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<document id="DF72C2B3B2BB42DBAB1D44FD5E322304" ID-DOI="10.1080/00222933.2020.1781949" ID-ISSN="1464-5262" ID-Zenodo-Dep="5753671" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1621624434080" checkinUser="carolina" docAuthor="Kazantsev, Sergey" docDate="2020" docId="E259B81CA66CFFDE6AA9B2E0A57D4D1A" docLanguage="en" docName="JNATHIST.54.17-18.1073-1080.pdf" docOrigin="Journal of Natural History 54 (17 - 18)" docSource="http://dx.doi.org/10.1080/00222933.2020.1781949" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Retromalisus Kazantsev 2020, gen. nov." docType="treatment" docVersion="4" lastPageNumber="1075" masterDocId="1E60C064A66EFFDA6B6FB047A1364969" masterDocTitle="Retromalisus damzeni, gen. et sp. nov., a second Baltic amber taxon of the extinct family Berendtimiridae (Insecta: Coleoptera)" masterLastPageNumber="1080" masterPageNumber="1073" pageNumber="1074" updateTime="1698940605863" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="A023925BC4D81BDD7C965B3FBD4BADB4">Retromalisus damzeni, gen. et sp. nov., a second Baltic amber taxon of the extinct family Berendtimiridae (Insecta: Coleoptera)</mods:title>
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<mods:namePart id="F7BE295D16B68E5E90A1936EA8FBAC33">Kazantsev, Sergey</mods:namePart>
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<mods:title id="A462D1B0C2B58C6F84951048D708A8E7">Journal of Natural History</mods:title>
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<mods:date id="D701C51531A21E366DA18BF6CF9B989B">2020</mods:date>
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<mods:number id="F2494B680FC109D8079221665A4BCBFA">17 - 18</mods:number>
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<treatment id="E259B81CA66CFFDE6AA9B2E0A57D4D1A" ID-DOI="http://doi.org/10.5281/zenodo.5753749" ID-Zenodo-Dep="5753749" LSID="urn:lsid:plazi:treatment:E259B81CA66CFFDE6AA9B2E0A57D4D1A" httpUri="http://treatment.plazi.org/id/E259B81CA66CFFDE6AA9B2E0A57D4D1A" lastPageId="4" lastPageNumber="1075" pageId="2" pageNumber="1074">
<subSubSection id="22EA5A81A66CFFD86AA9B2E0A2684BA8" box="[454,862,679,705]" pageId="2" pageNumber="1074" type="nomenclature">
<paragraph id="6A4F090AA66CFFD86AA9B2E0A2684BA8" blockId="2.[454,862,679,705]" box="[454,862,679,705]" pageId="2" pageNumber="1074">
<heading id="3107BE66A66CFFD86AA9B2E0A2684BA8" box="[454,862,679,705]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="1074" reason="2">
<taxonomicName id="ADF07289A66CFFD86AA9B2E0A3D04BA8" authority="Kazantsev" authorityName="Kazantsev" authorityYear="2020" box="[454,742,679,705]" class="Insecta" family="Berendtimiridae" genus="Retromalisus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1074" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5884D518A66CFFD86AA9B2E0A3534BA8" bold="true" box="[454,613,679,705]" italics="true" pageId="2" pageNumber="1074">Retromalisus</emphasis>
Kazantsev
</taxonomicName>
,
<taxonomicNameLabel id="43B76863A66CFFD8699FB2E0A2684BA8" box="[752,862,679,705]" pageId="2" pageNumber="1074" rank="genus">gen. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="22EA5A81A66CFFD86BCFB2B9A2FF4A71" box="[160,969,766,792]" pageId="2" pageNumber="1074" type="type_taxon">
<paragraph id="6A4F090AA66CFFD86BCFB2B9A2FF4A71" blockId="2.[160,969,766,792]" box="[160,969,766,792]" pageId="2" pageNumber="1074">
<emphasis id="5884D518A66CFFD86BCFB2B9A00F4A71" bold="true" box="[160,313,766,792]" italics="true" pageId="2" pageNumber="1074">
<typeStatus id="B54BB7A8A66CFFD86BCFB2B9A1EE4A71" box="[160,216,766,792]" pageId="2" pageNumber="1074">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="ADF07289A66CFFD86A25B2B8A3654A71" authority="Kazantsev, 2020" authorityName="Kazantsev" authorityYear="2020" box="[330,595,767,792]" class="Insecta" family="Berendtimiridae" genus="Retromalisus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1074" phylum="Arthropoda" rank="species" species="damzeni" status="sp. nov.">
<emphasis id="5884D518A66CFFD86A25B2B8A3654A71" bold="true" box="[330,595,767,792]" pageId="2" pageNumber="1074">Retromalisus damzeni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="43B76863A66CFFD86934B2B8A3824A71" box="[603,692,767,792]" pageId="2" pageNumber="1074" rank="species">sp. nov.</taxonomicNameLabel>
, by present designation.
