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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1137.97068" ID-Pensoft-Pub="1313-2970-1137-133" ID-Pensoft-UUID="818ADEB99A7A5D0C851421411FA5B421" ID-ZooBank="DFA4D0BDCBA649D487ACC741E6C216E9" ModsDocID="1313-2970-1137-133" checkinTime="1671892165935" checkinUser="pensoft" docAuthor="Caterino, Michael S." docDate="2022" docId="67BE608F976A50CEB559ADBB12F2355D" docLanguage="en" docName="ZooKeys 1137: 133-175" docOrigin="ZooKeys 1137" docPubDate="2022-12-22" docSource="http://dx.doi.org/10.3897/zookeys.1137.97068" docTitle="Euconnus attritus Caterino 2022, sp. nov." docType="treatment" docUuid="37927727-6073-4A5B-8B1C-9281D47B3699" docUuidSource="ZooBank" docVersion="1" id="818ADEB99A7A5D0C851421411FA5B421" lastPageNumber="133" masterDocId="818ADEB99A7A5D0C851421411FA5B421" masterDocTitle="First report of the Euconnus Thomson subgenus Cladoconnus Reitter in the New World, represented by thirteen new Appalachian species (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="175" masterPageNumber="133" pageNumber="133" updateTime="1671892165935" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>First report of the Euconnus Thomson subgenus Cladoconnus Reitter in the New World, represented by thirteen new Appalachian species (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Caterino, Michael S.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-2597-5707</mods:nameIdentifier>
<mods:affiliation>Dept of Plant &amp; Environmental Sciences, Clemson University, Clemson, SC 29634, USA</mods:affiliation>
<mods:nameIdentifier type="email">mcateri@clemson.edu</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-12-22</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1137</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>133</mods:start>
<mods:end>175</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1137.97068</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1137.97068</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1137-133</mods:identifier>
<mods:identifier type="ZooBank">DFA4D0BDCBA649D487ACC741E6C216E9</mods:identifier>
<mods:identifier type="Pensoft-UUID">818ADEB99A7A5D0C851421411FA5B421</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:zoobank.org:act:37927727-6073-4A5B-8B1C-9281D47B3699" httpUri="http://treatment.plazi.org/id/67BE608F976A50CEB559ADBB12F2355D" lastPageNumber="133" pageId="0" pageNumber="133">
<subSubSection pageId="0" pageNumber="133" type="nomenclature">
<paragraph pageId="0" pageNumber="133">
<taxonomicName LSID="https://zoobank.org/37927727-6073-4A5B-8B1C-9281D47B3699" authority="Caterino, 2022" authorityName="Caterino" authorityYear="2022" class="Insecta" family="Staphylinidae" genus="Euconnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Euconnus attritus" order="Coleoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="attritus" status="sp. nov.">Euconnus attritus</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="133">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="133" type="description">
<paragraph pageId="0" pageNumber="133">
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Map of collecting records for Euconnus (Cladoconnus) falcatus complex. E. (C.) falcatus (circles), E. (C.) cataloochee (stars), E. (C.) tusquitee (triangules), E. (C.) draco (hexagons), E. (C.) kilmeri (plus sign), and E. (C.) attritus (x)." figureDoi="10.3897/zookeys.1137.97068.figure11" httpUri="https://binary.pensoft.net/fig/786422" pageId="0" pageNumber="133">Figs 11</figureCitation>
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Aedeagus. A Euconnus draco B Euconnus tusquitee C Euconnus attritus." figureDoi="10.3897/zookeys.1137.97068.figure12" httpUri="https://binary.pensoft.net/fig/786423" pageId="0" pageNumber="133">, 12C</figureCitation>
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Habitus and character photos of Euconnus attritus A male, lateral view B male, dorsal view C female, lateral view D female, dorsal view E SEM of male antenna F SEM of venter of head." figureDoi="10.3897/zookeys.1137.97068.figure13" httpUri="https://binary.pensoft.net/fig/786424" pageId="0" pageNumber="133">, 13</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="133" type="type material">
