2329FFE7FCBB8FBAFC6BC89A" box="[869,1014,691,717]" degrees="132" direction="east" minutes="20.98" orientation="longitude" pageId="10" pageNumber="361" precision="9" value="132.34967">132°20.98’E</geoCoordinate>
, ~
<quantity id="F0549B7E2329FFE7FBCC8FBAFBC1C89A" box="[1042,1116,691,717]" metricMagnitude="2" metricUnit="m" metricValue="3.8" pageId="10" pageNumber="361" unit="m" value="380.0">
<elevation id="BC81D1A82329FFE7FBCC8FBAFBC1C89A" box="[1042,1116,691,717]" metricMagnitude="2" metricUnit="m" metricValue="3.8" pageId="10" pageNumber="361" unit="m" value="380.0">380 m</elevation>
</quantity>
alt., litter,
<date id="4312105B2329FFE7FB028FBAFAE8C89A" box="[1244,1397,691,717]" pageId="10" pageNumber="361" value="2016-07-24">
<collectingDate id="5356E9B32329FFE7FB028FBAFAE8C89A" box="[1244,1397,691,717]" pageId="10" pageNumber="361" value="2016-07-24">24 July 2016</collectingDate>
</date>
.
<collectorName id="9A59534D2329FFE7FA5F8FBAFE86C8A6" pageId="10" pageNumber="361">N. Kuznetsova</collectorName>
&amp;
<collectorName id="9A59534D2329FFE7FE9C8FDEFE5AC8A6" box="[322,455,727,753]" pageId="10" pageNumber="361">M. Potapov</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="87C43CC62329FFE7FDD78FDFFBC2C942" ID-GBIF-Occurrence="3053021334" collectingDate="2017-08-11" collectorName="N. Kuznetsova &amp; A. Kuprin &amp; A. Geraskina" country="Russia" elevation="500" latitude="44.022" location="Chuguevka" longLatPrecision="11" longitude="134.15033" pageId="10" pageNumber="361" specimenCount="1" specimenCount-juvenile="1" typeStatus="paratype">
<specimenCount id="21AAFD122329FFE7FDD78FDFFDE0C8A6" box="[521,637,726,753]" pageId="10" pageNumber="361" type="juvenile">
<emphasis id="05D8EA892329FFE7FDD78FDFFD8AC8A7" bold="true" box="[521,535,726,752]" pageId="10" pageNumber="361">1</emphasis>
juvenile
</specimenCount>
, same region, but ~
<quantity id="F0549B7E2329FFE7FC808FDEFC3AC8A6" box="[862,935,727,753]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="361" unit="km" value="30.0">30 km</quantity>
SE of
<location id="327360402329FFE7FC278FDEFBE6C8A6" LSID="urn:lsid:plazi:treatment:BF05878D2329FFE1FF498CBFFEB6CE16:327360402329FFE7FC278FDEFBE6C8A6" box="[1017,1147,727,753]" country="Russia" latitude="44.022" longLatPrecision="11" longitude="134.15033" name="Chuguevka" pageId="10" pageNumber="361">Chuguevka</location>
, mixed forest,
<geoCoordinate id="5298505C2329FFE7FAF88FDEFA01C8A6" box="[1318,1436,727,753]" degrees="44" direction="north" minutes="1.32" orientation="latitude" pageId="10" pageNumber="361" precision="9" value="44.022">44°1.32’N</geoCoordinate>
<geoCoordinate id="5298505C2329FFE7FF498FF2FE87C942" box="[151,282,763,789]" degrees="134" direction="east" minutes="9.02" orientation="longitude" pageId="10" pageNumber="361" precision="9" value="134.15033">134°9.02’E</geoCoordinate>
,
<quantity id="F0549B7E2329FFE7FEFB8FF3FEF7C942" box="[293,362,762,789]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="361" unit="m" value="500.0">
<elevation id="BC81D1A82329FFE7FEFB8FF3FEF7C942" box="[293,362,762,789]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="361" unit="m" value="500.0">500 m</elevation>
</quantity>
alt.,
<date id="4312105B2329FFE7FE7E8FF2FDCFC942" box="[416,594,763,789]" pageId="10" pageNumber="361" value="2017-08-11">
<collectingDate id="5356E9B32329FFE7FE7E8FF2FDCFC942" box="[416,594,763,789]" pageId="10" pageNumber="361" value="2017-08-11">11 August 2017</collectingDate>
</date>
.
<collectorName id="9A59534D2329FFE7FD848FF2FC9FC942" box="[602,770,763,789]" pageId="10" pageNumber="361">N. Kuznetsova</collectorName>
,
<collectorName id="9A59534D2329FFE7FCD58FF2FCE6C942" box="[779,891,763,789]" pageId="10" pageNumber="361">A. Kuprin</collectorName>
&amp;
<collectorName id="9A59534D2329FFE7FC458FF2FBB3C942" box="[923,1070,763,789]" pageId="10" pageNumber="361">A. Geraskina</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="87C43CC62329FFE7FBB48FF3FC13C90A" ID-GBIF-Occurrence="3053021357" collectingDate="2016-07-27" collectorName="N. Kuznetsova &amp; M. Potapov" country="Russia" elevation="100" latitude="43.114666" location="State Nature Reserve" longLatPrecision="11" longitude="131.48717" pageId="10" pageNumber="361" specimenCount="2" specimenCount-juvenile="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="21AAFD122329FFE7FBB48FF3FB29C942" box="[1130,1204,762,789]" pageId="10" pageNumber="361" type="male">
<emphasis id="05D8EA892329FFE7FBB48FF3FBE5C943" bold="true" box="[1130,1144,762,788]" pageId="10" pageNumber="361">1</emphasis>
male
</specimenCount>
and
<specimenCount id="21AAFD122329FFE7FB368FF3FAC4C942" box="[1256,1369,762,789]" pageId="10" pageNumber="361" type="juvenile">
<emphasis id="05D8EA892329FFE7FB368FF3FB6BC943" bold="true" box="[1256,1270,762,788]" pageId="10" pageNumber="361">1</emphasis>
juvenile
</specimenCount>
, same region, but «Kedrovaya Pad»
<location id="327360402329FFE7FE2F8E16FD7BC96E" LSID="urn:lsid:plazi:treatment:BF05878D2329FFE1FF498CBFFEB6CE16:327360402329FFE7FE2F8E16FD7BC96E" box="[497,742,799,825]" country="Russia" latitude="43.114666" longLatPrecision="11" longitude="131.48717" name="State Nature Reserve" pageId="10" pageNumber="361">State Nature Reserve</location>
, mixed coniferous-broadleaf forest,
<geoCoordinate id="5298505C2329FFE7FB598E16FB60C96E" box="[1159,1277,799,825]" degrees="43" direction="north" minutes="6.88" orientation="latitude" pageId="10" pageNumber="361" precision="9" value="43.114666">43°6.88’N</geoCoordinate>
<geoCoordinate id="5298505C2329FFE7FAD98E16FA05C96E" box="[1287,1432,799,825]" degrees="131" direction="east" minutes="29.23" orientation="longitude" pageId="10" pageNumber="361" precision="9" value="131.48717">131°29.23’E</geoCoordinate>
, ~
<quantity id="F0549B7E2329FFE7FF788E4AFF73C90A" box="[166,238,835,861]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="361" unit="m" value="100.0">
<elevation id="BC81D1A82329FFE7FF788E4AFF73C90A" box="[166,238,835,861]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="361" unit="m" value="100.0">100 m</elevation>
</quantity>
alt., litter,
<date id="4312105B2329FFE7FEB48E4AFD9DC90A" box="[362,512,835,861]" pageId="10" pageNumber="361" value="2016-07-27">
<collectingDate id="5356E9B32329FFE7FEB48E4AFD9DC90A" box="[362,512,835,861]" pageId="10" pageNumber="361" value="2016-07-27">27 July 2016</collectingDate>
</date>
.
