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<document id="211F8AF952D91D597DF2E88AAFBF040F" ID-CLB-Dataset="21062" ID-DOI="10.1080/00222933.2014.974706" ID-GBIF-Dataset="b2aec862-e219-4e84-af79-c6a721df5439" ID-ISSN="1464-5262" ID-Zenodo-Dep="4006372" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1598626848476" checkinUser="carolina" docAuthor="Gwiazdowicz, Dariusz J., Marchenko, Irina I. &amp; Teodorowicz, Ewa" docDate="2014" docId="03EA8788FFD9FFE7576CFEC1FC30FA6A" docLanguage="en" docName="JNATHIST.49.27-28.1659-1671.pdf" docOrigin="Journal of Natural History 49 (27)" docStyle="DocumentStyle:210803DF447D5DCB84349979EDD3507F.6:JNatHist.2013-2015.journal_article.2cover" docStyleId="210803DF447D5DCB84349979EDD3507F" docStyleName="JNatHist.2013-2015.journal_article.2cover" docStyleVersion="6" docTitle="Orthadenella coulsoni Gwiazdowicz &amp; Marchenko &amp; Teodorowicz 2014, sp. nov." docType="treatment" docVersion="7" lastPageNumber="1670" masterDocId="FFD3FFF0FFDBFFEC5566FFD6FF9AFFC9" masterDocTitle="Description of Orthadenella coulsoni sp. nov. (Acari: Mesostigmata: Melicharidae) from Siberia with a key to the females of Orthadenella" masterLastPageNumber="1671" masterPageNumber="1659" pageNumber="1661" updateTime="1698845720951" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="61AAB13C9CAC391F368713104D27F41E">Description of Orthadenella coulsoni sp. nov. (Acari: Mesostigmata: Melicharidae) from Siberia with a key to the females of Orthadenella</mods:title>
</mods:titleInfo>
<mods:name id="9D087E68A7FDC1DB3548CC794149504E" type="personal">
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<mods:namePart id="CC5C2D14D736200281542C88730C2E3F">Gwiazdowicz, Dariusz J.</mods:namePart>
<mods:affiliation id="F1F838E66EA0F76229B25FA5810EE49E">Faculty of Forestry, Poznan University of Life Sciences, Poznan, Poland;</mods:affiliation>
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<mods:namePart id="7D2C1EC43D4EA2882ABFB78B109639F9">Marchenko, Irina I.</mods:namePart>
<mods:affiliation id="B34ABDE3018B1F30F6424BACF0F4DFB7">Institute of Systematics and Ecology of Animals, Russian Academy of Sciences, Novosibirsk, Russia</mods:affiliation>
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<mods:namePart id="E5BBE66089F12A606BC26D4F5C1F928D">Teodorowicz, Ewa</mods:namePart>
<mods:affiliation id="045281DD58D872F8537B404D367CBDC6">Faculty of Forestry, Poznan University of Life Sciences, Poznan, Poland;</mods:affiliation>
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<mods:title id="39A6554F7F2190454D617F9FBAA43FC9">Journal of Natural History</mods:title>
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<mods:part id="7BFA700A534353B73000BA98B93C925F">
<mods:date id="B029CBC25FEB31B82E16BDBD06DD6EBC">2014</mods:date>
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<mods:number id="1205CACA9B502C0E1281272E7A3D293A">2014-11-06</mods:number>
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<mods:number id="C2CE748AA8964A0844CCBF9A610DEAD3">49</mods:number>
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<mods:number id="3C69EFF101B19EA87CD0C87D1328C71B">27</mods:number>
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<mods:start id="6CE1A8D8FF4D12F66303A93CD799434B">1659</mods:start>