</paragraph>
</subSubSection>
<subSubSection id="22EA5A81A66CFFD96BCFB303A0C74DD1" lastPageId="3" lastPageNumber="1075" pageId="2" pageNumber="1074" type="description">
<paragraph id="6A4F090AA66CFFD86BCFB303A27F4D47" blockId="2.[160,1156,836,1728]" pageId="2" pageNumber="1074">
<emphasis id="5884D518A66CFFD86BCFB303A01B4A37" bold="true" box="[160,301,836,862]" italics="true" pageId="2" pageNumber="1074">Description.</emphasis>
Adult male (?). Alate, flattened, elongate. Head moderately large, transverse, exposed, smoothly narrowed posteriorly, ventrally noticeably semi-circularly excavated medially. Eyes relatively small, spherical. Palps small, slender; ultimate palpomeres elongate, narrowed and pointed apically. Bases of antennae widely separated; antenna 11- segmented, moderately long, filiform; pedicel (antennomere 2) elongate, about as long as antennomere 3, antennomere 3 slightly shorter than antennomere 4; pubescence on antennomeres 315 short, dense and suberect (
<figureCitation id="F2CB158FA66CFFD869D4B452A20A4D47" box="[699,828,1045,1070]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStart-3="Figure 4" captionStartId-0="3.[160,225,1052,1075]" captionStartId-1="4.[160,225,849,872]" captionStartId-2="5.[160,225,907,930]" captionStartId-3="6.[160,225,1009,1032]" captionTargetBox-0="[163,1153,142,1020]" captionTargetBox-1="[163,1153,142,817]" captionTargetBox-2="[163,1153,142,875]" captionTargetBox-3="[166,1150,145,974]" captionTargetId-0="figure-176@3.[163,1153,142,1020]" captionTargetId-1="figure-225@4.[163,1153,142,817]" captionTargetId-2="figure-207@5.[163,1153,142,875]" captionTargetId-3="figure-31@6.[163,1153,142,977]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="Figure 1. General view of Retromalisus damzeni gen. et sp. nov., holotype male, dorsal view." captionText-1="Figure 2. General view of Retromalisus damzeni gen. et sp. nov., holotype male, ventral view." captionText-2="Figure 3. Anterior part of body of Retromalisus damzeni gen. et sp. nov., holotype male, ventral view." captionText-3="Figure 4. Head of Retromalisus damzeni gen. et sp. nov., holotype male, ventral view, contour." figureDoi-0="http://doi.org/10.5281/zenodo.5753673" figureDoi-1="http://doi.org/10.5281/zenodo.5753675" figureDoi-2="http://doi.org/10.5281/zenodo.5753677" figureDoi-3="http://doi.org/10.5281/zenodo.5753679" httpUri-0="https://zenodo.org/record/5753673/files/figure.png" httpUri-1="https://zenodo.org/record/5753675/files/figure.png" httpUri-2="https://zenodo.org/record/5753677/files/figure.png" httpUri-3="https://zenodo.org/record/5753679/files/figure.png" pageId="2" pageNumber="1074">Figures 14</figureCitation>
).
