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<document id="52F239518A9EB1CE24CD144B92961D24" ID-DOI="10.1080/00222933.2011.622057" ID-ISSN="1464-5262" ID-Zenodo-Dep="5204903" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629069748535" checkinUser="felipe" docAuthor="Back, Jinwook, Lee, Wonchoel &amp; Huys, Rony" docDate="2011" docId="03EA8794A131965DFDB7FB4DFE889B98" docLanguage="en" docName="JNatHist.45.47-48.2939-2964.pdf" docOrigin="Journal of Natural History 45 (47 - 48)" docSource="http://dx.doi.org/10.1080/00222933.2011.622057" docStyle="DocumentStyle:DE8A86AD4006EFA56F95B2AAEFBF2813.5:JNatHist.2008-2012.journal_article.2cover" docStyleId="DE8A86AD4006EFA56F95B2AAEFBF2813" docStyleName="JNatHist.2008-2012.journal_article.2cover" docStyleVersion="5" docTitle="Remanea naksanensis Back &amp; Lee &amp; Huys 2011, sp. nov." docType="treatment" docVersion="3" lastPageNumber="2948" masterDocId="FFD3FFECA1359656FFB8FFC9FFB49F5F" masterDocTitle="A new species of Remanea Klie, 1929 (Copepoda: Harpacticoida: Paramesochridae) with a redescription of the type species" masterLastPageNumber="2964" masterPageNumber="2939" pageNumber="2941" updateTime="1699059045482" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="CC04389E71751F01B290B125BF6A5519" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="838BAE60075769D5A36F0588C8E8ED69">A new species of Remanea Klie, 1929 (Copepoda: Harpacticoida: Paramesochridae) with a redescription of the type species</mods:title>
</mods:titleInfo>
<mods:name id="C4DC8E67BD8C3C2890916EA5404F7848" type="personal">
<mods:role id="6F6D308B900EA6EF4294EA7E6EC14DBC">
<mods:roleTerm id="A5ABB29C056DE3F6620D8A94A6B4D7F2">Author</mods:roleTerm>
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<mods:namePart id="5D72E18FB2122B813D7878DE6249E840">Back, Jinwook</mods:namePart>
</mods:name>
<mods:name id="32AF1250EB7C5E9B94E9A704FE3C04BD" type="personal">
<mods:role id="D668B14AA5DD1D54AFCE2E16F67E15CC">
<mods:roleTerm id="923AA0CB1C0FD00B6B5B7928EC4819D0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8F8D84DD0B4A3B015B867F703A805973">Lee, Wonchoel</mods:namePart>
</mods:name>
<mods:name id="710CA9435D3C0BE1ABD384CAC2C8B330" type="personal">
<mods:role id="543444AB144770F5284011C2EAA2F922">
<mods:roleTerm id="7FEE7A4A2F18322BBA0D168B75690836">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DC9CA17301DDF439AF3E767D05103440">Huys, Rony</mods:namePart>
</mods:name>
<mods:typeOfResource id="D10208DD34C2466B57118C0590EDE09A">text</mods:typeOfResource>
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<mods:titleInfo id="69C8BC1F434FD9E9AD322AD5E1850CF2">
<mods:title id="27AC479930ED72D9CFCAF81F57907B99">Journal of Natural History</mods:title>
</mods:titleInfo>
<mods:part id="9ED4D2B2337A22428CFA0781B54E4BBD">
<mods:date id="65C7E272A9D1B901D18F1C06C507E9D4">2011</mods:date>
<mods:detail id="0E3E12B0D339153C0008BD4DE7D3C3F2" type="pubDate">
<mods:number id="58A0D97C007DCFF59D05D030F9E686D6">2011-12-31</mods:number>
</mods:detail>
<mods:detail id="4DC2B555941B62DE96EAF7E723B27E11" type="volume">
<mods:number id="DBB40424045E4935882A9BC7DE60B549">45</mods:number>
</mods:detail>
<mods:detail id="A190BF3ED37B66CF16E21007074984B7" type="issue">
<mods:number id="4DB6DEEF1B0B90927FA30A29DB7D8762">47 - 48</mods:number>
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<mods:start id="AFD02E90A993EB5B75D0A9B654C4439B">2939</mods:start>
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<mods:url id="20B9A54812A7F74F0CD2713E8530F07D">http://dx.doi.org/10.1080/00222933.2011.622057</mods:url>
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<mods:classification id="FA3D45BD0D1643E5533CBC7166A58937">journal article</mods:classification>
<mods:identifier id="DB4F47B2672F1BE6E7E5D978B72A4B1C" type="DOI">10.1080/00222933.2011.622057</mods:identifier>
<mods:identifier id="D163D5993C145A8B601A1220222059BE" type="ISSN">1464-5262</mods:identifier>
<mods:identifier id="EE5DB7BD1336399652490F0EB2D637A9" type="Zenodo-Dep">5204903</mods:identifier>
</mods:mods>
<treatment id="03EA8794A131965DFDB7FB4DFE889B98" LSID="urn:lsid:plazi:treatment:03EA8794A131965DFDB7FB4DFE889B98" httpUri="http://treatment.plazi.org/id/03EA8794A131965DFDB7FB4DFE889B98" lastPageId="11" lastPageNumber="2948" pageId="4" pageNumber="2941">
<subSubSection id="C3596509A1319652FDB7FB4DFCED9BC4" box="[527,857,1155,1180]" pageId="4" pageNumber="2941" type="nomenclature">
<paragraph id="8BFC3682A1319652FDB7FB4DFCED9BC4" blockId="4.[443,940,1092,1211]" box="[527,857,1155,1180]" pageId="4" pageNumber="2941">
<heading id="D0B481EEA1319652FDB7FB4DFCED9BC4" box="[527,857,1155,1180]" centered="true" fontSize="10" level="2" pageId="4" pageNumber="2941" reason="3">
<taxonomicName id="4C434D01A1319652FDB7FB4DFD4A9BC3" authority="Back &amp; Lee &amp; Huys, 2011" authorityName="Back &amp; Lee &amp; Huys" authorityYear="2011" box="[527,766,1155,1180]" class="Hexanauplia" family="Paramesochridae" genus="Remanea" kingdom="Animalia" order="Harpacticoida" pageId="4" pageNumber="2941" phylum="Arthropoda" rank="species" species="naksanensis" status="sp. nov.">
<emphasis id="B937EA90A1319652FDB7FB4DFD4A9BC3" bold="true" box="[527,766,1155,1180]" italics="true" pageId="4" pageNumber="2941">Remanea naksanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20457EBA1319652FCBCFB4DFCED9BC4" box="[772,857,1156,1179]" pageId="4" pageNumber="2941" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3596509A131965DFDD1FB6AFE889B98" lastPageId="11" lastPageNumber="2948" pageId="4" pageNumber="2941" type="description">
<paragraph id="8BFC3682A1319652FDD1FB6AFD4A9BE4" blockId="4.[443,940,1092,1211]" box="[617,766,1187,1211]" pageId="4" pageNumber="2941">
(
<figureCitation id="13782A07A1319652FDC8FB6AFD419BE4" box="[624,757,1187,1211]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStart-3="Figure 4" captionStart-4="Figure 5" captionStart-5="Figure 6" captionStart-6="Figure 7" captionStartId-0="6.[211,278,1561,1583]" captionStartId-1="7.[143,210,1568,1590]" captionStartId-2="9.[143,210,1445,1467]" captionStartId-3="10.[211,278,1584,1606]" captionStartId-4="12.[211,278,1584,1606]" captionStartId-5="13.[143,210,1602,1624]" captionStartId-6="14.[211,278,1682,1704]" captionTargetBox-0="[211,1170,169,1528]" captionTargetBox-1="[143,1102,161,1535]" captionTargetBox-2="[143,1101,162,1412]" captionTargetBox-3="[213,1169,163,1551]" captionTargetBox-4="[211,1170,161,1551]" captionTargetBox-5="[143,1102,163,1568]" captionTargetBox-6="[211,1171,161,1649]" captionTargetId-0="figure-5@6.[211,1170,169,1528]" captionTargetId-1="figure-6@7.[143,1102,161,1535]" captionTargetId-2="figure-6@9.[143,1102,161,1412]" captionTargetId-3="figure-5@10.[211,1170,163,1551]" captionTargetId-4="figure-5@12.[211,1170,161,1551]" captionTargetId-5="figure-6@13.[143,1102,162,1569]" captionTargetId-6="figure-5@14.