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<treatment id="03EA8788FFD9FFE7576CFEC1FC30FA6A" ID-DOI="http://doi.org/10.5281/zenodo.4324243" ID-GBIF-Taxon="167371038" ID-Zenodo-Dep="4324243" LSID="urn:lsid:plazi:treatment:03EA8788FFD9FFE7576CFEC1FC30FA6A" httpUri="http://treatment.plazi.org/id/03EA8788FFD9FFE7576CFEC1FC30FA6A" lastPageId="11" lastPageNumber="1670" pageId="2" pageNumber="1661">
<subSubSection id="C3596515FFD9FFEE576CFEC1FCC4FEF9" box="[522,862,279,304]" pageId="2" pageNumber="1661" type="nomenclature">
<paragraph id="8BFC369EFFD9FFEE576CFEC1FCC4FEF9" blockId="2.[473,911,279,336]" box="[522,862,279,304]" pageId="2" pageNumber="1661">
<heading id="D0B481F2FFD9FFEE576CFEC1FCC4FEF9" box="[522,862,279,304]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="1661" reason="2">
<taxonomicName id="4C434D1DFFD9FFEE576CFEC1FD61FEE6" authority="Gwiazdowicz &amp; Marchenko &amp; Teodorowicz, 2014" authorityName="Gwiazdowicz &amp; Marchenko &amp; Teodorowicz" authorityYear="2014" box="[522,763,279,303]" class="Arachnida" family="Ascidae" genus="Orthadenella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="1661" phylum="Arthropoda" rank="species" species="coulsoni" status="sp. nov.">
<emphasis id="B937EA8CFFD9FFEE576CFEC1FD61FEE6" bold="true" box="[522,763,279,303]" italics="true" pageId="2" pageNumber="1661">Orthadenella coulsoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20457F7FFD9FFEE5662FECFFCC4FEF9" box="[772,862,281,304]" pageId="2" pageNumber="1661" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3596515FFD9FFEE54BFFEEEFC15FE99" box="[473,911,312,336]" pageId="2" pageNumber="1661" type="description">
<paragraph id="8BFC369EFFD9FFEE54BFFEEEFC15FE99" blockId="2.[473,911,279,336]" box="[473,911,312,336]" pageId="2" pageNumber="1661">
(
<figureCitation id="13782A1BFFD9FFEE5484FEEEFDD6FE99" box="[482,588,312,336]" captionStart="Figure 1" captionStartId="3.[144,211,1640,1662]" captionTargetBox="[211,1037,168,1608]" captionTargetId="figure@3.[211,1037,168,1608]" captionTargetPageId="3" captionText="Figure 1. Orthadenella coulsoni sp. nov., female: dorsal view of idiosoma." figureDoi="http://doi.org/10.5281/zenodo.4006374" httpUri="https://zenodo.org/record/4006374/files/figure.png" pageId="2" pageNumber="1661">Figures 1</figureCitation>
,
<figureCitation id="13782A1BFFD9FFEE573AFEEEFDF3FE99" box="[604,617,312,336]" captionStart="Figure 2" captionStartId="4.[212,279,1640,1662]" captionTargetBox="[263,1120,168,1608]" captionTargetId="figure@4.[263,1121,168,1608]" captionTargetPageId="4" captionText="Figure 2. Orthadenella coulsoni sp. nov., female: ventral view of idiosoma." figureDoi="http://doi.org/10.5281/zenodo.4006376" httpUri="https://zenodo.org/record/4006376/files/figure.png" pageId="2" pageNumber="1661">2</figureCitation>
,
<figureCitation id="13782A1BFFD9FFEE571FFEEEFD24FE99" box="[633,702,312,336]" captionStart="Figure 3" captionStartId="5.[144,211,1463,1485]" captionTargetBox="[145,1104,168,1431]" captionTargetId="figure@5.[144,1104,168,1431]" captionTargetPageId="5" captionText="Figure 3. Orthadenella coulsoni sp. nov., female: (A) tritosternum; (B) gnathosoma; (C) sperm access system; (D) epistomes; (E) chelicera; (F) dorsal of palptarsus; (G) ventral of palptarsus." figureDoi="http://doi.org/10.5281/zenodo.4006378" httpUri="https://zenodo.org/record/4006378/files/figure.png" pageId="2" pageNumber="1661">3AG</figureCitation>
