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<document ID-CLB-Dataset="24088" ID-DOI="http://dx.doi.org/10.3897/zookeys.910.39486" ID-GBIF-Dataset="fd47c8fc-f4b8-4924-9a45-51e09fcd5e5b" ID-PMC="PMC7026197" ID-Pensoft-Pub="1313-2970-910-1" ID-Pensoft-UUID="5E9B7F572E9952BD87FD845179195CB5" ID-PubMed="32099514" ID-ZooBank="2C11E2A17D5D42A080ECE5FC618FF47B" ModsDocID="1313-2970-910-1" checkinTime="1581415676306" checkinUser="pensoft" docAuthor="Silva, Marcos Santos &amp; Carbayo, Fernando" docDate="2020" docId="494E275D192851F0B259A3F7736F7F10" docLanguage="en" docName="ZooKeys 910: 1-42" docOrigin="ZooKeys 910" docSource="http://dx.doi.org/10.3897/zookeys.910.39486" docTitle="Pseudogeoplana schirchi" docType="treatment" docVersion="6" id="5E9B7F572E9952BD87FD845179195CB5" lastPageNumber="1" masterDocId="5E9B7F572E9952BD87FD845179195CB5" masterDocTitle="X-ray microcomputed tomography applied to the taxonomic study of rare material: redescriptions of seven of Schirch's Brazilian species of land planarians (Geoplanidae, Platyhelminthes)" masterLastPageNumber="42" masterPageNumber="1" pageNumber="1" updateTime="1732748294428" updateUser="ExternalLinkService">
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<mods:title id="FE50104B566F9DC0F441067AACE67C69">X-ray microcomputed tomography applied to the taxonomic study of rare material: redescriptions of seven of Schirch's Brazilian species of land planarians (Geoplanidae, Platyhelminthes)</mods:title>
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<mods:namePart id="48BF48A09AF216482042693A40546AE4">Silva, Marcos Santos</mods:namePart>
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<mods:namePart id="3FB2F3F9D444DBA87D7A97A1C8CB4F37">Carbayo, Fernando</mods:namePart>
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<treatment id="494E275D192851F0B259A3F7736F7F10" ID-GBIF-Taxon="162074486" LSID="urn:lsid:plazi:treatment:494E275D192851F0B259A3F7736F7F10" httpUri="http://treatment.plazi.org/id/494E275D192851F0B259A3F7736F7F10" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Geoplanidae" scope_phylum="Platyhelminthes" scope_subFamily="Rhynchodeminae">
<subSubSection id="CFEDB3F2CEFE1CC2D1C6148161F64868" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="E2905E51C605967AE8DD947FC70ADA15" pageId="0" pageNumber="1">
<taxonomicName id="15C9C551F400DF6991EB0713D42831E6" ID-CoL="4NX84" LSID="494E275D-1928-51F0-B259-A3F7736F7F10" authority="(Schirch, 1929)" baseAuthorityName="Schirch" baseAuthorityYear="1929" family="Geoplanidae" genus="Pseudogeoplana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudogeoplana schirchi" order="Tricladida" pageId="0" pageNumber="1" phylum="Platyhelminthes" rank="species" species="schirchi">Pseudogeoplana schirchi (Schirch, 1929)</taxonomicName>
<figureCitation id="8A5116B7D10D5DFAB38933947CD7B212" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Pseudogeoplana schirchi (Schirch 1929), Holotype. A Original illustration of Ps. schirchi by Schirch (1929) B dorsal view of fixed holotype." figureDoi="10.3897/zookeys.910.39486.figure25" httpUri="https://binary.pensoft.net/fig/378104" pageId="0" pageNumber="1">Figures 25</figureCitation>
<figureCitation id="A3B8E68875C2DC6BCDA5D5529B5C9F40" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Pseudogeoplana schirchi (Schirch 1929). Holotype. A Photomicrograph of a sagittal section of anterior extremity B virtual sagittal section showing pharyngeal region, pixel size: 9 µm." figureDoi="10.3897/zookeys.910.39486.figure26" httpUri="https://binary.pensoft.net/fig/378105" pageId="0" pageNumber="1">, 26</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="5E2C0D45559AAA89D4742EACEA133961" pageId="0" pageNumber="1" type="reference_group">
<paragraph id="2E2170B868F12DCD94452E38105ACE49" pageId="0" pageNumber="1">
<taxonomicName id="EFC0BC5A7BA7441481F1857670D7EC25" family="Geoplanidae" genus="Geoplana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geoplana maximiliani" order="Tricladida" pageId="0" pageNumber="1" phylum="Platyhelminthes" rank="species" species="maximiliani">Geoplana maximiliani</taxonomicName>
Schirch, 1929: 30; taf. 2, fig, 10. Type locality:
<normalizedToken id="5E04684B96316E1E106FC41B64653849" originalValue="Teresópolis">Teresopolis</normalizedToken>
, State of Rio de Janeiro, Brazil.