[211,1171,161,1649]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="9" captionTargetPageId-3="10" captionTargetPageId-4="12" captionTargetPageId-5="13" captionTargetPageId-6="14" captionText-0="Figure 1. Remanea naksanensis sp. nov. Female: (A) Habitus, dorsal; (B) habitus, lateral; (C) urosome (excluding P5-bearing somite), ventral. Scale bars in µm." captionText-1="Figure 2. Remanea naksanensis sp. nov. Female: (A) Antennule, ventral; (B) antenna; (C) pseudoperculum, anal somite and caudal rami, dorsal (inset showing full length of setae IV–V); (D) fifth pair of legs (P5); (E) P6 and genital field. Scale bars in µm." captionText-2="Figure 3. Remanea naksanensis sp. nov. Female: (A) Mandible; (B) maxillule; (C) maxilla [syncoxal endites (1–4) and endopod (5) shown in separate insets; arrow indicates minute process on distal coxal endite]; (D) maxilliped. Scale bars in µm." captionText-3="Figure 4. Remanea naksanensis sp. nov. Female: (A) P1, anterior; (B) P2, anterior; (C) P3, anterior; (D) P4, anterior. Scale bars in µm." captionText-4="Figure 5. Remanea naksanensis sp. nov. Male: (A) Habitus, dorsal; (B) antennule, ventral (arrows indicate position of partial surface sutures on swollen segment 5). Scale bars in µm." captionText-5="Figure 6. Remanea naksanensis sp. nov. Male: (A) P2, anterior; (B) P3, anterior; (C) left P5, anterior; (D) armature of left P6. Scale bars in µm." captionText-6="Figure 7. Remanea naksanensis sp. nov., scanning electron micrographs. Female: (A) Antennary exopod; (B) distal segment of mandibular palp; (C) maxilliped; (D) genital field and P6; (E) P5; (G) pseudoperculum; (H) caudal rami, dorsal. Male: (F) fifth antennulary segment with surface sutures arrowed. Scale bars in µm." figureDoi-0="http://doi.org/10.5281/zenodo.5204905" figureDoi-1="http://doi.org/10.5281/zenodo.5204907" figureDoi-2="http://doi.org/10.5281/zenodo.5204909" figureDoi-3="http://doi.org/10.5281/zenodo.5204911" figureDoi-4="http://doi.org/10.5281/zenodo.5204913" figureDoi-5="http://doi.org/10.5281/zenodo.5204915" figureDoi-6="http://doi.org/10.5281/zenodo.5204917" httpUri-0="https://zenodo.org/record/5204905/files/figure.png" httpUri-1="https://zenodo.org/record/5204907/files/figure.png" httpUri-2="https://zenodo.org/record/5204909/files/figure.png" httpUri-3="https://zenodo.org/record/5204911/files/figure.png" httpUri-4="https://zenodo.org/record/5204913/files/figure.png" httpUri-5="https://zenodo.org/record/5204915/files/figure.png" httpUri-6="https://zenodo.org/record/5204917/files/figure.png" pageId="4" pageNumber="2941">Figures 1–7</figureCitation>
)
</paragraph>
<paragraph id="8BFC3682A1319652FF6BFAC5FED69A7C" blockId="4.[211,354,1291,1315]" box="[211,354,1291,1315]" pageId="4" pageNumber="2941">
<emphasis id="B937EA90A1319652FF6BFAC5FED69A7C" box="[211,354,1291,1315]" italics="true" pageId="4" pageNumber="2941">Type locality</emphasis>
</paragraph>
<paragraph id="8BFC3682A1319652FF6BFAFAFE239AD4" blockId="4.[211,1172,1327,1419]" pageId="4" pageNumber="2941">
<materialsCitation id="3B2B3CDFA1319652FF6BFAFAFE279AD4" country="South Korea" latitude="38.105553" location="Naksan" longLatPrecision="19" longitude="128.6461" pageId="4" pageNumber="2941" specimenCount="1">
<location id="8E9C6059A1319652FF6BFAFAFE9F9A14" LSID="urn:lsid:plazi:treatment:03EA8794A131965DFDB7FB4DFE889B98:8E9C6059A1319652FF6BFAFAFE9F9A14" box="[211,299,1331,1355]" country="South Korea" latitude="38.105553" longLatPrecision="19" longitude="128.6461" name="Naksan" pageId="4" pageNumber="2941">Naksan</location>
sandy beach, east coast of
<collectingCountry id="F3547612A1319652FDDFFAFAFD4D9A14" box="[615,761,1331,1355]" name="South Korea" pageId="4" pageNumber="2941">South Korea</collectingCountry>
(
<geoCoordinate id="EE775045A1319652FCB1FAFAFC239A14" box="[777,919,1327,1355]" degrees="38" direction="north" minutes="06" orientation="latitude" pageId="4" pageNumber="2941" precision="15" seconds="20" value="38.105553">
38
<superScript id="7C369BCAA1319652FC9DFAE6FC9A9A1F" attach="none" box="[805,814,1327,1344]" fontSize="7" pageId="4" pageNumber="2941">◦</superScript>
06
<superScript id="7C369BCAA1319652FCF2FAE6FCFB9A1F" attach="none" box="[842,847,1327,1344]" fontSize="7" pageId="4" pageNumber="2941">′</superScript>
20
<superScript id="7C369BCAA1319652FCD3FAE6FCC29A1F" attach="left" box="[875,886,1327,1344]" fontSize="7" pageId="4" pageNumber="2941">′′</superScript>
N
</geoCoordinate>
,
<geoCoordinate id="EE775045A1319652FC1CFAFAFB8F9A14" box="[932,1083,1327,1355]" degrees="128" direction="east" minutes="38" orientation="longitude" pageId="4" pageNumber="2941" precision="15" seconds="46" value="128.6461">
128
<superScript id="7C369BCAA1319652FC74FAE6FC619A1F" attach="none" box="[972,981,1327,1344]" fontSize="7" pageId="4" pageNumber="2941">◦</superScript>
38
<superScript id="7C369BCAA1319652FC49FAE6FC429A1F" attach="left" box="[1009,1014,1327,1344]" fontSize="7" pageId="4" pageNumber="2941">′</superScript>
46
<superScript id="7C369BCAA1319652FBAAFAE6FBA99A1F" attach="left" box="[1042,1053,1327,1344]" fontSize="7" pageId="4" pageNumber="2941">′′</superScript>
E
</geoCoordinate>
); washings of sandy sediments from a brackish water system (salinity 4–7 psu) near a small unnamed stream
</materialsCitation>
.
</paragraph>
<paragraph id="8BFC3682A1319652FF6BFA12FE119AAC" blockId="4.[211,421,1499,1523]" box="[211,421,1499,1523]" pageId="4" pageNumber="2941">
<emphasis id="B937EA90A1319652FF6BFA12FE119AAC" box="[211,421,1499,1523]" italics="true" pageId="4" pageNumber="2941">Material examined</emphasis>
</paragraph>
<paragraph id="8BFC3682A1319652FF6BF9CBFB279925" blockId="4.[211,1171,1538,1658]" pageId="4" pageNumber="2941">
<materialsCitation id="3B2B3CDFA1319652FF6BF9CBFB279925" collectionCode="NIBR" country="South Korea" location="All" pageId="4" pageNumber="2941" specimenCode="NIBRV0000238524, NIBRV0000238525, NIBRV0000238526, NIBRV0000238527" specimenCount="17" specimenCount-female="15" specimenCount-male="2" typeStatus="holotype">
<typeStatus id="54F88820A1319652FF6BF9CBFE8A9945" box="[211,318,1538,1562]" pageId="4" pageNumber="2941" type="holotype">Holotype</typeStatus>
female (
<specimenCode id="DBE59EF9A1319652FE12F9CBFD3C9945" box="[426,648,1538,1562]" collectionCode="NIBRV" pageId="4" pageNumber="2941">NIBRV0000238524</specimenCode>
) dissected on six slides, and
<typeStatus id="54F88820A1319652FC55F9CAFBE49944" box="[1005,1104,1539,1563]" pageId="4" pageNumber="2941" type="paratype">paratype</typeStatus>
male (
<specimenCode id="DBE59EF9A1319652FF62F9EBFE0C9965" box="[218,440,1570,1594]" collectionCode="NIBRV" pageId="4" pageNumber="2941">NIBRV0000238525</specimenCode>
) dissected on four slides. Additional
<typeStatus id="54F88820A1319652FCCBF9EAFC549964" box="[883,992,1571,1595]" pageId="4" pageNumber="2941" type="paratype">paratypes</typeStatus>
represented by
<specimenCount id="9D45FD0BA1319652FF6BF98BFEFE9905" box="[211,330,1602,1626]" pageId="4" pageNumber="2941" type="female">14 females</specimenCount>
(
<specimenCode id="DBE59EF9A1319652FEE0F98BFD829905" box="[344,566,1602,1626]" collectionCode="NIBRV" pageId="4" pageNumber="2941">NIBRV0000238526</specimenCode>
) and
<specimenCount id="9D45FD0BA1319652FDCEF98BFD429905" box="[630,758,1602,1626]" pageId="4" pageNumber="2941" type="male">three males</specimenCount>
(
<specimenCode id="DBE59EF9A1319652FCBCF98BFC559905" box="[772,993,1602,1626]" collectionCode="NIBRV" pageId="4" pageNumber="2941">NIBRV0000238527</specimenCode>
) in ethanol.
