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<document id="2FD97FFE786F4BEE295785BB92085B47" ID-DOI="10.5852/ejt.2024.968.2737" ID-ISSN="2118-9773" ID-Zenodo-Dep="14186039" ID-ZooBank="C38A1E04-4541-4BFD-92E7-DB615BE736CF" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1732018292087" checkinUser="plazi" docAuthor="Schröder, Ove, Schächinger, Peter M., Bouchet, Philippe &amp; Haase, Martin" docDate="2024" docId="038387E6FFF3FFA7CB692934FAD6FE0D" docLanguage="en" docName="EJT.2024.968.275-294.pdf" docOrigin="European Journal of Taxonomy 968" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2737/12561" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Viriiella touaouroua Schröder, Schächinger, Bouchet &amp; Haase, 2024, gen. et sp. nov." docType="treatment" docUuid="18A13BF1-D46C-4A61-A0F8-17B86351C903" docUuidSource="ZooBank" docVersion="3" lastPageNumber="285" masterDocId="FFBAFF9EFFFBFFACC92A2A69FFC4FF80" masterDocTitle="A new genus and species of spring snails (Caenogastropoda, Tateidae) from the ultramafic South of New Caledonia" masterLastPageNumber="294" masterPageNumber="275" pageNumber="283" updateTime="1732212620015" updateUser="felipe" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="E4513B7A5B752AB42D187A51133066FE">A new genus and species of spring snails (Caenogastropoda, Tateidae) from the ultramafic South of New Caledonia</mods:title>
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<mods:namePart id="0519802F0D2671C18989C51DEDFDA577">Schröder, Ove</mods:namePart>
<mods:nameIdentifier id="4B0DF53D699BAAE4CA7694330E12CB26" type="LSID">8A198DAE-862A-43DC-8F57-E982BFFCBCA1</mods:nameIdentifier>
<mods:affiliation id="16BA607F42ACE446E76B4326BD3326E8">AG Vogelwarte, Zoological Institute and Museum, University of Greifswald, Soldmannstrasse 23, 17489 Greifswald, Germany.</mods:affiliation>
<mods:nameIdentifier id="A3283FD04F5DAA4640BB00090F038573" type="email">ove.schroeder@outlook.de</mods:nameIdentifier>
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<mods:namePart id="1ED222ED16309FF19310DD5D6CD15C73">Schächinger, Peter M.</mods:namePart>
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<mods:affiliation id="F1ADECF811FCC2E7A64E217B14320ED3">AG Vogelwarte, Zoological Institute and Museum, University of Greifswald, Soldmannstrasse 23, 17489 Greifswald, Germany.</mods:affiliation>
<mods:nameIdentifier id="252482F2398C2EF1EF925C8FBD3C8ACC" type="email">peter.schaechinger@uni-greifswald.de</mods:nameIdentifier>
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<mods:namePart id="D64470F8BB115D56CFEF53D2CBFB9CD2">Bouchet, Philippe</mods:namePart>
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<mods:affiliation id="FC0EE625922E0EDD6E898F04FDABF5E5">Muséum national d’Histoire naturelle, Institut Systématique Evolution Biodiversité (ISYEB), CNRS, Sorbonne Université, EPHE, Université des Antilles, Paris, France.</mods:affiliation>
<mods:nameIdentifier id="BF274AA3F9931F1502AA5CE86AE150CE" type="email">pbouchet@mnhn.fr</mods:nameIdentifier>
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<mods:namePart id="744094910313B65A5BD43557043C00CA">Haase, Martin</mods:namePart>
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<mods:affiliation id="920F24BB3FCAAC75BD60553567EC9817">AG Vogelwarte, Zoological Institute and Museum, University of Greifswald, Soldmannstrasse 23, 17489 Greifswald, Germany.</mods:affiliation>
<mods:nameIdentifier id="A27237DE3CF67FC8AEDEE0D788FA28F6" type="email">martin.haase@uni-greifswald.de</mods:nameIdentifier>
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<mods:title id="C905FAC43CD3CA980E811B7881C3C9D8">European Journal of Taxonomy</mods:title>
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<mods:date id="5ED7321B8AE884052637DA307D44ECA2">2024</mods:date>
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<mods:number id="91A9801A4E7AE2687F9D8740B604B2B9">2024-11-18</mods:number>
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<mods:number id="A6DC316F54071FDF1D72D68126219282">968</mods:number>
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<mods:url id="32C67D66B8D916B830650C58917C3906">https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2737/12561</mods:url>
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<mods:classification id="F71AA77E351C0A0678C9C164633F575D">journal article</mods:classification>
<mods:identifier id="0F4494A3543B59F1320A91F3654D427E" type="DOI">10.5852/ejt.2024.968.2737</mods:identifier>
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<treatment id="038387E6FFF3FFA7CB692934FAD6FE0D" LSID="urn:lsid:zoobank.org:act:18A13BF1-D46C-4A61-A0F8-17B86351C903" httpUri="http://treatment.plazi.org/id/038387E6FFF3FFA7CB692934FAD6FE0D" lastPageId="11" lastPageNumber="285" pageId="8" pageNumber="283">
<subSubSection id="C330657BFFF3FFA4CB692934FC35FCF8" box="[579,1009,861,888]" pageId="8" pageNumber="283" type="nomenclature">
<paragraph id="8B9536F0FFF3FFA4CB692934FC35FCF8" blockId="8.[368,1219,861,965]" box="[579,1009,861,888]" pageId="8" pageNumber="283">
<heading id="D0DD819CFFF3FFA4CB692934FC35FCF8" box="[579,1009,861,888]" centered="true" fontSize="11" level="2" pageId="8" pageNumber="283" reason="2">
<taxonomicName id="4C2A4D73FFF3FFA4CB692934FCFEFCF8" authorityName="Schröder &amp; Schächinger &amp; Bouchet &amp; Haase" authorityYear="2024" box="[579,826,861,888]" class="Gastropoda" family="Tateidae" genus="Viriiella" kingdom="Animalia" order="Littorinimorpha" pageId="8" pageNumber="283" phylum="Mollusca" rank="species" species="touaouroua" status="gen. et sp. nov.">
<emphasis id="B95EEAE2FFF3FFA4CB692934FCFEFCF8" bold="true" box="[579,826,861,888]" italics="true" pageId="8" pageNumber="283">Viriiella touaouroua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26D5799FFF3FFA4CA6B2937FC35FCF8" box="[833,1009,862,888]" pageId="8" pageNumber="283" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="C330657BFFF3FFA4C85A29EDFC88FC45" pageId="8" pageNumber="283" type="description">
<paragraph id="8B9536F0FFF3FFA4C85A29EDFB07FC1F" blockId="8.[368,1219,861,965]" box="[368,1219,900,927]" pageId="8" pageNumber="283">
<uri id="FFBB3AF2FFF3FFA4C85A29EDFB07FC1F" box="[368,1219,900,927]" pageId="8" pageNumber="283">
urn:lsid:zoobank.org:act:
<uuid id="FF8C0C25FFF3FFA4CBBD29EDFB07FC1F" box="[663,1219,900,927]" pageId="8" pageNumber="283">18A13BF1-D46C-4A61-A0F8-17B86351C903</uuid>