</paragraph>
<paragraph id="D0DBB366645A7BC0523BDED88155AF1C" pageId="0" pageNumber="1">
<taxonomicName id="3F479526301C56D0E5704A0BD29B834A" baseAuthorityName="Schirch" baseAuthorityYear="1929" family="Geoplanidae" genus="Pseudogeoplana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudogeoplana schirchi" order="Tricladida" pageId="0" pageNumber="1" phylum="Platyhelminthes" rank="species" species="schirchi">Pseudogeoplana schirchi</taxonomicName>
:
<bibRefCitation id="342869649AD214FF982A1871C6930A72" author="Ogren, RE" journalOrPublisher="Bulletin of Fuji Women's University Series" pageId="0" pageNumber="1" pagination="79 - 166" refId="B27" refString="Ogren, RE, Kawakatsu, M, 1990. Index to the species of the family Geoplanidae (Turbellaria, Tricladida, Terricola) Part I: Geoplaninae. Bulletin of Fuji Women's University Series 1 (28): 79 - 166" title="Index to the species of the family Geoplanidae (Turbellaria, Tricladida, Terricola) Part I: Geoplaninae." volume="1" year="1990">Ogren and Kawakatsu 1990</bibRefCitation>
: 159. nom. nov.
</paragraph>
</subSubSection>
<subSubSection id="BE9CDFD1C85C4FC40894B8A029F5A909" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="8F9C647DC894B740939322240BC16F90" pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph id="52B772C1B6334BA2A220E5331C6265FB" pageId="0" pageNumber="1">
Type material. Single holotype received on loan in 70 % ethanol and, subsequently, three-dimensional (3D) images and virtual sections of syntype MNRJ 217 were obtained by microcomputed tomography.
<emphasis id="06CFE9DD1FE13B21BD0118B1FBE69339" bold="true" pageId="0" pageNumber="1">MNRJ 217</emphasis>
,
<emphasis id="F65BE350E1D811AEDC5BFB623BAC5FF0" bold="true" pageId="0" pageNumber="1">holotype</emphasis>
, here designated by monotypy:
<normalizedToken id="7F8BD3A4131CBEBAC2CAF68EF48F3394" originalValue="Teresópolis">Teresopolis</normalizedToken>
, State of Rio de Janeiro, Brazil. P. Schirch
<emphasis id="65455962787FCF466B2DA692D56C7A96" italics="true" pageId="0" pageNumber="1">Coll</emphasis>
(unknown). Sagittal sections of anterior extremity on 97 slides; horizontal sections of pre-pharyngeal region on 15 slides. Remaining part of body preserved in 80 % ethanol.
</paragraph>
</subSubSection>
<subSubSection id="7035359B1C765E151E1BD547C48EE649" pageId="0" pageNumber="1" type="external aspect">
<paragraph id="7A70C747DBB623B972506F13462D03D0" pageId="0" pageNumber="1">External aspect.</paragraph>
<paragraph id="5098D262EEAEB355C66A7A0DC3401295" pageId="0" pageNumber="1">
Fixed holotype measured 45 mm long, 6 mm wide and 0.9 mm high. Elongated body, with parallel margins; anterior extremity tapers, posterior rounded. Dorsum convex, ventral side flat. Dorsum has yellow olive color, except in some parts where epithelium is lost (Fig.