<location id="8E9C6059A1319652FBC8F98BFB279905" LSID="urn:lsid:plazi:treatment:03EA8794A131965DFDB7FB4DFE889B98:8E9C6059A1319652FBC8F98BFB279905" box="[1136,1171,1602,1626]" country="South Korea" name="All" pageId="4" pageNumber="2941">All</location>
specimens were deposited in the
<collectionCode id="ED52AE47A1319652FD82F9AAFD379925" box="[570,643,1635,1658]" country="South Korea" httpUri="http://grbio.org/cool/r4iq-k5jx" name="National Institute of Biological Resources" pageId="4" pageNumber="2941">NIBR</collectionCode>
,
<collectingCountry id="F3547612A1319652FD35F9AAFD629925" box="[653,726,1635,1658]" name="South Korea" pageId="4" pageNumber="2941">Korea</collectingCountry>
;
<specimenCount id="9D45FD0BA1319652FD67F9AAFCC39925" box="[735,887,1634,1658]" pageId="4" pageNumber="2941" type="female">seven females</specimenCount>
and
<specimenCount id="9D45FD0BA1319652FC14F9ABFB959925" box="[940,1057,1634,1658]" pageId="4" pageNumber="2941" type="male">four males</specimenCount>
were used
</materialsCitation>
</paragraph>
<paragraph id="8BFC3682A1309653FF37FF6EFC559F80" blockId="5.[143,1103,167,255]" pageId="5" pageNumber="2942">
for scanning electron microscopy while
<specimenCount id="9D45FD0BA1309653FDF4FF6EFD779FE0" box="[588,707,167,191]" pageId="5" pageNumber="2942" type="female">12 females</specimenCount>
were used for DNA extraction. All specimens were collected from the
<typeStatus id="54F88820A1309653FDABFF01FDF79FBF" box="[531,579,200,224]" pageId="5" pageNumber="2942">type</typeStatus>
locality by J. Back on
<date id="FFFD1042A1309653FCF9FF0EFC699F80" box="[833,989,199,223]" pageId="5" pageNumber="2942" value="2010-04-23">23 April 2010</date>
.
</paragraph>
<paragraph id="8BFC3682A1309653FF02FF2EFBB49FA0" blockId="5.[143,1103,167,255]" box="[186,1024,231,255]" pageId="5" pageNumber="2942">DNA-barcode (mtCOI) sequences and traces were submitted to GenBank.</paragraph>
<paragraph id="8BFC3682A1309653FF37FEA9FECE9E28" blockId="5.[143,378,351,375]" box="[143,378,351,375]" pageId="5" pageNumber="2942">
<emphasis id="B937EA90A1309653FF37FEA9FECE9E28" box="[143,378,351,375]" italics="true" pageId="5" pageNumber="2942">Description of female</emphasis>
</paragraph>
<paragraph id="8BFC3682A1309653FF37FE4FFE3B9EA1" blockId="5.[143,1104,390,1690]" pageId="5" pageNumber="2942">
Body fusiform, slightly depressed dorsoventrally (
<figureCitation id="13782A07A1309653FD05FE4FFCE79EC1" box="[701,851,390,414]" captionStart="Figure 1" captionStartId="6.[211,278,1561,1583]" captionTargetBox="[211,1170,169,1528]" captionTargetId="figure-5@6.[211,1170,169,1528]" captionTargetPageId="6" captionText="Figure 1. Remanea naksanensis sp. nov. Female: (A) Habitus, dorsal; (B) habitus, lateral; (C) urosome (excluding P5-bearing somite), ventral. Scale bars in µm." figureDoi="http://doi.org/10.5281/zenodo.5204905" httpUri="https://zenodo.org/record/5204905/files/figure.png" pageId="5" pageNumber="2942">Figure 1A, B</figureCitation>
), with sensilla as illustrated. Total body length, 553 µm (
<emphasis id="B937EA90A1309653FD99FE6EFD9A9EE1" box="[545,558,423,446]" italics="true" pageId="5" pageNumber="2942">n</emphasis>
= 15, mean = 541 µm); largest width (153 µm) measured at posterior margin of cephalic shield. Body somites with well-developed arthrodial membranes.
</paragraph>
<paragraph id="8BFC3682A1309653FF02FDCFFE509D01" blockId="5.[143,1104,390,1690]" pageId="5" pageNumber="2942">Cephalothorax and somites bearing P2–P4 with crenulate posterior hyaline frill. Cephalothorax bell-shaped, with several sensilla; pleural areas weakly developed, posterolateral angles rounded.</paragraph>
<paragraph id="8BFC3682A1309653FF02FDAFFEDC9C02" blockId="5.[143,1104,390,1690]" pageId="5" pageNumber="2942">
Urosomites (
<figureCitation id="13782A07A1309653FEE8FDAFFE5A9D21" box="[336,494,614,638]" captionStart="Figure 1" captionStartId="6.[211,278,1561,1583]" captionTargetBox="[211,1170,169,1528]" captionTargetId="figure-5@6.[211,1170,169,1528]" captionTargetPageId="6" captionText="Figure 1. Remanea naksanensis sp. nov. Female: (A) Habitus, dorsal; (B) habitus, lateral; (C) urosome (excluding P5-bearing somite), ventral. Scale bars in µm." figureDoi="http://doi.org/10.5281/zenodo.5204905" httpUri="https://zenodo.org/record/5204905/files/figure.png" pageId="5" pageNumber="2942">Figure 1A–C</figureCitation>
), except for anal somite, with striations on hyaline frills dorsally and laterally. Fifth urosomite and anal somite without sensilla; the latter small, without anal operculum, but with well-developed flimsy serrate pseudoperculum arising from penultimate somite. Genital somite (second urosomite) and third urosomite completely fused forming genital double-somite; genital field located in posterior half of second urosomite (proximal half of genital double-somite), with copulatory pore positioned medially, and two small pores located on either side of copulatory pore.