,
<figureCitation id="13782A1BFFD9FFEE57A8FEEEFC89FE86" box="[718,787,312,336]" captionStart="Figure 4" captionStartId="6.[212,279,1409,1431]" captionTargetBox="[292,1092,168,1377]" captionTargetId="figure@6.[292,1092,168,1377]" captionTargetPageId="6" captionText="Figure 4. Orthadenella coulsoni sp. nov., female: (A) leg I; (B) leg II; (C) leg III; (D) leg IV." figureDoi="http://doi.org/10.5281/zenodo.4006380" httpUri="https://zenodo.org/record/4006380/files/figure.png" pageId="2" pageNumber="1661">4AD</figureCitation>
,
<figureCitation id="13782A1BFFD9FFEE5645FEEEFCAAFE99" box="[803,816,312,336]" captionStart="Figure 5" captionStartId="7.[144,211,1640,1662]" captionTargetBox="[196,1050,168,1608]" captionTargetId="figure@7.[196,1051,168,1608]" captionTargetPageId="7" captionText="Figure 5. Orthadenella coulsoni sp. nov., male: ventral view of idiosoma." figureDoi="http://doi.org/10.5281/zenodo.4006384" httpUri="https://zenodo.org/record/4006384/files/figure.png" pageId="2" pageNumber="1661">5</figureCitation>
,
<figureCitation id="13782A1BFFD9FFEE5627FEEEFC1CFE86" box="[833,902,312,336]" captionStart="Figure 6" captionStartId="8.[212,279,1581,1603]" captionTargetBox="[212,1172,168,1549]" captionTargetId="figure@8.[212,1172,168,1549]" captionTargetPageId="8" captionText="Figure 6. Orthadenella coulsoni sp. nov., male: (A) tritosternum; (B) gnathosoma; (C) chelicerae; (D) epistomes." figureDoi="http://doi.org/10.5281/zenodo.4006386" httpUri="https://zenodo.org/record/4006386/files/figure.png" pageId="2" pageNumber="1661">6AD</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3596515FFD9FFEE55B2FE42FEDAFD1C" pageId="2" pageNumber="1661" type="materials_examined">
<paragraph id="8BFC369EFFD9FFEE55B2FE42FEEAFE62" blockId="2.[212,1171,403,499]" box="[212,368,403,427]" pageId="2" pageNumber="1661">
<heading id="D0B481F2FFD9FFEE55B2FE42FEEAFE62" box="[212,368,403,427]" fontSize="10" level="3" pageId="2" pageNumber="1661" reason="5">
<emphasis id="B937EA8CFFD9FFEE55B2FE42FEEAFE62" box="[212,368,403,427]" italics="true" pageId="2" pageNumber="1661">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8BFC369EFFD9FFEE55B2FE6DFB1DFE3B" blockId="2.[212,1171,403,499]" pageId="2" pageNumber="1661">
<materialsCitation id="3B2B3CC3FFD9FFEE55B2FE6DFB1DFE3B" ID-GBIF-Occurrence="2849225301" collectingDate="2011-08-17" collectorName="I. I. Marchenko." county="Female" elevation="700" latitude="51.5" location="Shebalino District" longLatPrecision="1090" longitude="85.03333" municipality="North" pageId="2" pageNumber="1661" specimenCount="1" stateProvince="Altay" typeStatus="holotype">
<emphasis id="B937EA8CFFD9FFEE55B2FE6DFED8FE1A" box="[212,322,443,467]" italics="true" pageId="2" pageNumber="1661">
<typeStatus id="54F8883CFFD9FFEE55B2FE6DFEA4FE1A" box="[212,318,443,467]" pageId="2" pageNumber="1661" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCounty id="629D4E12FFD9FFEE542CFE6CFE3AFE1B" box="[330,416,442,466]" pageId="2" pageNumber="1661">Female</collectingCounty>
,
<collectingMunicipality id="6B98ACE4FFD9FFEE54CBFE6CFE68FE1B" box="[429,498,442,466]" pageId="2" pageNumber="1661">North</collectingMunicipality>