<collectorName id="9A59534D2329FFE7FDD48E4AFD2CC90A" box="[522,689,835,861]" pageId="10" pageNumber="361">N. Kuznetsova</collectorName>
&amp;
<collectorName id="9A59534D2329FFE7FD098E4AFCC7C90A" box="[727,858,835,861]" pageId="10" pageNumber="361">M. Potapov</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="87C43CC62329FFE7FC458E4BFD74C9F2" ID-GBIF-Occurrence="3053021331" collectingDate="2018-09-19" collectingDateMax="2018-09-20" collectingDateMin="2018-09-19" collectorName="A. Kuprin" country="Russia" county="Chuguev District" elevation="550" latitude="43.600666" location="Mount Oblachnaya" longLatPrecision="11" longitude="134.193" municipality="National Park" pageId="10" pageNumber="361" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="21AAFD122329FFE7FC458E4BFC62C90A" box="[923,1023,834,861]" pageId="10" pageNumber="361" type="female">
<emphasis id="05D8EA892329FFE7FC458E4BFC34C90B" bold="true" box="[923,937,834,860]" pageId="10" pageNumber="361">1</emphasis>
female
</specimenCount>
, same region,
<collectingCounty id="DE724E172329FFE7FB7D8E4AFAFAC90A" box="[1187,1383,835,861]" pageId="10" pageNumber="361">Chuguev District</collectingCounty>
,
<collectingMunicipality id="D777ACE12329FFE7FAAC8E4AFE8EC9D6" pageId="10" pageNumber="361">National Park</collectingMunicipality>
«Zov Tigra»,
<location id="327360402329FFE7FE648E6EFD0BC9D6" LSID="urn:lsid:plazi:treatment:BF05878D2329FFE1FF498CBFFEB6CE16:327360402329FFE7FE648E6EFD0BC9D6" box="[442,662,871,897]" country="Russia" county="Chuguev District" latitude="43.600666" longLatPrecision="11" longitude="134.193" municipality="National Park" name="Mount Oblachnaya" pageId="10" pageNumber="361">Mount Oblachnaya</location>
,
<location id="327360402329FFE7FD7D8E6EFC1CC9D6" LSID="urn:lsid:plazi:treatment:BF05878D2329FFE1FF498CBFFEB6CE16:327360402329FFE7FD7D8E6EFC1CC9D6" box="[675,897,871,897]" country="Russia" county="Chuguev District" latitude="43.600666" longLatPrecision="11" longitude="134.193" municipality="National Park" name="Ussuri River valley" pageId="10" pageNumber="361">Ussuri River valley</location>
,
<geoCoordinate id="5298505C2329FFE7FC518E6EFB8EC9D6" box="[911,1043,871,897]" degrees="43" direction="north" minutes="36.04" orientation="latitude" pageId="10" pageNumber="361" precision="9" value="43.600666">43°36.04’N</geoCoordinate>
<geoCoordinate id="5298505C2329FFE7FBC28E6FFB31C9D6" box="[1052,1196,870,897]" degrees="134" direction="east" minutes="11.58" orientation="longitude" pageId="10" pageNumber="361" precision="9" value="134.193">134°11.58’E</geoCoordinate>
, ~
<quantity id="F0549B7E2329FFE7FB0F8E6FFA87C9D6" box="[1233,1306,870,897]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="10" pageNumber="361" unit="m" value="550.0">
<elevation id="BC81D1A82329FFE7FB0F8E6FFA87C9D6" box="[1233,1306,870,897]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="10" pageNumber="361" unit="m" value="550.0">550 m</elevation>
</quantity>
alt., mixed forest, litter,
<date id="4312105B2329FFE7FEF58E82FDABC9F2" box="[299,566,907,933]" pageId="10" pageNumber="361" value="2018-09-19" valueMax="2018-09-20" valueMin="2018-09-19">
<collectingDate id="5356E9B32329FFE7FEF58E82FDABC9F2" box="[299,566,907,933]" pageId="10" pageNumber="361" value="2018-09-19" valueMax="2018-09-20" valueMin="2018-09-19">19–20 September 2018</collectingDate>
</date>
.
<collectorName id="9A59534D2329FFE7FD9E8E82FD29C9F2" box="[576,692,907,933]" pageId="10" pageNumber="361">A. Kuprin</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="87C43CC62329FFE7FD298E83FA0AC99E" ID-GBIF-Occurrence="3053021364" collectingDate="2006-06-29" collectorName="A. Brinev &amp; A. Polyakov" country="Russia" county="Territory" elevation="1950" location="Bureinskaya Rassoshina River" municipality="Ezop Mt. Range" pageId="10" pageNumber="361" specimenCount="3" specimenCount-female="3" stateProvince="Khabarovsk" typeStatus="paratype">
<specimenCount id="21AAFD122329FFE7FD298E83FCF5C9F2" box="[759,872,906,933]" pageId="10" pageNumber="361" type="female">
<emphasis id="05D8EA892329FFE7FD298E83FC98C9F3" bold="true" box="[759,773,906,932]" pageId="10" pageNumber="361">3</emphasis>
females
</specimenCount>
, northern part of
<collectingRegion id="F568F8792329FFE7FBEF8E82FB25C9F2" box="[1073,1208,907,933]" country="Russia" name="Khabarovsk" pageId="10" pageNumber="361">Khabarovsk</collectingRegion>
<collectingCounty id="DE724E172329FFE7FB1F8E82FAB8C9F2" box="[1217,1317,907,933]" pageId="10" pageNumber="361">Territory</collectingCounty>
,
<collectingMunicipality id="D777ACE12329FFE7FAEC8E82FF7CC99E" pageId="10" pageNumber="361">Ezop Mt. Range</collectingMunicipality>
,
<location id="327360402329FFE7FF338EA6FDD8C99E" LSID="urn:lsid:plazi:treatment:BF05878D2329FFE1FF498CBFFEB6CE16:327360402329FFE7FF338EA6FDD8C99E" box="[237,581,943,969]" country="Russia" county="Territory" municipality="Ezop Mt. Range" name="Bureinskaya Rassoshina River" pageId="10" pageNumber="361" stateProvince="Khabarovsk">Bureinskaya Rassoshina River</location>
, alpine belt,
<quantity id="F0549B7E2329FFE7FD0D8EA6FCF3C99E" box="[723,878,943,969]" metricMagnitude="3" metricUnit="m" metricValue="1.95" metricValueMax="2.0" metricValueMin="1.9" pageId="10" pageNumber="361" unit="m" value="1950.0" valueMax="2000.0" valueMin="1900.0">
<elevation id="BC81D1A82329FFE7FD0D8EA6FCF3C99E" box="[723,878,943,969]" metricMagnitude="3" metricUnit="m" metricValue="1.95" metricValueMax="2.0" metricValueMin="1.9" pageId="10" pageNumber="361" unit="m" value="1950.0" valueMax="2000.0" valueMin="1900.0">1900–2000 m</elevation>
</quantity>
alt.,
<date id="4312105B2329FFE7FC788EA6FBA2C99E" box="[934,1087,943,969]" pageId="10" pageNumber="361" value="2006-06-29">
<collectingDate id="5356E9B32329FFE7FC788EA6FBA2C99E" box="[934,1087,943,969]" pageId="10" pageNumber="361" value="2006-06-29">29 June 2006</collectingDate>
</date>
.
