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<document id="F15965B05F64C20CFB4BFD7248A4D3B0" ID-CLB-Dataset="186677" ID-DOI="10.11646/zootaxa.5205.5.3" ID-GBIF-Dataset="e1b0e133-4416-41dc-a9f8-69da1e114cb7" ID-ISSN="1175-5326" ID-Zenodo-Dep="7318489" ID-ZooBank="EEC036E7-C49C-4B42-BB4C-04BB8A6B8D8E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1668421213205" checkinUser="plazi" docAuthor="Prokofiev, Artem M." docDate="2022" docId="038387EAFFB9AC6312F11F8CFA4CF85E" docLanguage="en" docName="zootaxa.5205.5.3.pdf" docOrigin="Zootaxa 5205 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Maechidius addarcis Prokofiev 2022, new species" docType="treatment" docVersion="6" lastPageNumber="446" masterDocId="FFBAFF92FFB8AC6212661F55FB16FFA9" masterDocTitle="New species and records of Maechidius Macleay, 1819 from New Guinea (Coleoptera: Scarabaeidae: Melolonthinae)" masterLastPageNumber="462" masterPageNumber="445" pageNumber="446" updateTime="1699411288883" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="5B62D5A6EFB37F599BA24AF2A5BCF80A">New species and records of Maechidius Macleay, 1819 from New Guinea (Coleoptera: Scarabaeidae: Melolonthinae)</mods:title>
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<mods:namePart id="1B6E536FBE3BD3C5DE6B6561AC00F27D">Prokofiev, Artem M.</mods:namePart>
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<mods:date id="95C238506A0A7309ECE71FC1AB5A6369">2022</mods:date>
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<treatment id="038387EAFFB9AC6312F11F8CFA4CF85E" ID-DOI="http://doi.org/10.5281/zenodo.7318491" ID-GBIF-Taxon="204971262" ID-Zenodo-Dep="7318491" LSID="urn:lsid:plazi:treatment:038387EAFFB9AC6312F11F8CFA4CF85E" httpUri="http://treatment.plazi.org/id/038387EAFFB9AC6312F11F8CFA4CF85E" lastPageNumber="446" pageId="1" pageNumber="446">
<subSubSection id="C3306577FFB9AC6312F11F8CF9A7FF5A" box="[151,689,217,244]" pageId="1" pageNumber="446" type="nomenclature">
<paragraph id="8B9536FCFFB9AC6312F11F8CF9A7FF5A" blockId="1.[151,689,217,280]" box="[151,689,217,244]" pageId="1" pageNumber="446">
<heading id="D0DD8190FFB9AC6312F11F8CF9A7FF5A" bold="true" box="[151,689,217,244]" fontSize="11" level="1" pageId="1" pageNumber="446" reason="1">
<emphasis id="B95EEAEEFFB9AC6312F11F8CF9A7FF5A" bold="true" box="[151,689,217,244]" pageId="1" pageNumber="446">
<taxonomicName id="4C2A4D7FFFB9AC6312F11F8CF902FF5A" authority="Prokofiev" authorityName="Prokofiev" authorityYear="2022" box="[151,532,217,244]" class="Insecta" family="Melolonthidae" genus="Maechidius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="446" phylum="Arthropoda" rank="species" species="addarcis" status="sp. nov.">
<emphasis id="B95EEAEEFFB9AC6312F11F8CFA85FF5D" bold="true" box="[151,403,217,244]" italics="true" pageId="1" pageNumber="446">Maechidius addarcis</emphasis>
Prokofiev
</taxonomicName>
,
<taxonomicNameLabel id="A26D5795FFB9AC6310471F8FF9A7FF5A" box="[545,689,217,244]" pageId="1" pageNumber="446" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C3306577FFB9AC6312F11FABFBCFFEB1" box="[151,217,254,280]" pageId="1" pageNumber="446" type="description">
<paragraph id="8B9536FCFFB9AC6312F11FABFBCFFEB1" blockId="1.[151,689,217,280]" box="[151,217,254,280]" pageId="1" pageNumber="446">