<collectingRegion id="4987F87CFFD9FFEE549DFE6CFDAEFE1B" box="[507,564,442,466]" country="Russia" name="Altay" pageId="2" pageNumber="1661">Altai</collectingRegion>
(
<geoCoordinate id="EE775059FFD9FFEE5723FE6CFD2EFE1B" box="[581,692,442,467]" degrees="51" direction="north" minutes="30" orientation="latitude" pageId="2" pageNumber="1661" precision="925" value="51.5">
51° 30
<emphasis id="B937EA8CFFD9FFEE57EBFE6DFD09FE1A" bold="true" box="[653,659,443,467]" italics="true" pageId="2" pageNumber="1661"></emphasis>
N
</geoCoordinate>
,
<geoCoordinate id="EE775059FFD9FFEE57A7FE6CFCB6FE1B" box="[705,812,442,467]" degrees="85" direction="east" minutes="02" orientation="longitude" pageId="2" pageNumber="1661" precision="925" value="85.03333">
85° 02
<emphasis id="B937EA8CFFD9FFEE566CFE6DFC8AFE1A" bold="true" box="[778,784,443,467]" italics="true" pageId="2" pageNumber="1661"></emphasis>
E
</geoCoordinate>
),
<elevation id="006ED1ADFFD9FFEE5624FE6CFC28FE1B" box="[834,946,442,466]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="2" pageNumber="1661" unit="m" value="700.0">
<quantity id="4CBB9B7BFFD9FFEE5624FE6CFC13FE1B" box="[834,905,442,466]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="2" pageNumber="1661" unit="m" value="700.0">700 m</quantity>
asl
</elevation>
,
<location id="8E9C6045FFD9FFEE56D9FE6CFB15FE1B" LSID="urn:lsid:plazi:treatment:03EA8788FFD9FFE7576CFEC1FC30FA6A:8E9C6045FFD9FFEE56D9FE6CFB15FE1B" box="[959,1167,442,466]" county="Female" latitude="51.5" longLatPrecision="1090" longitude="85.03333" municipality="North" name="Shebalino District" pageId="2" pageNumber="1661" stateProvince="Altay">Shebalino District</location>
,
<taxonomicName id="4C434D1DFFD9FFEE55B2FE0DFEF1FE3A" box="[212,363,475,499]" class="Pinopsida" family="Pinaceae" genus="Larix" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="2" pageNumber="1661" phylum="Tracheophyta" rank="species" species="sibirica">
<emphasis id="B937EA8CFFD9FFEE55B2FE0DFEF1FE3A" box="[212,363,475,499]" italics="true" pageId="2" pageNumber="1661">Larix sibirica</emphasis>
</taxonomicName>
forest, on the hillside, in litter,
<date id="FFFD105EFFD9FFEE57B1FE0CFC16FE3B" box="[727,908,474,498]" pageId="2" pageNumber="1661" value="2011-08-17">
<collectingDate id="EFB9E9B6FFD9FFEE57B1FE0CFC16FE3B" box="[727,908,474,498]" pageId="2" pageNumber="1661" value="2011-08-17">17 August 2011</collectingDate>
</date>
, coll.
<collectorName id="26B65348FFD9FFEE56B4FE0DFB1DFE3B" box="[978,1159,474,498]" pageId="2" pageNumber="1661">I.I. Marchenko.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="8BFC369EFFD9FFEE55B2FDC8FEDAFD1C" blockId="2.[212,1172,541,725]" pageId="2" pageNumber="1661">
<materialsCitation id="3B2B3CC3FFD9FFEE55B2FDC8FBC6FDBC" ID-GBIF-Occurrence="2849225303" collectingDate="1999-06-10" collectorName="I. I. Marchenko" county="Two" elevation="1200" latitude="51.133335" location="Shebalino District" longLatPrecision="1092" longitude="85.566666" municipality="North" pageId="2" pageNumber="1661" specimenCount="11" specimenCount-female="11" stateProvince="Altay" typeStatus="paratype">
<emphasis id="B937EA8CFFD9FFEE55B2FDC8FED0FDFC" box="[212,330,542,565]" italics="true" pageId="2" pageNumber="1661">
