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<document id="C4D9415F238F2317148D418CAAD5DEB3" ID-DOI="10.3897/zoologia.37.e37384" ID-ISSN="1984-4689" ID-Zenodo-Dep="13174110" ID-ZooBank="95E87889-A1B4-47D0-BCF4-3E48140E7467" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722587265700" checkinUser="felipe" docAuthor="Pantoja, Paulo, Drago-Bisneto, Marcos &amp; Saturnino, Regiane" docDate="2020" docId="038387E7FFCE3B44FEF1554F25BFFDBE" docLanguage="en" docName="Zoologia.37.e37384.pdf" docOrigin="Zoologia (e 37384) 37" docSource="http://dx.doi.org/10.3897/zoologia.37.e37384" docStyle="DocumentStyle:9D87517C92C8E430512793D922E147D4.1:Zoologia.2017-2020.journal_article" docStyleId="9D87517C92C8E430512793D922E147D4" docStyleName="Zoologia.2017-2020.journal_article" docStyleVersion="1" docTitle="Berlandiella zabele Pantoja, Drago-Bisneto &amp; Saturnino, 2020, sp. nov." docType="treatment" docUuid="A284875E-3883-478C-9361-B2D39576009F" docUuidSource="ZooBank" docVersion="2" lastPageNumber="6" masterDocId="FFBAFF9FFFCC3B41FFC25049216FF873" masterDocTitle="Updates on Berlandiella (Araneae: Philodromidae): a new species, description of the male of B. querencia and new diagnosis for the genus" masterLastPageNumber="13" masterPageNumber="1" pageNumber="3" updateTime="1722620052173" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="C3479CD3E89F4081837C88D735F96DED">Updates on Berlandiella (Araneae: Philodromidae): a new species, description of the male of B. querencia and new diagnosis for the genus</mods:title>
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<mods:affiliation id="931DFB4CE5374B9118275C2A3F42E1F0">Laboratório de Aracnologia, Coordenação de Zoologia, Museu Paraense Emílio Goeldi. Avenida Perimetral 1901, &amp; Programa de Pós-Graduação em Zoologia, Museu Paraense Emílio Goeldi, Universidade Federal do Pará. Rua</mods:affiliation>
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<mods:namePart id="369C23898D1A920BBD5D4B271602951B">Drago-Bisneto, Marcos</mods:namePart>
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<taxonomicName id="4C2A4D72FFCE3B43FEF1554F2360FD52" ID-CoL="99264" authorityName="Pantoja &amp; Drago-Bisneto &amp; Saturnino" authorityYear="2020" box="[307,527,1286,1313]" class="Arachnida" family="Philodromidae" genus="Berlandiella" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="zabele" status="sp. nov.">
<emphasis id="B95EEAE3FFCE3B43FEF1554F2360FD52" box="[307,527,1286,1313]" italics="true" pageId="2" pageNumber="3">Berlandiella zabele</emphasis>
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http://zoobank.org/
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<figureCitation id="13112A74FFCE3B43FEB3550620A1FD10" box="[369,462,1359,1379]" captionStart-0="Figures 25" captionStart-1="Figures 610" captionStart-2="Figures 1116" captionStart-3="Figures 1722" captionStart-4="Figures 2326" captionStartId-0="2.[159,226,1184,1204]" captionStartId-1="3.[150,216,1093,1113]" captionStartId-2="4.[159,226,1056,1076]" captionStartId-3="5.[151,218,1059,1079]" captionStartId-4="6.[159,226,1185,1205]" captionTargetBox-0="[234,1340,197,1169]" captionTargetBox-1="[302,1281,202,1077]" captionTargetBox-2="[159,1436,197,1041]" captionTargetBox-3="[151,1428,197,1046]" captionTargetBox-4="[310,1286,197,1171]" captionTargetId-0="figure-1@2.[234,1343,197,1169]" captionTargetId-1="figure-0@3.[299,1282,197,1078]" captionTargetId-2="figure-0@4.[159,1436,197,1041]" captionTargetId-3="figure-0@5.[151,1428,197,1046]" captionTargetId-4="figure-0@6.[310,1286,197,1171]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionText-0="Figures 25. Berlandiella zabele sp. nov.: (2) female holotype habitus (MPEG.ARA 34482); (3) male paratype habitus (MPEG.ARA 34480); (4) epigynum, ventral; (5) epigynum, dorsal clarified. Scale bars: 23: 1 mm, 45: 0.2 mm." captionText-1="Figures 610. Genitalia of Berlandiella zabele sp. nov.: (6) epigynum ventral; (7) epigynum dorsal; (8) palp, ventral; (9) palp, retrolateral; (10) palp, dorsal. CD: copulatory ducts, CO: copulatory opening, Co: conductor, E: embolus, EG: epigynal groove, FD: fertilization duct, GH: glandular head, CP: cymbial process, RTA: retrolateral tibial apophysis, S: spermatheca, VTA: ventral tibial apophysis. Scale bars: 0.3 mm." captionText-2="Figures 1116. Palp of Berlandiella zabele sp. nov.: (11) ventral; (12) detail of apical portion of tegulum; (13) dorsal (arrow indicates scopula in apex of cymbium); (14) scopula in cymbium (indicated by arrow); (15) detail of CP (indicated by arrow) and RTA; (16) detail showing excavation in the RTA. CP: cymbial process, RTA: retrolateral tibial apophysis. Scale bars: 11 = 200 µm, 12, 15, 16 = 50 µm, 13 = 150 µm, 14 = 20 µm." captionText-3="Figures 1722. Carapace of Berlandiella zabele sp. nov. male: (17) dorsal; (18) anterodorsal; (19) detail of tegument with many sockets of setae (indicated by arrows); (20) strong setae in the cephalic region; (21) hairs; (22) scales in the lateral of carapace. Scale bars: 7, 20 = 30 µm, 18 = 300 µm, 19, 22 = 50 µm, 20 = 30 µm, 21 = 10µm." captionText-4="Figures 2326. Leg I of Berlandiella zabele sp. nov. male: (23) tarsus and apex of metatarsus with scopula (indicated by arrows); (24) different types of seta in the metatarsus (smaller rectangle show tenent barbs of scopula); (25) claw and claw tufts; (26) detail of tenet barbs of claw tufts. Scale bars: 23 = 150 µm, 24, 25 = 20 µm, 26 = 5 µm." pageId="2" pageNumber="3">Figs 226</figureCitation>
,
<figureCitation id="13112A74FFCE3B43FE1555062378FD10" box="[471,535,1359,1379]" captionStart="Figures 4043" captionStartId="9.[150,217,1192,1212]" captionTargetBox="[288,1271,197,1179]" captionTargetId="figure-383@9.[288,1271,197,1179]" captionTargetPageId="9" captionText="Figures 4043. Setae: (40) setae on the side tibia of B. querencia male; (41) pair of macrosetae in the venter of tibia I of B. zabele male; (42) trichobothrium on the dorsum of tarsus I of B. zabele male; (43) strong bristles on the side of the metatarsus of B. querencia female. Scale bars: 40, 43 = 50 µm, 41 = 100 µm, 42 = 20 µm." pageId="2" pageNumber="3">41, 43</figureCitation>
,
<figureCitation id="13112A74FFCE3B43FDE255062357FD10" box="[544,568,1359,1379]" captionStart="Figure 44" captionStartId="10.[159,217,950,970]" captionTargetBox="[275,1320,199,937]" captionTargetId="figure-546@10.[274,1322,198,935]" captionTargetPageId="10" captionText="Figure 44. Distribution map of Berlandiella zabele sp. nov. and B. querencia." pageId="2" pageNumber="3">44</figureCitation>
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Type-material.