</paragraph>
<paragraph id="8BFC3682A1309653FF02FCACFDC79B03" blockId="5.[143,1104,390,1690]" pageId="5" pageNumber="2942">
Caudal rami (
<figureCitation id="13782A07A1309653FEE2FCACFE6F9C22" box="[346,475,869,893]" captionStart="Figure 1" captionStartId="6.[211,278,1561,1583]" captionTargetBox="[211,1170,169,1528]" captionTargetId="figure-5@6.[211,1170,169,1528]" captionTargetPageId="6" captionText="Figure 1. Remanea naksanensis sp. nov. Female: (A) Habitus, dorsal; (B) habitus, lateral; (C) urosome (excluding P5-bearing somite), ventral. Scale bars in µm." figureDoi="http://doi.org/10.5281/zenodo.5204905" httpUri="https://zenodo.org/record/5204905/files/figure.png" pageId="5" pageNumber="2942">Figures 1C</figureCitation>
,
<figureCitation id="13782A07A1309653FE52FCACFDB89C22" box="[490,524,869,893]" captionStart="Figure 2" captionStartId="7.[143,210,1568,1590]" captionTargetBox="[143,1102,161,1535]" captionTargetId="figure-6@7.[143,1102,161,1535]" captionTargetPageId="7" captionText="Figure 2. Remanea naksanensis sp. nov. Female: (A) Antennule, ventral; (B) antenna; (C) pseudoperculum, anal somite and caudal rami, dorsal (inset showing full length of setae IV–V); (D) fifth pair of legs (P5); (E) P6 and genital field. Scale bars in µm." figureDoi="http://doi.org/10.5281/zenodo.5204907" httpUri="https://zenodo.org/record/5204907/files/figure.png" pageId="5" pageNumber="2942">2C</figureCitation>
,
<figureCitation id="13782A07A1309653FDA2FCACFD8A9C22" box="[538,574,869,893]" captionStart="Figure 7" captionStartId="14.[211,278,1682,1704]" captionTargetBox="[211,1171,161,1649]" captionTargetId="figure-5@14.[211,1171,161,1649]" captionTargetPageId="14" captionText="Figure 7. Remanea naksanensis sp. nov., scanning electron micrographs. Female: (A) Antennary exopod; (B) distal segment of mandibular palp; (C) maxilliped; (D) genital field and P6; (E) P5; (G) pseudoperculum; (H) caudal rami, dorsal. Male: (F) fifth antennulary segment with surface sutures arrowed. Scale bars in µm." figureDoi="http://doi.org/10.5281/zenodo.5204917" httpUri="https://zenodo.org/record/5204917/files/figure.png" pageId="5" pageNumber="2942">7H</figureCitation>
) rectangular, about 1.6 times as long as wide; with three transverse spinular rows dorsally; with spinules at bases of setae III (ventrally) and VII (dorsally), and around distal margin of rami (ventrally); secretory pores located dorsally near base of seta III and ventrally near base of seta IV; with seven setae, setae I–II located halfway down the outer margin; setae III–VII located in distal third of ramus; seta II short and pinnate; seta I bare; seta III plumose; seta IV well developed and plumose, seta V longest and pinnate in distal half; seta VI pinnate; dorsal seta VII bi-articulate at base, naked.
</paragraph>
<paragraph id="8BFC3682A1309653FF02FBACFECD9BC3" blockId="5.[143,1104,390,1690]" pageId="5" pageNumber="2942">
Rostrum (
<figureCitation id="13782A07A1309653FE94FBADFE159B23" box="[300,417,1124,1148]" captionStart="Figure 1" captionStartId="6.[211,278,1561,1583]" captionTargetBox="[211,1170,169,1528]" captionTargetId="figure-5@6.[211,1170,169,1528]" captionTargetPageId="6" captionText="Figure 1. Remanea naksanensis sp. nov. Female: (A) Habitus, dorsal; (B) habitus, lateral; (C) urosome (excluding P5-bearing somite), ventral. Scale bars in µm." figureDoi="http://doi.org/10.5281/zenodo.5204905" httpUri="https://zenodo.org/record/5204905/files/figure.png" pageId="5" pageNumber="2942">Figure 1A</figureCitation>
) well developed, with rounded tip, defined at base; with two small lateral sensilla.
</paragraph>
<paragraph id="8BFC3682A1309653FF02FB6AFF799AC4" blockId="5.[143,1104,390,1690]" pageId="5" pageNumber="2942">
Antennule (
<figureCitation id="13782A07A1309653FE85FB6AFE069BE4" box="[317,434,1187,1211]" captionStart="Figure 2" captionStartId="7.[143,210,1568,1590]" captionTargetBox="[143,1102,161,1535]" captionTargetId="figure-6@7.[143,1102,161,1535]" captionTargetPageId="7" captionText="Figure 2. Remanea naksanensis sp. nov. Female: (A) Antennule, ventral; (B) antenna; (C) pseudoperculum, anal somite and caudal rami, dorsal (inset showing full length of setae IV–V); (D) fifth pair of legs (P5); (E) P6 and genital field. Scale bars in µm." figureDoi="http://doi.org/10.5281/zenodo.5204907" httpUri="https://zenodo.org/record/5204907/files/figure.png" pageId="5" pageNumber="2942">Figure 2A</figureCitation>
) slender, eight-segmented; segment 1 with short row of long spinules along anterior margin and one pinnate seta; segments 1 and 2 similar in length; segment 4 with sub-cylindrical process bearing one bare seta fused basally to aesthetasc; armature formula: 1-[1 pinnate], 2-[7 pinnate + 1 spinulose + 3 bare], 3-[5 pinnate + 3 bare], 4-[3 pinnate + (1 + ae)], 5-[1 pinnate + 1 bare], 6-[3 pinnate + 1 spinulose + 1 bare], 7-[1 pinnate + 1 spinulose seta + bare], 8-[5 bare + (2 + ae)]. Acrothek consisting of apical aesthetasc and two basally fused bare setae.
</paragraph>
<paragraph id="8BFC3682A1309653FF02FA6AFDC899C5" blockId="5.[143,1104,390,1690]" pageId="5" pageNumber="2942">
Antenna (
<figureCitation id="13782A07A1309653FE91FA6AFE129AE4" box="[297,422,1443,1467]" captionStart="Figure 2" captionStartId="7.[143,210,1568,1590]" captionTargetBox="[143,1102,161,1535]" captionTargetId="figure-6@7.[143,1102,161,1535]" captionTargetPageId="7" captionText="Figure 2. Remanea naksanensis sp. nov. Female: (A) Antennule, ventral; (B) antenna; (C) pseudoperculum, anal somite and caudal rami, dorsal (inset showing full length of setae IV–V); (D) fifth pair of legs (P5); (E) P6 and genital field. Scale bars in µm." figureDoi="http://doi.org/10.5281/zenodo.5204907" httpUri="https://zenodo.org/record/5204907/files/figure.png" pageId="5" pageNumber="2942">Figures 2B</figureCitation>
,
<figureCitation id="13782A07A1309653FE08FA6AFE629AE4" box="[432,470,1443,1467]" captionStart="Figure 7" captionStartId="14.[211,278,1682,1704]" captionTargetBox="[211,1171,161,1649]" captionTargetId="figure-5@14.[211,1171,161,1649]" captionTargetPageId="14" captionText="Figure 7. Remanea naksanensis sp. nov., scanning electron micrographs. Female: (A) Antennary exopod; (B) distal segment of mandibular palp; (C) maxilliped; (D) genital field and P6; (E) P5; (G) pseudoperculum; (H) caudal rami, dorsal. Male: (F) fifth antennulary segment with surface sutures arrowed. Scale bars in µm." figureDoi="http://doi.org/10.5281/zenodo.5204917" httpUri="https://zenodo.org/record/5204917/files/figure.png" pageId="5" pageNumber="2942">7A</figureCitation>
): coxa and basis without surface ornamentation. Exopod two-segmented; exp-1 shorter than exp-2, the former with one pinnate seta distally; exp-2 with row of strong spinules apically and with four setae; armature formula: 1-[1 pinnate], 2-[3 pinnate + 1 bare]. Endopod two-segmented; enp-1 with lateral seta, without surface ornamentation; lateral armature of enp-2 consisting of two bare setae, one small spinulose element, and one pinnate seta; distal armature of enp-2 consisting of four geniculate and two bare setae (one small bare seta laterally and one long bare seta fused at base to largest geniculate seta).
</paragraph>
<caption id="DF3C660AA1339650FF6BF9D0FC239913" ID-DOI="http://doi.org/10.5281/zenodo.5204905" ID-Zenodo-Dep="5204905" httpUri="https://zenodo.org/record/5204905/files/figure.png" pageId="6" pageNumber="2943" startId="6.[211,278,1561,1583]" targetBox="[211,1170,169,1528]" targetPageId="6">
<paragraph id="8BFC3682A1339650FF6BF9D0FC239913" blockId="6.[211,1172,1561,1613]" pageId="6" pageNumber="2943">
Figure 1.
<taxonomicName id="4C434D01A1339650FEF9F9D3FDAA9970" authority="Back &amp; Lee &amp; Huys, 2011" authorityName="Back &amp; Lee &amp; Huys" authorityYear="2011" box="[321,542,1561,1583]" class="Hexanauplia" family="Paramesochridae" genus="Remanea" kingdom="Animalia" order="Harpacticoida" pageId="6" pageNumber="2943" phylum="Arthropoda" rank="species" species="naksanensis" status="sp. nov.">
<emphasis id="B937EA90A1339650FEF9F9D3FDAA9970" box="[321,542,1561,1583]" italics="true" pageId="6" pageNumber="2943">Remanea naksanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20457EBA1339650FD92F9D3FDC89970" box="[554,636,1562,1583]" pageId="6" pageNumber="2943" rank="species">sp. nov.</taxonomicNameLabel>
Female: (A) Habitus, dorsal; (B) habitus, lateral; (C) urosome (excluding P5-bearing somite), ventral. Scale bars in µm.