</uri>
</paragraph>
<paragraph id="8B9536F0FFF3FFA4CBCD29C2FC88FC45" blockId="8.[368,1219,861,965]" box="[743,844,939,965]" pageId="8" pageNumber="283">
<figureCitation id="13112A75FFF3FFA4CBCD29C2FC88FC45" box="[743,844,939,965]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStart-5="Fig" captionStartId-0="8.[189,232,1963,1989]" captionStartId-1="9.[189,232,2019,2045]" captionStartId-2="10.[191,234,1963,1989]" captionStartId-3="11.[189,232,1892,1918]" captionStartId-4="12.[189,232,1453,1479]" captionStartId-5="12.[399,442,1998,2024]" captionTargetBox-0="[189,1398,1320,1925]" captionTargetBox-1="[189,1398,1376,1981]" captionTargetBox-2="[296,1293,1177,1925]" captionTargetBox-3="[189,1398,1132,1855]" captionTargetBox-4="[352,1220,265,1415]" captionTargetBox-5="[189,1399,1590,1960]" captionTargetId-0="figure-273@8.[189,1399,1320,1925]" captionTargetId-1="figure-291@9.[189,1399,1376,1981]" captionTargetId-2="figure-306@10.[296,1293,1177,1925]" captionTargetId-3="figure-334@11.[189,1399,1132,1855]" captionTargetId-4="figure-1@12.[352,1222,265,1417]" captionTargetId-5="figure-2@12.[189,1399,1590,1960]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionTargetPageId-3="11" captionTargetPageId-4="12" captionTargetPageId-5="12" captionText-0="Fig. 3. Types of Viriiella touaouroua gen. et sp. nov. A. Holotype (MNHN-IM-2000-39460). B-D. Paratypes (MNHN-IM-2000-39461)." captionText-1="Fig. 4. SEM micrographs of paratypes of Viriiella touaouroua gen. et sp. nov. (MNHN-IM-2000-39461)." captionText-2="Fig. 5. Radula of Viriiella touaouroua gen. et sp. nov. Abbreviations: L=lateral teeth; M1 =inner marginal teeth; M2 =outer marginal teeth; R =rhachis (central) teeth." captionText-3="Fig. 6. Female genitalia of Viriiella touaouroua gen. et sp. nov. A. All parts opaque. B. Oviduct transparent revealing the seminal receptacle. Abbreviations: ac= anterior capsule gland; ag=albumen gland; bc =bursa copulatrix; od =oviduct; ov =ovary; pc =posterior capsule gland; rs =receptaculum semins; vc =ventral channel." captionText-4="Fig. 7. Male genitalia of Viriiella touaouroua gen. et sp. nov. Abbreviations: pe=penis; pr =prostate; pv =pallial vas deferens; te =testis; vd =vas deferens; vs= vesicula seminalis." captionText-5="Fig. 8. Penes of three males of Viriiella touaouroua gen. et sp. nov." figureDoi-0="http://doi.org/10.5281/zenodo.14186045" figureDoi-1="http://doi.org/10.5281/zenodo.14186047" figureDoi-2="http://doi.org/10.5281/zenodo.14186049" figureDoi-3="http://doi.org/10.5281/zenodo.14186051" figureDoi-4="http://doi.org/10.5281/zenodo.14186053" figureDoi-5="http://doi.org/10.5281/zenodo.14186055" httpUri-0="https://zenodo.org/record/14186045/files/figure.png" httpUri-1="https://zenodo.org/record/14186047/files/figure.png" httpUri-2="https://zenodo.org/record/14186049/files/figure.png" httpUri-3="https://zenodo.org/record/14186051/files/figure.png" httpUri-4="https://zenodo.org/record/14186053/files/figure.png" httpUri-5="https://zenodo.org/record/14186055/files/figure.png" pageId="8" pageNumber="283">Figs 3–8</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C330657BFFF3FFA4C9972986FB67FB01" pageId="8" pageNumber="283" type="diagnosis">
<paragraph id="8B9536F0FFF3FFA4C9972986FEF2FB89" blockId="8.[189,310,1007,1033]" box="[189,310,1007,1033]" pageId="8" pageNumber="283">
<heading id="D0DD819CFFF3FFA4C9972986FEF2FB89" bold="true" box="[189,310,1007,1033]" fontSize="11" level="3" pageId="8" pageNumber="283" reason="3">
<emphasis id="B95EEAE2FFF3FFA4C9972986FEF2FB89" bold="true" box="[189,310,1007,1033]" pageId="8" pageNumber="283">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8B9536F0FFF3FFA4C9972E48FB67FB01" blockId="8.[189,1398,1057,1154]" pageId="8" pageNumber="283">Shell small, cylindrical to cylindro-conical, 1.8 to 2 times as high as wide, not more than four rather flat whorls; aperture semi-lunar, not extending far beyond the outline of the spire, lip continuous, slightly thickened, orthocline; umbilicus narrow; smooth protoconch with about 1.25 whorls.</paragraph>
</subSubSection>
<subSubSection id="C330657BFFF3FFA4C9972EC5FB54FB78" box="[189,1168,1196,1273]" pageId="8" pageNumber="283" type="etymology">
<paragraph id="8B9536F0FFF3FFA4C9972EC5FE80FB46" blockId="8.[189,324,1196,1222]" box="[189,324,1196,1222]" pageId="8" pageNumber="283">
<heading id="D0DD819CFFF3FFA4C9972EC5FE80FB46" bold="true" box="[189,324,1196,1222]" fontSize="11" level="3" pageId="8" pageNumber="283" reason="3">
<emphasis id="B95EEAE2FFF3FFA4C9972EC5FE80FB46" bold="true" box="[189,324,1196,1222]" pageId="8" pageNumber="283">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8B9536F0FFF3FFA4C9972EB7FB54FB78" blockId="8.[189,1168,1246,1273]" box="[189,1168,1246,1273]" pageId="8" pageNumber="283">
The epithetum refers to Touaourou, the name of the community of the
<typeStatus id="54918852FFF3FFA4CAD32EB6FBE8FB79" box="[1017,1068,1247,1273]" pageId="8" pageNumber="283">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<caption id="DF556678FFF3FFA4C9972DC2FD76F868" ID-DOI="http://doi.org/10.5281/zenodo.14186045" ID-Zenodo-Dep="14186045" httpUri="https://zenodo.org/record/14186045/files/figure.png" pageId="8" pageNumber="283" startId="8.[189,232,1963,1989]" targetBox="[189,1398,1320,1925]" targetPageId="8">
<paragraph id="8B9536F0FFF3FFA4C9972DC2FD76F868" blockId="8.[189,1399,1963,2025]" pageId="8" pageNumber="283">
<emphasis id="B95EEAE2FFF3FFA4C9972DC2FED1F845" bold="true" box="[189,277,1963,1989]" pageId="8" pageNumber="283">Fig. 3.</emphasis>
Types of
<taxonomicName id="4C2A4D73FFF3FFA4C88D2DC2FD64F845" authorityName="Schröder &amp; Schächinger &amp; Bouchet &amp; Haase" authorityYear="2024" box="[423,672,1963,1989]" class="Gastropoda" family="Tateidae" genus="Viriiella" kingdom="Animalia" order="Littorinimorpha" pageId="8" pageNumber="283" phylum="Mollusca" rank="species" species="touaouroua" status="gen. et sp. nov.">
<emphasis id="B95EEAE2FFF3FFA4C88D2DC2FD64F845" box="[423,672,1963,1989]" italics="true" pageId="8" pageNumber="283">Viriiella touaouroua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26D5799FFF3FFA4CB9B2DC5FCBBF846" box="[689,895,1964,1990]" pageId="8" pageNumber="283" rank="species">gen. et sp. nov.</taxonomicNameLabel>
<emphasis id="B95EEAE2FFF3FFA4CABA2DC2FC61F845" bold="true" box="[912,933,1963,1989]" pageId="8" pageNumber="283">A</emphasis>
. Holotype (MNHN-IM-2000-39460).