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from
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<date id="FF941031FFCE3B43FEFF55F020A5FDB8" box="[317,458,1464,1484]" pageId="2" pageNumber="3" value="2005-06-24">
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leg. (
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.
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).
<typeStatus id="54918853FFCE3B43FF35559E2032FD99" box="[247,349,1495,1514]" pageId="2" pageNumber="3" type="paratype">Paratypes</typeStatus>
.
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,
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from
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,
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<date id="FF941031FFCE3B43FD0755BD21BDFE56" pageId="2" pageNumber="3" value="2007-01-29">
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et al. leg. (
<collectionCode id="ED3BAE34FFCE3B43FE03565B236FFE56" box="[449,512,1554,1573]" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="2" pageNumber="3">MPEG</collectionCode>
.
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);
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, same collection data as for preceding,
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<collectingDate id="EFD0E9D9FFCE3B43FD3056792194FE12" pageId="2" pageNumber="3" value="2007-01-26">26 Jan. 2007</collectingDate>
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,
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</materialsCitation>
<materialsCitation id="3B423CACFFCE3B43FEE25607231AFE0C" collectingDate="2007-01-24" collectingDateMax="2007-01-29" collectingDateMin="2005-06-24" collectionCode="LS" collectorName="Carvalho" country="Brazil" county="Parque Nacional de Sete Cidades" latitude="-4.0944166" location="Carvalho" longLatPrecision="1" longitude="-41.731472" municipality="Brasileira" pageId="2" pageNumber="3" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<collectorName id="26DF5327FFCE3B43FEE256072014FE12" box="[288,379,1614,1633]" pageId="2" pageNumber="3">Carvalho</collectorName>
et al. leg. (CHNUFPI 3398);
<specimenCount id="9D2CFD78FFCE3B43FD49560723A0FE12" box="[651,719,1614,1633]" count="1" pageId="2" pageNumber="3" type="male">1 male</specimenCount>
, same collection data as for preceding,
<date id="FF941031FFCE3B43FE185625233AFE0C" box="[474,597,1644,1663]" pageId="2" pageNumber="3" value="2007-01-24">
<collectingDate id="EFD0E9D9FFCE3B43FE185625233AFE0C" box="[474,597,1644,1663]" pageId="2" pageNumber="3" value="2007-01-24">24 Jan. 2007</collectingDate>
</date>
,
<collectionCode id="ED3BAE34FFCE3B43FD9C5625231AFE0C" box="[606,629,1644,1663]" country="United Kingdom" name="Linnean Society of London" pageId="2" pageNumber="3">LS</collectionCode>
</materialsCitation>
<materialsCitation id="3B423CACFFCE3B43FDB85625226DFEC9" collectingDate="2007-01-22" collectingDateMax="2007-01-29" collectingDateMin="2005-06-24" collectionCode="MPEG, LS" collectorName="Carvalho" country="Brazil" county="Parque Nacional de Sete Cidades" latitude="-4.0944166" location="Carvalho" longLatPrecision="1" longitude="-41.731472" municipality="Brasileira" pageId="2" pageNumber="3" specimenCode="ARA 15585, ARA 15579" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<location id="8EF5602AFFCE3B43FDB8562523BAFE0C" LSID="urn:lsid:plazi:treatment:038387E7FFCE3B44FEF1554F25BFFDBE:8EF5602AFFCE3B43FDB8562523BAFE0C" box="[634,725,1644,1663]" name="Carvalho" pageId="2" pageNumber="3">Carvalho</location>
et al. leg. (
<collectionCode id="ED3BAE34FFCE3B43FF0C56C32062FEEE" box="[206,269,1674,1693]" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="2" pageNumber="3">MPEG</collectionCode>
.
<specimenCode id="DB8C9E8AFFCE3B43FECD56C320E9FEEF" box="[271,390,1673,1693]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="2" pageNumber="3">ARA 15585</specimenCode>
);
<specimenCount id="9D2CFD78FFCE3B43FE5A56C3209EFEEE" box="[408,497,1674,1693]" count="1" pageId="2" pageNumber="3" type="female">1 female</specimenCount>
, same collection data as for preceding,
<date id="FF941031FFCE3B43FECC56E120E3FEC8" box="[270,396,1704,1723]" pageId="2" pageNumber="3" value="2007-01-22">
<collectingDate id="EFD0E9D9FFCE3B43FECC56E120E3FEC8" box="[270,396,1704,1723]" pageId="2" pageNumber="3" value="2007-01-22">22 Jan. 2007</collectingDate>
</date>
,
<collectionCode id="ED3BAE34FFCE3B43FE5556E120C1FEC8" box="[407,430,1704,1723]" country="United Kingdom" name="Linnean Society of London" pageId="2" pageNumber="3">LS</collectionCode>
<collectorName id="26DF5327FFCE3B43FE7656EE2360FEC9" box="[436,527,1703,1722]" pageId="2" pageNumber="3">Carvalho</collectorName>
leg. (
<collectionCode id="ED3BAE34FFCE3B43FD8656E123ECFEC8" box="[580,643,1704,1723]" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="2" pageNumber="3">MPEG</collectionCode>
.
<specimenCode id="DB8C9E8AFFCE3B43FD4756E12394FEC9" box="[645,763,1703,1723]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="2" pageNumber="3">ARA 15579</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B9536F1FFCE3B43FF19568C2553FD99" blockId="2.[157,780,1404,1783]" lastBlockId="2.[815,1438,1285,1783]" pageId="2" pageNumber="3">
Diagnosis. The female of
<taxonomicName id="4C2A4D72FFCE3B43FE27568C23F7FEAB" authorityName="Pantoja &amp; Drago-Bisneto &amp; Saturnino" authorityYear="2020" box="[485,664,1733,1752]" class="Arachnida" family="Philodromidae" genus="Berlandiella" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="zabele" status="sp. nov.">
<emphasis id="B95EEAE3FFCE3B43FE27568C23F7FEAB" box="[485,664,1733,1752]" italics="true" pageId="2" pageNumber="3">Berlandiella zabele</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26D5798FFCE3B43FD63568F239CFEAA" box="[673,755,1734,1753]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
is similar to
<taxonomicName id="4C2A4D72FFCE3B43FEC556AD200EFE85" authorityName="Mello-Leitao" authorityYear="1929" box="[263,353,1764,1782]" class="Arachnida" family="Philodromidae" genus="Berlandiella" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="magna">
<emphasis id="B95EEAE3FFCE3B43FEC556AD200EFE85" box="[263,353,1764,1782]" italics="true" pageId="2" pageNumber="3">B. magna</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2A4D72FFCE3B43FE5756AD2368FE85" authorityName="Lise &amp; Silva" authorityYear="2011" box="[405,519,1764,1782]" class="Arachnida" family="Philodromidae" genus="Berlandiella" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="querencia">