<collectorName id="9A59534D2329FFE7FB968EA6FB2BC99E" box="[1096,1206,943,969]" pageId="10" pageNumber="361">A. Brinev</collectorName>
&amp;
<collectorName id="9A59534D2329FFE7FB068EA6FAF9C99E" box="[1240,1380,943,969]" pageId="10" pageNumber="361">A. Polyakov</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="87C43CC62329FFE7FF498EDBFB83CE46" ID-GBIF-Occurrence="3053021363" collectingDate="2018-07" collectionCode="MSPU" collectorName="L. Lobkova" country="Russia" latitude="52.966667" location="Khalatyrsky" longLatPrecision="1080" longitude="158.81667" municipality="Petropavlovsk" pageId="10" pageNumber="361" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Kamchatka" typeStatus="paratype">
<specimenCount id="21AAFD122329FFE7FF498EDBFE98C9BA" box="[151,261,978,1005]" pageId="10" pageNumber="361" type="female">
<emphasis id="05D8EA892329FFE7FF498EDBFF38C9BB" bold="true" box="[151,165,978,1004]" pageId="10" pageNumber="361">2</emphasis>
females
</specimenCount>
and
<specimenCount id="21AAFD122329FFE7FEE08EDBFE13C9BA" box="[318,398,978,1005]" pageId="10" pageNumber="361" type="male">
<emphasis id="05D8EA892329FFE7FEE08EDBFED1C9BB" bold="true" box="[318,332,978,1004]" pageId="10" pageNumber="361">1</emphasis>
male
</specimenCount>
,
<collectingRegion id="F568F8792329FFE7FE458EDAFD80C9BA" box="[411,541,979,1005]" country="Russia" name="Kamchatka" pageId="10" pageNumber="361">Kamchatka</collectingRegion>
, environs of
<collectingMunicipality id="D777ACE12329FFE7FD6D8EDAFCCBC9BA" box="[691,854,979,1005]" pageId="10" pageNumber="361">Petropavlovsk</collectingMunicipality>
,
<location id="327360402329FFE7FCBD8EDAFC71C9BA" LSID="urn:lsid:plazi:treatment:BF05878D2329FFE1FF498CBFFEB6CE16:327360402329FFE7FCBD8EDAFC71C9BA" box="[867,1004,979,1005]" country="Russia" latitude="52.966667" longLatPrecision="1080" longitude="158.81667" municipality="Petropavlovsk" name="Khalatyrsky" pageId="10" pageNumber="361" stateProvince="Kamchatka">Khalatyrsky</location>
beach [~
<geoCoordinate id="5298505C2329FFE7FB8B8EDBFB27C9BA" box="[1109,1210,978,1005]" degrees="52" direction="north" minutes="58" orientation="latitude" pageId="10" pageNumber="361" precision="925" value="52.966667">52°58’N</geoCoordinate>
,
<geoCoordinate id="5298505C2329FFE7FB198EDBFAAAC9BA" box="[1223,1335,978,1005]" degrees="158" direction="east" minutes="49" orientation="longitude" pageId="10" pageNumber="361" precision="925" value="158.81667">158°49’E</geoCoordinate>
], seaside meadow, under
<taxonomicName id="F0AC4D182329FFE7FE978EFEFE02CE47" box="[329,415,1015,1040]" class="Liliopsida" family="Poaceae" genus="Leymus" kingdom="Plantae" order="Poales" pageId="10" pageNumber="361" phylum="Tracheophyta" rank="genus">
<emphasis id="05D8EA892329FFE7FE978EFEFE02CE47" box="[329,415,1015,1040]" italics="true" pageId="10" pageNumber="361">Leymus</emphasis>
</taxonomicName>
sp., sand,
<date id="4312105B2329FFE7FDCB8EFEFD1BCE46" box="[533,646,1015,1041]" pageId="10" pageNumber="361" value="2018-07">
<collectingDate id="5356E9B32329FFE7FDCB8EFEFD1BCE46" box="[533,646,1015,1041]" pageId="10" pageNumber="361" value="2018-07">July 2018</collectingDate>
</date>
.
<collectorName id="9A59534D2329FFE7FD4E8EFEFC89CE46" box="[656,788,1015,1041]" pageId="10" pageNumber="361">L. Lobkova</collectorName>
leg. [
<collectionCode id="51BDAE5E2329FFE7FC8F8EFEFC3DCE46" box="[849,928,1015,1041]" pageId="10" pageNumber="361">MSPU</collectionCode>
collection]
</materialsCitation>
</paragraph>
<paragraph id="3713369B2329FFE7FF198913FD72CE2A" blockId="10.[151,1437,546,2013]" pageId="10" pageNumber="361">
<emphasis id="05D8EA892329FFE7FF198913FE0ACE63" bold="true" box="[199,407,1050,1076]" pageId="10" pageNumber="361">
Other material.
<specimenCount id="21AAFD122329FFE7FE578913FE0ACE63" box="[393,407,1050,1076]" pageId="10" pageNumber="361" type="female">1</specimenCount>
</emphasis>
female,
<collectingCountry id="4FBB760B2329FFE7FE268912FDA6CE62" box="[504,571,1051,1077]" name="Japan" pageId="10" pageNumber="361">Japan</collectingCountry>
, Honshu Island,
<collectingRegion id="F568F8792329FFE7FD298912FC56CE62" box="[759,971,1051,1077]" country="Japan" name="Nagano" pageId="10" pageNumber="361">Nagano Prefecture</collectingRegion>
, E Chino city, Kitayama, surroundings of Mugikusa Hutte,
<geoCoordinate id="5298505C2329FFE7FE858936FE4CCE0E" box="[347,465,1087,1113]" degrees="36" direction="north" minutes="2.72" orientation="latitude" pageId="10" pageNumber="361" precision="9" value="36.045334">36°2.72’N</geoCoordinate>
<geoCoordinate id="5298505C2329FFE7FE078936FDF7CE0E" box="[473,618,1087,1113]" degrees="138" direction="east" minutes="21.92" orientation="longitude" pageId="10" pageNumber="361" precision="9" value="138.36533">138°21.92’E</geoCoordinate>
,
<quantity id="F0549B7E2329FFE7FDA88937FD56CE0E" box="[630,715,1086,1113]" metricMagnitude="3" metricUnit="m" metricValue="2.15" pageId="10" pageNumber="361" unit="m" value="2150.0">2150 m</quantity>
alt., mixed forest (
<emphasis id="05D8EA892329FFE7FC7C8936FBA1CE0F" box="[930,1084,1087,1113]" italics="true" pageId="10" pageNumber="361">
<taxonomicName id="F0AC4D182329FFE7FC7C8936FC73CE0E" box="[930,1006,1087,1113]" class="Magnoliopsida" family="Betulaceae" genus="Betula" kingdom="Plantae" order="Fagales" pageId="10" pageNumber="361" phylum="Tracheophyta" rank="genus">Betula</taxonomicName>
,
<taxonomicName id="F0AC4D182329FFE7FC248936FBA1CE0F" box="[1018,1084,1087,1112]" class="Pinopsida" family="Pinaceae" genus="Tsuga" kingdom="Plantae" order="Pinales" pageId="10" pageNumber="361" phylum="Tracheophyta" rank="genus">Tsuga</taxonomicName>
</emphasis>
,
<taxonomicName id="F0AC4D182329FFE7FB948936FB15CE0E" box="[1098,1160,1087,1113]" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="10" pageNumber="361" phylum="Tracheophyta" rank="genus">
<emphasis id="05D8EA892329FFE7FB948936FB15CE0E" box="[1098,1160,1087,1113]" italics="true" pageId="10" pageNumber="361">Abies</emphasis>
</taxonomicName>
), litter,
<date id="4312105B2329FFE7FB3D8936FA04CE0E" box="[1251,1433,1087,1113]" pageId="10" pageNumber="361" value="2016-08-10">10 August 2016</date>
. M. Potapov &amp; N. Kuznetsova leg. [
<collectionCode id="51BDAE5E2329FFE7FDFF896AFDEDCE2A" box="[545,624,1123,1149]" pageId="10" pageNumber="361">MSPU</collectionCode>
collection]
</paragraph>
</subSubSection>