</paragraph>
<paragraph id="6A4F090AA66CFFD86BAFB470A2674CC2" blockId="2.[160,1156,836,1728]" pageId="2" pageNumber="1074">
Pronotum transverse, but distinctly narrower than elytra, trapezoidal, with slightly convex anterior and posterior margins and small acute posterior angles with noticeable incision in front of them; disk medially with obscure longitudinal rib, laterally with slightly more pronounced transverse carinae reaching lateral margins; margins narrowly bordered (
<figureCitation id="F2CB158FA66CFFD86BC7B485A01F4DB2" box="[168,297,1218,1243]" captionStart-0="Figure 3" captionStart-1="Figure 4" captionStart-2="Figures 56" captionStartId-0="5.[160,225,907,930]" captionStartId-1="6.[160,225,1009,1032]" captionStartId-2="6.[160,235,1500,1523]" captionTargetBox-0="[163,1153,142,875]" captionTargetBox-1="[166,1150,145,974]" captionTargetBox-2="[165,1150,1092,1466]" captionTargetId-0="figure-207@5.[163,1153,142,875]" captionTargetId-1="figure-31@6.[163,1153,142,977]" captionTargetId-2="figure-50@6.[163,1153,1090,1468]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="6" captionText-0="Figure 3. Anterior part of body of Retromalisus damzeni gen. et sp. nov., holotype male, ventral view." captionText-1="Figure 4. Head of Retromalisus damzeni gen. et sp. nov., holotype male, ventral view, contour." captionText-2="Figures 56. Details of Retromalisus damzeni gen. et sp. nov., holotype male: 5 pronotum, dorsal view, contour; 6 abdomen, ventral view, contour." figureDoi-0="http://doi.org/10.5281/zenodo.5753677" figureDoi-1="http://doi.org/10.5281/zenodo.5753679" figureDoi-2="http://doi.org/10.5281/zenodo.5753681" httpUri-0="https://zenodo.org/record/5753677/files/figure.png" httpUri-1="https://zenodo.org/record/5753679/files/figure.png" httpUri-2="https://zenodo.org/record/5753681/files/figure.png" pageId="2" pageNumber="1074">Figures 35</figureCitation>
). Prosternum short, T-shaped, with slightly convex medially anterior margin and short bifurcate intercoxal process (
<figureCitation id="F2CB158FA66CFFD8691BB4A2A3CD4D97" box="[628,763,1253,1278]" captionStart-0="Figure 3" captionStart-1="Figure 4" captionStartId-0="5.[160,225,907,930]" captionStartId-1="6.[160,225,1009,1032]" captionTargetBox-0="[163,1153,142,875]" captionTargetBox-1="[166,1150,145,974]" captionTargetId-0="figure-207@5.[163,1153,142,875]" captionTargetId-1="figure-31@6.[163,1153,142,977]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="Figure 3. Anterior part of body of Retromalisus damzeni gen. et sp. nov., holotype male, ventral view." captionText-1="Figure 4. Head of Retromalisus damzeni gen. et sp. nov., holotype male, ventral view, contour." figureDoi-0="http://doi.org/10.5281/zenodo.5753677" figureDoi-1="http://doi.org/10.5281/zenodo.5753679" httpUri-0="https://zenodo.org/record/5753677/files/figure.png" httpUri-1="https://zenodo.org/record/5753679/files/figure.png" pageId="2" pageNumber="1074">Figures 34</figureCitation>
). Scutellum triangular (
<figureCitation id="F2CB158FA66CFFD86F7BB4A2A5404D97" box="[1044,1142,1253,1278]" captionStart="Figure 1" captionStartId="3.[160,225,1052,1075]" captionTargetBox="[163,1153,142,1020]" captionTargetId="figure-176@3.[163,1153,142,1020]" captionTargetPageId="3" captionText="Figure 1. General view of Retromalisus damzeni gen. et sp. nov., holotype male, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5753673" httpUri="https://zenodo.org/record/5753673/files/figure.png" pageId="2" pageNumber="1074">Figure 1</figureCitation>