<emphasis id="B95EEAE3FFCE3B43FE5756AD2368FE85" box="[405,519,1764,1782]" italics="true" pageId="2" pageNumber="3">B. querencia</emphasis>
</taxonomicName>
by the long and ellipsoid spermathecae (
<figureCitation id="13112A74FFCE3B43FC0D554C2546FD6A" box="[975,1065,1285,1305]" captionStart-0="Figures 25" captionStart-1="Figures 610" captionStartId-0="2.[159,226,1184,1204]" captionStartId-1="3.[150,216,1093,1113]" captionTargetBox-0="[234,1340,197,1169]" captionTargetBox-1="[302,1281,202,1077]" captionTargetId-0="figure-1@2.[234,1343,197,1169]" captionTargetId-1="figure-0@3.[299,1282,197,1078]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="Figures 25. Berlandiella zabele sp. nov.: (2) female holotype habitus (MPEG.ARA 34482); (3) male paratype habitus (MPEG.ARA 34480); (4) epigynum, ventral; (5) epigynum, dorsal clarified. Scale bars: 23: 1 mm, 45: 0.2 mm." captionText-1="Figures 610. Genitalia of Berlandiella zabele sp. nov.: (6) epigynum ventral; (7) epigynum dorsal; (8) palp, ventral; (9) palp, retrolateral; (10) palp, dorsal. CD: copulatory ducts, CO: copulatory opening, Co: conductor, E: embolus, EG: epigynal groove, FD: fertilization duct, GH: glandular head, CP: cymbial process, RTA: retrolateral tibial apophysis, S: spermatheca, VTA: ventral tibial apophysis. Scale bars: 0.3 mm." pageId="2" pageNumber="3">Figs 47</figureCitation>
), but are distinguished from those species by having its copulation opening more widely separated from each other and located on the posterior edge of the epigynal groove (
<figureCitation id="13112A74FFCE3B43FC1C55162575FD00" box="[990,1050,1375,1395]" captionStart="Figures 25" captionStartId="2.[159,226,1184,1204]" captionTargetBox="[234,1340,197,1169]" captionTargetId="figure-1@2.[234,1343,197,1169]" captionTargetPageId="2" captionText="Figures 25. Berlandiella zabele sp. nov.: (2) female holotype habitus (MPEG.ARA 34482); (3) male paratype habitus (MPEG.ARA 34480); (4) epigynum, ventral; (5) epigynum, dorsal clarified. Scale bars: 23: 1 mm, 45: 0.2 mm." pageId="2" pageNumber="3">Figs 4</figureCitation>
,
<figureCitation id="13112A74FFCE3B43FBE75516255CFD01" box="[1061,1075,1375,1394]" captionStart="Figures 610" captionStartId="3.[150,216,1093,1113]" captionTargetBox="[302,1281,202,1077]" captionTargetId="figure-0@3.[299,1282,197,1078]" captionTargetPageId="3" captionText="Figures 610. Genitalia of Berlandiella zabele sp. nov.: (6) epigynum ventral; (7) epigynum dorsal; (8) palp, ventral; (9) palp, retrolateral; (10) palp, dorsal. CD: copulatory ducts, CO: copulatory opening, Co: conductor, E: embolus, EG: epigynal groove, FD: fertilization duct, GH: glandular head, CP: cymbial process, RTA: retrolateral tibial apophysis, S: spermatheca, VTA: ventral tibial apophysis. Scale bars: 0.3 mm." pageId="2" pageNumber="3">6</figureCitation>
). The male resembles
<taxonomicName id="4C2A4D72FFCE3B43FAD2552924EEFD01" authorityName="Lise &amp; Silva" authorityYear="2011" box="[1296,1409,1376,1394]" class="Arachnida" family="Philodromidae" genus="Berlandiella" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="querencia">
<emphasis id="B95EEAE3FFCE3B43FAD2552924EEFD01" box="[1296,1409,1376,1394]" italics="true" pageId="2" pageNumber="3">B. querencia</emphasis>
</taxonomicName>
in possessing a cymbial process (CP), but differs by the position of the CP, closest to the cymbium (
<figureCitation id="13112A74FFCE3B43FBB455D225B9FDDD" box="[1142,1238,1435,1455]" captionStart="Figures 610" captionStartId="3.[150,216,1093,1113]" captionTargetBox="[302,1281,202,1077]" captionTargetId="figure-0@3.[299,1282,197,1078]" captionTargetPageId="3" captionText="Figures 610. Genitalia of Berlandiella zabele sp. nov.: (6) epigynum ventral; (7) epigynum dorsal; (8) palp, ventral; (9) palp, retrolateral; (10) palp, dorsal. CD: copulatory ducts, CO: copulatory opening, Co: conductor, E: embolus, EG: epigynal groove, FD: fertilization duct, GH: glandular head, CP: cymbial process, RTA: retrolateral tibial apophysis, S: spermatheca, VTA: ventral tibial apophysis. Scale bars: 0.3 mm." pageId="2" pageNumber="3">Figs 810</figureCitation>
,
<figureCitation id="13112A74FFCE3B43FB2055D22427FDDD" box="[1250,1352,1435,1454]" captionStart="Figures 1116" captionStartId="4.[159,226,1056,1076]" captionTargetBox="[159,1436,197,1041]" captionTargetId="figure-0@4.[159,1436,197,1041]" captionTargetPageId="4" captionText="Figures 1116. Palp of Berlandiella zabele sp. nov.: (11) ventral; (12) detail of apical portion of tegulum; (13) dorsal (arrow indicates scopula in apex of cymbium); (14) scopula in cymbium (indicated by arrow); (15) detail of CP (indicated by arrow) and RTA; (16) detail showing excavation in the RTA. CP: cymbial process, RTA: retrolateral tibial apophysis. Scale bars: 11 = 200 µm, 12, 15, 16 = 50 µm, 13 = 150 µm, 14 = 20 µm." pageId="2" pageNumber="3">13, 15, 16</figureCitation>
), by the long retrolateral tibial apophysis (RTA), excavated in the distal portion (
<figureCitation id="13112A74FFCE3B43FC4A559E255FFD99" box="[904,1072,1495,1515]" captionStart-0="Figures 610" captionStart-1="Figures 1116" captionStartId-0="3.[150,216,1093,1113]" captionStartId-1="4.[159,226,1056,1076]" captionTargetBox-0="[302,1281,202,1077]" captionTargetBox-1="[159,1436,197,1041]" captionTargetId-0="figure-0@3.[299,1282,197,1078]" captionTargetId-1="figure-0@4.[159,1436,197,1041]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="Figures 610. Genitalia of Berlandiella zabele sp. nov.: (6) epigynum ventral; (7) epigynum dorsal; (8) palp, ventral; (9) palp, retrolateral; (10) palp, dorsal. CD: copulatory ducts, CO: copulatory opening, Co: conductor, E: embolus, EG: epigynal groove, FD: fertilization duct, GH: glandular head, CP: cymbial process, RTA: retrolateral tibial apophysis, S: spermatheca, VTA: ventral tibial apophysis. Scale bars: 0.3 mm." captionText-1="Figures 1116. Palp of Berlandiella zabele sp. nov.: (11) ventral; (12) detail of apical portion of tegulum; (13) dorsal (arrow indicates scopula in apex of cymbium); (14) scopula in cymbium (indicated by arrow); (15) detail of CP (indicated by arrow) and RTA; (16) detail showing excavation in the RTA. CP: cymbial process, RTA: retrolateral tibial apophysis. Scale bars: 11 = 200 µm, 12, 15, 16 = 50 µm, 13 = 150 µm, 14 = 20 µm." pageId="2" pageNumber="3">Figs 811, 15, 16</figureCitation>
).