<subSubSection id="7FB665102329FFE7FF19898FFCCACF66" pageId="10" pageNumber="361" type="diagnosis">
<paragraph id="3713369B2329FFE7FF19898FFCCACF66" blockId="10.[151,1437,546,2013]" pageId="10" pageNumber="361">
<emphasis id="05D8EA892329FFE7FF19898FFEDFCEF7" bold="true" box="[199,322,1158,1184]" pageId="10" pageNumber="361">Diagnosis.</emphasis>
Medium sized species. PAO rounded, slightly larger than nearest ocellus. Buccal cone elongate, complete set of labral setae (4/2334 totally), labium with seta L and labial organites additionally to 12 ordinary setae, 5 setae in perilabial area. Dorsal chaetotaxy of the
<taxonomicName id="F0AC4D182329FFE7FCC089C6FC1DCEBE" authorityName="Borner" authorityYear="1901" box="[798,896,1231,1257]" class="Entognatha" family="Neanuridae" genus="Pseudachorutes" kingdom="Animalia" order="Collembola" pageId="10" pageNumber="361" phylum="Arthropoda" rank="species" species="parvulus">
<emphasis id="05D8EA892329FFE7FCC089C6FC1DCEBE" box="[798,896,1231,1257]" italics="true" pageId="10" pageNumber="361">parvulus</emphasis>
</taxonomicName>
<typeStatus id="E81788392329FFE7FC5689C6FC26CEBE" box="[904,955,1231,1257]" pageId="10" pageNumber="361">type</typeStatus>
, Th. I with 3+3 setae, most of dorsal setae on abdominal tip more or less clearly modified, sensilla short (about as long as ordinary setae). Mucro with lateral lamella almost reaching tip. Each anal valve with two hr-setae.
</paragraph>
</subSubSection>
<subSubSection id="7FB665102329FFE1FF198833FC05CBA5" lastPageId="12" lastPageNumber="363" pageId="10" pageNumber="361" type="description">
<paragraph id="3713369B2329FFE7FF198833FB34CF2E" blockId="10.[151,1437,546,2013]" pageId="10" pageNumber="361">
<emphasis id="05D8EA892329FFE7FF198833FEC4CF03" bold="true" box="[199,345,1338,1364]" pageId="10" pageNumber="361">Description.</emphasis>
Length (without antennae) up to
<quantity id="F0549B7E2329FFE7FD0D8833FCA4CF02" box="[723,825,1338,1365]" metricMagnitude="-3" metricUnit="m" metricValue="1.52" pageId="10" pageNumber="361" unit="mm" value="1.52">1.52 mm</quantity>
, but usually smaller, holotype—
<quantity id="F0549B7E2329FFE7FB748832FA9CCF02" box="[1194,1281,1339,1365]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="10" pageNumber="361" unit="mm" value="1.4">1.4 mm</quantity>
. Colour greyblue, not especially dark. Tegument granulation rather coarse, especially in larger specimens.
</paragraph>
<paragraph id="3713369B2329FFE7FF19888AFB00CC7A" blockId="10.[151,1437,546,2013]" pageId="10" pageNumber="361">
Antennae slightly shorter than head, Ant. III–IV fused dorsally, ventral separation well marked. Ant. IV with simple or slightly divided apical vesicle, external ms, subapical or and seta i present; dorsal side of Ant. IV with six clearly differentiated, curved sensilla (S1–S4, S7–S8), ventral side with utmost few pointed, truncate or slightly clavate sensilliform setae (
<figureCitation id="AF972A1E2329FFE7FE1C88E6FD8ACC5E" box="[450,535,1519,1545]" captionStart="FIGURES 31–39" captionStartId="11.[151,264,1906,1930]" captionTargetBox="[241,1346,192,1874]" captionTargetId="figure-20@11.[232,1356,181,1882]" captionTargetPageId="11" captionText="FIGURES 31–39. Pseudachorutes variabilis sp. nov. (31–35, 37–39) and P. subcrassus (36): 31, 33, head chaetotaxy (two different specimen); 32, 34, chaetotaxy of Abd. V–VI (two different specimen); 34a, shape of setae (Japanese specimen); 35–36, labium; 37, tip of tibiotarsus with unguis; 38, right and left mandibles; 39, Ant. III–IV (ventral side). Scales: Figs 31–34 = 0.1 mm, 35–39 = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.4574825" httpUri="https://zenodo.org/record/4574825/files/figure.png" pageId="10" pageNumber="361">Fig. 39</figureCitation>
). Antennal organ of Ant. III typical, inner sensilla small, sgv slightly curved and clearly longer than sgd, ventral ms present. Ant. I–II with usual 7 and 12 setae, respectively.
</paragraph>
<paragraph id="3713369B2329FFE7FF198B3EFDE8CD52" blockId="10.[151,1437,546,2013]" pageId="10" pageNumber="361">
Head with 8+8 subequal ocelli. PAO rounded or slightly elliptic, consisting of 8–12 vesicles, slightly larger than nearest ocellus B, ratio 1.2–1.3: 1. Buccal cone elongate. Maxilla styliform with tiny apical teeth, lamellae not clearly seen. Mandibles thin and membranous, usually with four apical teeth (
<figureCitation id="AF972A1E2329FFE7FBE48B76FB0DCCCE" box="[1082,1168,1663,1689]" captionStart="FIGURES 31–39" captionStartId="11.[151,264,1906,1930]" captionTargetBox="[241,1346,192,1874]" captionTargetId="figure-20@11.[232,1356,181,1882]" captionTargetPageId="11" captionText="FIGURES 31–39. Pseudachorutes variabilis sp. nov. (31–35, 37–39) and P. subcrassus (36): 31, 33, head chaetotaxy (two different specimen); 32, 34, chaetotaxy of Abd. V–VI (two different specimen); 34a, shape of setae (Japanese specimen); 35–36, labium; 37, tip of tibiotarsus with unguis; 38, right and left mandibles; 39, Ant. III–IV (ventral side). Scales: Figs 31–34 = 0.1 mm, 35–39 = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.4574825" httpUri="https://zenodo.org/record/4574825/files/figure.png" pageId="10" pageNumber="361">Fig. 38</figureCitation>
). Distal edge of labrum pointed, number of labral setae as follows: 4/2334. Main part of labium with four proximal ordinary setae, distal seta L on tiny papilla and two labial organites; submentum and mentum with usual set of four setae each, i.e. 4+4 (
<figureCitation id="AF972A1E2329FFE7FAB18BCEFF2BCD52" captionStart="FIGURES 31–39" captionStartId="11.[151,264,1906,1930]" captionTargetBox="[241,1346,192,1874]" captionTargetId="figure-20@11.[232,1356,181,1882]" captionTargetPageId="11" captionText="FIGURES 31–39. Pseudachorutes variabilis sp. nov. (31–35, 37–39) and P. subcrassus (36): 31, 33, head chaetotaxy (two different specimen); 32, 34, chaetotaxy of Abd. V–VI (two different specimen); 34a, shape of setae (Japanese specimen); 35–36, labium; 37, tip of tibiotarsus with unguis; 38, right and left mandibles; 39, Ant. III–IV (ventral side). Scales: Figs 31–34 = 0.1 mm, 35–39 = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.4574825" httpUri="https://zenodo.org/record/4574825/files/figure.png" pageId="10" pageNumber="361">Fig. 35</figureCitation>
). Perilabial area usually with 5+5 setae.