<emphasis id="B95EEAE2FFF3FFA4C9972DA7FF34F868" bold="true" box="[189,240,1998,2024]" pageId="8" pageNumber="283">B-D</emphasis>
. Paratypes (MNHN-IM-2000-39461).
</paragraph>
</caption>
<subSubSection id="C330657BFFF2FFA5C9972B60FAA8FDA9" pageId="9" pageNumber="284" type="materials_examined">
<paragraph id="8B9536F0FFF2FFA5C9972B60FEA8FEA2" blockId="9.[189,364,264,291]" box="[189,364,264,291]" pageId="9" pageNumber="284">
<heading id="D0DD819CFFF2FFA5C9972B60FEA8FEA2" bold="true" box="[189,364,264,291]" fontSize="11" level="3" pageId="9" pageNumber="284" reason="3">
<emphasis id="B95EEAE2FFF2FFA5C9972B60FEA8FEA2" bold="true" box="[189,364,264,291]" pageId="9" pageNumber="284">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8B9536F0FFF2FFA5C9C02B53FBAEFE40" blockId="9.[189,1399,314,448]" pageId="9" pageNumber="284">
<materialsCitation id="3B423CADFFF2FFA5C9C02B53FBAEFE40" collectingDate="2016-11-03" collectorName="Bouchet &amp; Haase" country="New Caledonia" county="Yate" latitude="-22.184889" location="small stream draining a swamp below a slope dominated by Niauli trees on the western bank of the road" longLatPrecision="1" longitude="166.97083" municipality="Touaourou" pageId="9" pageNumber="284" specimenCode="MNHN-IM-2000-39460" specimenCount="1" stateProvince="Province Sud" typeStatus="holotype">
<emphasis id="B95EEAE2FFF2FFA5C9C02B53FE98FED4" bold="true" box="[234,348,314,340]" pageId="9" pageNumber="284">
<typeStatus id="54918852FFF2FFA5C9C02B53FE98FED4" box="[234,348,314,340]" pageId="9" pageNumber="284" type="holotype">Holotype</typeStatus>
</emphasis>
(
<figureCitation id="13112A75FFF2FFA5C8462B53FE0DFED4" box="[364,457,314,340]" captionStart="Fig" captionStartId="8.[189,232,1963,1989]" captionTargetBox="[189,1398,1320,1925]" captionTargetId="figure-273@8.[189,1399,1320,1925]" captionTargetPageId="8" captionText="Fig. 3. Types of Viriiella touaouroua gen. et sp. nov. A. Holotype (MNHN-IM-2000-39460). B-D. Paratypes (MNHN-IM-2000-39461)." figureDoi="http://doi.org/10.5281/zenodo.14186045" httpUri="https://zenodo.org/record/14186045/files/figure.png" pageId="9" pageNumber="284">Fig. 3A</figureCitation>
)
<collectingCountry id="F33D7660FFF2FFA5C9972B09FE77FEF9" box="[189,435,351,378]" name="New Caledonia" pageId="9" pageNumber="284">NEW CALEDONIA</collectingCountry>
•
<collectingRegion id="49EEF812FFF2FFA5C8FF2B36FDBCFEF9" box="[469,632,351,377]" country="New Caledonia" name="Province Sud" pageId="9" pageNumber="284">Province Sud</collectingRegion>
, district of
<collectingCounty id="62F44E7CFFF2FFA5CA222B36FC84FEF9" box="[776,832,351,377]" pageId="9" pageNumber="284">Yaté</collectingCounty>
, community
<collectingMunicipality id="6BF1AC8AFFF2FFA5CAF52B09FBA5FEFA" box="[991,1121,352,378]" pageId="9" pageNumber="284">Touaourou</collectingMunicipality>
,
<location id="8EF5602BFFF2FFA5CD5A2B36FB7EFE1C" LSID="urn:lsid:plazi:treatment:038387E6FFF3FFA7CB692934FAD6FE0D:8EF5602BFFF2FFA5CD5A2B36FB7EFE1C" country="New Caledonia" county="Yate" latitude="-22.184889" longLatPrecision="1" longitude="166.97083" municipality="Touaourou" name="small stream draining a swamp below a slope dominated by Niauli trees on the western bank of the road" pageId="9" pageNumber="284" stateProvince="Province Sud">small stream draining a swamp below a slope dominated by Niauli trees on the western bank of the road</location>
;
<geoCoordinate id="EE1E5037FFF2FFA5CDE02BE8FAB5FE1D" box="[1226,1393,385,413]" degrees="22" direction="south" minutes="11" orientation="latitude" pageId="9" pageNumber="284" precision="1" seconds="05.6" value="-22.184889">22°11ʹ05.6ʺ S</geoCoordinate>
,
<geoCoordinate id="EE1E5037FFF2FFA5C9972BCDFEB7FE40" box="[189,371,420,448]" degrees="166" direction="east" minutes="58" orientation="longitude" pageId="9" pageNumber="284" precision="1" seconds="15.0" value="166.97083">166°58ʹ15.0ʺ E</geoCoordinate>
;
<date id="FF941030FFF2FFA5C8552BCFFDD7FE40" box="[383,531,422,448]" pageId="9" pageNumber="284" value="2016-11-03">
<collectingDate id="EFD0E9D8FFF2FFA5C8552BCFFDD7FE40" box="[383,531,422,448]" pageId="9" pageNumber="284" value="2016-11-03">3 Nov. 2016</collectingDate>
</date>
;
<collectorName id="26DF5326FFF2FFA5CB352BCFFD45FE40" box="[543,641,422,448]" pageId="9" pageNumber="284">Bouchet</collectorName>
and
<collectorName id="26DF5326FFF2FFA5CB932BCFFCC5FE40" box="[697,769,422,448]" pageId="9" pageNumber="284">Haase</collectorName>
leg.;
<specimenCode id="DB8C9E8BFFF2FFA5CA692BCFFBA2FE40" box="[835,1126,422,448]" pageId="9" pageNumber="284">MNHN-IM-2000-39460</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8B9536F0FFF2FFA5C9C02B83FAA8FDA9" blockId="9.[189,1388,490,554]" pageId="9" pageNumber="284">
<materialsCitation id="3B423CADFFF2FFA5C9C02B83FAACFDA9" collectingDate="2016-11-03" collectorName="Bouchet &amp; Haase" country="New Caledonia" county="Yate" latitude="-22.184889" location="NEW CALEDONIA" longLatPrecision="1" longitude="166.97083" municipality="Touaourou" pageId="9" pageNumber="284" specimenCode="MNHN-IM-2000-39461" specimenCount="50" stateProvince="Province Sud" typeStatus="paratype">
<emphasis id="B95EEAE2FFF2FFA5C9C02B83FEA3FD84" bold="true" box="[234,359,490,516]" pageId="9" pageNumber="284">