</paragraph>
<paragraph id="8B9536F1FFCE3B42FCAF55BC24F8FD9B" blockId="2.[815,1438,1285,1783]" lastBlockId="3.[807,1431,1254,1781]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
Description. Female (
<typeStatus id="54918853FFCE3B43FB9055BD25DEFE7B" box="[1106,1201,1524,1544]" pageId="2" pageNumber="3" type="holotype">holotype</typeStatus>
MPEG.ARA 34482). Carapace slightly longer than wide, wider at the level of legs III, narrowed anteriorly, with many sockets of bristles; median region yellow with a brown central spot and wide dark brown bands on each side, formed by a tangle of dark spots. Clypeus vertical with many sockets of bristles, with a median brown spot and irregular spots on lateral. Chelicerae yellow with two teeth on the promargin of the fang furrow, with a large dark brown spot on the anterior face. Labium orange with rounded apex, reaching almost to the middle of the endites. Endites pale yellow, excavated diagonally. Sternum pale yellow. Legs covered by various types of setae and with many robust macrosetae on tibiae and metatarsi. Left and right tibiae I with 0220 ventral macrosetae, left metatarsus I with 220 and right metatarsus I with 221 ventral macrosetae. Left and right tibia II with 0220 ventral macrosetae and right metatarsus II with 221 ventral macrosetae. Tarsi and metatarsi with scopula. Long and thin trichobothria in tarsi and metatarsi, with distal and proximal plate limit well differentiated; the distal margin of the trichobothrial hood is well defined; hood entire, differentiated with transverse ridges; alveolus notched; distal plate smooth, slightly elevated. Femura yellow with brown spots, more abundant in prolateral sides of femora I-III, and a small white spot at the apex; patellae I-III yellow with brown spots; patella IV yellow; tibiae and metatarsi I-III yellow, shaded with brown, but tibia and metatarsus III darker; tibia IV yellow with a basal dark spot on prolateral side; tarsi I-IV yellow, with a dark ring at their bases. Tarsal claw pectinate and conspicuous with claw apex tooth curved and seven short and robust teeth. Palp yellow with brown spots. Abdomen yellow, longer than wide, densely covered by setae sockets, the dorsum covered by a wide brown spot with two longitudinal yellow bands around the heart mark; laterals yellow shaded with brown and the venter yellow stippled with brown. Triangular epigynal groove, shallow, demarcated by sclerotized edges, small and widely spaced copulatory openings, elliptical and elongated spermathecae, thickly covered with punctuations.
</paragraph>
<caption id="DF556679FFCF3B42FF54540C24F9FCE6" pageId="3" pageNumber="4" startId="3.[150,216,1093,1113]" targetBox="[302,1281,202,1077]" targetPageId="3" targetType="figure">
<paragraph id="8B9536F1FFCF3B42FF54540C24F9FCE6" blockId="3.[150,1430,1093,1173]" pageId="3" pageNumber="4">
Figures 610. Genitalia of
<taxonomicName id="4C2A4D72FFCF3B42FE41540C2349FC2A" authorityName="Pantoja &amp; Drago-Bisneto &amp; Saturnino" authorityYear="2020" box="[387,550,1093,1113]" class="Arachnida" family="Philodromidae" genus="Berlandiella" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="zabele" status="sp. nov.">
<emphasis id="B95EEAE3FFCF3B42FE41540C2349FC2A" box="[387,550,1093,1113]" italics="true" pageId="3" pageNumber="4">Berlandiella zabele</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26D5798FFCF3B42FDE9540C2319FC2A" box="[555,630,1093,1113]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
: (6) epigynum ventral; (7) epigynum dorsal; (8) palp, ventral; (9) palp, retrolateral; (10) palp, dorsal. CD: copulatory ducts, CO: copulatory opening, Co: conductor, E: embolus, EG: epigynal groove, FD: fertilization duct, GH: glandular head, CP: cymbial process, RTA: retrolateral tibial apophysis, S: spermatheca, VTA: ventral tibial apophysis. Scale bars: 0.3 mm.
</paragraph>
</caption>
<paragraph id="8B9536F1FFCF3B45FCA755BA2088FD67" blockId="3.[807,1431,1254,1781]" lastBlockId="4.[157,780,1221,1778]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
Measurements. Total length 3.72. Carapace 1.85 long, 1.62 wide, 0.59 high. Chelicerae 0.57 long, 0.33 wide. Labium 0.30 long, 0.23 wide. Endites 0.43 long, 0.20 wide. Sternum 0.96 long, 0.89 wide. Clypeus 0.27 high. Abdomen 2.58 long, 1.66 wide, 1.22 high. Leg: I. femur 1.54; patella 0.72; tibia 1.27; metatarsus 1.04; tarsus 0.50; total length 5.07; II. 1.68; 0.72; 1.39; 1.13; 0.76; 5.68; III. 1.42; 0.63; 1.07; 0.97; 0.51; 4.60; IV. 1.17; 0.63; 1.12; 1.18; 0.61; 4.71. Leg formula II-I-IV-III. Eye diameters and eye interdistances. AME 0.08, ALE 0.09, PME 0.07, PLE 0.08, AME-AME 0.15, AME-ALE 0.08, PME-PME 0.22, PME-PLE 0.27. MOQ 0.25 long in frontal view,
<quantity id="4CD29B14FFC83B45FE2954AA235BFC85" box="[491,564,1251,1270]" metricMagnitude="-3" metricUnit="m" metricValue="5.587999999999999" pageId="4" pageNumber="5" unit="in" value="0.22">0.22 in</quantity>
dorsal view, anterior width 0.30, posterior width 0.36.
</paragraph>
<caption id="DF556679FFC83B45FF5D54692011FCFE" pageId="4" pageNumber="5" startId="4.[159,226,1056,1076]" targetBox="[159,1436,197,1041]" targetPageId="4" targetType="figure">
<paragraph id="8B9536F1FFC83B45FF5D54692011FCFE" blockId="4.[159,1437,1055,1165]" pageId="4" pageNumber="5">
Figures 1116. Palp of
<taxonomicName id="4C2A4D72FFC83B45FEBD54562347FC40" authorityName="Pantoja &amp; Drago-Bisneto &amp; Saturnino" authorityYear="2020" box="[383,552,1055,1075]" class="Arachnida" family="Philodromidae" genus="Berlandiella" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="zabele" status="sp. nov.">
<emphasis id="B95EEAE3FFC83B45FEBD54562347FC40" box="[383,552,1055,1075]" italics="true" pageId="4" pageNumber="5">Berlandiella zabele</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26D5798FFC83B45FDF2546923EFFC47" box="[560,640,1056,1076]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
: (11) ventral; (12) detail of apical portion of tegulum; (13) dorsal (arrow indicates scopula in apex of cymbium); (14) scopula in cymbium (indicated by arrow); (15) detail of CP (indicated by arrow) and RTA; (16) detail showing excavation in the RTA. CP: cymbial process, RTA: retrolateral tibial apophysis. Scale bars: 11 = 200 µm, 12, 15, 16 = 50 µm, 13 = 150 µm, 14 = 20 µm.