<figureCitation id="13112A79FFB9AC6312F11FABFBCFFEB1" box="[151,217,254,280]" captionStart="FIGURE 1" captionStartId="2.[152,255,1102,1126]" captionTargetBox="[157,1427,189,1072]" captionTargetId="figure-27@2.[151,1436,181,1078]" captionTargetPageId="2" captionText="FIGURE 1. Maechidius addarcis Prokofiev, new species. (A) dorsal view, holotype; (B) protibia, holotype; (C) aedeagus, lateral view, holotype; (D) parameres, dorsal view, holotype; (E) spiculum gastrale, dorsal view, holotype; (F) ditto, lateral view; (G) protibia, female paratype. Scale bars: (A, B, G) 1.0 mm; (CF) 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7318493" httpUri="https://zenodo.org/record/7318493/files/figure.png" pageId="1" pageNumber="446">Fig. 1</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3306577FFB9AC6312F11E14F8DFFED6" pageId="1" pageNumber="446" type="materials_examined">
<paragraph id="8B9536FCFFB9AC6312F11E14F8DFFED6" blockId="1.[151,1437,320,2039]" pageId="1" pageNumber="446">
<emphasis id="B95EEAEEFFB9AC6312F11E14FA5EFEF3" bold="true" box="[151,328,320,347]" pageId="1" pageNumber="446">Type material.</emphasis>
<materialsCitation id="3B423CA1FFB9AC6313341E14FA17FED6" ID-GBIF-Occurrence="3965147310" collectingDate="1921-11" collectingDateMax="1939-01" collectingDateMin="1921-11" collectionCode="BMNH" collectorName="R. Neill Rothschild" country="Guinea-Bissau" location="New" municipality="Doron" pageId="1" pageNumber="446" specimenCount="1" typeStatus="holotype">
<emphasis id="B95EEAEEFFB9AC6313341E14FAA9FEF2" bold="true" box="[338,447,321,347]" pageId="1" pageNumber="446">
<typeStatus id="5491885EFFB9AC6313341E14FAA9FEF2" box="[338,447,321,347]" pageId="1" pageNumber="446" type="holotype">Holotype</typeStatus>
</emphasis>
, male (
<figureCitation id="13112A79FFB9AC63107E1E15F963FEF2" box="[536,629,320,347]" captionStart="FIGURE 1" captionStartId="2.[152,255,1102,1126]" captionTargetBox="[157,1427,189,1072]" captionTargetId="figure-27@2.[151,1436,181,1078]" captionTargetPageId="2" captionText="FIGURE 1. Maechidius addarcis Prokofiev, new species. (A) dorsal view, holotype; (B) protibia, holotype; (C) aedeagus, lateral view, holotype; (D) parameres, dorsal view, holotype; (E) spiculum gastrale, dorsal view, holotype; (F) ditto, lateral view; (G) protibia, female paratype. Scale bars: (A, B, G) 1.0 mm; (CF) 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7318493" httpUri="https://zenodo.org/record/7318493/files/figure.png" pageId="1" pageNumber="446">Fig. 1A</figureCitation>
): “
<location id="8EF56027FFB9AC6310F11E14F9D8FEF2" LSID="urn:lsid:plazi:treatment:038387EAFFB9AC6312F11F8CFA4CF85E:8EF56027FFB9AC6310F11E14F9D8FEF2" box="[663,718,321,347]" country="Guinea-Bissau" municipality="Doron" name="New" pageId="1" pageNumber="446">New</location>
<collectingCountry id="F33D766CFFB9AC6310BE1E15F83FFEF3" box="[728,809,320,346]" name="Guinea-Bissau" pageId="1" pageNumber="446">Guinea</collectingCountry>
/
<collectingMunicipality id="6BF1AC86FFB9AC6311231E14F89BFEF2" box="[837,909,321,347]" pageId="1" pageNumber="446">Doron</collectingMunicipality>
<date id="FF94103CFFB9AC6311F01E15F8FDFEF3" box="[918,1003,320,346]" pageId="1" pageNumber="446" value="1921-11">
<collectingDate id="EFD0E9D4FFB9AC6311F01E15F8FDFEF3" box="[918,1003,320,346]" pageId="1" pageNumber="446" value="1921-11">xi.1921</collectingDate>
</date>
/
<collectorName id="26DF532AFFB9AC6316611E14FFF3FEF3" box="[1031,1253,320,347]" pageId="1" pageNumber="446">R. Neill Rothschild</collectorName>
/ B.M.