<paragraph pageId="0" pageNumber="133">Type material.</paragraph>
<paragraph pageId="0" pageNumber="133">
<emphasis bold="true" italics="true" pageId="0" pageNumber="133">Holotype</emphasis>
♂, deposited in FMNH: &quot;USA:NC: Mitchell Co.,
<geoCoordinate degrees="36.0931" direction="north" orientation="latitude" precision="5" value="36.0931">36.0931°N</geoCoordinate>
,
<geoCoordinate degrees="82.1453" direction="west" orientation="longitude" precision="5" value="-82.1453">82.1453°W</geoCoordinate>
, Roan High Bluff, 6225' viii.15.2018, M.Caterino, sifted
<taxonomicName class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" lsidName="Abies" order="Pinales" pageId="0" pageNumber="133" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="133">Abies</emphasis>
</taxonomicName>
litter&quot; / &quot;[QR code] CLEMSON-ENT CUAC000003146&quot; / &quot;Caterino DNA Voucher Extraction MSC3360 Morphosp.RHB.A.012&quot;;
<emphasis bold="true" pageId="0" pageNumber="133">
<emphasis italics="true" pageId="0" pageNumber="133">Paratypes</emphasis>
(62
</emphasis>
, CUAC, FMNH, CNCI, UNHC) - 3 ♀, same data as type; 18 ♀, 16 ♂: &quot;USA:NC: Mitchell Co.,
<geoCoordinate degrees="36.0933" direction="north" orientation="latitude" precision="5" value="36.0933">36.0933°N</geoCoordinate>
,
<geoCoordinate degrees="82.1447" direction="west" orientation="longitude" precision="5" value="-82.1447">82.1447°W</geoCoordinate>
, Roan High Bluff, 6251' viii.15.2018, M.Caterino, sifted
<taxonomicName class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" lsidName="Abies" order="Pinales" pageId="0" pageNumber="133" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="133">Abies</emphasis>
</taxonomicName>
litter&quot;; 1 ♀, 1 ♂: &quot;USA:NC: Mitchell Co.,
<geoCoordinate degrees="36.0999" direction="north" orientation="latitude" precision="5" value="36.0999">36.0999°N</geoCoordinate>
,
<geoCoordinate degrees="82.1345" direction="west" orientation="longitude" precision="5" value="-82.1345">82.1345°W</geoCoordinate>
, Roan High Bluff, 6146', viii.15.2018, M.Caterino,
<taxonomicName class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" lsidName="Rhododendron" order="Ericales" pageId="0" pageNumber="133" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="133">Rhododendron</emphasis>
</taxonomicName>
litter&quot;; 2 ♀, 12 ♂: &quot;USA: NC: Mitchell Co.,
<geoCoordinate degrees="36.1041" direction="north" orientation="latitude" precision="5" value="36.1041">36.1041°N</geoCoordinate>
,
<geoCoordinate degrees="82.1223" direction="west" orientation="longitude" precision="5" value="-82.1223">82.1223°W</geoCoordinate>
, PisgahNF, vi.8.2020, Roan High Knob, 6276', M. Caterino, conifer litter&quot;; 5 ♀, 4 ♂: &quot;USA: NC: Mitchell Co.,
<geoCoordinate degrees="36.1045" direction="north" orientation="latitude" precision="5" value="36.1045">36.1045°N</geoCoordinate>
,
<geoCoordinate degrees="82.1224" direction="west" orientation="longitude" precision="5" value="-82.1224">82.1224°W</geoCoordinate>
, PisgahNF, vi.8.2020, Roan High Knob, 6286', M. Caterino, conifer litter&quot;.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="133" type="materials_examined">
<paragraph pageId="0" pageNumber="133">Other material.</paragraph>
<paragraph pageId="0" pageNumber="133">
(52) -
<materialsCitation collectingDate="2020-08-05" county="Unicoi Co." elevation="1613" location="Cherokee National Forest" municipality="Big Bald" specimenCount="6" specimenCount-female="1" specimenCount-male="5">
<emphasis bold="true" pageId="0" pageNumber="133">TN</emphasis>
:
<collectingCounty>Unicoi Co.</collectingCounty>
,
<location LSID="urn:lsid:plazi:treatment:67BE608F976A50CEB559ADBB12F2355D:557CAA87A2BADBF60C0B419A1253E6D7" county="Unicoi Co." municipality="Big Bald" name="Cherokee National Forest">Cherokee National Forest</location>
,
<collectingMunicipality>Big Bald</collectingMunicipality>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.6128492" metricValueMax="1.6294608" metricValueMin="1.5962376" unit="ft" value="5291.5" valueMax="5346.0" valueMin="5237.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.6128492" metricValueMax="1.6294608" metricValueMin="1.5962376" unit="ft" value="5291.5" valueMax="5346.0" valueMin="5237.0">5237-5346ft</elevation>