</paragraph>
</caption>
<paragraph id="8B9536F1FFC83B45FF195556240CFD96" blockId="4.[157,780,1221,1778]" lastBlockId="4.[815,1438,1221,1778]" pageId="4" pageNumber="5">
Male (
<typeStatus id="54918853FFC83B45FED85557201EFD41" box="[282,369,1310,1330]" pageId="4" pageNumber="5" type="paratype">paratype</typeStatus>
MPEG.ARA 34480). Shape and bristles of carapace (
<figureCitation id="13112A74FFC83B45FEE5557520F6FD23" box="[295,409,1340,1360]" captionStart="Figures 1722" captionStartId="5.[151,218,1059,1079]" captionTargetBox="[151,1428,197,1046]" captionTargetId="figure-0@5.[151,1428,197,1046]" captionTargetPageId="5" captionText="Figures 1722. Carapace of Berlandiella zabele sp. nov. male: (17) dorsal; (18) anterodorsal; (19) detail of tegument with many sockets of setae (indicated by arrows); (20) strong setae in the cephalic region; (21) hairs; (22) scales in the lateral of carapace. Scale bars: 7, 20 = 30 µm, 18 = 300 µm, 19, 22 = 50 µm, 20 = 30 µm, 21 = 10µm." pageId="4" pageNumber="5">Figs 1722</figureCitation>
) as in female; median region dark yellow with a brown central spot and wide dark brown bands on each side, formed by a tangle of dark spots. Clypeus as in female. Chelicerae as in female. Shape of labium and endites as in female; labium brown and endites pale yellow with some brown spots. Sternum pale yellow, mottled with brown. Legs covered by various types of setae and abundant macrosetae in tibiae and metatarsi (
<figureCitation id="13112A74FFC83B45FE50564420B3FE52" box="[402,476,1549,1569]" captionStart="Figures 2326" captionStartId="6.[159,226,1185,1205]" captionTargetBox="[310,1286,197,1171]" captionTargetId="figure-0@6.[310,1286,197,1171]" captionTargetPageId="6" captionText="Figures 2326. Leg I of Berlandiella zabele sp. nov. male: (23) tarsus and apex of metatarsus with scopula (indicated by arrows); (24) different types of seta in the metatarsus (smaller rectangle show tenent barbs of scopula); (25) claw and claw tufts; (26) detail of tenet barbs of claw tufts. Scale bars: 23 = 150 µm, 24, 25 = 20 µm, 26 = 5 µm." pageId="4" pageNumber="5">Figs 24</figureCitation>
,
<figureCitation id="13112A74FFC83B45FE2A5647236BFE52" box="[488,516,1550,1569]" captionStart="Figures 4043" captionStartId="9.[150,217,1192,1212]" captionTargetBox="[288,1271,197,1179]" captionTargetId="figure-383@9.[288,1271,197,1179]" captionTargetPageId="9" captionText="Figures 4043. Setae: (40) setae on the side tibia of B. querencia male; (41) pair of macrosetae in the venter of tibia I of B. zabele male; (42) trichobothrium on the dorsum of tarsus I of B. zabele male; (43) strong bristles on the side of the metatarsus of B. querencia female. Scale bars: 40, 43 = 50 µm, 41 = 100 µm, 42 = 20 µm." pageId="4" pageNumber="5">42</figureCitation>
). Tarsi and metatarsi with dense scopula (
<figureCitation id="13112A74FFC83B45FEF6566220F0FE4C" box="[308,415,1579,1599]" captionStart="Figures 1722" captionStartId="5.[151,218,1059,1079]" captionTargetBox="[151,1428,197,1046]" captionTargetId="figure-0@5.[151,1428,197,1046]" captionTargetPageId="5" captionText="Figures 1722. Carapace of Berlandiella zabele sp. nov. male: (17) dorsal; (18) anterodorsal; (19) detail of tegument with many sockets of setae (indicated by arrows); (20) strong setae in the cephalic region; (21) hairs; (22) scales in the lateral of carapace. Scale bars: 7, 20 = 30 µm, 18 = 300 µm, 19, 22 = 50 µm, 20 = 30 µm, 21 = 10µm." pageId="4" pageNumber="5">Figs 17, 18</figureCitation>
). Arrangement of the ventral macrosetae of the tarsi and metatarsi I-II as in female. Trichobothria (
<figureCitation id="13112A74FFC83B45FF64562E2186FE09" box="[166,233,1639,1659]" captionStart="Figures 4043" captionStartId="9.[150,217,1192,1212]" captionTargetBox="[288,1271,197,1179]" captionTargetId="figure-383@9.[288,1271,197,1179]" captionTargetPageId="9" captionText="Figures 4043. Setae: (40) setae on the side tibia of B. querencia male; (41) pair of macrosetae in the venter of tibia I of B. zabele male; (42) trichobothrium on the dorsum of tarsus I of B. zabele male; (43) strong bristles on the side of the metatarsus of B. querencia female. Scale bars: 40, 43 = 50 µm, 41 = 100 µm, 42 = 20 µm." pageId="4" pageNumber="5">Fig. 42</figureCitation>
) as in female. Femura yellow shaded with brown, with a transversal white stripe at the apex and yellow longitudinal stripe on retrolateral side; dorsum of the patellae yellow with a longitudinal brown stripe in the middle and a transversal white stripe at the apex, venter shaded with brown; base of the tibiae shaded with brown, the remaining yellow stippled with brown; base and apex of the tarsi shaded with brown, the remaining yellow stippled with brown; metatarsi yellow with a small brown spot at the base. Abdomen brown, oval, wider than long with many brown bristles curved backwards on the dorsum and sides; anteroventral region yellow. Palpal cymbium dark yellow covered by many setae, including scopula in the apex (
<figureCitation id="13112A74FFC83B45FA8D55312225FDDA" captionStart="Figures 1116" captionStartId="4.[159,226,1056,1076]" captionTargetBox="[159,1436,197,1041]" captionTargetId="figure-0@4.[159,1436,197,1041]" captionTargetPageId="4" captionText="Figures 1116. Palp of Berlandiella zabele sp. nov.: (11) ventral; (12) detail of apical portion of tegulum; (13) dorsal (arrow indicates scopula in apex of cymbium); (14) scopula in cymbium (indicated by arrow); (15) detail of CP (indicated by arrow) and RTA; (16) detail showing excavation in the RTA. CP: cymbial process, RTA: retrolateral tibial apophysis. Scale bars: 11 = 200 µm, 12, 15, 16 = 50 µm, 13 = 150 µm, 14 = 20 µm." pageId="4" pageNumber="5">Figs 13, 14</figureCitation>
), cymbial process present; elongated and robust bulb with a small and irregular membranous conductor below the embolus, which is short, sharp and straight; RTA semi-translucent.
</paragraph>
<paragraph id="8B9536F1FFC83B44FCAF55B92340FCAD" blockId="4.[815,1438,1221,1778]" lastBlockId="5.[149,773,1167,1784]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
Measurements. Total length 3.72. Carapace 1.49 long, 1.51 wide, 0.51 high. Chelicerae 0.44 long, 0.25 wide. Labium 0.17 long, 0.22 wide. Endites 0.37 long, 0.17 wide. Sternum 0.82 long, 0.75 wide. Clypeus 0.17 high. Abdomen 1.41 long, 1.00 wide, 0.66 high. Legs: I. femur 1.24; patella 0.59; tibia 1.03; metatarsus 0.89; tarsus 0.63; total length 4.38; II. 1.50; 0.60; 1.25; 1.00; 0.68; 5,03; III. 1.24; 0.50; 0.97; 0.76; 0.50; 3.97; IV. 1.35; 0.54; 1.08; 1.05; 0.51; 4.53. Leg formula: II-IV-I-III. Eye diameters and eye interdistances. AME 0.08, ALE 0.07, PME 0.06, PLE 0.10, AME-AME 0.12, AME-ALE 0.05, PME-PME 0.20, PME-PLE 0.18. MOQ 0.21 long in frontal view,
<quantity id="4CD29B14FFC93B44FDBA54E423D0FCB3" box="[632,703,1197,1216]" metricMagnitude="-3" metricUnit="m" metricValue="4.826" pageId="5" pageNumber="6" unit="in" value="0.19">0.19 in</quantity>
dorsal view, anterior width 0.24, posterior 0.32.