<typeStatus id="54918852FFF2FFA5C9C02B83FEA3FD84" box="[234,359,490,516]" pageId="9" pageNumber="284" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<figureCitation id="13112A75FFF2FFA5C85D2B83FE38FD84" box="[375,508,490,516]" captionStart="Fig" captionStartId="8.[189,232,1963,1989]" captionTargetBox="[189,1398,1320,1925]" captionTargetId="figure-273@8.[189,1399,1320,1925]" captionTargetPageId="8" captionText="Fig. 3. Types of Viriiella touaouroua gen. et sp. nov. A. Holotype (MNHN-IM-2000-39460). B-D. Paratypes (MNHN-IM-2000-39461)." figureDoi="http://doi.org/10.5281/zenodo.14186045" httpUri="https://zenodo.org/record/14186045/files/figure.png" pageId="9" pageNumber="284">Figs 3B–D</figureCitation>
,
<figureCitation id="13112A75FFF2FFA5CB222B83FDDCFD84" box="[520,536,490,516]" captionStart="Fig" captionStartId="9.[189,232,2019,2045]" captionTargetBox="[189,1398,1376,1981]" captionTargetId="figure-291@9.[189,1399,1376,1981]" captionTargetPageId="9" captionText="Fig. 4. SEM micrographs of paratypes of Viriiella touaouroua gen. et sp. nov. (MNHN-IM-2000-39461)." figureDoi="http://doi.org/10.5281/zenodo.14186047" httpUri="https://zenodo.org/record/14186047/files/figure.png" pageId="9" pageNumber="284">4</figureCitation>
)
<collectingCountry id="F33D7660FFF2FFA5C9972879FE74FDAA" box="[189,432,528,554]" name="New Caledonia" pageId="9" pageNumber="284">NEW CALEDONIA</collectingCountry>
•&gt;
<specimenCount id="9D2CFD79FFF2FFA5C8CB2866FD40FDA9" box="[481,644,527,554]" count="50" pageId="9" pageNumber="284" type="generic">50 specimens</specimenCount>
; same collection data as for holotype;
<specimenCode id="DB8C9E8BFFF2FFA5CD6C2866FAACFDA9" box="[1094,1384,527,553]" pageId="9" pageNumber="284">MNHN-IM-2000-39461</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C330657BFFF2FFA6C997283AFE1CFE30" lastPageId="10" lastPageNumber="285" pageId="9" pageNumber="284" type="description">
<paragraph id="8B9536F0FFF2FFA5C997283AFE8AFDED" blockId="9.[189,334,595,621]" box="[189,334,595,621]" pageId="9" pageNumber="284">
<heading id="D0DD819CFFF2FFA5C997283AFE8AFDED" bold="true" box="[189,334,595,621]" fontSize="11" level="3" pageId="9" pageNumber="284" reason="3">
<emphasis id="B95EEAE2FFF2FFA5C997283AFE8AFDED" bold="true" box="[189,334,595,621]" pageId="9" pageNumber="284">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B9536F0FFF2FFA5C99728ECFCABFC88" blockId="9.[189,1399,645,777]" pageId="9" pageNumber="284">
<smallCapsWord id="8D73A02CFFF2FFA5C99728ECFEC5FD1E" baselines="664,665" box="[189,257,645,671]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Shell" pageId="9" pageNumber="284">SHELL</smallCapsWord>
(
<figureCitation id="13112A75FFF2FFA5C83828ECFEB8FD1F" box="[274,380,645,671]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="8.[189,232,1963,1989]" captionStartId-1="9.[189,232,2019,2045]" captionTargetBox-0="[189,1398,1320,1925]" captionTargetBox-1="[189,1398,1376,1981]" captionTargetId-0="figure-273@8.[189,1399,1320,1925]" captionTargetId-1="figure-291@9.[189,1399,1376,1981]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="Fig. 3. Types of Viriiella touaouroua gen. et sp. nov. A. Holotype (MNHN-IM-2000-39460). B-D. Paratypes (MNHN-IM-2000-39461)." captionText-1="Fig. 4. SEM micrographs of paratypes of Viriiella touaouroua gen. et sp. nov. (MNHN-IM-2000-39461)." figureDoi-0="http://doi.org/10.5281/zenodo.14186045" figureDoi-1="http://doi.org/10.5281/zenodo.14186047" httpUri-0="https://zenodo.org/record/14186045/files/figure.png" httpUri-1="https://zenodo.org/record/14186047/files/figure.png" pageId="9" pageNumber="284">Figs 3–4</figureCitation>
;
<tableCitation id="C6A8034BFFF2FFA5C8A328ECFE26FD1F" box="[393,482,645,671]" captionStart="Table 2" captionStartId="14.[189,259,265,291]" captionTargetPageId="14" captionText="Table 2. Measurements of 22 shells. Measurements in mm. Abbreviations: AH = aperture height; AW =aperture width; BWW= body whorl width; cv = coefficient of variation; max = maximum; min = minimum; sd = standard deviation; SH = shell height; SW =shell width; W= number of whorls." pageId="9" pageNumber="284">Table 2</tableCitation>
). Small (&lt;
<quantity id="4CD29B15FFF2FFA5CB4828ECFD79FD1F" box="[610,701,645,671]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="9" pageNumber="284" unit="mm" value="1.3">1.3 mm</quantity>
high), cylindrical to cylindro-conical, 1.8 to 2 times higher than wide, not more than four rather flat whorls; aperture semi-lunar, not extending far beyond the outline of the spire, lip continuous, slightly thickened, palatal thinner, apical angle acute, orthocline; umbilicus narrow; smooth protoconch with 1–1.25 whorls.
</paragraph>
<paragraph id="8B9536F0FFF2FFA5C997295BFB3BFCCD" blockId="9.[189,1279,818,845]" box="[189,1279,818,845]" pageId="9" pageNumber="284">
<smallCapsWord id="8D73A02CFFF2FFA5C997295BFE86FCCB" baselines="837,838" box="[189,322,818,844]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Operculum" pageId="9" pageNumber="284">OPERCULUM</smallCapsWord>
(N=6). Ovate, thin, yellow, paucispiral, nucleus excentric, without smear or peg.