</quantity>
,
<collectingDate value="2020-08-05">5-Aug-2020</collectingDate>
(
<specimenCount type="female">1 ♀</specimenCount>
,
<specimenCount type="male">5 ♂</specimenCount>
)
</materialsCitation>
;
<materialsCitation collectingDate="2022-04-21" county="Avery Co." elevation="1617" location="Grandfather Mt." specimenCount="9" specimenCount-female="4" specimenCount-male="5">
<emphasis bold="true" pageId="0" pageNumber="133">NC</emphasis>
:
<collectingCounty>Avery Co.</collectingCounty>
,
<location LSID="urn:lsid:plazi:treatment:67BE608F976A50CEB559ADBB12F2355D:5664E4D652A6312C61E244518F145F7B" county="Avery Co." name="Grandfather Mt.">Grandfather Mt.</location>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.6169639999999998" metricValueMax="1.636776" metricValueMin="1.597152" unit="ft" value="5305.0" valueMax="5370.0" valueMin="5240.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.6169639999999998" metricValueMax="1.636776" metricValueMin="1.597152" unit="ft" value="5305.0" valueMax="5370.0" valueMin="5240.0">5240-5370ft.</elevation>
</quantity>
,
<collectingDate value="2022-04-21">21-Apr-2022</collectingDate>
(
<specimenCount type="female">4 ♀</specimenCount>
,
<specimenCount type="male">5 ♂</specimenCount>
)
</materialsCitation>
;
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" location="Caldwell Co. Grandfather Mt. State Park" municipality="Calloway Peak" specimenCount="25" specimenCount-female="12" specimenCount-male="13">
<location LSID="urn:lsid:plazi:treatment:67BE608F976A50CEB559ADBB12F2355D:E5F98DCE33FAE47FA424B4115F3B0BA7" municipality="Calloway Peak" name="Caldwell Co. Grandfather Mt. State Park">Caldwell Co. Grandfather Mt. State Park</location>
,
<collectingMunicipality>Calloway Peak</collectingMunicipality>
,
<collectingDate value="2020-10-06">6-Oct-2020</collectingDate>
&amp;
<collectingDate value="2021-05-17">
17-May-
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.2095344110000001" metricValueMax="1.8028920000000004" metricValueMin="0.6161768220000001" unit="ft" value="3968.28875" valueMax="5915.0" valueMin="2021.5775">2021, 5775-5915ft.</quantity>
</collectingDate>
(
<specimenCount type="female">12 ♀</specimenCount>
,
<specimenCount type="male">13 ♂</specimenCount>
)
</materialsCitation>
;
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="Nuwati Tr." elevation="1277" location="Caldwell Co. Grandfather Mt. State Park" specimenCount="1" specimenCount-male="1">
<location LSID="urn:lsid:plazi:treatment:67BE608F976A50CEB559ADBB12F2355D:9A4C7FB56DFE5C90EB1D786CB4A5BCA3" name="Caldwell Co. Grandfather Mt. State Park">Caldwell Co. Grandfather Mt. State Park</location>
,
<collectorName>Nuwati Tr.</collectorName>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.277112" unit="ft" value="4190.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.277112" unit="ft" value="4190.0">4190ft.</elevation>
</quantity>
(
<specimenCount type="male">1 ♂</specimenCount>
)
</materialsCitation>
;
<materialsCitation collectingDate="2020-06-15" collectorName="Woody Ridge Tr." county="Yancey Co." elevation="1596" location="Pisgah National Forest" specimenCount="9" specimenCount-female="5" specimenCount-male="4">
<collectingCounty>Yancey Co.</collectingCounty>
,
<location LSID="urn:lsid:plazi:treatment:67BE608F976A50CEB559ADBB12F2355D:F2D16B51898CADF2006CD92AA0019542" county="Yancey Co." name="Pisgah National Forest">Pisgah National Forest</location>
,
<collectorName>Woody Ridge Tr.</collectorName>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.5960851999999999" metricValueMax="1.6419576" metricValueMin="1.5502128" unit="ft" value="5236.5" valueMax="5387.0" valueMin="5086.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.5960851999999999" metricValueMax="1.6419576" metricValueMin="1.5502128" unit="ft" value="5236.5" valueMax="5387.0" valueMin="5086.0">5086-5387ft.</elevation>
</quantity>
,
<collectingDate value="2020-06-15">15-Jun-2020</collectingDate>
(
<specimenCount type="female">5 ♀</specimenCount>
,
<specimenCount type="male">4 ♂</specimenCount>
)
</materialsCitation>
;