</paragraph>
<paragraph id="3713369B2329FFE1FF198A06FBA4CA82" blockId="10.[151,1437,546,2013]" lastBlockId="12.[151,1437,151,1089]" lastPageId="12" lastPageNumber="363" pageId="10" pageNumber="361">
Most of dorsal setae, especially on abdominal tip modified. Degree of these modifications in different specimens highly variable: from truncated or slightly capitate setae (
<figureCitation id="AF972A1E2329FFE7FCC18A3AFC3DCD1A" box="[799,928,1843,1869]" captionStart="FIGURES 31–39" captionStartId="11.[151,264,1906,1930]" captionTargetBox="[241,1346,192,1874]" captionTargetId="figure-20@11.[232,1356,181,1882]" captionTargetPageId="11" captionText="FIGURES 31–39. Pseudachorutes variabilis sp. nov. (31–35, 37–39) and P. subcrassus (36): 31, 33, head chaetotaxy (two different specimen); 32, 34, chaetotaxy of Abd. V–VI (two different specimen); 34a, shape of setae (Japanese specimen); 35–36, labium; 37, tip of tibiotarsus with unguis; 38, right and left mandibles; 39, Ant. III–IV (ventral side). Scales: Figs 31–34 = 0.1 mm, 35–39 = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.4574825" httpUri="https://zenodo.org/record/4574825/files/figure.png" pageId="10" pageNumber="361">Figs 31–32</figureCitation>
) to distally widened and flattened ones (
<figureCitation id="AF972A1E2329FFE7FAB48A3AFE85CD26" captionStart="FIGURES 31–39" captionStartId="11.[151,264,1906,1930]" captionTargetBox="[241,1346,192,1874]" captionTargetId="figure-20@11.[232,1356,181,1882]" captionTargetPageId="11" captionText="FIGURES 31–39. Pseudachorutes variabilis sp. nov. (31–35, 37–39) and P. subcrassus (36): 31, 33, head chaetotaxy (two different specimen); 32, 34, chaetotaxy of Abd. V–VI (two different specimen); 34a, shape of setae (Japanese specimen); 35–36, labium; 37, tip of tibiotarsus with unguis; 38, right and left mandibles; 39, Ant. III–IV (ventral side). Scales: Figs 31–34 = 0.1 mm, 35–39 = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.4574825" httpUri="https://zenodo.org/record/4574825/files/figure.png" pageId="10" pageNumber="361">Figs 33–34, 34a</figureCitation>
), sensilla short (about as long as ordinary setae or even shorter), their number as usual, i.e. 22/11111. Main characteristics: head with unpaired d0, but without a0, setae sd2 at level or slightly posterior to d2 (
<figureCitation id="AF972A1E2329FFE7FA9E8A72FF2BCDEE" captionStart="FIGURES 31–39" captionStartId="11.[151,264,1906,1930]" captionTargetBox="[241,1346,192,1874]" captionTargetId="figure-20@11.[232,1356,181,1882]" captionTargetPageId="11" captionText="FIGURES 31–39. Pseudachorutes variabilis sp. nov. (31–35, 37–39) and P. subcrassus (36): 31, 33, head chaetotaxy (two different specimen); 32, 34, chaetotaxy of Abd. V–VI (two different specimen); 34a, shape of setae (Japanese specimen); 35–36, labium; 37, tip of tibiotarsus with unguis; 38, right and left mandibles; 39, Ant. III–IV (ventral side). Scales: Figs 31–34 = 0.1 mm, 35–39 = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.4574825" httpUri="https://zenodo.org/record/4574825/files/figure.png" pageId="10" pageNumber="361">Figs 31, 33</figureCitation>
), Th. I usually with 3+3 setae (but one female with abnormal 3+4 also seen), Th. II with a2-setae, dorso-external group with two setae (a3–a4) in front of p3–p4, only lateral m6 (S) in m-row, lateral ms present. Th. III identical, but without a2 and ms. Abd. I–III with only one seta (a3) in front of p3–p4, setae of m-row absent.Abd. IV with 9+9 setae in a–p rows, setae of m-row absent. Abd. V without p2 as usual (
<figureCitation id="AF972A1E232FFFE1FC748DB2FBB0CA82" box="[938,1069,187,213]" captionStart="FIGURES 31–39" captionStartId="11.[151,264,1906,1930]" captionTargetBox="[241,1346,192,1874]" captionTargetId="figure-20@11.[232,1356,181,1882]" captionTargetPageId="11" captionText="FIGURES 31–39. Pseudachorutes variabilis sp. nov. (31–35, 37–39) and P. subcrassus (36): 31, 33, head chaetotaxy (two different specimen); 32, 34, chaetotaxy of Abd. V–VI (two different specimen); 34a, shape of setae (Japanese specimen); 35–36, labium; 37, tip of tibiotarsus with unguis; 38, right and left mandibles; 39, Ant. III–IV (ventral side). Scales: Figs 31–34 = 0.1 mm, 35–39 = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.4574825" httpUri="https://zenodo.org/record/4574825/files/figure.png" pageId="12" pageNumber="363">Figs 32, 34</figureCitation>
).
</paragraph>
<caption id="63D366132328FFE6FF498A7BFE1ACDA1" ID-DOI="http://doi.org/10.5281/zenodo.4574825" ID-Zenodo-Dep="4574825" httpUri="https://zenodo.org/record/4574825/files/figure.png" pageId="11" pageNumber="362" startId="11.[151,264,1906,1930]" targetBox="[241,1346,192,1874]" targetPageId="11">
<paragraph id="3713369B2328FFE6FF498A7BFE1ACDA1" blockId="11.[151,1437,1906,2039]" pageId="11" pageNumber="362">
<emphasis id="05D8EA892328FFE6FF498A7BFECBCDDD" bold="true" box="[151,342,1906,1930]" pageId="11" pageNumber="362">FIGURES 31–39.</emphasis>
<taxonomicName id="F0AC4D182328FFE6FE818A7AFDF7CDDC" authority="Babenko &amp; Kuznetsova &amp; Nakamori &amp; Shveenkova, 2021" authorityName="Babenko &amp; Kuznetsova &amp; Nakamori &amp; Shveenkova" authorityYear="2021" box="[351,618,1907,1931]" class="Entognatha" family="Neanuridae" genus="Pseudachorutes" kingdom="Animalia" order="Collembola" pageId="11" pageNumber="362" phylum="Arthropoda" rank="species" species="variabilis" status="sp. nov.">
<emphasis id="05D8EA892328FFE6FE818A7AFDF7CDDC" box="[351,618,1907,1931]" italics="true" pageId="11" pageNumber="362">Pseudachorutes variabilis</emphasis>
</taxonomicName>
<emphasis id="05D8EA892328FFE6FDAD8A7AFD58CDDC" bold="true" box="[627,709,1907,1931]" pageId="11" pageNumber="362">
<taxonomicNameLabel id="1EEB57F22328FFE6FDAD8A7AFD58CDDC" box="[627,709,1907,1931]" pageId="11" pageNumber="362" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(31–35, 37–39) and
<taxonomicName id="F0AC4D182328FFE6FC7C8A7AFBB1CDDC" authorityName="Tullberg" authorityYear="1871" box="[930,1068,1907,1931]" class="Entognatha" family="Neanuridae" genus="Pseudachorutes" kingdom="Animalia" order="Collembola" pageId="11" pageNumber="362" phylum="Arthropoda" rank="species" species="subcrassus">
<emphasis id="05D8EA892328FFE6FC7C8A7AFBB1CDDC" box="[930,1068,1907,1931]" italics="true" pageId="11" pageNumber="362">P. subcrassus</emphasis>
</taxonomicName>
(36): 31, 33, head chaetotaxy (two different specimen); 32, 34, chaetotaxy of Abd. V–VI (two different specimen); 34a, shape of setae (Japanese specimen); 35–36, labium; 37, tip of tibiotarsus with unguis; 38, right and left mandibles; 39, Ant. III–IV (ventral side). Scales: Figs 31–34 = 0.1 mm, 35–39 = 0.01 mm.