<date id="FF94103CFFB9AC6317231E14FE87FEF2" box="[1349,1425,321,347]" bridgedPair="-" pageId="1" pageNumber="446" value="1939-01">
<collectingDate id="EFD0E9D4FFB9AC6317231E14FE87FEF2" box="[1349,1425,321,347]" bridgedPair="-" pageId="1" pageNumber="446" value="1939-01">1939-I</collectingDate>
</date>
” (
<collectionCode id="ED3BAE39FFB9AC6312F91E30FBEFFED6" box="[159,249,357,383]" pageId="1" pageNumber="446">BMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B423CA1FFB9AC63136A1E30F8D3FED6" ID-GBIF-Occurrence="3965147312" box="[268,965,356,383]" collectingDate="1921-11" collectingDateMax="1939-01" collectingDateMin="1921-11" collectionCode="BMNH" collectorName="R. Neill Rothschild" country="Guinea-Bissau" location="New" municipality="Doron" pageId="1" pageNumber="446" specimenCount="3" specimenCount-female="3" typeStatus="paratype">
<emphasis id="B95EEAEEFFB9AC63136A1E30FA92FED6" bold="true" box="[268,388,357,383]" pageId="1" pageNumber="446">
<typeStatus id="5491885EFFB9AC63136A1E30FA92FED6" box="[268,388,357,383]" pageId="1" pageNumber="446" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="9D2CFD75FFB9AC6313F41E30F917FED7" box="[402,513,356,383]" pageId="1" pageNumber="446" type="female">3 females</specimenCount>
, same data as for the holotype (
<collectionCode id="ED3BAE39FFB9AC6311051E30F8ABFED6" box="[867,957,357,383]" pageId="1" pageNumber="446">BMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3306577FFB9AC6312A11EDDFF07F97F" pageId="1" pageNumber="446" type="description">
<paragraph id="8B9536FCFFB9AC6312A11EDDFF96F98A" blockId="1.[151,1437,320,2039]" pageId="1" pageNumber="446">
<emphasis id="B95EEAEEFFB9AC6312A11EDDF90DFE0A" bold="true" box="[199,539,392,419]" pageId="1" pageNumber="446">
Description of the
<typeStatus id="5491885EFFB9AC6313CB1EDCF900FE0A" box="[429,534,393,419]" pageId="1" pageNumber="446" type="holotype">holotype</typeStatus>
.
</emphasis>
Total body length
<quantity id="4CD29B19FFB9AC63109D1EDDF840FE0A" box="[763,854,392,419]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="1" pageNumber="446" unit="mm" value="8.5">8.5 mm</quantity>
, greatest width
<quantity id="4CD29B19FFB9AC6316771EDCFF7BFE0A" box="[1041,1133,393,419]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="1" pageNumber="446" unit="mm" value="4.2">4.2 mm</quantity>
. Dark reddish-brown with legs more reddish; antennae and palpi reddish; setation pale. Body (including dorsal surface of head but not labroclypeus) is almost completely covered with microscopical velvety pubescence. Antennae 9-segmented, club 3-lamellate. Labroclypeus broadly concave anteriorly, its lateral margins strongly bisinuate in both dorsal and lateral views; anterolateral angles small and pointed, triangular, weakly reflexed; canthus angulate. Dorsal surface of labroclypeus coarsely punctured, punctures deep, annular, with dense microscopical velvety pubescence; intervening spaces variable, smooth and glossy. Canthus is tightly covered with dense microscopical pubescence, with few moderately long erect setae. Underside of labroclypeus rather densely setose. Frons flat; punctures similar to those on labroclypeus, but much densely arranged; intervening spaces very narrow; setae erect to semi-erect, short, commensurable with diameter of punctures. Pronotum transverse with anterior margin concave and bisinuate; anterolateral angles acute; sides of pronotum broadly rounded in anterior two-thirds and deeply concave in posterior third; basal margin strongly convex; posterolateral angles almost straight. Lateral margins of pronotum crenulate, with about 20 poorly expressed crenulae, better developed along middle widest part of pronotum; a very short seta (often worn) present between every two crenulae. Pronotal punctures large, annular to hexagonal; intervening spaces very narrow; setation (largely worn) similar to that on head and elytra. Hypomeron separated from prosternum by moderately high emarginate carina bearing long setae; antennal pockets rather deep; punctures large, annular; intervening spaces generally smaller than punctures; setae moderately long