). Metaventrite transverse, conspicuously widening posteriorly (
<figureCitation id="F2CB158FA66CFFD86831B540A28A4C49" box="[862,956,1287,1312]" captionStart="Figure 2" captionStartId="4.[160,225,849,872]" captionTargetBox="[163,1153,142,817]" captionTargetId="figure-225@4.[163,1153,142,817]" captionTargetPageId="4" captionText="Figure 2. General view of Retromalisus damzeni gen. et sp. nov., holotype male, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5753675" httpUri="https://zenodo.org/record/5753675/files/figure.png" pageId="2" pageNumber="1074">Figure 2</figureCitation>
). Elytra elongate, almost parallel-sided, only slightly narrowed in the middle, shining, with 10 regular longitudinal rows of round punctures (eight in distal fifth) and fully sclerotised narrow epipleures attaining to elytral apices. Metathoracic wings fully developed; both anterior and posterior edges with a fringe of short hairs (
<figureCitation id="F2CB158FA66CFFD869ACB5D5A2734CC2" box="[707,837,1426,1451]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStartId-0="3.[160,225,1052,1075]" captionStartId-1="4.[160,225,849,872]" captionTargetBox-0="[163,1153,142,1020]" captionTargetBox-1="[163,1153,142,817]" captionTargetId-0="figure-176@3.[163,1153,142,1020]" captionTargetId-1="figure-225@4.[163,1153,142,817]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="Figure 1. General view of Retromalisus damzeni gen. et sp. nov., holotype male, dorsal view." captionText-1="Figure 2. General view of Retromalisus damzeni gen. et sp. nov., holotype male, ventral view." figureDoi-0="http://doi.org/10.5281/zenodo.5753673" figureDoi-1="http://doi.org/10.5281/zenodo.5753675" httpUri-0="https://zenodo.org/record/5753673/files/figure.png" httpUri-1="https://zenodo.org/record/5753675/files/figure.png" pageId="2" pageNumber="1074">Figures 12</figureCitation>
).
</paragraph>
<paragraph id="6A4F090AA66CFFD86BAFB5F2A2784F31" blockId="2.[160,1156,836,1728]" pageId="2" pageNumber="1074">
Legs relatively long and slender; front and middle coxae elongate, apices of hind coxae elongate and separated; trochanters elongate, apically overlaying bases of femurs; femurs and tibiae straight, narrow, subequal in length; tibial spurs small but well visible; tarsi 5-segmented, tarsomeres 13 narrow, without plantar pad, tarsomere 4 incised to the base and conspicuously widened; claws simple (
<figureCitation id="F2CB158FA66CFFD869AFB678A2774F31" box="[704,833,1599,1624]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStartId-0="3.[160,225,1052,1075]" captionStartId-1="4.[160,225,849,872]" captionTargetBox-0="[163,1153,142,1020]" captionTargetBox-1="[163,1153,142,817]" captionTargetId-0="figure-176@3.[163,1153,142,1020]" captionTargetId-1="figure-225@4.[163,1153,142,817]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="Figure 1. General view of Retromalisus damzeni gen. et sp. nov., holotype male, dorsal view." captionText-1="Figure 2. General view of Retromalisus damzeni gen. et sp. nov., holotype male, ventral view." figureDoi-0="http://doi.org/10.5281/zenodo.5753673" figureDoi-1="http://doi.org/10.5281/zenodo.5753675" httpUri-0="https://zenodo.org/record/5753673/files/figure.png" httpUri-1="https://zenodo.org/record/5753675/files/figure.png" pageId="2" pageNumber="1074">Figures 12</figureCitation>
).