<figureCitation id="D7F75F64BB90F85084B053948FBDDD96" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Pseudogeoplana schirchi (Schirch 1929), Holotype. A Original illustration of Ps. schirchi by Schirch (1929) B dorsal view of fixed holotype." figureDoi="10.3897/zookeys.910.39486.figure25" httpUri="https://binary.pensoft.net/fig/378104" pageId="0" pageNumber="1">25B</figureCitation>
). Ventral side brown grey with some darker regions. Relative position of mouth: body length, 58 %.
</paragraph>
<caption id="05E56BBFC61649F41244CC4F92E6A7C6" doi="10.3897/zookeys.910.39486.figure25" httpUri="https://binary.pensoft.net/fig/378104" pageId="0" pageNumber="1" start="Figure 25" startId="F25">
<paragraph id="BB1934D7552F8BEAC30193D778BC068A" pageId="0" pageNumber="1">
<emphasis id="0C7FE02523D1A36DF44D65A3D9B8D39A" bold="true" pageId="0" pageNumber="1">Figure 25.</emphasis>
<taxonomicName id="1A0415CA941AF80FEC154A38FC50BD36" baseAuthorityName="Schirch" baseAuthorityYear="1929" family="Geoplanidae" genus="Pseudogeoplana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudogeoplana schirchi" order="Tricladida" pageId="0" pageNumber="1" phylum="Platyhelminthes" rank="species" species="schirchi">
<emphasis id="499A575C397D9FC894673D9EF760CE34" italics="true" pageId="0" pageNumber="1">Pseudogeoplana schirchi</emphasis>
</taxonomicName>
(
<bibRefCitation id="B0321C220C4E1E03ED90A76B014DDAC5" author="Schirch, PF" journalOrPublisher="Stain Technology" pageId="0" pageNumber="1" refId="B31" refString="Schirch, PF, 1929. Sobre as planarias terrestres do Brasil. Boletim do Museu Nacional, Rio de Janeiro 5, 27-38." title="Sobre as planarias terrestres do Brasil. Boletim do Museu Nacional, Rio de Janeiro 5, 27 - 38." year="1929">Schirch 1929</bibRefCitation>
), Holotype.
<emphasis id="ED7370E22B6CFA1186B26E6A86C2BDB1" bold="true" pageId="0" pageNumber="1">A</emphasis>
Original illustration of
<taxonomicName id="D805F87B48FB67A92F8043040EC23288" lsidName="Ps. schirchi" pageId="0" pageNumber="1" rank="species" species="schirchi">
<emphasis id="1AE709565FE850DFBCC3C81AACDF8797" italics="true" pageId="0" pageNumber="1">Ps. schirchi</emphasis>
</taxonomicName>
by
<bibRefCitation id="A76679B7B4FF00A7CA838754B1F7865E" author="Schirch, PF" journalOrPublisher="Stain Technology" pageId="0" pageNumber="1" refId="B31" refString="Schirch, PF, 1929. Sobre as planarias terrestres do Brasil. Boletim do Museu Nacional, Rio de Janeiro 5, 27-38." title="Sobre as planarias terrestres do Brasil. Boletim do Museu Nacional, Rio de Janeiro 5, 27 - 38." year="1929">Schirch (1929)</bibRefCitation>
<emphasis id="C5D4D244E69371C3725FFC5F4616BBF8" bold="true" pageId="0" pageNumber="1">B</emphasis>
dorsal view of fixed holotype.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="5713F355E5DAF09467CDB956968FE734" pageId="0" pageNumber="1" type="internal morphology">
<paragraph id="3ABF2222095A6C5043D68DDE6B4C0A55" pageId="0" pageNumber="1">Internal morphology.</paragraph>
<paragraph id="C9554F4CA4C0FE85884814A1034710E1" pageId="0" pageNumber="1">
Creeping sole comprised ~88 % body width. Histological sections of holotype are of poor quality. Cutaneous musculature comprises three layers: a subepithelial circular layer, followed by a diagonal layer of decussate fibers and a clearly distinguishable innermost longitudinal layer. Muscle fibers of the longitudinal layer (90
<normalizedToken id="B97D449CD9C51E44C220A92595316172" originalValue="µm">µm</normalizedToken>
thick dorsally; 70
<normalizedToken id="AEE234B4E55B512363B3D0FBA4968B4C" originalValue="µm">µm</normalizedToken>
thick ventrally) arranged into bundles (Fig.