<typeStatus id="54F8883CFFD9FFEE55B2FDC8FEDDFDFC" box="[212,327,542,565]" pageId="2" pageNumber="1661" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectingCounty id="629D4E12FFD9FFEE5434FDC8FE1FFDFC" box="[338,389,542,565]" pageId="2" pageNumber="1661">Two</collectingCounty>
females, the same data as holotype;
<specimenCount id="9D45FD17FFD9FFEE564EFDCBFC38FDFC" box="[808,930,541,565]" pageId="2" pageNumber="1661" type="female">11 females</specimenCount>
,
<collectingMunicipality id="6B98ACE4FFD9FFEE56C9FDCBFC6EFDFC" box="[943,1012,541,565]" pageId="2" pageNumber="1661">North</collectingMunicipality>
<collectingRegion id="4987F87CFFD9FFEE569AFDCBFBAFFDFC" box="[1020,1077,541,565]" country="Russia" name="Altay" pageId="2" pageNumber="1661">Altai</collectingRegion>
(
<geoCoordinate id="EE775059FFD9FFEE5120FDCBFF76FD9C" degrees="51" direction="north" minutes="08" orientation="latitude" pageId="2" pageNumber="1661" precision="925" value="51.133335">
51° 08
<emphasis id="B937EA8CFFD9FFEE51E8FDCBFB0EFDFC" bold="true" box="[1166,1172,541,565]" italics="true" pageId="2" pageNumber="1661"></emphasis>
N
</geoCoordinate>
,
<geoCoordinate id="EE775059FFD9FFEE5466FDEBFEE2FD9C" box="[256,376,573,597]" degrees="85" direction="east" minutes="34" orientation="longitude" pageId="2" pageNumber="1661" precision="925" value="85.566666">
85° 34
<emphasis id="B937EA8CFFD9FFEE5429FDEBFECFFD9C" bold="true" box="[335,341,573,597]" italics="true" pageId="2" pageNumber="1661"></emphasis>
E
</geoCoordinate>
),
<elevation id="006ED1ADFFD9FFEE54F3FDEBFDB9FD9C" box="[405,547,573,597]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="1661" unit="m" value="1200.0">
<quantity id="4CBB9B7BFFD9FFEE54F3FDEBFE6AFD9C" box="[405,496,573,597]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="1661" unit="m" value="1200.0">1200 m</quantity>
asl
</elevation>
.,
<location id="8E9C6045FFD9FFEE575DFDEBFC88FD9C" LSID="urn:lsid:plazi:treatment:03EA8788FFD9FFE7576CFEC1FC30FA6A:8E9C6045FFD9FFEE575DFDEBFC88FD9C" box="[571,786,573,597]" county="Two" latitude="51.133335" longLatPrecision="1092" longitude="85.566666" municipality="North" name="Shebalino District" pageId="2" pageNumber="1661" stateProvince="Altay">Shebalino District</location>
, environs of
<location id="8E9C6045FFD9FFEE56DDFDEBFB15FD9C" LSID="urn:lsid:plazi:treatment:03EA8788FFD9FFE7576CFEC1FC30FA6A:8E9C6045FFD9FFEE56DDFDEBFB15FD9C" box="[955,1167,573,597]" county="Two" latitude="51.133335" longLatPrecision="1092" longitude="85.566666" municipality="North" name="Topuchaja village" pageId="2" pageNumber="1661" stateProvince="Altay">Topuchaja village</location>
, swamped forest of
<taxonomicName id="4C434D1DFFD9FFEE54D5FD88FDD1FDBC" authorityName="hort. ex Gordon" authorityYear="1858" box="[435,587,605,629]" class="Pinopsida" family="Pinaceae" genus="Picea" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="2" pageNumber="1661" phylum="Tracheophyta" rank="species" species="sibirica">
<emphasis id="B937EA8CFFD9FFEE54D5FD88FDD1FDBC" box="[435,587,605,629]" italics="true" pageId="2" pageNumber="1661">Picea sibirica</emphasis>
</taxonomicName>
, in litter,
<date id="FFFD105EFFD9FFEE57A2FD8BFCFAFDBC" box="[708,864,605,629]" pageId="2" pageNumber="1661" value="1999-06-10">
<collectingDate id="EFB9E9B6FFD9FFEE57A2FD8BFCFAFDBC" box="[708,864,605,629]" pageId="2" pageNumber="1661" value="1999-06-10">10 June 1999</collectingDate>
</date>
, coll.