</paragraph>
<paragraph id="6A4F090AA66CFFD86BAFB625A51D4FA9" blockId="2.[160,1156,836,1728]" pageId="2" pageNumber="1074">
Abdomen with eight ventrites first medially completely interrupted by metacoxae; penultimate ventrite broadly semi-circularly incised; ultimate ventrite largely invaginated, its exposed portion transverse, rather deeply semicircularly incised (
<figureCitation id="F2CB158FA66CFFD868F3B6E0A5354FA9" box="[924,1027,1703,1728]" captionStart="Figure 2" captionStartId="4.[160,225,849,872]" captionTargetBox="[163,1153,142,817]" captionTargetId="figure-225@4.[163,1153,142,817]" captionTargetPageId="4" captionText="Figure 2. General view of Retromalisus damzeni gen. et sp. nov., holotype male, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5753675" httpUri="https://zenodo.org/record/5753675/files/figure.png" pageId="2" pageNumber="1074">Figures 2</figureCitation>
,
<figureCitation id="F2CB158FA66CFFD86F7FB6E0A5284FA9" box="[1040,1054,1703,1728]" captionStart="Figures 56" captionStartId="6.[160,235,1500,1523]" captionTargetBox="[165,1150,1092,1466]" captionTargetId="figure-50@6.[163,1153,1090,1468]" captionTargetPageId="6" captionText="Figures 56. Details of Retromalisus damzeni gen. et sp. nov., holotype male: 5 pronotum, dorsal view, contour; 6 abdomen, ventral view, contour." figureDoi="http://doi.org/10.5281/zenodo.5753681" httpUri="https://zenodo.org/record/5753681/files/figure.png" pageId="2" pageNumber="1074">6</figureCitation>
).
</paragraph>
<caption id="3E8F5982A66DFFD96BCFB45BA5024D5A" ID-DOI="http://doi.org/10.5281/zenodo.5753673" ID-Zenodo-Dep="5753673" box="[160,1076,1052,1075]" httpUri="https://zenodo.org/record/5753673/files/figure.png" pageId="3" pageNumber="1075" startId="3.[160,225,1052,1075]" targetBox="[163,1153,142,1020]" targetPageId="3">
<paragraph id="6A4F090AA66DFFD96BCFB45BA5024D5A" blockId="3.[160,1076,1052,1075]" box="[160,1076,1052,1075]" pageId="3" pageNumber="1075">
<emphasis id="5884D518A66DFFD96BCFB45BA1CD4D5A" bold="true" box="[160,251,1052,1075]" pageId="3" pageNumber="1075">Figure 1.</emphasis>
General view of
<taxonomicName id="ADF07289A66DFFD96ACBB45BA3404D5A" authority="Kazantsev, 2020" authorityName="Kazantsev" authorityYear="2020" box="[420,630,1052,1075]" class="Insecta" family="Berendtimiridae" genus="Retromalisus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1075" phylum="Arthropoda" rank="species" species="damzeni" status="gen. et sp. nov.">
<emphasis id="5884D518A66DFFD96ACBB45BA3404D5A" box="[420,630,1052,1075]" italics="true" pageId="3" pageNumber="1075">Retromalisus damzeni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="43B76863A66DFFD96912B45BA22F4D5A" box="[637,793,1052,1075]" pageId="3" pageNumber="1075" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, holotype male, dorsal view.
</paragraph>
</caption>
<paragraph id="6A4F090AA66DFFD96BCFB43BA0C74DD1" blockId="3.[160,1155,1148,1208]" pageId="3" pageNumber="1075">
<emphasis id="5884D518A66DFFD96BCFB43BA1CB4DFF" bold="true" box="[160,253,1148,1174]" italics="true" pageId="3" pageNumber="1075">Female.</emphasis>
Unknown (the shape of the last abdominal segments suggests that the studied specimen is probably a male).