<materialsCitation collectingDate="2021-10-19" county="Yancey Co." elevation="1920" location="Pisgah National Forest" municipality="Celo Knob" specimenCount="1" specimenCount-male="1">
<collectingCounty>Yancey Co.</collectingCounty>
,
<location LSID="urn:lsid:plazi:treatment:67BE608F976A50CEB559ADBB12F2355D:9AB0BDFC061ED27C661BD4467957675C" county="Yancey Co." municipality="Celo Knob" name="Pisgah National Forest">Pisgah National Forest</location>
,
<collectingMunicipality>Celo Knob</collectingMunicipality>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.92024" unit="ft" value="6300.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.92024" unit="ft" value="6300.0">6300ft.</elevation>
</quantity>
,
<collectingDate value="2021-10-19">19-Oct-2021</collectingDate>
(
<specimenCount type="male">1 ♂</specimenCount>
)
</materialsCitation>
;
<materialsCitation collectingDate="2022-03-01" county="Madison Co." elevation="1445" location="Pisgah National Forest" municipality="Camp Creek" specimenCount="1" specimenCount-male="1">
<collectingCounty>Madison Co.</collectingCounty>
,
<location LSID="urn:lsid:plazi:treatment:67BE608F976A50CEB559ADBB12F2355D:9C8AF9CE3C8AF939AE1ADCB901A2DAF8" county="Madison Co." municipality="Camp Creek" name="Pisgah National Forest">Pisgah National Forest</location>
,
<collectingMunicipality>Camp Creek</collectingMunicipality>
Bald,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.4450568" unit="ft" value="4741.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.4450568" unit="ft" value="4741.0">4741ft.</elevation>
</quantity>
,
<collectingDate value="2022-03-01">1-Mar-2022</collectingDate>
(
<specimenCount type="male">1 ♂</specimenCount>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="133" type="description">
<paragraph pageId="0" pageNumber="133">Diagnostic description.</paragraph>
<paragraph pageId="0" pageNumber="133">
This species is extremely similar in external morphology to many of the preceding, and is also best distinguished by male genitalic characters. However, it does exhibit a few unusual characters. The male antennomeres VIII-IX are slightly enlarged, but lack carinae on their inner/anterior edges (Figs
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Habitus and character photos of Euconnus attritus A male, lateral view B male, dorsal view C female, lateral view D female, dorsal view E SEM of male antenna F SEM of venter of head." figureDoi="10.3897/zookeys.1137.97068.figure13" httpUri="https://binary.pensoft.net/fig/786424" pageId="0" pageNumber="133">13A, E</figureCitation>
); the
<normalizedToken originalValue="females">female's</normalizedToken>
antennal club is shorter, and essentially trimerous (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Habitus and character photos of Euconnus attritus A male, lateral view B male, dorsal view C female, lateral view D female, dorsal view E SEM of male antenna F SEM of venter of head." figureDoi="10.3897/zookeys.1137.97068.figure13" httpUri="https://binary.pensoft.net/fig/786424" pageId="0" pageNumber="133">13D</figureCitation>
), though antennomere VIII is slightly enlarged relative to VII; males and females flightless; median lobe of aedeagus (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Aedeagus. A Euconnus draco B Euconnus tusquitee C Euconnus attritus." figureDoi="10.3897/zookeys.1137.97068.figure12" httpUri="https://binary.pensoft.net/fig/786423" pageId="0" pageNumber="133">12C</figureCitation>
) relatively long, nearly as long as basal bulb, narrowed to bluntly rounded, but not knobbed apex; shoulders of aedeagus sloped; parameres weakly curved, apices tapered, bearing three (rarely two) long straight setae; compressor plate symmetrical, parallel-sided to near apex, then narrowed to subacute apex, nearly as long as median lobe; endophallus with single long, upper sclerite, strongly sinuate, apically acute, bearing a sharp secondary process near its midpoint; the right side of the compressor plate sclerotized in a linear band, appearing as a second similar linear sclerite; actual second upper sclerite is much shorter, bent strongly at thickened midpoint, tapering to a thin, acute tip; lower endophallic sclerite a similar-length, trifid claw-like process on a sinuate stem originating near basal orifice.