</paragraph>
<paragraph id="8B9536F0FFF2FFA5C997291EFE43FC33" blockId="9.[189,1399,886,947]" pageId="9" pageNumber="284">
<smallCapsWord id="8D73A02CFFF2FFA5C997291EFEEBFC0F" baselines="906,906" box="[189,303,887,913]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="External" pageId="9" pageNumber="284">EXTERNAL</smallCapsWord>
<smallCapsWord id="8D73A02CFFF2FFA5C81D2915FE59FC0F" baselines="906" box="[311,413,892,911]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="features" pageId="9" pageNumber="284">FEATURES</smallCapsWord>
(N=30). Epidermis without pigment; eyes black (
<figureCitation id="13112A75FFF2FFA5CADF291FFBF9FC10" box="[1013,1085,886,912]" captionStart="Fig" captionStartId="8.[189,232,1963,1989]" captionTargetBox="[189,1398,1320,1925]" captionTargetId="figure-273@8.[189,1399,1320,1925]" captionTargetPageId="8" captionText="Fig. 3. Types of Viriiella touaouroua gen. et sp. nov. A. Holotype (MNHN-IM-2000-39460). B-D. Paratypes (MNHN-IM-2000-39461)." figureDoi="http://doi.org/10.5281/zenodo.14186045" httpUri="https://zenodo.org/record/14186045/files/figure.png" pageId="9" pageNumber="284">Fig. 3</figureCitation>
), small; cephalic tentacles without ciliation.
</paragraph>
<paragraph id="8B9536F0FFF2FFA5C99729B7FBCDFC78" blockId="9.[189,1033,989,1016]" box="[189,1033,989,1016]" pageId="9" pageNumber="284">
<smallCapsWord id="8D73A02CFFF2FFA5C99729B7FEDEFC76" baselines="1009,1009" box="[189,282,990,1016]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Mantle" pageId="9" pageNumber="284">MANTLE</smallCapsWord>
<smallCapsWord id="8D73A02CFFF2FFA5C808298AFEA9FC76" baselines="1009" box="[290,365,995,1014]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="cavity" pageId="9" pageNumber="284">CAVITY</smallCapsWord>
(N =2). Ctenidium with 10 filaments; osphradium ovate.
</paragraph>
<paragraph id="8B9536F0FFF2FFA5C9972E4BFA95FB02" blockId="9.[189,1399,1057,1154]" pageId="9" pageNumber="284">
<smallCapsWord id="8D73A02CFFF2FFA5C9972E4BFEEAFBBA" baselines="1077,1077" box="[189,302,1058,1084]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Digestive" pageId="9" pageNumber="284">DIGESTIVE</smallCapsWord>
<smallCapsWord id="8D73A02CFFF2FFA5C8132E4EFE4EFBBA" baselines="1077" box="[313,394,1063,1082]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="system" pageId="9" pageNumber="284">SYSTEM</smallCapsWord>
. Radula formula R: 4–5 1 4–5 / 1 1, L: 3–4 1 6–7, M1: 17–19, M2: 14–15 (N= 1;
<figureCitation id="13112A75FFF2FFA5C9972E2CFEC7FBE0" box="[189,259,1093,1120]" captionStart="Fig" captionStartId="10.[191,234,1963,1989]" captionTargetBox="[296,1293,1177,1925]" captionTargetId="figure-306@10.[296,1293,1177,1925]" captionTargetPageId="10" captionText="Fig. 5. Radula of Viriiella touaouroua gen. et sp. nov. Abbreviations: L=lateral teeth; M1 =inner marginal teeth; M2 =outer marginal teeth; R =rhachis (central) teeth." figureDoi="http://doi.org/10.5281/zenodo.14186049" httpUri="https://zenodo.org/record/14186049/files/figure.png" pageId="9" pageNumber="284">Fig. 5</figureCitation>
); stomach with two equally sized chambers, proximal without caecum (N=5); pallial rectal loop in females wide adjoining the pallial oviduct, narrower in males, not adjoining the prostate (N=30).
</paragraph>
<paragraph id="8B9536F0FFF2FFA5C9972EC5FD42FAB0" blockId="9.[188,1399,1196,1328]" pageId="9" pageNumber="284">
<smallCapsWord id="8D73A02CFFF2FFA5C9972EC5FED0FB45" baselines="1215,1216" box="[189,276,1196,1222]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Female" pageId="9" pageNumber="284">FEMALE</smallCapsWord>
<smallCapsWord id="8D73A02CFFF2FFA5C8302ED8FE43FB44" baselines="1215" box="[282,391,1201,1220]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="genitalia" pageId="9" pageNumber="284">GENITALIA</smallCapsWord>
(N =3;
<figureCitation id="13112A75FFF2FFA5C8CB2EC5FDE2FB46" box="[481,550,1196,1222]" captionStart="Fig" captionStartId="11.[189,232,1892,1918]" captionTargetBox="[189,1398,1132,1855]" captionTargetId="figure-334@11.[189,1399,1132,1855]" captionTargetPageId="11" captionText="Fig. 6. Female genitalia of Viriiella touaouroua gen. et sp. nov. A. All parts opaque. B. Oviduct transparent revealing the seminal receptacle. Abbreviations: ac= anterior capsule gland; ag=albumen gland; bc =bursa copulatrix; od =oviduct; ov =ovary; pc =posterior capsule gland; rs =receptaculum semins; vc =ventral channel." figureDoi="http://doi.org/10.5281/zenodo.14186051" httpUri="https://zenodo.org/record/14186051/files/figure.png" pageId="9" pageNumber="284">Fig. 6</figureCitation>
; Supp. file 1). Ovary a simple sack, starting at 1.5 whorls below the apex, comprising about ⅓ whorls, not reaching stomach; receptaculum seminis elongate, small with very short duct; bursa copulatrix large, elongate, club-shaped, reaching far behind albumen gland; anterior capsule gland much smaller than posterior one.
</paragraph>
<caption id="DF556678FFF2FFA5C9972D8AFAB3F87D" ID-DOI="http://doi.org/10.5281/zenodo.14186047" ID-Zenodo-Dep="14186047" box="[189,1399,2019,2045]" httpUri="https://zenodo.org/record/14186047/files/figure.png" pageId="9" pageNumber="284" startId="9.[189,232,2019,2045]" targetBox="[189,1398,1376,1981]" targetPageId="9">
<paragraph id="8B9536F0FFF2FFA5C9972D8AFAB3F87D" blockId="9.[189,1399,2019,2045]" box="[189,1399,2019,2045]" pageId="9" pageNumber="284">
<emphasis id="B95EEAE2FFF2FFA5C9972D8AFECDF87D" bold="true" box="[189,265,2019,2045]" pageId="9" pageNumber="284">Fig. 4.</emphasis>
SEM micrographs of paratypes of
<taxonomicName id="4C2A4D73FFF2FFA5CBB32D8AFC42F87D" authorityName="Schröder &amp; Schächinger &amp; Bouchet &amp; Haase" authorityYear="2024" box="[665,902,2019,2045]" class="Gastropoda" family="Tateidae" genus="Viriiella" kingdom="Animalia" order="Littorinimorpha" pageId="9" pageNumber="284" phylum="Mollusca" rank="species" species="touaouroua" status="gen. et sp. nov.">
<emphasis id="B95EEAE2FFF2FFA5CBB32D8AFC42F87D" box="[665,902,2019,2045]" italics="true" pageId="9" pageNumber="284">Viriiella touaouroua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26D5799FFF2FFA5CAA72D8AFBFCF87D" box="[909,1080,2019,2045]" pageId="9" pageNumber="284" rank="species">gen. et sp. nov.</taxonomicNameLabel>
(MNHN-IM-2000-39461).