</paragraph>
</subSubSection>
<subSubSection id="22EA5A81A66DFFD96BCFB4A3A0444C2A" pageId="3" pageNumber="1075" type="etymology">
<paragraph id="6A4F090AA66DFFD96BCFB4A3A0444C2A" blockId="3.[160,1156,1252,1347]" pageId="3" pageNumber="1075">
<emphasis id="5884D518A66DFFD96BCFB4A3A0154D97" bold="true" box="[160,291,1252,1278]" italics="true" pageId="3" pageNumber="1075">Etymology.</emphasis>
The name of the new genus is a combination of retro, the Latin for backward, and the genus name
<taxonomicName id="ADF07289A66DFFD96AB1B540A3774C49" box="[478,577,1287,1312]" class="Insecta" family="Omalisidae" genus="Omalisus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1075" phylum="Arthropoda" rank="genus">
<emphasis id="5884D518A66DFFD96AB1B540A3774C49" box="[478,577,1287,1312]" italics="true" pageId="3" pageNumber="1075">Omalisus</emphasis>
</taxonomicName>
, alluding to its certain similarity to the latter taxon. Gender masculine.
</paragraph>
</subSubSection>
<subSubSection id="22EA5A81A66DFFD96BCFB529A3CA4F5F" pageId="3" pageNumber="1075" type="diagnosis">
<paragraph id="6A4F090AA66DFFD96BCFB529A3CA4F5F" blockId="3.[160,1155,1390,1590]" pageId="3" pageNumber="1075">
<emphasis id="5884D518A66DFFD96BCFB529A0804CE1" bold="true" box="[160,438,1390,1416]" italics="true" pageId="3" pageNumber="1075">Differential diagnosis.</emphasis>
<taxonomicName id="ADF07289A66DFFD96AA9B528A3664CE1" authority="Kazantsev, 2020" authorityName="Kazantsev" authorityYear="2020" box="[454,592,1391,1416]" class="Insecta" family="Berendtimiridae" genus="Retromalisus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1075" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5884D518A66DFFD96AA9B528A3664CE1" box="[454,592,1391,1416]" italics="true" pageId="3" pageNumber="1075">Retromalisus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="43B76863A66DFFD96904B528A3DF4CE1" box="[619,745,1391,1416]" pageId="3" pageNumber="1075" rank="genus">gen. nov.</taxonomicNameLabel>
, is easily distinguishable from
<taxonomicName id="ADF07289A66DFFD96BCFB5D5A0034CC2" baseAuthorityName="Winkler" baseAuthorityYear="1987" box="[160,309,1426,1451]" class="Insecta" family="Berendtimiridae" genus="Berendtimirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1075" phylum="Arthropoda" rank="genus">
<emphasis id="5884D518A66DFFD96BCFB5D5A0034CC2" box="[160,309,1426,1451]" italics="true" pageId="3" pageNumber="1075">Berendtimirus</emphasis>
</taxonomicName>
, the only other member of the family, by the smaller eyes, shorter scapus (antennomere 1), narrower mandibles, shorter antennae and 10 longitudinal rows of elytral punctures (vs. nine in
<taxonomicName id="ADF07289A66DFFD96A96B590A3B84C99" baseAuthorityName="Winkler" baseAuthorityYear="1987" box="[505,654,1495,1520]" class="Insecta" family="Berendtimiridae" genus="Berendtimirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1075" phylum="Arthropoda" rank="genus">
<emphasis id="5884D518A66DFFD96A96B590A3B84C99" box="[505,654,1495,1520]" italics="true" pageId="3" pageNumber="1075">Berendtimirus</emphasis>
</taxonomicName>
) (
<figureCitation id="F2CB158FA66DFFD969C6B590A2184C99" box="[681,814,1495,1520]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStart-3="Figure 4" captionStartId-0="3.[160,225,1052,1075]" captionStartId-1="4.[160,225,849,872]" captionStartId-2="5.[160,225,907,930]" captionStartId-3="6.[160,225,1009,1032]" captionTargetBox-0="[163,1153,142,1020]" captionTargetBox-1="[163,1153,142,817]" captionTargetBox-2="[163,1153,142,875]" captionTargetBox-3="[166,1150,145,974]" captionTargetId-0="figure-176@3.[163,1153,142,1020]" captionTargetId-1="figure-225@4.[163,1153,142,817]" captionTargetId-2="figure-207@5.[163,1153,142,875]" captionTargetId-3="figure-31@6.