</paragraph>
</caption>
<paragraph id="3713369B232FFFE1FF198DD6FC4BCBDE" blockId="12.[151,1437,151,1089]" pageId="12" pageNumber="363">Thoracic sterna without setae. Ventral tube with 4+4 setae, no seta on sternum of Abd. I, Abd. II with (3)4+4 ventral setae, Abd. III with 5–7 setae. Tenaculum with 3+3 teeth as usual. Furca not especially long. Manubrium with 8–9+8–9 setae on main part, 4 setae on each basolateral lobe and 2 basal setae in line. Dorsal side of dens with six setae and uniform coarse granulation, hyaline field on its ventral side rather large. Mucro with upturned tip and lateral lamella almost reaching tip. Each anal valve with two tiny hr-setae.</paragraph>
<paragraph id="3713369B232FFFE1FF198C9AFC05CBA5" blockId="12.[151,1437,151,1089]" pageId="12" pageNumber="363">
Legs I–III with 1, 2, 2 setae on upper subcoxae, 0, 2, 2 setae on lower subcoxae, 3, 7, 7 setae on coxae, 6, 6, 6 on trochanters, 13, 12, 11 setae on femora and 19, 19, 18 setae on tibiotarsi. Unguis with clear inner tooth in mid part of inner edge and pair of tiny lateral ones in basal half (
<figureCitation id="AF972A1E232FFFE1FCE98CD1FC16CBA5" box="[823,907,472,498]" captionStart="FIGURES 31–39" captionStartId="11.[151,264,1906,1930]" captionTargetBox="[241,1346,192,1874]" captionTargetId="figure-20@11.[232,1356,181,1882]" captionTargetPageId="11" captionText="FIGURES 31–39. Pseudachorutes variabilis sp. nov. (31–35, 37–39) and P. subcrassus (36): 31, 33, head chaetotaxy (two different specimen); 32, 34, chaetotaxy of Abd. V–VI (two different specimen); 34a, shape of setae (Japanese specimen); 35–36, labium; 37, tip of tibiotarsus with unguis; 38, right and left mandibles; 39, Ant. III–IV (ventral side). Scales: Figs 31–34 = 0.1 mm, 35–39 = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.4574825" httpUri="https://zenodo.org/record/4574825/files/figure.png" pageId="12" pageNumber="363">Fig. 37</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="7FB66510232FFFE1FF198CF2FD8BC980" pageId="12" pageNumber="363" type="etymology">
<paragraph id="3713369B232FFFE1FF198CF2FA85C842" blockId="12.[151,1437,151,1089]" box="[199,1304,507,533]" pageId="12" pageNumber="363">
<emphasis id="05D8EA89232FFFE1FF198CF2FED3C842" bold="true" box="[199,334,507,533]" pageId="12" pageNumber="363">Etymology.</emphasis>
The name of the new species reflects a high variability of the shape of its dorsal setae.
</paragraph>
<paragraph id="3713369B232FFFE1FF198F14FD8BC980" blockId="12.[151,1437,151,1089]" pageId="12" pageNumber="363">
<emphasis id="05D8EA89232FFFE1FF198F14FEA0C860" bold="true" box="[199,317,541,567]" pageId="12" pageNumber="363">Affinities.</emphasis>
The main diagnostic features of
<taxonomicName id="F0AC4D18232FFFE1FD788F17FCB2C860" authority="Babenko &amp; Kuznetsova &amp; Nakamori &amp; Shveenkova, 2021" authorityName="Babenko &amp; Kuznetsova &amp; Nakamori &amp; Shveenkova" authorityYear="2021" box="[678,815,541,568]" class="Entognatha" family="Neanuridae" genus="Pseudachorutes" kingdom="Animalia" order="Collembola" pageId="12" pageNumber="363" phylum="Arthropoda" rank="species" species="variabilis" status="sp. nov.">
<emphasis id="05D8EA89232FFFE1FD788F17FD2AC860" box="[678,695,542,567]" italics="true" pageId="12" pageNumber="363">P</emphasis>
.
<emphasis id="05D8EA89232FFFE1FD1A8F14FCB2C860" box="[708,815,541,567]" italics="true" pageId="12" pageNumber="363">variabilis</emphasis>
</taxonomicName>
<emphasis id="05D8EA89232FFFE1FCEB8F14FC13C860" bold="true" box="[821,910,541,567]" pageId="12" pageNumber="363">
<taxonomicNameLabel id="1EEB57F2232FFFE1FCEB8F14FC13C860" box="[821,910,541,567]" pageId="12" pageNumber="363" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are identical to those of
<taxonomicName id="F0AC4D18232FFFE1FB428F17FF4CC80D" authority="Tullberg, 1871" authorityName="Tullberg" authorityYear="1871" class="Entognatha" family="Neanuridae" genus="Pseudachorutes" kingdom="Animalia" order="Collembola" pageId="12" pageNumber="363" phylum="Arthropoda" rank="species" species="subcrassus">
<emphasis id="05D8EA89232FFFE1FB428F17FAACC860" box="[1180,1329,541,567]" italics="true" pageId="12" pageNumber="363">P. subcrassus</emphasis>
<bibRefCitation id="533D4B6A232FFFE1FAE88F14FF4CC80D" author="Tullberg, T." pageId="12" pageNumber="363" pagination="143 - 155" refId="ref31822" refString="Tullberg, T. (1871) F orteckning ofver Svenska Podurider. Ofversigt af Kongliga Vetenskaps-Akademiens Forhandlingar, 28, 143 - 155." type="journal article" year="1871">Tullberg, 1871</bibRefCitation>
</taxonomicName>
, which probably has a predominantly European range. Both these species are characterized by similar sensory equipment of Ant. IV, labrum with 4/2334 setae, labium with seta L and two organites additionally to 12 ordinary setae, practically identical mandibular shape, unguis with a clear inner tooth and mucro of similar shape. Both species also have the same
<typeStatus id="E8178839232FFFE1FE748FA0FE47C894" box="[426,474,681,707]" pageId="12" pageNumber="363">type</typeStatus>
of dorsal chaetotaxy with 3+3 setae on Th. I, Th. II with a2 setae, two setae in front of p3–p4 on Th. II–III and one a
<quantity id="F0549B7E232FFFE1FDDB8FC2FDB2C8B2" box="[517,559,715,741]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="12" pageNumber="363" unit="in" value="3.0">3 in</quantity>
front of p3–p4 on Abd. I–III. In addition to the differences in the shape of the dorsal setae, which is a highly variable feature in
<taxonomicName id="F0AC4D18232FFFE1FD628FE7FCD8C950" authority="Babenko &amp; Kuznetsova &amp; Nakamori &amp; Shveenkova, 2021" authorityName="Babenko &amp; Kuznetsova &amp; Nakamori &amp; Shveenkova" authorityYear="2021" box="[700,837,749,776]" class="Entognatha" family="Neanuridae" genus="Pseudachorutes" kingdom="Animalia" order="Collembola" pageId="12" pageNumber="363" phylum="Arthropoda" rank="species" species="variabilis" status="sp. nov.">
<emphasis id="05D8EA89232FFFE1FD628FE7FD50C950" box="[700,717,750,775]" italics="true" pageId="12" pageNumber="363">P</emphasis>
.