<figureCitation id="0BE0E950EEED2CFF8DA39027C66649DB" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Pseudogeoplana schirchi (Schirch 1929). Holotype. A Photomicrograph of a sagittal section of anterior extremity B virtual sagittal section showing pharyngeal region, pixel size: 9 µm." figureDoi="10.3897/zookeys.910.39486.figure26" httpUri="https://binary.pensoft.net/fig/378105" pageId="0" pageNumber="1">26A</figureCitation>
). No muscle modifications at the anterior extremity of the body.
</paragraph>
<paragraph id="5EAA5FC42BFE1A74C16914DCF69EBE99" pageId="0" pageNumber="1">
Mouth situated at a distance from the root of the pharynx equivalent to 56 % of the pharyngeal pocket length (Fig.
<figureCitation id="38D45546D9A2F5357223C9495B56457B" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Pseudogeoplana schirchi (Schirch 1929). Holotype. A Photomicrograph of a sagittal section of anterior extremity B virtual sagittal section showing pharyngeal region, pixel size: 9 µm." figureDoi="10.3897/zookeys.910.39486.figure26" httpUri="https://binary.pensoft.net/fig/378105" pageId="0" pageNumber="1">26B</figureCitation>
). Pharynx cylindrical, occupying ~90 % of pharyngeal pocket. Esophagus, apparently absent. Copulatory apparatus not developed, neither are testes or ovaries.
</paragraph>
<caption id="F7CA5A2390A444EB05AC301729291225" doi="10.3897/zookeys.910.39486.figure26" httpUri="https://binary.pensoft.net/fig/378105" pageId="0" pageNumber="1" start="Figure 26" startId="F26">
<paragraph id="73E9A6293F87F52FDBF6719E79661607" pageId="0" pageNumber="1">
<emphasis id="20884FA8BDAC436C437C5FD8437EEE85" bold="true" pageId="0" pageNumber="1">Figure 26.</emphasis>
<taxonomicName id="46AB80EC07C12BC190EA52D343C48D75" baseAuthorityName="Schirch" baseAuthorityYear="1929" family="Geoplanidae" genus="Pseudogeoplana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudogeoplana schirchi" order="Tricladida" pageId="0" pageNumber="1" phylum="Platyhelminthes" rank="species" species="schirchi">
<emphasis id="8D0DEFAD33FDA70EE1F8B43EADB9E26F" italics="true" pageId="0" pageNumber="1">Pseudogeoplana schirchi</emphasis>
</taxonomicName>
(
<bibRefCitation id="E6C1CF50AADF6844DB195D134FA9CAD6" author="Schirch, PF" journalOrPublisher="Stain Technology" pageId="0" pageNumber="1" refId="B31" refString="Schirch, PF, 1929. Sobre as planarias terrestres do Brasil. Boletim do Museu Nacional, Rio de Janeiro 5, 27-38." title="Sobre as planarias terrestres do Brasil. Boletim do Museu Nacional, Rio de Janeiro 5, 27 - 38." year="1929">Schirch 1929</bibRefCitation>
). Holotype.
<emphasis id="42D2DC19D7D5A664C8F4D6260333C051" bold="true" pageId="0" pageNumber="1">A</emphasis>
Photomicrograph of a sagittal section of anterior extremity
<emphasis id="D0CCAAB6AFCD39094908A43CF1F72229" bold="true" pageId="0" pageNumber="1">B</emphasis>
virtual sagittal section showing pharyngeal region, pixel size: 9
<normalizedToken id="99D3721F179C4EAEF121D75597C1ECD6" originalValue="µm">µm</normalizedToken>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="E0E8D819D0E6C95C66733A88CB38202D" pageId="0" pageNumber="1" type="remarks">
<paragraph id="FABC5EB24A2C792860C462F6312F5350" pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph id="359CD260050C3FEA724F5A48F16D1B03" pageId="0" pageNumber="1">
Schirch did not provide any measurement or description of the specimen he studied but illustrated its dorsal aspect (Fig.