[163,1153,142,977]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="Figure 1. General view of Retromalisus damzeni gen. et sp. nov., holotype male, dorsal view." captionText-1="Figure 2. General view of Retromalisus damzeni gen. et sp. nov., holotype male, ventral view." captionText-2="Figure 3. Anterior part of body of Retromalisus damzeni gen. et sp. nov., holotype male, ventral view." captionText-3="Figure 4. Head of Retromalisus damzeni gen. et sp. nov., holotype male, ventral view, contour." figureDoi-0="http://doi.org/10.5281/zenodo.5753673" figureDoi-1="http://doi.org/10.5281/zenodo.5753675" figureDoi-2="http://doi.org/10.5281/zenodo.5753677" figureDoi-3="http://doi.org/10.5281/zenodo.5753679" httpUri-0="https://zenodo.org/record/5753673/files/figure.png" httpUri-1="https://zenodo.org/record/5753675/files/figure.png" httpUri-2="https://zenodo.org/record/5753677/files/figure.png" httpUri-3="https://zenodo.org/record/5753679/files/figure.png" pageId="3" pageNumber="1075">Figures 14</figureCitation>
). The number of ventrites in
<taxonomicName id="ADF07289A66DFFD96BCFB5BDA01C4F7A" authority="Kazantsev, 2020" authorityName="Kazantsev" authorityYear="2020" box="[160,298,1530,1555]" class="Insecta" family="Berendtimiridae" genus="Retromalisus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1075" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5884D518A66DFFD96BCFB5BDA01C4F7A" box="[160,298,1530,1555]" italics="true" pageId="3" pageNumber="1075">Retromalisus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="43B76863A66DFFD96A5FB5BDA0A14F7A" box="[304,407,1530,1555]" pageId="3" pageNumber="1075" rank="genus">gen. nov.</taxonomicNameLabel>
(eight) is also noticeably greater than that in
<taxonomicName id="ADF07289A66DFFD968E6B5BDA5284F7A" baseAuthorityName="Winkler" baseAuthorityYear="1987" box="[905,1054,1530,1555]" class="Insecta" family="Berendtimiridae" genus="Berendtimirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1075" phylum="Arthropoda" rank="genus">
<emphasis id="5884D518A66DFFD968E6B5BDA5284F7A" box="[905,1054,1530,1555]" italics="true" pageId="3" pageNumber="1075">Berendtimirus</emphasis>
</taxonomicName>
(six), this difference addressed in the Discussion section below.
</paragraph>
</subSubSection>
<subSubSection id="22EA5A81A66DFFDE6BCFB626A57D4D1A" lastPageId="4" lastPageNumber="1076" pageId="3" pageNumber="1075" type="discussion">
<paragraph id="6A4F090AA66DFFDE6BCFB626A57D4D1A" blockId="3.[160,1155,1633,1728]" lastBlockId="4.[160,1155,941,1139]" lastPageId="4" lastPageNumber="1076" pageId="3" pageNumber="1075">
<emphasis id="5884D518A66DFFD96BCFB626A0344F12" bold="true" box="[160,258,1633,1659]" italics="true" pageId="3" pageNumber="1075">Remark.</emphasis>
<taxonomicName id="ADF07289A66DFFD96A7EB625A0AE4F12" authority="Kazantsev, 2020" authorityName="Kazantsev" authorityYear="2020" box="[273,408,1634,1659]" class="Insecta" family="Berendtimiridae" genus="Retromalisus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1075" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5884D518A66DFFD96A7EB625A0AE4F12" box="[273,408,1634,1659]" italics="true" pageId="3" pageNumber="1075">Retromalisus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="43B76863A66DFFD96ACDB625A33A4F12" box="[418,524,1634,1659]" pageId="3" pageNumber="1075" rank="genus">gen. nov.</taxonomicNameLabel>
cannot be referred to
<taxonomicName id="ADF07289A66DFFD9687BB625A2AF4F12" authorityName="Imhoff" authorityYear="1856" box="[788,921,1634,1659]" class="Insecta" family="Cantharidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1075" phylum="Arthropoda" rank="family">Cantharidae</taxonomicName>
due to its complete sclerotised elytral epipleures reaching elytral apices, while in all studied cantharids the epipleures are short, developed mostly at the humeral area (e.g.