</paragraph>
</caption>
<caption id="DF3C660AA1329651FF37F9E9FC3C992F" ID-DOI="http://doi.org/10.5281/zenodo.5204907" ID-Zenodo-Dep="5204907" httpUri="https://zenodo.org/record/5204907/files/figure.png" pageId="7" pageNumber="2944" startId="7.[143,210,1568,1590]" targetBox="[143,1102,161,1535]" targetPageId="7">
<paragraph id="8BFC3682A1329651FF37F9E9FC3C992F" blockId="7.[143,1103,1568,1648]" pageId="7" pageNumber="2944">
Figure 2.
<taxonomicName id="4C434D01A1329651FEB9F9E8FE569969" authority="Back &amp; Lee &amp; Huys, 2011" authorityName="Back &amp; Lee &amp; Huys" authorityYear="2011" box="[257,482,1568,1590]" class="Hexanauplia" family="Paramesochridae" genus="Remanea" kingdom="Animalia" order="Harpacticoida" pageId="7" pageNumber="2944" phylum="Arthropoda" rank="species" species="naksanensis" status="sp. nov.">
<emphasis id="B937EA90A1329651FEB9F9E8FE569969" box="[257,482,1568,1590]" italics="true" pageId="7" pageNumber="2944">Remanea naksanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20457EBA1329651FE4AF9E8FDFC9969" box="[498,584,1569,1590]" pageId="7" pageNumber="2944" rank="species">sp. nov.</taxonomicNameLabel>
Female: (A) Antennule, ventral; (B) antenna; (C) pseudoperculum, anal somite and caudal rami, dorsal (inset showing full length of setae IV–V); (D) fifth pair of legs (P5); (E) P6 and genital field. Scale bars in µm.
</paragraph>
</caption>
<paragraph id="8BFC3682A13D965EFF46FF6EFBC39E00" blockId="8.[211,1172,167,1690]" pageId="8" pageNumber="2945">
Mandible (
<figureCitation id="13782A07A13D965EFEC0FF6EFE4C9FE0" box="[376,504,167,191]" captionStart="Figure 3" captionStartId="9.[143,210,1445,1467]" captionTargetBox="[143,1101,162,1412]" captionTargetId="figure-6@9.[143,1102,161,1412]" captionTargetPageId="9" captionText="Figure 3. Remanea naksanensis sp. nov. Female: (A) Mandible; (B) maxillule; (C) maxilla [syncoxal endites (1–4) and endopod (5) shown in separate insets; arrow indicates minute process on distal coxal endite]; (D) maxilliped. Scale bars in µm." figureDoi="http://doi.org/10.5281/zenodo.5204909" httpUri="https://zenodo.org/record/5204909/files/figure.png" pageId="8" pageNumber="2945">Figures 3A</figureCitation>
,
<figureCitation id="13782A07A13D965EFDBAFF6EFD919FE0" box="[514,549,167,191]" captionStart="Figure 7" captionStartId="14.[211,278,1682,1704]" captionTargetBox="[211,1171,161,1649]" captionTargetId="figure-5@14.[211,1171,161,1649]" captionTargetPageId="14" captionText="Figure 7. Remanea naksanensis sp. nov., scanning electron micrographs. Female: (A) Antennary exopod; (B) distal segment of mandibular palp; (C) maxilliped; (D) genital field and P6; (E) P5; (G) pseudoperculum; (H) caudal rami, dorsal. Male: (F) fifth antennulary segment with surface sutures arrowed. Scale bars in µm." figureDoi="http://doi.org/10.5281/zenodo.5204917" httpUri="https://zenodo.org/record/5204917/files/figure.png" pageId="8" pageNumber="2945">7B</figureCitation>
): coxa well developed, with process; gnathobase with six small blunt teeth and one pinnate seta at dorsal corner. Palp biramous; basis elongate, with four pinnate setae; exopod one-segmented, with three inner and two distal pinnate setae, and ornamented as figured; endopod two-segmented; enp-1 as long as enp-2, with two bare distal setae; enp-2 with seven setae (two bare setae, three distal setae fused at base, and one apical long seta and one slender bare seta confluent at base).
</paragraph>
<paragraph id="8BFC3682A13D965EFF46FEAEFE239D41" blockId="8.[211,1172,167,1690]" pageId="8" pageNumber="2945">
Maxillule (
<figureCitation id="13782A07A13D965EFEC5FEAEFE439E20" box="[381,503,359,383]" captionStart="Figure 3" captionStartId="9.[143,210,1445,1467]" captionTargetBox="[143,1101,162,1412]" captionTargetId="figure-6@9.[143,1102,161,1412]" captionTargetPageId="9" captionText="Figure 3. Remanea naksanensis sp. nov. Female: (A) Mandible; (B) maxillule; (C) maxilla [syncoxal endites (1–4) and endopod (5) shown in separate insets; arrow indicates minute process on distal coxal endite]; (D) maxilliped. Scale bars in µm." figureDoi="http://doi.org/10.5281/zenodo.5204909" httpUri="https://zenodo.org/record/5204909/files/figure.png" pageId="8" pageNumber="2945">Figure 3B</figureCitation>
): praecoxa subquadrate, without ornamentation, arthrite well developed, with five strong spines and one seta apically, three bare lateral elements, and two surface setae. Coxa with five pinnate setae. Basis with five setae. Exopod onesegmented, with four slender bare setae of different lengths and with lateral row of spinules. Endopod one-segmented, longer than exopod, with three apical, and two bare lateral setae.
</paragraph>
<paragraph id="8BFC3682A13D965EFF46FDEFFBA29DA2" blockId="8.[211,1172,167,1690]" pageId="8" pageNumber="2945">
Maxilla (
<figureCitation id="13782A07A13D965EFEDAFDEFFE639D61" box="[354,471,550,574]" captionStart="Figure 3" captionStartId="9.[143,210,1445,1467]" captionTargetBox="[143,1101,162,1412]" captionTargetId="figure-6@9.[143,1102,161,1412]" captionTargetPageId="9" captionText="Figure 3. Remanea naksanensis sp. nov. Female: (A) Mandible; (B) maxillule; (C) maxilla [syncoxal endites (1–4) and endopod (5) shown in separate insets; arrow indicates minute process on distal coxal endite]; (D) maxilliped. Scale bars in µm." figureDoi="http://doi.org/10.5281/zenodo.5204909" httpUri="https://zenodo.org/record/5204909/files/figure.png" pageId="8" pageNumber="2945">Figure 3C</figureCitation>
): syncoxa with four endites. First endite small, with one pinnate and one bipinnate setae; second endite with two pinnate setae; third endite with two pinnate and one bare setae; distal endite close to third endite, with one pinnate lateral seta, two pinnate distal elements, and with one small process proximally (arrowed in
<figureCitation id="13782A07A13D965EFF6BFD6CFEF09DE2" box="[211,324,677,701]" captionStart="Figure 3" captionStartId="9.[143,210,1445,1467]" captionTargetBox="[143,1101,162,1412]" captionTargetId="figure-6@9.[143,1102,161,1412]" captionTargetPageId="9" captionText="Figure 3. Remanea naksanensis sp. nov. Female: (A) Mandible; (B) maxillule; (C) maxilla [syncoxal endites (1–4) and endopod (5) shown in separate insets; arrow indicates minute process on distal coxal endite]; (D) maxilliped. Scale bars in µm." figureDoi="http://doi.org/10.5281/zenodo.5204909" httpUri="https://zenodo.org/record/5204909/files/figure.png" pageId="8" pageNumber="2945">Figure 3C</figureCitation>
– inset 4). Allobasis with one bare seta, one stout claw-like element, and one bare accompanying seta. Endopod two-segmented; enp-1 rectangular, with five slender setae; enp-2 with one pinnate claw, two apical, and one naked lateral setae.