<collectorName id="26B65348FFD9FFEE56CFFD88FBC6FDBC" box="[937,1116,605,629]" pageId="2" pageNumber="1661">I.I. Marchenko</collectorName>
</materialsCitation>
;
<materialsCitation id="3B2B3CC3FFD9FFEE510DFD8BFEDAFD1C" ID-GBIF-Occurrence="2849225302" collectingDate="1964-07-16" collectorName="S. K. Stebaeva." county="Central" elevation="2000" latitude="50.2" location="river Kuraika" longLatPrecision="1098" longitude="88.05" municipality="Kurai Ridge" pageId="2" pageNumber="1661" specimenCount="1" stateProvince="Altay" typeStatus="paratype">
five females and eight males,
<collectingCounty id="629D4E12FFD9FFEE5492FDABFDD2FD5C" box="[500,584,637,661]" pageId="2" pageNumber="1661">Central</collectingCounty>
<collectingRegion id="4987F87CFFD9FFEE5736FDABFD13FD5C" box="[592,649,637,661]" country="Russia" name="Altay" pageId="2" pageNumber="1661">Altai</collectingRegion>
(
<geoCoordinate id="EE775059FFD9FFEE57FCFDABFC90FD5C" box="[666,778,637,661]" degrees="50" direction="north" minutes="12" orientation="latitude" pageId="2" pageNumber="1661" precision="925" value="50.2">
50° 12
<emphasis id="B937EA8CFFD9FFEE5785FDABFD73FD5C" bold="true" box="[739,745,637,661]" italics="true" pageId="2" pageNumber="1661"></emphasis>
N
</geoCoordinate>
,
<geoCoordinate id="EE775059FFD9FFEE567EFDABFC19FD5C" box="[792,899,637,661]" degrees="88" direction="east" minutes="03" orientation="longitude" pageId="2" pageNumber="1661" precision="925" value="88.05">
88° 03
<emphasis id="B937EA8CFFD9FFEE5606FDABFCFCFD5C" bold="true" box="[864,870,637,661]" italics="true" pageId="2" pageNumber="1661"></emphasis>
E
</geoCoordinate>
),
<collectingMunicipality id="6B98ACE4FFD9FFEE56FFFDABFBB2FD5C" box="[921,1064,637,661]" pageId="2" pageNumber="1661">Kurai Ridge</collectingMunicipality>
foothills,
<elevation id="006ED1ADFFD9FFEE55B2FD4BFEC5FD7C" box="[212,351,669,693]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="1661" unit="m" value="2000.0">
<quantity id="4CBB9B7BFFD9FFEE55B2FD4BFEB7FD7C" box="[212,301,669,693]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="1661" unit="m" value="2000.0">2000 m</quantity>
asl
</elevation>
., floodplain
<location id="8E9C6045FFD9FFEE549FFD4BFD06FD7C" LSID="urn:lsid:plazi:treatment:03EA8788FFD9FFE7576CFEC1FC30FA6A:8E9C6045FFD9FFEE549FFD4BFD06FD7C" box="[505,668,669,693]" county="Central" latitude="50.2" longLatPrecision="1098" longitude="88.05" municipality="Kurai Ridge" name="river Kuraika" pageId="2" pageNumber="1661" stateProvince="Altay">river Kuraika</location>
, shrub, in litter,
<date id="FFFD105EFFD9FFEE5614FD4BFB8AFD7C" box="[882,1040,669,693]" pageId="2" pageNumber="1661" value="1964-07-16">
<collectingDate id="EFB9E9B6FFD9FFEE5614FD4BFB8AFD7C" box="[882,1040,669,693]" pageId="2" pageNumber="1661" value="1964-07-16">16 July 1964</collectingDate>
</date>
, coll.