</paragraph>
<caption doi="10.3897/zookeys.1137.97068.figure13" httpUri="https://binary.pensoft.net/fig/786424" pageId="0" pageNumber="133" start="Figure 13" startId="F13">
<paragraph pageId="0" pageNumber="133">
<emphasis bold="true" pageId="0" pageNumber="133">Figure 13.</emphasis>
Habitus and character photos of
<taxonomicName authorityName="Caterino" authorityYear="2022" class="Insecta" family="Staphylinidae" genus="Euconnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Euconnus attritus" order="Coleoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="attritus">
<emphasis italics="true" pageId="0" pageNumber="133">Euconnus attritus</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="133">A</emphasis>
male, lateral view
<emphasis bold="true" pageId="0" pageNumber="133">B</emphasis>
male, dorsal view
<emphasis bold="true" pageId="0" pageNumber="133">C</emphasis>
female, lateral view
<emphasis bold="true" pageId="0" pageNumber="133">D</emphasis>
female, dorsal view
<emphasis bold="true" pageId="0" pageNumber="133">E</emphasis>
SEM of male antenna
<emphasis bold="true" pageId="0" pageNumber="133">F</emphasis>
SEM of venter of head.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="133" type="distribution">
<paragraph pageId="0" pageNumber="133">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="133">
This species is known only from northeast of the Asheville depression, although there it is moderately widespread, occurring in all the major ranges: the Black Mts, Roan Highlands, Grandfather Mountain, Big Bald, and Camp Creek Bald. Its known localities span an elevation range of 3800-6300 ft. There is a CUAC specimen labelled 'Toxaway
<normalizedToken originalValue="Mountain">Mountain'</normalizedToken>
(in the more southerly Nantahala Mts.) that is almost certainly mislabeled, as that site was visited the same day as the Big Bald locality.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="133" type="remarks">
<paragraph pageId="0" pageNumber="133">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="133">
Of the pale species of
<taxonomicName authorityName="Reitter" authorityYear="1909" class="Insecta" family="Staphylinidae" genus="Cladoconnus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Cladoconnus" order="Coleoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="133">Cladoconnus</emphasis>
</taxonomicName>
, this species is only sympatric with
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="E. adversus" order="Coleoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="adversus">
<emphasis italics="true" pageId="0" pageNumber="133">E. adversus</emphasis>
</taxonomicName>
(it is also sympatric with the larger and darker
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="E. vetustus" order="Coleoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="vetustus">
<emphasis italics="true" pageId="0" pageNumber="133">E. vetustus</emphasis>
</taxonomicName>
and
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="E. vexillus" order="Coleoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="vexillus">
<emphasis italics="true" pageId="0" pageNumber="133">E. vexillus</emphasis>
</taxonomicName>
). Males of
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="E. adversus" order="Coleoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="adversus">
<emphasis italics="true" pageId="0" pageNumber="133">E. adversus</emphasis>
</taxonomicName>
have very conspicuous antennal carinae, which are completely lacking in
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="E. attritus" order="Coleoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="attritus">
<emphasis italics="true" pageId="0" pageNumber="133">E. attritus</emphasis>
</taxonomicName>
. Females, however, will be indistinguishable. Males of
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="E. attritus" order="Coleoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="attritus">
<emphasis italics="true" pageId="0" pageNumber="133">E. attritus</emphasis>
</taxonomicName>
have very distinctive endophallic sclerites, particularly the long, sinuate left upper sclerite, with its secondary median spike.
</paragraph>
<paragraph pageId="0" pageNumber="133">
The name of this species suggests that the subgenus-typical male antennal carinae are 'worn
<normalizedToken originalValue="away">away'</normalizedToken>
.
</paragraph>
</subSubSection>
</treatment>
</document>