</paragraph>
</caption>
<paragraph id="8B9536F0FFF1FFA6C9972B60FE1CFE30" blockId="10.[187,1400,265,432]" pageId="10" pageNumber="285">
<smallCapsWord id="8D73A02CFFF1FFA6C9972B60FF3BFEA2" baselines="284,285" box="[189,255,265,291]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Male" pageId="10" pageNumber="285">MALE</smallCapsWord>
<smallCapsWord id="8D73A02CFFF1FFA6C82F2B67FEB6FEA1" baselines="284" box="[261,370,270,289]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="genitalia" pageId="10" pageNumber="285">GENITALIA</smallCapsWord>
(N=5;
<figureCitation id="13112A75FFF1FFA6C8E12B60FDF6FEA3" box="[459,562,265,291]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="12.[189,232,1453,1479]" captionStartId-1="12.[399,442,1998,2024]" captionTargetBox-0="[352,1220,265,1415]" captionTargetBox-1="[189,1399,1590,1960]" captionTargetId-0="figure-1@12.[352,1222,265,1417]" captionTargetId-1="figure-2@12.[189,1399,1590,1960]" captionTargetPageId-0="12" captionTargetPageId-1="12" captionText-0="Fig. 7. Male genitalia of Viriiella touaouroua gen. et sp. nov. Abbreviations: pe=penis; pr =prostate; pv =pallial vas deferens; te =testis; vd =vas deferens; vs= vesicula seminalis." captionText-1="Fig. 8. Penes of three males of Viriiella touaouroua gen. et sp. nov." figureDoi-0="http://doi.org/10.5281/zenodo.14186053" figureDoi-1="http://doi.org/10.5281/zenodo.14186055" httpUri-0="https://zenodo.org/record/14186053/files/figure.png" httpUri-1="https://zenodo.org/record/14186055/files/figure.png" pageId="10" pageNumber="285">Figs 7–8</figureCitation>
; Supp. file 2). Testis lobate, starting 0.75 whorls below apex, comprising up to 0.75 whorls, almost reaching stomach; vesicula seminalis below testis fairly small; vas deferens proceeding along stomach reaching kidney-shaped prostate ventrally, distal vas deferens leaving prostate at anterior tip, initially straight then undulating before reaching penis; penis simple, elongate, slender, simple, distal end blunt.
</paragraph>
</subSubSection>
<subSubSection id="C330657BFFF1FFA7C9972BB3FAD6FE0D" lastPageId="11" lastPageNumber="286" pageId="10" pageNumber="285" type="discussion">
<paragraph id="8B9536F0FFF1FFA6C9972BB3FEEBFE74" blockId="10.[189,303,474,500]" box="[189,303,474,500]" pageId="10" pageNumber="285">
<heading id="D0DD819CFFF1FFA6C9972BB3FEEBFE74" bold="true" box="[189,303,474,500]" fontSize="11" level="3" pageId="10" pageNumber="285" reason="3">
<emphasis id="B95EEAE2FFF1FFA6C9972BB3FEEBFE74" bold="true" box="[189,303,474,500]" pageId="10" pageNumber="285">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B9536F0FFF1FFA6C9972865FC59FCC0" blockId="10.[185,1399,524,832]" pageId="10" pageNumber="285">
<taxonomicName id="4C2A4D73FFF1FFA6C9972865FE6CFDA6" authorityName="Schröder &amp; Schächinger &amp; Bouchet &amp; Haase" authorityYear="2024" box="[189,424,524,550]" class="Gastropoda" family="Tateidae" genus="Viriiella" kingdom="Animalia" order="Littorinimorpha" pageId="10" pageNumber="285" phylum="Mollusca" rank="species" species="touaouroua" status="gen. et sp. nov.">
<emphasis id="B95EEAE2FFF1FFA6C9972865FE6CFDA6" box="[189,424,524,550]" italics="true" pageId="10" pageNumber="285">Viriiella touaouroua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26D5799FFF1FFA6C8812865FD94FDA6" box="[427,592,524,550]" pageId="10" pageNumber="285" rank="species">gen. et sp.nov.</taxonomicNameLabel>
is the smallest freshwater gastropod so far known from
<collectingCountry id="F33D7660FFF1FFA6CD972865FAB7FDA6" box="[1213,1395,524,550]" name="New Caledonia" pageId="10" pageNumber="285">New Caledonia</collectingCountry>
.
<taxonomicName id="4C2A4D73FFF1FFA6C9932846FD2EFDC9" authority="Haase &amp; Bouchet, 1998" authorityName="Haase &amp; Bouchet" authorityYear="1998" box="[185,746,559,586]" class="Gastropoda" family="Tateidae" genus="Leiorhagium" kingdom="Animalia" order="Littorinimorpha" pageId="10" pageNumber="285" phylum="Mollusca" rank="species" species="granum">
<emphasis id="B95EEAE2FFF1FFA6C9932846FE72FDC9" box="[185,438,559,585]" italics="true" pageId="10" pageNumber="285">Leiorhagium granum</emphasis>
<bibRefCitation id="EFBB4B01FFF1FFA6C8EF2859FD2EFDC9" author="Haase M. &amp; Bouchet P." box="[453,746,559,586]" pageId="10" pageNumber="285" pagination="43 - 129" refId="ref7965" refString="Haase M. &amp; Bouchet P. 1998. Radiation of crenobiontic gastropods on an ancient continental island: the Hemistomia - clade in New Caledonia (Gastropoda: Hydrobiidae). Hydrobiologia 367: 43 - 129. https: // doi. org / 10.1023 / A: 1003219931171" type="journal article" year="1998">Haase &amp; Bouchet, 1998</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C2A4D73FFF1FFA6CA052859FB37FDC9" authority="Haase &amp; Bouchet, 1998" authorityName="Haase &amp; Bouchet" authorityYear="1998" box="[815,1267,559,586]" class="Gastropoda" family="Tateidae" genus="Leiorhagium" kingdom="Animalia" order="Littorinimorpha" pageId="10" pageNumber="285" phylum="Mollusca" rank="species" species="granulum">
<emphasis id="B95EEAE2FFF1FFA6CA052859FC07FDC9" box="[815,963,559,585]" italics="true" pageId="10" pageNumber="285">L. granulum</emphasis>
<bibRefCitation id="EFBB4B01FFF1FFA6CAE42859FB37FDC9" author="Haase M. &amp; Bouchet P." box="[974,1267,559,586]" pageId="10" pageNumber="285" pagination="43 - 129" refId="ref7965" refString="Haase M. &amp; Bouchet P. 1998. Radiation of crenobiontic gastropods on an ancient continental island: the Hemistomia - clade in New Caledonia (Gastropoda: Hydrobiidae). Hydrobiologia 367: 43 - 129. https: // doi. org / 10.1023 / A: 1003219931171" type="journal article" year="1998">Haase &amp; Bouchet, 1998</bibRefCitation>
</taxonomicName>
, both only known by their shells, thus with tentative generic allocation, measure at least
<quantity id="4CD29B15FFF1FFA6CD65283BFB6DFDED" box="[1103,1193,594,621]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="285" unit="mm" value="1.5">1.5 mm</quantity>
in height and are much more conical (
<bibRefCitation id="EFBB4B01FFF1FFA6C886281FFD06FD10" author="Haase M. &amp; Bouchet P." box="[428,706,629,656]" pageId="10" pageNumber="285" pagination="43 - 129" refId="ref7965" refString="Haase M. &amp; Bouchet P. 1998. Radiation of crenobiontic gastropods on an ancient continental island: the Hemistomia - clade in New Caledonia (Gastropoda: Hydrobiidae). Hydrobiologia 367: 43 - 129. https: // doi. org / 10.1023 / A: 1003219931171" type="journal article" year="1998">Haase &amp; Bouchet 1998</bibRefCitation>
). The
<typeStatus id="54918852FFF1FFA6CA20281FFCF9FD10" box="[778,829,630,656]" pageId="10" pageNumber="285">type</typeStatus>
locality lies in a narrow strip of alluvial deposits between the coast and a range of ultramafic hills (see https://georep.nc/explorateur-cartographique provided by the Gouvernement de la Nouvelle Calédonie). These slope colluviums come from the dismantling of the peridotite massif located directly to the West of the
<typeStatus id="54918852FFF1FFA6CADC2889FBEDFD7A" box="[1014,1065,736,762]" pageId="10" pageNumber="285">type</typeStatus>
locality and form an aquifer at the foot of the massif (probably temporary and perched), which is fed by both the drainage of the massif and also by the creeks of the area (L. Russ, pers. com.).