<figureCitation id="5B0626BDA834FDF970405DDA93601442" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Pseudogeoplana schirchi (Schirch 1929), Holotype. A Original illustration of Ps. schirchi by Schirch (1929) B dorsal view of fixed holotype." figureDoi="10.3897/zookeys.910.39486.figure25" httpUri="https://binary.pensoft.net/fig/378104" pageId="0" pageNumber="1">25A</figureCitation>
). The specimen presents parallel body sides, whereas the figure shows a slightly lanceolate specimen. This situation casts some doubts on the labeling (as is the case of
<taxonomicName id="396179E94BE41C5BEF63DA215A68C7F5" lsidName="Ps. blaseri" pageId="0" pageNumber="1" rank="species" species="blaseri">
<emphasis id="27BD615EAB00FEA2088E53BEFCABCF66" italics="true" pageId="0" pageNumber="1">Ps. blaseri</emphasis>
</taxonomicName>
, see above). The specimen was apparently dehydrated for an uncertain period. This might explain the differences in body shape.
</paragraph>
<paragraph id="9834DA32E19628E06B828051113807AA" pageId="0" pageNumber="1">
Most features diagnosing the subfamily
<taxonomicName id="99BD44B969CF182B9F4E310B6DDB61DE" authorityName="Stimpson" authorityYear="1857" lsidName="" pageId="0" pageNumber="1" rank="subFamily" subFamily="Geoplaninae">Geoplaninae</taxonomicName>
were observed in the holotype, namely, creeping sole covering most of ventral surface, mouth posterior to mid-body, and a well-developed cutaneous musculature organized into bundles. The immaturity of the specimen makes it impossible to know the position of the testes.
</paragraph>
<paragraph id="A89FD25ACB28F42478A17C0B4322705C" pageId="0" pageNumber="1">
The specimen collected in
<normalizedToken id="B128236A12613EBC287288EF54BF2DFF" originalValue="Teresópolis">Teresopolis</normalizedToken>
was identified by Schirch as
<taxonomicName id="F9A5719D4C1F90F92C7F969C17BB25A2" family="Geoplanidae" genus="Geoplana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geoplana maximiliani" order="Tricladida" pageId="0" pageNumber="1" phylum="Platyhelminthes" rank="species" species="maximiliani">
<emphasis id="19B9C23C053AD0CD6892031AA2310945" italics="true" pageId="0" pageNumber="1">Geoplana maximiliani</emphasis>
</taxonomicName>
Schultze &amp;
<normalizedToken id="5160F5507596096B75BFD671F2D87880" originalValue="Müller">Mueller</normalizedToken>
, 1857 (currently placed in
<taxonomicName id="92D079E19524BC97634F615FEA342AD8" authorityName="Ogren &amp; Kawakatsu" authorityYear="1990" family="Geoplanidae" genus="Pseudogeoplana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudogeoplana" order="Tricladida" pageId="0" pageNumber="1" phylum="Platyhelminthes" rank="genus">
<emphasis id="5D58F45333F6505E01B70DB1AA97E4CC" italics="true" pageId="0" pageNumber="1">Pseudogeoplana</emphasis>
</taxonomicName>
), originally from Blumenau, State of Santa Catarina, Brazil.