</paragraph>
<paragraph id="8BFC3682A13D965EFF46FCCCFCC89C22" blockId="8.[211,1172,167,1690]" pageId="8" pageNumber="2945">
Maxilliped (
<figureCitation id="13782A07A13D965EFE3FFCCCFDBD9C42" box="[391,521,773,797]" captionStart="Figure 3" captionStartId="9.[143,210,1445,1467]" captionTargetBox="[143,1101,162,1412]" captionTargetId="figure-6@9.[143,1102,161,1412]" captionTargetPageId="9" captionText="Figure 3. Remanea naksanensis sp. nov. Female: (A) Mandible; (B) maxillule; (C) maxilla [syncoxal endites (1–4) and endopod (5) shown in separate insets; arrow indicates minute process on distal coxal endite]; (D) maxilliped. Scale bars in µm." figureDoi="http://doi.org/10.5281/zenodo.5204909" httpUri="https://zenodo.org/record/5204909/files/figure.png" pageId="8" pageNumber="2945">Figures 3D</figureCitation>
,
<figureCitation id="13782A07A13D965EFDADFCCCFD829C42" box="[533,566,773,797]" captionStart="Figure 7" captionStartId="14.[211,278,1682,1704]" captionTargetBox="[211,1171,161,1649]" captionTargetId="figure-5@14.[211,1171,161,1649]" captionTargetPageId="14" captionText="Figure 7. Remanea naksanensis sp. nov., scanning electron micrographs. Female: (A) Antennary exopod; (B) distal segment of mandibular palp; (C) maxilliped; (D) genital field and P6; (E) P5; (G) pseudoperculum; (H) caudal rami, dorsal. Male: (F) fifth antennulary segment with surface sutures arrowed. Scale bars in µm." figureDoi="http://doi.org/10.5281/zenodo.5204917" httpUri="https://zenodo.org/record/5204917/files/figure.png" pageId="8" pageNumber="2945">7C</figureCitation>
) three-segmented. Syncoxa with two rows of spinules. Basis elongate, with rows of spinules halfway down inner and outer margin. Endopod indistinctly segmented, with one bare seta proximally, one stout claw medially, two stout claw-like bi-articulate setae, and one small seta distally.
</paragraph>
<paragraph id="8BFC3682A13D965EFF46FC4CFD859B23" blockId="8.[211,1172,167,1690]" pageId="8" pageNumber="2945">
P1 (
<figureCitation id="13782A07A13D965EFE93FC4CFE109CC2" box="[299,420,901,925]" captionStart="Figure 4" captionStartId="10.[211,278,1584,1606]" captionTargetBox="[213,1169,163,1551]" captionTargetId="figure-5@10.[211,1170,163,1551]" captionTargetPageId="10" captionText="Figure 4. Remanea naksanensis sp. nov. Female: (A) P1, anterior; (B) P2, anterior; (C) P3, anterior; (D) P4, anterior. Scale bars in µm." figureDoi="http://doi.org/10.5281/zenodo.5204911" httpUri="https://zenodo.org/record/5204911/files/figure.png" pageId="8" pageNumber="2945">Figure 4A</figureCitation>
): coxa and basis with spinules as figured; the latter with one outer bare and one inner pinnate seta. Exopod three-segmented, shorter than endopod; exp- 1 longest, with some spinules along outer margin and one pinnate outer spine; exp- 2 somewhat swollen distally, with one pinnate outer spine and one spinulose inner seta; exp-3 with six spines/setae. Endopod prehensile; enp-1 elongate and approximately as long as exopod, with row of spinules along outer margin and one long plumose inner seta; enp-2 small, slightly longer than wide, apically with two strong claw-like setae and one small pinnate element.
</paragraph>
<paragraph id="8BFC3682A13D965EFF46FB4DFDB19AC4" blockId="8.[211,1172,167,1690]" pageId="8" pageNumber="2945">
P2–P4 (
<figureCitation id="13782A07A13D965EFEEFFB4DFE479BC3" box="[343,499,1156,1180]" captionStart="Figure 4" captionStartId="10.[211,278,1584,1606]" captionTargetBox="[213,1169,163,1551]" captionTargetId="figure-5@10.[211,1170,163,1551]" captionTargetPageId="10" captionText="Figure 4. Remanea naksanensis sp. nov. Female: (A) P1, anterior; (B) P2, anterior; (C) P3, anterior; (D) P4, anterior. Scale bars in µm." figureDoi="http://doi.org/10.5281/zenodo.5204911" httpUri="https://zenodo.org/record/5204911/files/figure.png" pageId="8" pageNumber="2945">Figure 4B–D</figureCitation>
): coxae with row of long (P2) or slender (P3–P4) spinules on outer distal corner. Basis with long inner spinules and with rows of spinules on anterior surface. Exopod three-segmented; exp-1 with one strong pinnate outer spine; exp-2 shortest, with one strong pinnate outer spine and one bare inner seta; exp-3 with two pinnate outer spines, two plumose apical setae, and one bare (P2–P3) or plumose (P4) inner seta. Endopod two-segmented; enp-1 with one plumose inner seta; enp-2 as long as enp-1 (P2) or about 1.6 times as long as enp-1 (P3–P4), with one bare (P2) or pinnate (P3–P4) outer spine and two pinnate apical setae; with one pore close to inner distal corner, except for P4.
</paragraph>
<paragraph id="8BFC3682A13D965EFF46FA6AFCF89A85" blockId="8.[211,1172,167,1690]" pageId="8" pageNumber="2945">
Armature formula as follows: exopod: P2, 0.1.122; P3, 0.1.122; P4, 0.1.122; endopod: P2, 1.021; P3, 1.021 (
<quantity id="4CBB9B67A13D965EFD88FA0BFD3E9A85" box="[560,650,1474,1498]" metricMagnitude="-2" metricUnit="m" metricValue="2.5908" pageId="8" pageNumber="2945" unit="in" value="1.02">1.020 in</quantity>
male); P4, 1.021.
</paragraph>
<paragraph id="8BFC3682A13D965EFF46FA2BFE8F99C5" blockId="8.[211,1172,167,1690]" pageId="8" pageNumber="2945">
P5 (
<figureCitation id="13782A07A13D965EFE95FA2BFE009AA5" box="[301,436,1506,1530]" captionStart="Figure 2" captionStartId="7.[143,210,1568,1590]" captionTargetBox="[143,1102,161,1535]" captionTargetId="figure-6@7.[143,1102,161,1535]" captionTargetPageId="7" captionText="Figure 2. Remanea naksanensis sp. nov. Female: (A) Antennule, ventral; (B) antenna; (C) pseudoperculum, anal somite and caudal rami, dorsal (inset showing full length of setae IV–V); (D) fifth pair of legs (P5); (E) P6 and genital field. Scale bars in µm." figureDoi="http://doi.org/10.5281/zenodo.5204907" httpUri="https://zenodo.org/record/5204907/files/figure.png" pageId="8" pageNumber="2945">Figures 2D</figureCitation>
,
<figureCitation id="13782A07A13D965EFE7BFA2BFE569AA5" box="[451,482,1506,1530]" captionStart="Figure 7" captionStartId="14.[211,278,1682,1704]" captionTargetBox="[211,1171,161,1649]" captionTargetId="figure-5@14.[211,1171,161,1649]" captionTargetPageId="14" captionText="Figure 7. Remanea naksanensis sp. nov., scanning electron micrographs. Female: (A) Antennary exopod; (B) distal segment of mandibular palp; (C) maxilliped; (D) genital field and P6; (E) P5; (G) pseudoperculum; (H) caudal rami, dorsal. Male: (F) fifth antennulary segment with surface sutures arrowed. Scale bars in µm." figureDoi="http://doi.org/10.5281/zenodo.5204917" httpUri="https://zenodo.org/record/5204917/files/figure.png" pageId="8" pageNumber="2945">7E</figureCitation>
) with medially fused baseoendopods and discrete exopods. Baseoendopod with short pinnate outer basal seta; endopodal lobes shorter than exopods, each lobe with two apical pinnate setae of almost equal length, with long spinules along inner margin. Exopod well developed, with rows of spinules along inner and outer margins, with two pinnate apical setae and two spinulose short outer elements.
</paragraph>
<caption id="DF3C660AA13C965FFF37FA6CFD7E9AAA" ID-DOI="http://doi.org/10.5281/zenodo.5204909" ID-Zenodo-Dep="5204909" httpUri="https://zenodo.org/record/5204909/files/figure.png" pageId="9" pageNumber="2946" startId="9.[143,210,1445,1467]" targetBox="[143,1101,162,1412]" targetPageId="9">
<paragraph id="8BFC3682A13C965FFF37FA6CFD7E9AAA" blockId="9.[143,1103,1445,1526]" pageId="9" pageNumber="2946">
Figure 3.