</paragraph>
<paragraph id="8B9536F0FFF1FFA6C9972902FE24FC08" blockId="10.[189,480,875,904]" box="[189,480,875,904]" pageId="10" pageNumber="285">
<emphasis id="B95EEAE2FFF1FFA6C9972902FE24FC08" bold="true" box="[189,480,875,904]" italics="true" pageId="10" pageNumber="285">Phylogenetic analyses</emphasis>
</paragraph>
<paragraph id="8B9536F0FFF1FFA7C99729F6FAD6FE0D" blockId="10.[188,1399,927,1129]" lastBlockId="11.[188,1399,265,397]" lastPageId="11" lastPageNumber="286" pageId="10" pageNumber="285">
Both ML and BA yielded almost identical topologies (
<figureCitation id="13112A75FFF1FFA6CA1529F6FC43FC39" box="[831,903,927,953]" captionStart="Fig" captionStartId="13.[189,232,1892,1918]" captionTargetBox="[335,1221,746,1807]" captionTargetId="figure-173@13.[332,1232,743,1843]" captionTargetPageId="13" captionText="Fig. 9. Phylogenetic analyses. Maximum likelihood (ML) tree and section of Bayesian tree (BA). Support values are bootstrap support values/posterior probabilities; the latter are omitted for the New Caledonian taxa in the ML tree, the former in the Bayesian tree fragment. Scale bars=0.1 substitutions/ site." figureDoi="http://doi.org/10.5281/zenodo.14186057" httpUri="https://zenodo.org/record/14186057/files/figure.png" pageId="10" pageNumber="285">Fig. 9</figureCitation>
). The ingroup, i.e.,
<taxonomicName id="4C2A4D73FFF1FFA6CD5A29F6FB12FC39" authorityName="Thiele" authorityYear="1925" box="[1136,1238,927,953]" class="Gastropoda" family="Tateidae" kingdom="Animalia" order="Littorinimorpha" pageId="10" pageNumber="285" phylum="Mollusca" rank="family">Tateidae</taxonomicName>
, received full support. A clade containing
<taxonomicName id="4C2A4D73FFF1FFA6CB2B29ABFC02FC5D" authority="Ponder &amp; Haase, 2005" authorityName="Ponder &amp; Haase" authorityYear="2005" box="[513,966,962,989]" class="Gastropoda" family="Tateidae" genus="Sulawesidrobia" kingdom="Animalia" order="Littorinimorpha" pageId="10" pageNumber="285" phylum="Mollusca" rank="genus">
<emphasis id="B95EEAE2FFF1FFA6CB2B29ABFD72FC5C" box="[513,694,962,988]" italics="true" pageId="10" pageNumber="285">Sulawesidrobia</emphasis>
Ponder &amp;
<bibRefCitation id="EFBB4B01FFF1FFA6CA1F29AAFC02FC5D" author="Haase M. &amp; Gargominy O. &amp; Fontaine B." box="[821,966,962,989]" pageId="10" pageNumber="285" pagination="145 - 163" refId="ref8061" refString="Haase M., Gargominy O. &amp; Fontaine B. 2005. Rissooidean freshwater gastropods from the middle of the Pacific: the genus Fluviopupa on the Austral Islands (Caenogastropoda). Molluscan Research 25: 145 - 163." type="journal article" year="2005">Haase, 2005</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C2A4D73FFF1FFA6CAD629AAFA87FC5C" authority="Tenison-Woods, 1879" authorityName="Tenison-Woods" authorityYear="1879" box="[1020,1347,962,988]" class="Gastropoda" family="Tateidae" genus="Tatea" kingdom="Animalia" order="Littorinimorpha" pageId="10" pageNumber="285" phylum="Mollusca" rank="genus">
<emphasis id="B95EEAE2FFF1FFA6CAD629AAFBF8FC5C" box="[1020,1084,963,988]" italics="true" pageId="10" pageNumber="285">Tatea</emphasis>
Tenison-Woods, 1879
</taxonomicName>
was sister group to all remaining taxa, which formed two larger clades, both with two subclades. One clade contained all species of
<taxonomicName id="4C2A4D73FFF1FFA6C8FA2E61FD93FBA2" authorityName="Pilsbry" authorityYear="1911" box="[464,599,1032,1058]" class="Gastropoda" family="Tateidae" genus="Fluviopupa" kingdom="Animalia" order="Littorinimorpha" pageId="10" pageNumber="285" phylum="Mollusca" rank="genus">
<emphasis id="B95EEAE2FFF1FFA6C8FA2E61FD93FBA2" box="[464,599,1032,1058]" italics="true" pageId="10" pageNumber="285">Fluviopupa</emphasis>
</taxonomicName>
and all species from
<collectingCountry id="F33D7660FFF1FFA6CA672E60FC28FBA2" box="[845,1004,1032,1059]" name="New Zealand" pageId="10" pageNumber="285">New Zealand</collectingCountry>
, while in the other one, Australian genera were sister to all species from
<collectingCountry id="F33D7660FFF1FFA6CB542E45FCFDFBC6" box="[638,825,1068,1094]" name="New Caledonia" pageId="10" pageNumber="285">New Caledonia</collectingCountry>
. Within clades, there were only two differences: 1) in the relationships of the
<collectingCountry id="F33D7660FFF1FFA6CB302E26FD7CFBE9" box="[538,696,1103,1129]" name="New Zealand" pageId="10" pageNumber="285">New Zealand</collectingCountry>
genera
<taxonomicName id="4C2A4D73FFF1FFA6CA332E26FB84FBE9" authority="Haase, 2008" authorityName="Haase" authorityYear="2008" box="[793,1088,1103,1129]" class="Gastropoda" family="Tateidae" genus="Halopyrgus" kingdom="Animalia" order="Littorinimorpha" pageId="10" pageNumber="285" phylum="Mollusca" rank="genus">
<emphasis id="B95EEAE2FFF1FFA6CA332E26FC66FBE9" box="[793,930,1103,1129]" italics="true" pageId="10" pageNumber="285">Halopyrgus</emphasis>