<emphasis id="05D8EA89232FFFE1FD048FE4FCD8C950" box="[730,837,749,775]" italics="true" pageId="12" pageNumber="363">variabilis</emphasis>
</taxonomicName>
<emphasis id="05D8EA89232FFFE1FC928FE4FC38C950" bold="true" box="[844,933,749,775]" pageId="12" pageNumber="363">
<taxonomicNameLabel id="1EEB57F2232FFFE1FC928FE4FC38C950" box="[844,933,749,775]" pageId="12" pageNumber="363" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, these species differ in a number of small but constant characters. For instance, labrum in
<taxonomicName id="F0AC4D18232FFFE1FD538E18FC84C97D" authority="Babenko &amp; Kuznetsova &amp; Nakamori &amp; Shveenkova, 2021" authorityName="Babenko &amp; Kuznetsova &amp; Nakamori &amp; Shveenkova" authorityYear="2021" box="[653,793,784,811]" class="Entognatha" family="Neanuridae" genus="Pseudachorutes" kingdom="Animalia" order="Collembola" pageId="12" pageNumber="363" phylum="Arthropoda" rank="species" species="variabilis" status="sp. nov.">
<emphasis id="05D8EA89232FFFE1FD538E18FD03C97D" box="[653,670,785,810]" italics="true" pageId="12" pageNumber="363">P</emphasis>
.
<emphasis id="05D8EA89232FFFE1FD708E19FC84C97D" box="[686,793,784,810]" italics="true" pageId="12" pageNumber="363">variabilis</emphasis>
</taxonomicName>
<emphasis id="05D8EA89232FFFE1FCFF8E19FCE1C97D" bold="true" box="[801,892,784,810]" pageId="12" pageNumber="363">
<taxonomicNameLabel id="1EEB57F2232FFFE1FCFF8E19FCE1C97D" box="[801,892,784,810]" pageId="12" pageNumber="363" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is pointed distally but rounded in
<taxonomicName id="F0AC4D18232FFFE1FADB8E18FA01C97D" authorityName="Tullberg" authorityYear="1871" box="[1285,1436,784,810]" class="Entognatha" family="Neanuridae" genus="Pseudachorutes" kingdom="Animalia" order="Collembola" pageId="12" pageNumber="363" phylum="Arthropoda" rank="species" species="subcrassus">
<emphasis id="05D8EA89232FFFE1FADB8E18FA01C97D" box="[1285,1436,784,810]" italics="true" pageId="12" pageNumber="363">P. subcrassus</emphasis>
</taxonomicName>
(cf.
<figureCitation id="AF972A1E232FFFE1FF1B8E3AFE88C91A" box="[197,277,818,845]" captionStart="FIGURES 31–39" captionStartId="11.[151,264,1906,1930]" captionTargetBox="[241,1346,192,1874]" captionTargetId="figure-20@11.[232,1356,181,1882]" captionTargetPageId="11" captionText="FIGURES 31–39. Pseudachorutes variabilis sp. nov. (31–35, 37–39) and P. subcrassus (36): 31, 33, head chaetotaxy (two different specimen); 32, 34, chaetotaxy of Abd. V–VI (two different specimen); 34a, shape of setae (Japanese specimen); 35–36, labium; 37, tip of tibiotarsus with unguis; 38, right and left mandibles; 39, Ant. III–IV (ventral side). Scales: Figs 31–34 = 0.1 mm, 35–39 = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.4574825" httpUri="https://zenodo.org/record/4574825/files/figure.png" pageId="12" pageNumber="363">Fig. 35</figureCitation>
and
<figureCitation id="AF972A1E232FFFE1FE938E3AFE3DC919" box="[333,416,819,846]" captionStart="FIGURES 31–39" captionStartId="11.[151,264,1906,1930]" captionTargetBox="[241,1346,192,1874]" captionTargetId="figure-20@11.[232,1356,181,1882]" captionTargetPageId="11" captionText="FIGURES 31–39. Pseudachorutes variabilis sp. nov. (31–35, 37–39) and P. subcrassus (36): 31, 33, head chaetotaxy (two different specimen); 32, 34, chaetotaxy of Abd. V–VI (two different specimen); 34a, shape of setae (Japanese specimen); 35–36, labium; 37, tip of tibiotarsus with unguis; 38, right and left mandibles; 39, Ant. III–IV (ventral side). Scales: Figs 31–34 = 0.1 mm, 35–39 = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.4574825" httpUri="https://zenodo.org/record/4574825/files/figure.png" pageId="12" pageNumber="363">Fig. 36</figureCitation>
), lateral teeth on unguis is located in basal part in
<taxonomicName id="F0AC4D18232FFFE1FC068E3AFBFEC91A" authority="Babenko &amp; Kuznetsova &amp; Nakamori &amp; Shveenkova, 2021" authorityName="Babenko &amp; Kuznetsova &amp; Nakamori &amp; Shveenkova" authorityYear="2021" box="[984,1123,819,845]" class="Entognatha" family="Neanuridae" genus="Pseudachorutes" kingdom="Animalia" order="Collembola" pageId="12" pageNumber="363" phylum="Arthropoda" rank="species" species="variabilis" status="sp. nov.">
<emphasis id="05D8EA89232FFFE1FC068E3AFC74C91B" box="[984,1001,819,844]" italics="true" pageId="12" pageNumber="363">P</emphasis>
.
<emphasis id="05D8EA89232FFFE1FC268E3AFBFEC91A" box="[1016,1123,819,845]" italics="true" pageId="12" pageNumber="363">variabilis</emphasis>
</taxonomicName>
<emphasis id="05D8EA89232FFFE1FBB58E3AFB58C91A" bold="true" box="[1131,1221,819,845]" pageId="12" pageNumber="363">
<taxonomicNameLabel id="1EEB57F2232FFFE1FBB58E3AFB58C91A" box="[1131,1221,819,845]" pageId="12" pageNumber="363" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, but subapically in
<emphasis id="05D8EA89232FFFE1FF498E5FFEAAC938" box="[151,311,853,879]" italics="true" pageId="12" pageNumber="363">
<taxonomicName id="F0AC4D18232FFFE1FF498E5FFEAFC938" authorityName="Tullberg" authorityYear="1871" box="[151,306,853,879]" class="Entognatha" family="Neanuridae" genus="Pseudachorutes" kingdom="Animalia" order="Collembola" pageId="12" pageNumber="363" phylum="Arthropoda" rank="species" species="subcrassus">P. subcrassus</taxonomicName>
.