<collectorName id="26B65348FFD9FFEE5107FD4BFEDAFD1C" pageId="2" pageNumber="1661">S.K. Stebaeva.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3596515FFD9FFE655B2FCCFFCABFB74" lastPageId="10" lastPageNumber="1669" pageId="2" pageNumber="1661" type="diagnosis">
<paragraph id="8BFC369EFFD9FFEE55B2FCCFFEDAFCF9" blockId="2.[212,1142,793,856]" box="[212,320,793,816]" pageId="2" pageNumber="1661">
<heading id="D0B481F2FFD9FFEE55B2FCCFFEDAFCF9" box="[212,320,793,816]" fontSize="10" level="3" pageId="2" pageNumber="1661" reason="5">
<emphasis id="B937EA8CFFD9FFEE55B2FCCFFEDAFCF9" box="[212,320,793,816]" italics="true" pageId="2" pageNumber="1661">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BFC369EFFD9FFEE55B2FC96FBECFC91" blockId="2.[212,1142,793,856]" box="[212,1142,832,856]" pageId="2" pageNumber="1661">
<emphasis id="B937EA8CFFD9FFEE55B2FC96FEBEFC91" box="[212,292,832,856]" italics="true" pageId="2" pageNumber="1661">Female</emphasis>
(
<emphasis id="B937EA8CFFD9FFEE5453FC97FEC7FC91" box="[309,349,833,856]" italics="true" pageId="2" pageNumber="1661">n =</emphasis>
19). Idiosoma oval,
<quantity id="4CBB9B7BFFD9FFEE572DFC96FD4CFC91" box="[587,726,832,856]" metricMagnitude="-1" metricUnit="m" metricValue="4.1" metricValueMax="4.2" metricValueMin="4.0" pageId="2" pageNumber="1661" unit="mm" value="410.0" valueMax="420.0" valueMin="400.0">400420 µm</quantity>
in length and
<quantity id="4CBB9B7BFFD9FFEE5618FC96FB92FC91" box="[894,1032,832,856]" metricMagnitude="-1" metricUnit="m" metricValue="2.325" metricValueMax="2.4" metricValueMin="2.25" pageId="2" pageNumber="1661" unit="mm" value="232.5" valueMax="240.0" valueMin="225.0">225240 µm</quantity>
in width.
</paragraph>
<paragraph id="8BFC369EFFD9FFEE55B2FC55FED7FAB0" blockId="2.[212,1172,899,1401]" pageId="2" pageNumber="1661">
<emphasis id="B937EA8CFFD9FFEE55B2FC55FEBCFC52" box="[212,294,899,923]" italics="true" pageId="2" pageNumber="1661">Dorsal.</emphasis>
Holodorsal shield bearing 43 pairs of simple setae and among them 20 are situated on the opisthonotal posterior part. The shortest setae are given (in µm):
<emphasis id="B937EA8CFFD9FFEE5119FC75FB0EFC72" box="[1151,1172,931,955]" italics="true" pageId="2" pageNumber="1661">j2</emphasis>
(78),
<emphasis id="B937EA8CFFD9FFEE5447FC15FEAFFC12" box="[289,309,963,987]" italics="true" pageId="2" pageNumber="1661">j3</emphasis>
(1011),
<emphasis id="B937EA8CFFD9FFEE54CCFC15FE59FC12" box="[426,451,963,987]" italics="true" pageId="2" pageNumber="1661">z1</emphasis>
(57),
<emphasis id="B937EA8CFFD9FFEE577BFC15FDAFFC12" box="[541,565,963,987]" italics="true" pageId="2" pageNumber="1661">z2</emphasis>
(1012),
<emphasis id="B937EA8CFFD9FFEE57CCFC15FD58FC12" box="[682,706,963,987]" italics="true" pageId="2" pageNumber="1661">s1</emphasis>
(67),
<emphasis id="B937EA8CFFD9FFEE567DFC15FCACFC12" box="[795,822,963,987]" italics="true" pageId="2" pageNumber="1661">J5</emphasis>
(710); the longest are those labelled
<emphasis id="B937EA8CFFD9FFEE5453FC35FED0FC32" box="[309,330,995,1019]" italics="true" pageId="2" pageNumber="1661">j1</emphasis>
(2022),
<emphasis id="B937EA8CFFD9FFEE54DEFC35FE43FC32" box="[440,473,995,1019]" italics="true" pageId="2" pageNumber="1661">Z5</emphasis>