in anterior half, becoming short, not surpassing diameter of puncture in posterior half. Scutellar shield apically rounded, with dense and rather coarse annular puncturation; setae probably worn. Elytra weakly widened posteriad, broadly rounded apically, with four weak longitudinal carinae splitting to a series of sparsely sitting granules (two rows between sutural margin and humeral umbo, third row running from humerus toward apical umbo, fourth one extending forward from apical umbo along lateral margin of elytron); sutural joint not elevated. Punctures of elytral disc narrow, incision-shaped; intervening spaces generally smaller than length of punctures; setae adpressed to semi-erect, not or hardly surpassing length of corresponding punctures. Pygidium flat, with shallow annular punctures; intervening spaces very narrow; setae semi-erect, similar to those on elytra. Thorax and abdominal ventrites with small and shallow annular punctures; intervening spaces smaller than diameter of punctures; setae adpressed, generally exceeding diameter of punctures in length on thorax and not surpassing it on abdominal ventrites. Ventral profile of abdomen slightly convex. Protibia (
<figureCitation id="13112A79FFB9AC6313041A01FAAEFAC6" box="[354,440,1364,1391]" captionStart="FIGURE 1" captionStartId="2.[152,255,1102,1126]" captionTargetBox="[157,1427,189,1072]" captionTargetId="figure-27@2.[151,1436,181,1078]" captionTargetPageId="2" captionText="FIGURE 1. Maechidius addarcis Prokofiev, new species. (A) dorsal view, holotype; (B) protibia, holotype; (C) aedeagus, lateral view, holotype; (D) parameres, dorsal view, holotype; (E) spiculum gastrale, dorsal view, holotype; (F) ditto, lateral view; (G) protibia, female paratype. Scale bars: (A, B, G) 1.0 mm; (CF) 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7318493" httpUri="https://zenodo.org/record/7318493/files/figure.png" pageId="1" pageNumber="446">Fig. 1B</figureCitation>
) weakly widened distally, with two longitudinal carinae dorsally and two strong apical teeth directed outward; basal tooth weak. Protibial spur short, straight, somewhat inclined ventrally. Metatibial spurs short and straight, pointed; lower spur thicker and twice longer than upper one. Tarsomeres not elongated, setae not forming brush-like patches along their ventral margin. Tarsal claws with pulvilli. Aedeagus and spiculum gastrale, as on
<figureCitation id="13112A79FFB9AC6312BC1AB1FA4DFA56" box="[218,347,1508,1535]" captionStart="FIGURE 1" captionStartId="2.[152,255,1102,1126]" captionTargetBox="[157,1427,189,1072]" captionTargetId="figure-27@2.[151,1436,181,1078]" captionTargetPageId="2" captionText="FIGURE 1. Maechidius addarcis Prokofiev, new species. (A) dorsal view, holotype; (B) protibia, holotype; (C) aedeagus, lateral view, holotype; (D) parameres, dorsal view, holotype; (E) spiculum gastrale, dorsal view, holotype; (F) ditto, lateral view; (G) protibia, female paratype. Scale bars: (A, B, G) 1.0 mm; (CF) 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7318493" httpUri="https://zenodo.org/record/7318493/files/figure.png" pageId="1" pageNumber="446">Figs. 1CF</figureCitation>
; parameres symmetrical, with tips rectangular, crossing each other in default position. Spiculum gastrale flat along most of its length, only the base of its shaft strongly reflexed (
<figureCitation id="13112A79FFB9AC631678195DFF64F98A" box="[1054,1138,1544,1571]" captionStart="FIGURE 1" captionStartId="2.[152,255,1102,1126]" captionTargetBox="[157,1427,189,1072]" captionTargetId="figure-27@2.[151,1436,181,1078]" captionTargetPageId="2" captionText="FIGURE 1. Maechidius addarcis Prokofiev, new species. (A) dorsal view, holotype; (B) protibia, holotype; (C) aedeagus, lateral view, holotype; (D) parameres, dorsal view, holotype; (E) spiculum gastrale, dorsal view, holotype; (F) ditto, lateral view; (G) protibia, female paratype. Scale bars: (A, B, G) 1.0 mm; (CF) 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7318493" httpUri="https://zenodo.org/record/7318493/files/figure.png" pageId="1" pageNumber="446">Fig. 1F</figureCitation>
).