</emphasis>
Kaprus’ &amp;
<bibRefCitation id="533D4B6A232FFFE1FE638E5CFDFCC938" author="Weiner, W. M." box="[445,609,853,879]" pageId="12" pageNumber="363" pagination="1 - 23" refId="ref30668" refString="Kaprus', I. J. &amp; Weiner, W. M. (2009) The genus Pseudachorutes Tullberg, 1871 (Collembola, Neanuridae) in the Ukraine with descriptions of new species. Zootaxa, 2166 (1), 1 - 23. https: // doi. org / 10.11646 / zootaxa. 2166.1.1" type="journal article" year="2009">Weiner (2009)</bibRefCitation>
suggested using the relative position of the setae d2 and sd2–sd3 on the head as an additional criterion for distinguishing between
<taxonomicName id="F0AC4D18232FFFE1FCF28E70FC3DC9C5" box="[812,928,888,914]" class="Entognatha" family="Neanuridae" genus="Pseudachorutes" kingdom="Animalia" order="Collembola" pageId="12" pageNumber="363" phylum="Arthropoda" rank="species" species="parvula">
<emphasis id="05D8EA89232FFFE1FCF28E70FC3DC9C5" box="[812,928,888,914]" italics="true" pageId="12" pageNumber="363">P. parvula</emphasis>
</taxonomicName>
and
<taxonomicName id="F0AC4D18232FFFE1FC078E70FBECC9C5" authorityName="Tullberg" authorityYear="1871" box="[985,1137,888,914]" class="Entognatha" family="Neanuridae" genus="Pseudachorutes" kingdom="Animalia" order="Collembola" pageId="12" pageNumber="363" phylum="Arthropoda" rank="species" species="subcrassus">
<emphasis id="05D8EA89232FFFE1FC078E70FBECC9C5" box="[985,1137,888,914]" italics="true" pageId="12" pageNumber="363">P. subcrassus</emphasis>
</taxonomicName>
, i.e. d2 at the level of sd2 (
<taxonomicName id="F0AC4D18232FFFE1FF7F8E92FE9EC9E2" authorityName="Borner" authorityYear="1901" box="[161,259,923,949]" class="Entognatha" family="Neanuridae" genus="Pseudachorutes" kingdom="Animalia" order="Collembola" pageId="12" pageNumber="363" phylum="Arthropoda" rank="species" species="parvulus">
<emphasis id="05D8EA89232FFFE1FF7F8E92FE9EC9E2" box="[161,259,923,949]" italics="true" pageId="12" pageNumber="363">parvulus</emphasis>
</taxonomicName>
) or sd3 (
<taxonomicName id="F0AC4D18232FFFE1FEB78E92FE79C9E2" authorityName="Tullberg" authorityYear="1871" box="[361,484,923,949]" class="Entognatha" family="Neanuridae" genus="Pseudachorutes" kingdom="Animalia" order="Collembola" pageId="12" pageNumber="363" phylum="Arthropoda" rank="species" species="subcrassus">
<emphasis id="05D8EA89232FFFE1FEB78E92FE79C9E2" box="[361,484,923,949]" italics="true" pageId="12" pageNumber="363">subcrassus</emphasis>
</taxonomicName>
). The position of these setae in
<taxonomicName id="F0AC4D18232FFFE1FC948E92FC49C9E2" authority="Babenko &amp; Kuznetsova &amp; Nakamori &amp; Shveenkova, 2021" authorityName="Babenko &amp; Kuznetsova &amp; Nakamori &amp; Shveenkova" authorityYear="2021" box="[842,980,923,949]" class="Entognatha" family="Neanuridae" genus="Pseudachorutes" kingdom="Animalia" order="Collembola" pageId="12" pageNumber="363" phylum="Arthropoda" rank="species" species="variabilis" status="sp. nov.">
<emphasis id="05D8EA89232FFFE1FC948E92FCC6C9E3" box="[842,859,923,948]" italics="true" pageId="12" pageNumber="363">P</emphasis>
.
<emphasis id="05D8EA89232FFFE1FCB78E92FC49C9E2" box="[873,980,923,949]" italics="true" pageId="12" pageNumber="363">variabilis</emphasis>
</taxonomicName>
<emphasis id="05D8EA89232FFFE1FC028E92FBA8C9E2" bold="true" box="[988,1077,923,949]" pageId="12" pageNumber="363">
<taxonomicNameLabel id="1EEB57F2232FFFE1FC028E92FBA8C9E2" box="[988,1077,923,949]" pageId="12" pageNumber="363" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="AF972A1E232FFFE1FB9B8E92FB0BC9E2" box="[1093,1174,923,949]" captionStart="FIGURES 31–39" captionStartId="11.[151,264,1906,1930]" captionTargetBox="[241,1346,192,1874]" captionTargetId="figure-20@11.[232,1356,181,1882]" captionTargetPageId="11" captionText="FIGURES 31–39. Pseudachorutes variabilis sp. nov. (31–35, 37–39) and P. subcrassus (36): 31, 33, head chaetotaxy (two different specimen); 32, 34, chaetotaxy of Abd. V–VI (two different specimen); 34a, shape of setae (Japanese specimen); 35–36, labium; 37, tip of tibiotarsus with unguis; 38, right and left mandibles; 39, Ant. III–IV (ventral side). Scales: Figs 31–34 = 0.1 mm, 35–39 = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.4574825" httpUri="https://zenodo.org/record/4574825/files/figure.png" pageId="12" pageNumber="363">Fig. 31</figureCitation>
) is more similar to that of
<taxonomicName id="F0AC4D18232FFFE1FF688EB7FEAEC980" authorityName="Borner" authorityYear="1901" box="[182,307,957,983]" class="Entognatha" family="Neanuridae" genus="Pseudachorutes" kingdom="Animalia" order="Collembola" pageId="12" pageNumber="363" phylum="Arthropoda" rank="species" species="parvulus">
<emphasis id="05D8EA89232FFFE1FF688EB7FEAEC980" box="[182,307,957,983]" italics="true" pageId="12" pageNumber="363">P. parvulus</emphasis>
</taxonomicName>
, than
<emphasis id="05D8EA89232FFFE1FEA68EB7FD8BC980" box="[376,534,957,983]" italics="true" pageId="12" pageNumber="363">
<taxonomicName id="F0AC4D18232FFFE1FEA68EB7FD8FC980" authorityName="Tullberg" authorityYear="1871" box="[376,530,957,983]" class="Entognatha" family="Neanuridae" genus="Pseudachorutes" kingdom="Animalia" order="Collembola" pageId="12" pageNumber="363" phylum="Arthropoda" rank="species" species="subcrassus">P. subcrassus</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="7FB66510232FFFE1FF198EE9FEB6CE16" pageId="12" pageNumber="363" type="distribution">
<paragraph id="3713369B232FFFE1FF198EE9FEB6CE16" blockId="12.[151,1437,151,1089]" pageId="12" pageNumber="363">
<emphasis id="05D8EA89232FFFE1FF198EE9FEFCC9AD" bold="true" box="[199,353,992,1018]" pageId="12" pageNumber="363">Distribution.</emphasis>
The species appears to be widespread in the studied region; it was found in three State Nature Reserves (Sikhote-Alin, Ussuri and Kedrovaya Pad), in the northern part of the Khabarovsk Territory, Kamchatka and in
<collectingCountry id="4FBB760B232FFFE1FF3D892EFEBBCE16" box="[227,294,1063,1089]" name="Japan" pageId="12" pageNumber="363">Japan</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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