</paragraph>
<caption id="DF556679FFC93B44FF55546A239BFC00" pageId="5" pageNumber="6" startId="5.[151,218,1059,1079]" targetBox="[151,1428,197,1046]" targetPageId="5" targetType="figure">
<paragraph id="8B9536F1FFC93B44FF55546A239BFC00" blockId="5.[151,1429,1058,1139]" pageId="5" pageNumber="6">
Figures 1722. Carapace of
<taxonomicName id="4C2A4D72FFC93B44FE5E546B232CFC45" authorityName="Pantoja &amp; Drago-Bisneto &amp; Saturnino" authorityYear="2020" box="[412,579,1058,1078]" class="Arachnida" family="Philodromidae" genus="Berlandiella" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="zabele" status="sp. nov.">
<emphasis id="B95EEAE3FFC93B44FE5E546B232CFC45" box="[412,579,1058,1078]" italics="true" pageId="5" pageNumber="6">Berlandiella zabele</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26D5798FFC93B44FD8B546A23FDFC44" box="[585,658,1059,1079]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
male: (17) dorsal; (18) anterodorsal; (19) detail of tegument with many sockets of setae (indicated by arrows); (20) strong setae in the cephalic region; (21) hairs; (22) scales in the lateral of carapace. Scale bars: 7, 20 = 30 µm, 18 = 300 µm, 19, 22 = 50 µm, 20 = 30 µm, 21 = 10µm.
</paragraph>
</caption>
<paragraph id="8B9536F1FFC93B44FF1654A0226BFEA9" blockId="5.[149,773,1167,1784]" pageId="5" pageNumber="6">
<materialsCitation id="3B423CACFFC93B44FF1654A0226BFD69" collectingDate="2007-02-02" collectionCode="LS" country="Brazil" county="Material" latitude="-4.0944166" location="Parque Nacional de Sete Cidades" longLatPrecision="1" longitude="-41.731472" municipality="Piracuruca" pageId="5" pageNumber="6" specimenCount="1" stateProvince="Piaui">
<collectingCounty id="62F44E7DFFC93B44FF1654A0204DFC8F" box="[212,290,1257,1276]" pageId="5" pageNumber="6">Material</collectingCounty>
examined.
<collectingCountry id="F33D7661FFC93B44FE4F54A020B8FC8F" box="[397,471,1257,1276]" name="Brazil" pageId="5" pageNumber="6">BRAZIL</collectingCountry>
.
<emphasis id="B95EEAE3FFC93B44FE1954A02366FC8F" box="[475,521,1257,1276]" italics="true" pageId="5" pageNumber="6">
<collectingRegion id="49EEF813FFC93B44FE1954A02366FC8F" box="[475,521,1257,1276]" country="Brazil" name="Piaui" pageId="5" pageNumber="6">Piauí</collectingRegion>
</emphasis>
:
<collectingMunicipality id="6BF1AC8BFFC93B44FDD654A02316FC8F" box="[532,633,1257,1276]" pageId="5" pageNumber="6">Piracuruca</collectingMunicipality>
,
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,
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,
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,
<date id="FF941031FFC93B44FDA8554E238AFD69" box="[618,741,1287,1306]" pageId="5" pageNumber="6" value="2007-02-02">
<collectingDate id="EFD0E9D9FFC93B44FDA8554E238AFD69" box="[618,741,1287,1306]" pageId="5" pageNumber="6" value="2007-02-02">02 Feb. 2007</collectingDate>
</date>
,
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</materialsCitation>
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<location id="8EF5602AFFC93B44FF54556C2181FD4B" LSID="urn:lsid:plazi:treatment:038387E7FFCE3B44FEF1554F25BFFDBE:8EF5602AFFC93B44FF54556C2181FD4B" box="[150,238,1317,1336]" name="Carvalho" pageId="5" pageNumber="6">Carvalho</location>
et al. leg. (
<specimenCount id="9D2CFD78FFC93B44FE97556C20C9FD4B" box="[341,422,1317,1336]" count="1" pageId="5" pageNumber="6" type="female">1 female</specimenCount>
,
<collectionCode id="ED3BAE34FFC93B44FE6D556C2084FD4B" box="[431,491,1317,1336]" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="5" pageNumber="6">MPEG</collectionCode>
.
<specimenCode id="DB8C9E8AFFC93B44FE2C556C2331FD4B" box="[494,606,1317,1336]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="5" pageNumber="6">ARA 15576</specimenCode>
); same collection data as for preceding,
<date id="FF941031FFC93B44FE9E550A20B6FD25" box="[348,473,1347,1366]" pageId="5" pageNumber="6" value="2007-05-24">
<collectingDate id="EFD0E9D9FFC93B44FE9E550A20B6FD25" box="[348,473,1347,1366]" pageId="5" pageNumber="6" value="2007-05-24">24 May.2007</collectingDate>
</date>
,
<collectionCode id="ED3BAE34FFC93B44FE22550A2099FD25" box="[480,502,1347,1366]" country="United Kingdom" name="Linnean Society of London" pageId="5" pageNumber="6">LS</collectionCode>
</materialsCitation>
<materialsCitation id="3B423CACFFC93B44FE38550A23EAFDE2" collectingDate="2007-01-22" collectionCode="MPEG, LS" collectorName="Carvalho" country="Brazil" county="Material" latitude="-4.0944166" location="Carvalho" longLatPrecision="1" longitude="-41.731472" municipality="Piracuruca" pageId="5" pageNumber="6" specimenCode="ARA 15577, ARA 15578" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Piaui">
<location id="8EF5602AFFC93B44FE38550A233EFD25" LSID="urn:lsid:plazi:treatment:038387E7FFCE3B44FEF1554F25BFFDBE:8EF5602AFFC93B44FE38550A233EFD25" box="[506,593,1347,1366]" name="Carvalho" pageId="5" pageNumber="6">Carvalho</location>
et al. leg.(
<specimenCount id="9D2CFD78FFC93B44FD72550A2390FD25" box="[688,767,1347,1366]" count="1" pageId="5" pageNumber="6" type="female">1 female</specimenCount>
,
<collectionCode id="ED3BAE34FFC93B44FF57552821BEFD07" box="[149,209,1377,1396]" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="5" pageNumber="6">MPEG</collectionCode>
.
<specimenCode id="DB8C9E8AFFC93B44FF165528202BFD00" box="[212,324,1376,1396]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="5" pageNumber="6">ARA 15577</specimenCode>
); same collection data as for preceding,
<date id="FF941031FFC93B44FD7C552821A6FDE1" pageId="5" pageNumber="6" value="2007-01-22">
<collectingDate id="EFD0E9D9FFC93B44FD7C552821A6FDE1" pageId="5" pageNumber="6" value="2007-01-22">22 Jan. 2007</collectingDate>
</date>
,
<collectionCode id="ED3BAE34FFC93B44FF1055362186FDE1" box="[210,233,1407,1426]" country="United Kingdom" name="Linnean Society of London" pageId="5" pageNumber="6">LS</collectionCode>
<collectorName id="26DF5327FFC93B44FF2C55372029FDE2" box="[238,326,1406,1425]" pageId="5" pageNumber="6">Carvalho</collectorName>
leg. (
<specimenCount id="9D2CFD78FFC93B44FEB8553620AAFDE2" box="[378,453,1406,1426]" count="2" pageId="5" pageNumber="6" type="male">2 males</specimenCount>
,
<collectionCode id="ED3BAE34FFC93B44FE0C55362365FDE1" box="[462,522,1407,1426]" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="5" pageNumber="6">MPEG</collectionCode>
.