<bibRefCitation id="EFBB4B01FFF1FFA6CA802E26FB84FBE9" author="Haase M." box="[938,1088,1103,1129]" pageId="10" pageNumber="285" pagination="99 - 159" refId="ref7915" refString="Haase M. 2008. The radiation of hydrobiid gastropods in New Zealand: a revision including the description of new species based on morphology and mtDNA sequence information. Systematics and Biodiversity 6: 99 - 159. https: // doi. org / 10.1017 / S 1477200007002630" type="journal article" year="2008">Haase, 2008</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C2A4D73FFF1FFA7CD652E39FF3EFEA4" authority="Stimpson, 1865" authorityName="Stimpson" authorityYear="1865" class="Gastropoda" family="Tateidae" genus="Potamopyrgus" kingdom="Animalia" lastPageId="11" lastPageNumber="286" order="Littorinimorpha" pageId="10" pageNumber="285" phylum="Mollusca" rank="genus">
<emphasis id="B95EEAE2FFF1FFA6CD652E39FB3DFBE9" box="[1103,1273,1104,1129]" italics="true" pageId="10" pageNumber="285">Potamopyrgus</emphasis>
Stimpson, 1865
</taxonomicName>
, and
<taxonomicName id="4C2A4D73FFF0FFA7C81E2B60FDA3FEA3" authority="Haase, 2008" authorityName="Haase" authorityYear="2008" box="[308,615,265,292]" class="Gastropoda" family="Tateidae" genus="Sororipyrgus" kingdom="Animalia" order="Littorinimorpha" pageId="11" pageNumber="286" phylum="Mollusca" rank="genus">
<emphasis id="B95EEAE2FFF0FFA7C81E2B60FE09FEA3" box="[308,461,265,291]" italics="true" pageId="11" pageNumber="286">Sororipyrgus</emphasis>
<bibRefCitation id="EFBB4B01FFF0FFA7C8FE2B63FDA3FEA3" author="Haase M." box="[468,615,265,292]" pageId="11" pageNumber="286" pagination="99 - 159" refId="ref7915" refString="Haase M. 2008. The radiation of hydrobiid gastropods in New Zealand: a revision including the description of new species based on morphology and mtDNA sequence information. Systematics and Biodiversity 6: 99 - 159. https: // doi. org / 10.1017 / S 1477200007002630" type="journal article" year="2008">Haase, 2008</bibRefCitation>
</taxonomicName>
; 2) and most relevant for the present account, the relationship of the new genus
<taxonomicName id="4C2A4D73FFF0FFA7C86A2B45FE64FEC6" authorityName="Schröder &amp; Schächinger &amp; Bouchet &amp; Haase" authorityYear="2024" box="[320,416,300,326]" class="Gastropoda" family="Tateidae" genus="Viriiella" kingdom="Animalia" order="Littorinimorpha" pageId="11" pageNumber="286" phylum="Mollusca" rank="genus">
<emphasis id="B95EEAE2FFF0FFA7C86A2B45FE64FEC6" box="[320,416,300,326]" italics="true" pageId="11" pageNumber="286">Viriiella</emphasis>
</taxonomicName>
. In ML,
<taxonomicName id="4C2A4D73FFF0FFA7CB222B44FD69FEC6" authorityName="Schröder &amp; Schächinger &amp; Bouchet &amp; Haase" authorityYear="2024" box="[520,685,301,326]" class="Gastropoda" family="Tateidae" genus="Viriiella" kingdom="Animalia" order="Littorinimorpha" pageId="11" pageNumber="286" phylum="Mollusca" rank="species" species="touaouroua" status="gen. et sp. nov.">
<emphasis id="B95EEAE2FFF0FFA7CB222B44FD69FEC6" box="[520,685,301,326]" italics="true" pageId="11" pageNumber="286">V. touaouroua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26D5799FFF0FFA7CB9E2B44FCA7FEC7" box="[692,867,301,327]" pageId="11" pageNumber="286" rank="species">gen. et sp. nov.</taxonomicNameLabel>
was sister species of
<taxonomicName id="4C2A4D73FFF0FFA7CD742B44FE22FEEA" authority="(Haase &amp; Bouchet 1998)" authorityName="KJ" baseAuthorityName="Haase &amp; Bouchet" baseAuthorityYear="1998" class="Gastropoda" family="Tateidae" genus="Crosseana" kingdom="Animalia" order="Littorinimorpha" pageId="11" pageNumber="286" phylum="Mollusca" rank="species" species="melanosoma">
<emphasis id="B95EEAE2FFF0FFA7CD742B44FAB2FEC6" box="[1118,1398,300,327]" italics="true" pageId="11" pageNumber="286">Crosseana melanosoma</emphasis>
(
<bibRefCitation id="EFBB4B01FFF0FFA7C9EF2B39FE19FEEA" author="Haase M. &amp; Bouchet P." box="[197,477,335,362]" pageId="11" pageNumber="286" pagination="43 - 129" refId="ref7965" refString="Haase M. &amp; Bouchet P. 1998. Radiation of crenobiontic gastropods on an ancient continental island: the Hemistomia - clade in New Caledonia (Gastropoda: Hydrobiidae). Hydrobiologia 367: 43 - 129. https: // doi. org / 10.1023 / A: 1003219931171" type="journal article" year="1998">Haase &amp; Bouchet 1998</bibRefCitation>
)
</taxonomicName>
, in BA of
<taxonomicName id="4C2A4D73FFF0FFA7CB4B2B26FB50FEEA" authority="Haase &amp; Bouchet, 1998" authorityName="Haase &amp; Bouchet" authorityYear="1998" box="[609,1172,335,362]" class="Gastropoda" family="Tateidae" genus="Leiorhagium" kingdom="Animalia" order="Littorinimorpha" pageId="11" pageNumber="286" phylum="Mollusca" rank="species" species="kavuneva">
<emphasis id="B95EEAE2FFF0FFA7CB4B2B26FCABFEE9" box="[609,879,335,361]" italics="true" pageId="11" pageNumber="286">Leiorhagium kavuneva</emphasis>
<bibRefCitation id="EFBB4B01FFF0FFA7CA5C2B39FB50FEEA" author="Haase M. &amp; Bouchet P." box="[886,1172,335,362]" pageId="11" pageNumber="286" pagination="43 - 129" refId="ref7965" refString="Haase M. &amp; Bouchet P. 1998. Radiation of crenobiontic gastropods on an ancient continental island: the Hemistomia - clade in New Caledonia (Gastropoda: Hydrobiidae). Hydrobiologia 367: 43 - 129. https: // doi. org / 10.1023 / A: 1003219931171" type="journal article" year="1998">Haase &amp; Bouchet, 1998</bibRefCitation>
</taxonomicName>
. Both relationships were only insignificantly supported. Some of the deeper nodes were not well supported, either.
</paragraph>
</subSubSection>
</treatment>
</document>
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