</paragraph>
<paragraph id="8B9536FCFFB9AC6312A11978FF07F97F" blockId="1.[151,1437,320,2039]" pageId="1" pageNumber="446">
<emphasis id="B95EEAEEFFB9AC6312A11978FA26F9EE" bold="true" box="[199,304,1581,1607]" pageId="1" pageNumber="446">Females.</emphasis>
Total body length 8.0
<quantity id="4CD29B19FFB9AC63104B1979F994F9EE" box="[557,642,1580,1607]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="1" pageNumber="446" unit="mm" value="9.5">9.5 mm</quantity>
, greatest width 4.0
<quantity id="4CD29B19FFB9AC6311381979F8A2F9EE" box="[862,948,1580,1607]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="1" pageNumber="446" unit="mm" value="4.5">4.5 mm</quantity>
. Secondary sexual differences not expressed except the comparatively shorter antennal lamellae, more pronounced basal tooth of protibia (
<figureCitation id="13112A79FFB9AC6316DA1905FE05F9C2" box="[1212,1299,1616,1643]" captionStart="FIGURE 1" captionStartId="2.[152,255,1102,1126]" captionTargetBox="[157,1427,189,1072]" captionTargetId="figure-27@2.[151,1436,181,1078]" captionTargetPageId="2" captionText="FIGURE 1. Maechidius addarcis Prokofiev, new species. (A) dorsal view, holotype; (B) protibia, holotype; (C) aedeagus, lateral view, holotype; (D) parameres, dorsal view, holotype; (E) spiculum gastrale, dorsal view, holotype; (F) ditto, lateral view; (G) protibia, female paratype. Scale bars: (A, B, G) 1.0 mm; (CF) 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7318493" httpUri="https://zenodo.org/record/7318493/files/figure.png" pageId="1" pageNumber="446">Fig. 1G</figureCitation>
) and longer spurs on all tibiae. Metatibial spurs only slightly different in length, weakly clavate at tips. Sides of pronotum before posterolateral angles variably excavated. Longitudinal carinae on elytra variably developed, from continuous along almost whole length of elytra to expressed at their base and near umbones only.