<specimenCode id="DB8C9E8AFFC93B44FDCF55362311FDE2" box="[525,638,1406,1426]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="5" pageNumber="6">ARA 15578</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B423CACFFC93B44FD4C553623A7FD98" collectingDate="2006-12-02" collectingDateMax="2007-01-24" collectingDateMin="2006-12-02" collectionCode="MPEG, LS" collectorName="Carvalho" country="Brazil" county="Material" latitude="-4.0944166" location="Carvalho" longLatPrecision="1" longitude="-41.731472" municipality="Piracuruca" pageId="5" pageNumber="6" specimenCode="ARA 15580, ARA 15581" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Piaui">
same collection data as for preceding,
<date id="FF941031FFC93B44FE5655D5237CFDDC" box="[404,531,1436,1456]" pageId="5" pageNumber="6" value="2006-12-02">
<collectingDate id="EFD0E9D9FFC93B44FE5655D5237CFDDC" box="[404,531,1436,1456]" pageId="5" pageNumber="6" value="2006-12-02">02 Dec. 2006</collectingDate>
</date>
, NF Lo-Man-Hung leg. (
<specimenCount id="9D2CFD78FFC93B44FD3B55D521B7FDBE" count="1" pageId="5" pageNumber="6" type="female">1 female</specimenCount>
,
<collectionCode id="ED3BAE34FFC93B44FF2355F32072FDBE" box="[225,285,1466,1485]" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="5" pageNumber="6">MPEG</collectionCode>
.
<specimenCode id="DB8C9E8AFFC93B44FEE255F320FFFDBE" box="[288,400,1466,1485]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="5" pageNumber="6">ARA 15580</specimenCode>
); same collection data as for preceding,
<date id="FF941031FFC93B44FF545591207FFD98" box="[150,272,1496,1515]" pageId="5" pageNumber="6" value="2007-01-24">
<collectingDate id="EFD0E9D9FFC93B44FF545591207FFD98" box="[150,272,1496,1515]" pageId="5" pageNumber="6" value="2007-01-24">24 Jan. 2007</collectingDate>
</date>
,
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<collectorName id="26DF5327FFC93B44FEF6559120E3FD98" box="[308,396,1496,1515]" pageId="5" pageNumber="6">Carvalho</collectorName>
leg. (
<specimenCount id="9D2CFD78FFC93B44FE7D55912366FD98" box="[447,521,1496,1515]" count="2" pageId="5" pageNumber="6" type="male">2 males</specimenCount>
,
<collectionCode id="ED3BAE34FFC93B44FDD055912321FD98" box="[530,590,1496,1515]" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="5" pageNumber="6">MPEG</collectionCode>
.
<specimenCode id="DB8C9E8AFFC93B44FD93559123AEFD98" box="[593,705,1496,1515]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="5" pageNumber="6">ARA 15581</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B423CACFFC93B44FD135591233CFE7A" collectingDate="2007-01-26" collectingDateMax="2007-01-24" collectingDateMin="2006-12-02" collectionCode="LS" collectorName="Carvalho" country="Brazil" county="Material" latitude="-4.0944166" location="Carvalho" longLatPrecision="1" longitude="-41.731472" municipality="Piracuruca" pageId="5" pageNumber="6" specimenCount="1" stateProvince="Piaui">
same collection data as for preceding,
<date id="FF941031FFC93B44FE0255BF235AFE7A" box="[448,565,1526,1545]" pageId="5" pageNumber="6" value="2007-01-26">
<collectingDate id="EFD0E9D9FFC93B44FE0255BF235AFE7A" box="[448,565,1526,1545]" pageId="5" pageNumber="6" value="2007-01-26">26 Jan.2007</collectingDate>
</date>
,
<collectionCode id="ED3BAE34FFC93B44FDFF55BF233CFE7A" box="[573,595,1526,1545]" country="United Kingdom" name="Linnean Society of London" pageId="5" pageNumber="6">LS</collectionCode>
</materialsCitation>
<materialsCitation id="3B423CACFFC93B44FD9555BF205CFE36" collectingDate="2007-02-02" collectingDateMax="2007-01-24" collectingDateMin="2006-12-02" collectionCode="MPEG, LS" collectorName="Carvalho" country="Brazil" county="Material" latitude="-4.0944166" location="Carvalho" longLatPrecision="1" longitude="-41.731472" municipality="Piracuruca" pageId="5" pageNumber="6" specimenCode="ARA 15582" specimenCount="1" specimenCount-male="1" stateProvince="Piaui">
<location id="8EF5602AFFC93B44FD9555BF23C0FE7A" LSID="urn:lsid:plazi:treatment:038387E7FFCE3B44FEF1554F25BFFDBE:8EF5602AFFC93B44FD9555BF23C0FE7A" box="[599,687,1526,1545]" name="Carvalho" pageId="5" pageNumber="6">Carvalho</location>
et al.leg. (
<specimenCount id="9D2CFD78FFC93B44FF5C565D21B2FE54" box="[158,221,1556,1575]" count="1" pageId="5" pageNumber="6" type="male">1 male</specimenCount>
,
<collectionCode id="ED3BAE34FFC93B44FF27565D204EFE54" box="[229,289,1556,1575]" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="5" pageNumber="6">MPEG</collectionCode>
.
<specimenCode id="DB8C9E8AFFC93B44FEE6565D20FCFE54" box="[292,403,1556,1575]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="5" pageNumber="6">ARA 15582</specimenCode>
); same collection data as for preceding,
<date id="FF941031FFC93B44FF57567B207CFE36" box="[149,275,1586,1605]" pageId="5" pageNumber="6" value="2007-02-02">
<collectingDate id="EFD0E9D9FFC93B44FF57567B207CFE36" box="[149,275,1586,1605]" pageId="5" pageNumber="6" value="2007-02-02">02 Feb. 2007</collectingDate>
</date>
,
<collectionCode id="ED3BAE34FFC93B44FEDE567B205CFE36" box="[284,307,1586,1605]" country="United Kingdom" name="Linnean Society of London" pageId="5" pageNumber="6">LS</collectionCode>
</materialsCitation>
<materialsCitation id="3B423CACFFC93B44FEFA567B226EFEF3" collectingDate="2007-01-22" collectingDateMax="2007-01-24" collectingDateMin="2006-12-02" collectionCode="MPEG, FM" collectorName="Oliveira-Neto" country="Brazil" county="Material" latitude="-4.0993056" location="Carvalho" longLatPrecision="1" longitude="-41.71686" municipality="Piracuruca" pageId="5" pageNumber="6" specimenCode="ARA 15583, ARA 34478" specimenCount="2" specimenCount-male="2" stateProvince="Piaui">
<location id="8EF5602AFFC93B44FEFA567B20FFFE36" LSID="urn:lsid:plazi:treatment:038387E7FFCE3B44FEF1554F25BFFDBE:8EF5602AFFC93B44FEFA567B20FFFE36" box="[312,400,1586,1605]" latitude="-4.0993056" longLatPrecision="1" longitude="-41.71686" name="Carvalho" pageId="5" pageNumber="6">Carvalho</location>
et al. leg. (
<specimenCount id="9D2CFD78FFC93B44FE3E567B2352FE36" box="[508,573,1586,1605]" count="1" pageId="5" pageNumber="6" type="male">1 male</specimenCount>
,
<collectionCode id="ED3BAE34FFC93B44FD85567B23ECFE36" box="[583,643,1586,1605]" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="5" pageNumber="6">MPEG</collectionCode>
.