<taxonomicName id="4C434D01A13C965FFF44FA6FFE6C9AE4" authority="Back &amp; Lee &amp; Huys, 2011" authorityName="Back &amp; Lee &amp; Huys" authorityYear="2011" box="[252,472,1445,1467]" class="Hexanauplia" family="Paramesochridae" genus="Remanea" kingdom="Animalia" order="Harpacticoida" pageId="9" pageNumber="2946" phylum="Arthropoda" rank="species" species="naksanensis" status="sp. nov.">
<emphasis id="B937EA90A13C965FFF44FA6FFE6C9AE4" box="[252,472,1445,1467]" italics="true" pageId="9" pageNumber="2946">Remanea naksanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20457EBA13C965FFE5CFA6FFD819AE4" box="[484,565,1446,1467]" pageId="9" pageNumber="2946" rank="species">sp. nov.</taxonomicNameLabel>
Female: (A) Mandible; (B) maxillule; (C) maxilla [syncoxal endites (1–4) and endopod (5) shown in separate insets; arrow indicates minute process on distal coxal endite]; (D) maxilliped. Scale bars in µm.
</paragraph>
</caption>
<caption id="DF3C660AA13F965CFF6BF9F9FD3B993C" ID-DOI="http://doi.org/10.5281/zenodo.5204911" ID-Zenodo-Dep="5204911" httpUri="https://zenodo.org/record/5204911/files/figure.png" pageId="10" pageNumber="2947" startId="10.[211,278,1584,1606]" targetBox="[213,1169,163,1551]" targetPageId="10">
<paragraph id="8BFC3682A13F965CFF6BF9F9FD3B993C" blockId="10.[211,1171,1584,1635]" pageId="10" pageNumber="2947">
Figure 4.
<taxonomicName id="4C434D01A13F965CFE87F9F8FDAE9919" authority="Back &amp; Lee &amp; Huys, 2011" authorityName="Back &amp; Lee &amp; Huys" authorityYear="2011" box="[319,538,1584,1606]" class="Hexanauplia" family="Paramesochridae" genus="Remanea" kingdom="Animalia" order="Harpacticoida" pageId="10" pageNumber="2947" phylum="Arthropoda" rank="species" species="naksanensis" status="sp. nov.">
<emphasis id="B937EA90A13F965CFE87F9F8FDAE9919" box="[319,538,1584,1606]" italics="true" pageId="10" pageNumber="2947">Remanea naksanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20457EBA13F965CFD9CF9F9FDC0991A" box="[548,628,1584,1605]" pageId="10" pageNumber="2947" rank="species">sp. nov.</taxonomicNameLabel>
Female: (A) P1, anterior; (B) P2, anterior; (C) P3, anterior; (D) P4, anterior. Scale bars in µm.
</paragraph>
</caption>
<paragraph id="8BFC3682A13E965DFF02FF6EFB969F80" blockId="11.[143,1103,167,223]" pageId="11" pageNumber="2948">
P6 (
<figureCitation id="13782A07A13E965DFF5EFF6EFED19FE0" box="[230,357,167,191]" captionStart="Figure 2" captionStartId="7.[143,210,1568,1590]" captionTargetBox="[143,1102,161,1535]" captionTargetId="figure-6@7.[143,1102,161,1535]" captionTargetPageId="7" captionText="Figure 2. Remanea naksanensis sp. nov. Female: (A) Antennule, ventral; (B) antenna; (C) pseudoperculum, anal somite and caudal rami, dorsal (inset showing full length of setae IV–V); (D) fifth pair of legs (P5); (E) P6 and genital field. Scale bars in µm." figureDoi="http://doi.org/10.5281/zenodo.5204907" httpUri="https://zenodo.org/record/5204907/files/figure.png" pageId="11" pageNumber="2948">Figures 2E</figureCitation>
,
<figureCitation id="13782A07A13E965DFEC9FF6EFE219FE0" box="[369,405,167,191]" captionStart="Figure 7" captionStartId="14.[211,278,1682,1704]" captionTargetBox="[211,1171,161,1649]" captionTargetId="figure-5@14.[211,1171,161,1649]" captionTargetPageId="14" captionText="Figure 7. Remanea naksanensis sp. nov., scanning electron micrographs. Female: (A) Antennary exopod; (B) distal segment of mandibular palp; (C) maxilliped; (D) genital field and P6; (E) P5; (G) pseudoperculum; (H) caudal rami, dorsal. Male: (F) fifth antennulary segment with surface sutures arrowed. Scale bars in µm." figureDoi="http://doi.org/10.5281/zenodo.5204917" httpUri="https://zenodo.org/record/5204917/files/figure.png" pageId="11" pageNumber="2948">7D</figureCitation>
) represented by narrow transverse plate, armed with one bare inner seta and two pinnate outer elements of different lengths, the former longest.
</paragraph>
<paragraph id="8BFC3682A13E965DFF37FED7FED39E6A" blockId="11.[143,359,285,309]" box="[143,359,285,309]" pageId="11" pageNumber="2948">
<emphasis id="B937EA90A13E965DFF37FED7FED39E6A" box="[143,359,285,309]" italics="true" pageId="11" pageNumber="2948">Description of male</emphasis>
</paragraph>
<paragraph id="8BFC3682A13E965DFF37FE8DFBA39E83" blockId="11.[143,1104,324,1050]" pageId="11" pageNumber="2948">
Slightly smaller and more slender than female, body length 498 µm (
<emphasis id="B937EA90A13E965DFC1DFE8CFC069E03" box="[933,946,325,348]" italics="true" pageId="11" pageNumber="2948">n</emphasis>
= 4, mean = 488 µm) (
<figureCitation id="13782A07A13E965DFF44FEADFEC69E23" box="[252,370,356,380]" captionStart="Figure 5" captionStartId="12.[211,278,1584,1606]" captionTargetBox="[211,1170,161,1551]" captionTargetId="figure-5@12.[211,1170,161,1551]" captionTargetPageId="12" captionText="Figure 5. Remanea naksanensis sp. nov. Male: (A) Habitus, dorsal; (B) antennule, ventral (arrows indicate position of partial surface sutures on swollen segment 5). Scale bars in µm." figureDoi="http://doi.org/10.5281/zenodo.5204913" httpUri="https://zenodo.org/record/5204913/files/figure.png" pageId="11" pageNumber="2948">Figure 5A</figureCitation>
). Largest width (92 µm) measured at posterior margin of cephalic shield. Cephalothorax (
<figureCitation id="13782A07A13E965DFE21FE4DFDBA9EC3" box="[409,526,388,412]" captionStart="Figure 5" captionStartId="12.[211,278,1584,1606]" captionTargetBox="[211,1170,161,1551]" captionTargetId="figure-5@12.[211,1170,161,1551]" captionTargetPageId="12" captionText="Figure 5. Remanea naksanensis sp. nov. Male: (A) Habitus, dorsal; (B) antennule, ventral (arrows indicate position of partial surface sutures on swollen segment 5). Scale bars in µm." figureDoi="http://doi.org/10.5281/zenodo.5204913" httpUri="https://zenodo.org/record/5204913/files/figure.png" pageId="11" pageNumber="2948">Figure 5A</figureCitation>
) slightly more angular and with more sensilla than in female. General body shape and ornamentation as in female except for separation of genital somite; additional sexual dimorphism in antennule, P2, P3, P5 and P6.