</paragraph>
</subSubSection>
<subSubSection id="C3306577FFB9AC6312A119B5F89DF807" pageId="1" pageNumber="446" type="diagnosis">
<paragraph id="8B9536FCFFB9AC6312A119B5F89DF807" blockId="1.[151,1437,320,2039]" pageId="1" pageNumber="446">
<emphasis id="B95EEAEEFFB9AC6312A119B5FADBF953" bold="true" box="[199,461,1760,1786]" pageId="1" pageNumber="446">Differential diagnosis.</emphasis>
This species is keyed to the couplet
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<bibRefCitation id="EFBB4B0DFFB9AC6311FB19B5FF28F952" author="Telnov, D." box="[925,1086,1760,1787]" pageId="1" pageNumber="446" pagination="1 - 210" refId="ref11840" refString="Telnov, D. (2020) A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the IndoAustralian transition zone, and the first record of the tribe west of Wallace's Line. European Journal of Taxonomy, 721, 1 - 210. https: // doi. org / 10.5852 / ejt. 2020.721.1127" type="journal article" year="2020">Telnov (2020)</bibRefCitation>
, but can be easily distinguished from both
<taxonomicName id="4C2A4D7FFFB9AC6313731850F94FF8B6" authority="Heller, 1895" authorityName="Heller" authorityYear="1895" box="[277,601,1796,1823]" class="Insecta" family="Melolonthidae" genus="Maechidius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="446" phylum="Arthropoda" rank="species" species="subcostatus">
<emphasis id="B95EEAEEFFB9AC6313731850FAD6F8B7" box="[277,448,1796,1822]" italics="true" pageId="1" pageNumber="446">M. subcostatus</emphasis>
Heller, 1895
</taxonomicName>
and
<taxonomicName id="4C2A4D7FFFB9AC6310F11850F8BEF8B6" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[663,936,1796,1823]" class="Insecta" family="Melolonthidae" genus="Maechidius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="446" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="B95EEAEEFFB9AC6310F11850F81FF8B7" box="[663,777,1796,1822]" italics="true" pageId="1" pageNumber="446">M. similis</emphasis>
<bibRefCitation id="EFBB4B0DFFB9AC6311751851F8BEF8B6" author="Telnov, D." box="[787,936,1796,1823]" pageId="1" pageNumber="446" pagination="1 - 210" refId="ref11840" refString="Telnov, D. (2020) A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the IndoAustralian transition zone, and the first record of the tribe west of Wallace's Line. European Journal of Taxonomy, 721, 1 - 210. https: // doi. org / 10.5852 / ejt. 2020.721.1127" type="journal article" year="2020">Telnov, 2020</bibRefCitation>
</taxonomicName>
by dense coating of microscopical velvety pubescence and in the unique shape of parameres with tips crossing each other in default position. It is superficially somewhat similar to the members of the former genus
<taxonomicName id="4C2A4D7FFFB9AC6311641819F8ACF8CF" authorityName="Frey" authorityYear="1969" box="[770,954,1868,1894]" class="Insecta" family="Melolonthidae" genus="Paramaechidius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="446" phylum="Arthropoda" rank="genus">
<emphasis id="B95EEAEEFFB9AC6311641819F8ACF8CF" box="[770,954,1868,1894]" italics="true" pageId="1" pageNumber="446">Paramaechidius</emphasis>
</taxonomicName>
or
<taxonomicName id="4C2A4D7FFFB9AC6311871818FF78F8CF" authorityName="Heller" authorityYear="1910" box="[993,1134,1868,1894]" class="Insecta" family="Melolonthidae" genus="Maechidius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="446" phylum="Arthropoda" rank="species" species="pauxillus">
<emphasis id="B95EEAEEFFB9AC6311871818FF78F8CF" box="[993,1134,1868,1894]" italics="true" pageId="1" pageNumber="446">M. pauxillus</emphasis>
</taxonomicName>
group as of
<bibRefCitation id="EFBB4B0DFFB9AC63169A1819FE8AF8CE" author="Telnov, D." box="[1276,1436,1868,1895]" pageId="1" pageNumber="446" pagination="1 - 210" refId="ref11840" refString="Telnov, D. (2020) A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the IndoAustralian transition zone, and the first record of the tribe west of Wallace's Line. European Journal of Taxonomy, 721, 1 - 210. https: // doi. org / 10.5852 / ejt. 2020.721.1127" type="journal article" year="2020">Telnov (2020)</bibRefCitation>
but can be easily differentiated from them by absence of the clavate setae on the dorsum, presence of the tarsal pulvilli in both sexes, the dorsally flat frons and different aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="C3306577FFB9AC6312A118ECFA4CF85E" pageId="1" pageNumber="446" type="etymology">
<paragraph id="8B9536FCFFB9AC6312A118ECFA4CF85E" blockId="1.[151,1437,320,2039]" pageId="1" pageNumber="446">
<emphasis id="B95EEAEEFFB9AC6312A118ECFA58F87A" bold="true" box="[199,334,1977,2003]" pageId="1" pageNumber="446">Etymology.</emphasis>
The species epithet is created from the Latin words ad (to) and arx (tip), due to the conjoining tips of the parameres.
</paragraph>
</subSubSection>
</treatment>
</document>