<specimenCode id="DB8C9E8AFFC93B44FD44567B2398FE37" box="[646,759,1585,1605]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="5" pageNumber="6">ARA 15583</specimenCode>
); same collection data as for preceding,
<geoCoordinate id="EE1E5036FFC93B44FDC856062314FE11" box="[522,635,1615,1634]" degrees="04" direction="south" minutes="5" orientation="latitude" pageId="5" pageNumber="6" precision="1" seconds="57.5" value="-4.0993056">04°557.5”S</geoCoordinate>
,
<geoCoordinate id="EE1E5036FFC93B44FD475619226FFE10" box="[645,768,1616,1635]" degrees="41" direction="west" minutes="43" orientation="longitude" pageId="5" pageNumber="6" precision="1" seconds="0.7" value="-41.71686">41°430.7”W</geoCoordinate>
,
<date id="FF941031FFC93B44FF5A5624207BFEF3" box="[152,276,1645,1665]" pageId="5" pageNumber="6" value="2007-01-22">
<collectingDate id="EFD0E9D9FFC93B44FF5A5624207BFEF3" box="[152,276,1645,1665]" pageId="5" pageNumber="6" value="2007-01-22">22 Jan. 2007</collectingDate>
</date>
,
<collectionCode id="ED3BAE34FFC93B44FEDC56272052FEF2" box="[286,317,1646,1665]" country="China" name="Department of Nature, Fujian Province Museum" pageId="5" pageNumber="6">FM</collectionCode>
<collectorName id="26DF5327FFC93B44FE86562420AAFEF3" box="[324,453,1645,1664]" pageId="5" pageNumber="6">Oliveira-Neto</collectorName>
leg. (
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,
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.
<specimenCode id="DB8C9E8AFFC93B44FD4A56242395FEF3" box="[648,762,1645,1664]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="5" pageNumber="6">ARA 34478</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B423CACFFC93B44FF5A56C223FBFEED" box="[152,660,1675,1695]" collectingDate="2007-01-29" collectionCode="LS" country="Brazil" latitude="-4.135611" location="Brasileira" longLatPrecision="1" longitude="-41.719" pageId="5" pageNumber="6" specimenCount="1" stateProvince="Piaui">
<location id="8EF5602AFFC93B44FF5A56C22180FEED" LSID="urn:lsid:plazi:treatment:038387E7FFCE3B44FEF1554F25BFFDBE:8EF5602AFFC93B44FF5A56C22180FEED" box="[152,239,1675,1694]" country="Brazil" latitude="-4.135611" longLatPrecision="1" longitude="-41.719" name="Brasileira" pageId="5" pageNumber="6" stateProvince="Piaui">Brasileira</location>
,
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,
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,
<date id="FF941031FFC93B44FDC056C22318FEED" box="[514,631,1675,1695]" pageId="5" pageNumber="6" value="2007-01-29">
<collectingDate id="EFD0E9D9FFC93B44FDC056C22318FEED" box="[514,631,1675,1695]" pageId="5" pageNumber="6" value="2007-01-29">29 Jan. 2007</collectingDate>
</date>
,
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<materialsCitation id="3B423CACFFC93B44FD5A56C22097FEA8" collectingDate="2007-01-28" collectionCode="MPEG, LS" collectorName="Carvalho" country="Brazil" latitude="-4.135611" location="Carvalho" longLatPrecision="1" longitude="-41.719" pageId="5" pageNumber="6" specimenCode="ARA 34479" specimenCount="1" specimenCount-male="1" stateProvince="Piaui">
<location id="8EF5602AFFC93B44FD5A56C22380FEED" LSID="urn:lsid:plazi:treatment:038387E7FFCE3B44FEF1554F25BFFDBE:8EF5602AFFC93B44FD5A56C22380FEED" box="[664,751,1675,1694]" name="Carvalho" pageId="5" pageNumber="6">Carvalho</location>
et al. leg. (
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,
<collectionCode id="ED3BAE34FFC93B44FEE756E0200EFECF" box="[293,353,1705,1724]" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="5" pageNumber="6">MPEG</collectionCode>
.
<specimenCode id="DB8C9E8AFFC93B44FEA156E020BDFECF" box="[355,466,1705,1724]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="5" pageNumber="6">ARA 34479</specimenCode>
); same collection data as for preceding,
<date id="FF941031FFC93B44FF23568E2037FEA9" box="[225,344,1735,1754]" pageId="5" pageNumber="6" value="2007-01-28">
<collectingDate id="EFD0E9D9FFC93B44FF23568E2037FEA9" box="[225,344,1735,1754]" pageId="5" pageNumber="6" value="2007-01-28">28 Jan. 2007</collectingDate>
</date>
,
<collectionCode id="ED3BAE34FFC93B44FE9D568E2019FEA9" box="[351,374,1735,1754]" country="United Kingdom" name="Linnean Society of London" pageId="5" pageNumber="6">LS</collectionCode>
<collectorName id="26DF5327FFC93B44FEBB568E20BEFEA9" box="[377,465,1735,1754]" pageId="5" pageNumber="6">Carvalho</collectorName>
leg.
</materialsCitation>
<materialsCitation id="3B423CACFFC93B44FE39568E226FFEA9" box="[507,768,1735,1754]" collectionCode="MPEG" country="Brazil" pageId="5" pageNumber="6" specimenCode="ARA 34481" specimenCount="1" specimenCount-male="1" stateProvince="Piaui">
(
<specimenCount id="9D2CFD78FFC93B44FDC6568E232CFEA9" box="[516,579,1735,1754]" count="1" pageId="5" pageNumber="6" type="male">1 male</specimenCount>
,
<collectionCode id="ED3BAE34FFC93B44FD89568E23E8FEA9" box="[587,647,1735,1754]" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="5" pageNumber="6">MPEG</collectionCode>
.
<specimenCode id="DB8C9E8AFFC93B44FD4B568E2395FEA9" box="[649,762,1735,1754]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="5" pageNumber="6">ARA 34481</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B9536F1FFC93B44FF1656AC226BFE8B" blockId="5.[149,773,1167,1784]" box="[212,772,1764,1784]" pageId="5" pageNumber="6">
Distribution. Known only from the
<typeStatus id="54918853FFC93B44FDF256AC2334FE8B" box="[560,603,1765,1784]" pageId="5" pageNumber="6">type</typeStatus>
locality (
<figureCitation id="13112A74FFC93B44FD7656AD2396FE8B" box="[692,761,1764,1784]" captionStart="Figure 44" captionStartId="10.[159,217,950,970]" captionTargetBox="[275,1320,199,937]" captionTargetId="figure-546@10.[274,1322,198,935]" captionTargetPageId="10" captionText="Figure 44. Distribution map of Berlandiella zabele sp. nov. and B. querencia." pageId="5" pageNumber="6">Fig. 44</figureCitation>
).
</paragraph>
<paragraph id="8B9536F1FFC93B44FCA754C624FBFD25" blockId="5.[807,1430,1167,1486]" pageId="5" pageNumber="6">
Etymology. The specific epithet refers to a mythological character known by indigenous people of
<collectingRegion id="49EEF813FFC93B44FB1754E42466FCB3" box="[1237,1289,1197,1216]" country="Brazil" name="Piaui" pageId="5" pageNumber="6">Piauí</collectingRegion>
. According to indigenous mythology, Zabelê, a member of the Amanajós tribe, fell in love with Metara, a member of an enemy tribe. When the romance of Zabelê and Metara was discovered, tribal warfare result- ed in the death of both lovers. The god Tupã pitied the two lovers and so transformed them into birds that fly together for eternity.
</paragraph>
<paragraph id="8B9536F1FFC93B44FCA7552925BFFDBE" blockId="5.[807,1430,1167,1486]" pageId="5" pageNumber="6">
Remarks. The general color, legs especially, vary in the abundance of brown spots and some specimens may be darker or lighter than the
<typeStatus id="54918853FFC93B44FC2755D42551FDC3" box="[997,1086,1437,1456]" pageId="5" pageNumber="6" type="paratype">paratype</typeStatus>
. The pattern of ventral macrosetae of the legs also varies between individuals.
</paragraph>
</subSubSection>
</treatment>
</document>