</paragraph>
<paragraph id="8BFC3682A13E965DFF02FE2DFCC69D24" blockId="11.[143,1104,324,1050]" pageId="11" pageNumber="2948">
Antennule (
<figureCitation id="13782A07A13E965DFEFBFE2DFE739EA3" box="[323,455,484,508]" captionStart="Figure 5" captionStartId="12.[211,278,1584,1606]" captionTargetBox="[211,1170,161,1551]" captionTargetId="figure-5@12.[211,1170,161,1551]" captionTargetPageId="12" captionText="Figure 5. Remanea naksanensis sp. nov. Male: (A) Habitus, dorsal; (B) antennule, ventral (arrows indicate position of partial surface sutures on swollen segment 5). Scale bars in µm." figureDoi="http://doi.org/10.5281/zenodo.5204913" httpUri="https://zenodo.org/record/5204913/files/figure.png" pageId="11" pageNumber="2948">Figures 5B</figureCitation>
,
<figureCitation id="13782A07A13E965DFE60FE2DFE4E9EA3" box="[472,506,484,508]" captionStart="Figure 7" captionStartId="14.[211,278,1682,1704]" captionTargetBox="[211,1171,161,1649]" captionTargetId="figure-5@14.[211,1171,161,1649]" captionTargetPageId="14" captionText="Figure 7. Remanea naksanensis sp. nov., scanning electron micrographs. Female: (A) Antennary exopod; (B) distal segment of mandibular palp; (C) maxilliped; (D) genital field and P6; (E) P5; (G) pseudoperculum; (H) caudal rami, dorsal. Male: (F) fifth antennulary segment with surface sutures arrowed. Scale bars in µm." figureDoi="http://doi.org/10.5281/zenodo.5204917" httpUri="https://zenodo.org/record/5204917/files/figure.png" pageId="11" pageNumber="2948">7F</figureCitation>
): eight-segmented, subchirocer; segment 6 swollen, largest. Aesthetascs on sixth and eighth segments. Sixth segment with surface suture dorsoanteriorly (arrowed in
<figureCitation id="13782A07A13E965DFE76FDEAFDF69D64" box="[462,578,547,571]" captionStart="Figure 7" captionStartId="14.[211,278,1682,1704]" captionTargetBox="[211,1171,161,1649]" captionTargetId="figure-5@14.[211,1171,161,1649]" captionTargetPageId="14" captionText="Figure 7. Remanea naksanensis sp. nov., scanning electron micrographs. Female: (A) Antennary exopod; (B) distal segment of mandibular palp; (C) maxilliped; (D) genital field and P6; (E) P5; (G) pseudoperculum; (H) caudal rami, dorsal. Male: (F) fifth antennulary segment with surface sutures arrowed. Scale bars in µm." figureDoi="http://doi.org/10.5281/zenodo.5204917" httpUri="https://zenodo.org/record/5204917/files/figure.png" pageId="11" pageNumber="2948">Figure 7F</figureCitation>
). Armature formula: 1-[1 pinnate], 2-[4 bare + 1 pinnate], 3-[5 bare + 3 pinnate], 4-[3 bare + 3 pinnate], 5-[2 bare + 1 pinnate], 6-[5 bare + 6 pinnate + (1 + ae)], 7-[4 bare], 8-[7 bare + (2 + ae)].
</paragraph>
<paragraph id="8BFC3682A13E965DFF02FD4AFD0B9DA4" blockId="11.[143,1104,324,1050]" pageId="11" pageNumber="2948">
P2 (
<figureCitation id="13782A07A13E965DFF54FD4AFED39DC4" box="[236,359,643,667]" captionStart="Figure 6" captionStartId="13.[143,210,1602,1624]" captionTargetBox="[143,1102,163,1568]" captionTargetId="figure-6@13.[143,1102,162,1569]" captionTargetPageId="13" captionText="Figure 6. Remanea naksanensis sp. nov. Male: (A) P2, anterior; (B) P3, anterior; (C) left P5, anterior; (D) armature of left P6. Scale bars in µm." figureDoi="http://doi.org/10.5281/zenodo.5204915" httpUri="https://zenodo.org/record/5204915/files/figure.png" pageId="11" pageNumber="2948">Figure 6A</figureCitation>
) Basis with surface spinules and small outer seta. Exopod as in female. Enp-1 as in female; enp-2 armed with two long and one small bare setae apically, the latter smaller than homologous element in female. General shape, segmentation, and armature formula as in female.
</paragraph>
<paragraph id="8BFC3682A13E965DFF02FCCAFBE49C05" blockId="11.[143,1104,324,1050]" pageId="11" pageNumber="2948">
P3 (
<figureCitation id="13782A07A13E965DFF50FCCAFEEB9C44" box="[232,351,771,795]" captionStart="Figure 6" captionStartId="13.[143,210,1602,1624]" captionTargetBox="[143,1102,163,1568]" captionTargetId="figure-6@13.[143,1102,162,1569]" captionTargetPageId="13" captionText="Figure 6. Remanea naksanensis sp. nov. Male: (A) P2, anterior; (B) P3, anterior; (C) left P5, anterior; (D) armature of left P6. Scale bars in µm." figureDoi="http://doi.org/10.5281/zenodo.5204915" httpUri="https://zenodo.org/record/5204915/files/figure.png" pageId="11" pageNumber="2948">Figure 6B</figureCitation>
). Coxa, basis, and exopod as in female. Endopod two-segmented; enp-1 with rows of long spinules along outer margin and with one inner plumose seta. Enp-2 twice as long as enp-1, with two apical setae and well developed outer apophysis.
</paragraph>
<paragraph id="8BFC3682A13E965DFF02FCABFE259C85" blockId="11.[143,1104,324,1050]" pageId="11" pageNumber="2948">
P5 (
<figureCitation id="13782A07A13E965DFF5CFCABFEED9C25" box="[228,345,866,890]" captionStart="Figure 6" captionStartId="13.[143,210,1602,1624]" captionTargetBox="[143,1102,163,1568]" captionTargetId="figure-6@13.[143,1102,162,1569]" captionTargetPageId="13" captionText="Figure 6. Remanea naksanensis sp. nov. Male: (A) P2, anterior; (B) P3, anterior; (C) left P5, anterior; (D) armature of left P6. Scale bars in µm." figureDoi="http://doi.org/10.5281/zenodo.5204915" httpUri="https://zenodo.org/record/5204915/files/figure.png" pageId="11" pageNumber="2948">Figure 6C</figureCitation>
): baseoendopods confluent, forming large transverse plate; unarmed except for outer slender bare seta on either side. Exopod ovate, with row of spinules along inner margin; with two outer spinulose setae, one long apical bare seta and one small naked inner seta.
</paragraph>
<paragraph id="8BFC3682A13E965DFF02FC2BFC4E9B45" blockId="11.[143,1104,324,1050]" pageId="11" pageNumber="2948">
P6 (
<figureCitation id="13782A07A13E965DFF5FFC2BFED69CA5" box="[231,354,994,1018]" captionStart="Figure 6" captionStartId="13.[143,210,1602,1624]" captionTargetBox="[143,1102,163,1568]" captionTargetId="figure-6@13.[143,1102,162,1569]" captionTargetPageId="13" captionText="Figure 6. Remanea naksanensis sp. nov. Male: (A) P2, anterior; (B) P3, anterior; (C) left P5, anterior; (D) armature of left P6. Scale bars in µm." figureDoi="http://doi.org/10.5281/zenodo.5204915" httpUri="https://zenodo.org/record/5204915/files/figure.png" pageId="11" pageNumber="2948">Figure 6D</figureCitation>
): pair of P6 symmetrical, fused medially. Each P6 with one outer naked lateral seta proximally and one pinnate seta close to distal outer corner.
</paragraph>
<paragraph id="8BFC3682A13E965DFF37FBAEFEB39B20" blockId="11.[143,263,1127,1151]" box="[143,263,1127,1151]" pageId="11" pageNumber="2948">
<emphasis id="B937EA90A13E965DFF37FBAEFEB39B20" box="[143,263,1127,1151]" italics="true" pageId="11" pageNumber="2948">Etymology</emphasis>
</paragraph>
<paragraph id="8BFC3682A13E965DFF37FB46FE889B98" blockId="11.[143,1103,1167,1223]" pageId="11" pageNumber="2948">
<materialsCitation id="3B2B3CDFA13E965DFF37FB46FE8C9B98" country="South Korea" location="Naksan" pageId="11" pageNumber="2948" specimenCount="1">
The specific name refers to the
<typeStatus id="54F88820A13E965DFE55FB59FDA99BF7" box="[493,541,1168,1192]" pageId="11" pageNumber="2948">type</typeStatus>
locality, the
<location id="8E9C6059A13E965DFD16FB46FCB29BF8" LSID="urn:lsid:plazi:treatment:03EA8794A131965DFDB7FB4DFE889B98:8E9C6059A13E965DFD16FB46FCB29BF8" box="[686,774,1167,1191]" country="South Korea" name="Naksan" pageId="11" pageNumber="2948">Naksan</location>
sandy beach area on the east coast of
<collectingCountry id="F3547612A13E965DFF57FB79FE8C9B98" box="[239,312,1200,1223]" name="South Korea" pageId="11" pageNumber="2948">Korea</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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