<bibRefCitation id="0E6174FBA66DFFD9681DB6E0A52B4FA9" author="Kazantsev SV" box="[882,1053,1703,1728]" pageId="3" pageNumber="1075" pagination="73 - 248" refId="ref2528" refString="Kazantsev SV. 2005. Morphology of Lycidae with some considerations on evolution of the Coleoptera. Elytron. 17 - 18 (2004) et Coleopterological Monographs, 3: 73 - 248." type="journal article" year="2005">Kazantsev 2005</bibRefCitation>
). Neither can it be placed in
<taxonomicName id="ADF07289A66AFFDE6A13B3EAA34F4AAF" authorityName="Lacordaire" authorityYear="1857" box="[380,633,941,966]" class="Insecta" family="Elateridae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="1076" phylum="Arthropoda" rank="subFamily" subFamily="Omalisinae">Omalisinae (Elateridae)</taxonomicName>
due to the short prosternum with short intercoxal process and wide and deeply incised penultimate tarsomeres, as omalisines are characterised by the long prosterna with long intercoxal processes and narrow tarsi (e.g.
<bibRefCitation id="0E6174FBA66AFFDE6BCFB452A0BF4D47" author="Bocak L &amp; Brlik M" box="[160,393,1045,1070]" pageId="4" pageNumber="1076" pagination="189 - 212" refId="ref2390" refString="Bocak L, Brlik M. 2008. Revision of the family Omalisidae (Coleoptera, Elateroidea). Insect Syst Evol. 39 (2): 189 - 212. doi: 10.1163 / 187631208788784101" type="journal article" year="2008">Bocák and Brlik 2008</bibRefCitation>
). The shape of the ultimate ventrite in
<taxonomicName id="ADF07289A66AFFDE682FB452A2F14D47" authority="Kazantsev, 2020" authorityName="Kazantsev" authorityYear="2020" box="[832,967,1045,1070]" class="Insecta" family="Berendtimiridae" genus="Retromalisus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="1076" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5884D518A66AFFDE682FB452A2F14D47" box="[832,967,1045,1070]" italics="true" pageId="4" pageNumber="1076">Retromalisus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="43B76863A66AFFDE68BFB452A50E4D47" box="[976,1080,1045,1070]" pageId="4" pageNumber="1076" rank="genus">gen. nov.</taxonomicNameLabel>
is also quite unusual for males of the
<taxonomicName id="ADF07289A66AFFDE696FB470A34F4D39" authorityName="Lacordaire" authorityYear="1857" box="[512,633,1079,1104]" class="Insecta" family="Elateridae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="1076" phylum="Arthropoda" rank="subFamily" subFamily="Omalisinae">Omalisinae</taxonomicName>
lineage, where the ultimate ventrite is always elongated and much narrower than the penultimate one (e.g.
<bibRefCitation id="0E6174FBA66AFFDE6838B41DA5094D1A" author="Bocak L &amp; Brlik M" box="[855,1087,1114,1139]" pageId="4" pageNumber="1076" pagination="189 - 212" refId="ref2390" refString="Bocak L, Brlik M. 2008. Revision of the family Omalisidae (Coleoptera, Elateroidea). Insect Syst Evol. 39 (2): 189 - 212. doi: 10.1163 / 187631208788784101" type="journal article" year="2008">Bocák and Brlik 2008</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>