<bibRefCitation id="49A65905B89FD8F51A4251C3FCD6F769" DOI="https://doi.org/10.11606/issn.2526-3382.bffclzoologia.1959.120326" author="Froehlich, CG" journalOrPublisher="Zoologia" pageId="0" pageNumber="1" pagination="223 - 225" refId="B15" refString="Froehlich, CG, 1959. On Geoplanids from Brazil. Boletim da Faculdade de Filosofia, Ciencias e Letras, Universidade de Sao Paulo. Zoologia 22 (22): 223 - 225, DOI: https://doi.org/10.11606/issn.2526-3382.bffclzoologia.1959.120326" title="On Geoplanids from Brazil. Boletim da Faculdade de Filosofia, Ciencias e Letras, Universidade de Sao Paulo." url="https://doi.org/10.11606/issn.2526-3382.bffclzoologia.1959.120326" volume="22" year="1959">Froehlich (1959)</bibRefCitation>
considered the shape and color pattern of
<normalizedToken id="B6D98FBA0E57196E3C3A02324DEBAD7C" originalValue="Schirchs">Schirch's</normalizedToken>
specimen as compatible with Schultze and
<normalizedToken id="00C13F43AF12DB9CC1846BC979850E09" originalValue="Müllers">Mueller's</normalizedToken>
species, but refrained from confirming its conspecificity because the internal morphology of the specimens from both localities was unknown, plus they were collected far from each other (
<bibRefCitation id="669CC752BEB47CF05E1FDC5AD9236561" DOI="https://doi.org/10.11606/issn.2526-3382.bffclzoologia.1959.120326" author="Froehlich, CG" journalOrPublisher="Zoologia" pageId="0" pageNumber="1" pagination="223 - 225" refId="B15" refString="Froehlich, CG, 1959. On Geoplanids from Brazil. Boletim da Faculdade de Filosofia, Ciencias e Letras, Universidade de Sao Paulo. Zoologia 22 (22): 223 - 225, DOI: https://doi.org/10.11606/issn.2526-3382.bffclzoologia.1959.120326" title="On Geoplanids from Brazil. Boletim da Faculdade de Filosofia, Ciencias e Letras, Universidade de Sao Paulo." url="https://doi.org/10.11606/issn.2526-3382.bffclzoologia.1959.120326" volume="22" year="1959">Froehlich 1959</bibRefCitation>
).
<bibRefCitation id="04F046AF09EEEEAE722FF7AF26B4CC9D" author="Ogren, RE" journalOrPublisher="Bulletin of Fuji Women's University Series" pageId="0" pageNumber="1" pagination="79 - 166" refId="B27" refString="Ogren, RE, Kawakatsu, M, 1990. Index to the species of the family Geoplanidae (Turbellaria, Tricladida, Terricola) Part I: Geoplaninae. Bulletin of Fuji Women's University Series 1 (28): 79 - 166" title="Index to the species of the family Geoplanidae (Turbellaria, Tricladida, Terricola) Part I: Geoplaninae." volume="1" year="1990">Ogren and Kawakatsu (1990)</bibRefCitation>
endorsed
<normalizedToken id="054658DEBBDA7886F399060F67697F88" originalValue="Froehlichs">Froehlich's</normalizedToken>
opinion and gave a new name, namely
<taxonomicName id="A0CF736745B0CA243B51374CF264D733" baseAuthorityName="Schirch" baseAuthorityYear="1929" family="Geoplanidae" genus="Pseudogeoplana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudogeoplana schirchi" order="Tricladida" pageId="0" pageNumber="1" phylum="Platyhelminthes" rank="species" species="schirchi">
<emphasis id="130772DFD8BF8C4E07F7C2092B544B9A" italics="true" pageId="0" pageNumber="1">Pseudogeoplana schirchi</emphasis>
</taxonomicName>
, to the specimen from
<normalizedToken id="38F43D54F1593F504EBBA88C04B3F332" originalValue="Teresópolis">Teresopolis</normalizedToken>
to avoid further confusion. As no further morphological details could be observed, the species will remain in
<taxonomicName id="50F42A43622111D424DDA57F66CBE6CD" authorityName="Ogren &amp; Kawakatsu" authorityYear="1990" family="Geoplanidae" genus="Pseudogeoplana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudogeoplana" order="Tricladida" pageId="0" pageNumber="1" phylum="Platyhelminthes" rank="genus">
<emphasis id="C9B873C350D781B7990DC2EAA40B49AB" italics="true" pageId="0" pageNumber="1">Pseudogeoplana</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>