(3841),
<emphasis id="B937EA8CFFD9FFEE572EFC35FDF2FC32" box="[584,616,995,1019]" italics="true" pageId="2" pageNumber="1661">Z4</emphasis>
(2526),
<emphasis id="B937EA8CFFD9FFEE57BEFC35FD6EFC32" box="[728,756,995,1019]" italics="true" pageId="2" pageNumber="1661">S4</emphasis>
(2527) and
<emphasis id="B937EA8CFFD9FFEE56F7FC35FC37FC32" box="[913,941,995,1019]" italics="true" pageId="2" pageNumber="1661">S5</emphasis>
(2527). Other seta in
<emphasis id="B937EA8CFFD9FFEE5596FBD5FF6DFBD3" box="[240,247,1027,1050]" italics="true" pageId="2" pageNumber="1661">j</emphasis>
,
<emphasis id="B937EA8CFFD9FFEE5463FBD5FE8BFBD3" box="[261,273,1027,1050]" italics="true" pageId="2" pageNumber="1661">z</emphasis>
and
<emphasis id="B937EA8CFFD9FFEE542EFBD5FEC8FBD3" box="[328,338,1027,1050]" italics="true" pageId="2" pageNumber="1661">s</emphasis>
rows, on the podonotal part, are of a median length from
<quantity id="4CBB9B7BFFD9FFEE56B9FBD4FBF9FBD2" box="[991,1123,1026,1051]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" metricValueMax="1.7" metricValueMin="1.2" pageId="2" pageNumber="1661" unit="mm" value="14.5" valueMax="17.0" valueMin="12.0">12 to 17 µm</quantity>
and are as follows:
<emphasis id="B937EA8CFFD9FFEE541DFBF4FE0AFBF3" box="[379,400,1058,1082]" italics="true" pageId="2" pageNumber="1661">j4</emphasis>
(1315),
<emphasis id="B937EA8CFFD9FFEE549FFBF4FD97FBF3" box="[505,525,1058,1082]" italics="true" pageId="2" pageNumber="1661">j5</emphasis>
(1517),
<emphasis id="B937EA8CFFD9FFEE5711FBF4FD11FBF3" box="[631,651,1058,1082]" italics="true" pageId="2" pageNumber="1661">j6</emphasis>
(1517),
<emphasis id="B937EA8CFFD9FFEE5792FBF4FC97FBF3" box="[756,781,1058,1082]" italics="true" pageId="2" pageNumber="1661">z3</emphasis>
(1215),
<emphasis id="B937EA8CFFD9FFEE5611FBF4FC0AFBF3" box="[887,912,1058,1082]" italics="true" pageId="2" pageNumber="1661">z4</emphasis>
(1215),
<emphasis id="B937EA8CFFD9FFEE569FFBF4FB88FBF3" box="[1017,1042,1058,1082]" italics="true" pageId="2" pageNumber="1661">z5</emphasis>
(1215),
<emphasis id="B937EA8CFFD9FFEE511DFBF4FB0EFBF3" box="[1147,1172,1058,1082]" italics="true" pageId="2" pageNumber="1661">z6</emphasis>
(1215),
<emphasis id="B937EA8CFFD9FFEE5451FB94FED4FB93" box="[311,334,1090,1114]" italics="true" pageId="2" pageNumber="1661">s2</emphasis>
(1012),
<emphasis id="B937EA8CFFD9FFEE54DFFB94FE4AFB93" box="[441,464,1090,1114]" italics="true" pageId="2" pageNumber="1661">s3</emphasis>
(1215),
<emphasis id="B937EA8CFFD9FFEE575DFB94FDC8FB93" box="[571,594,1090,1114]" italics="true" pageId="2" pageNumber="1661">s4</emphasis>
(1215),
<emphasis id="B937EA8CFFD9FFEE57DBFB94FD4EFB93" box="[701,724,1090,1114]" italics="true" pageId="2" pageNumber="1661">s5</emphasis>
(1215),
<emphasis id="B937EA8CFFD9FFEE5659FB94FCCDFB93" box="[831,855,1090,1114]" italics="true" pageId="2" pageNumber="1661">s6</emphasis>
(1215), and slightly longer in
<emphasis id="B937EA8CFFD9FFEE5592FBB5FE9BFBB3" box="[244,257,1123,1146]" italics="true" pageId="2" pageNumber="1661">J</emphasis>
,
<emphasis id="B937EA8CFFD9FFEE5474FBB5FEBFFBB3" box="[274,293,1123,1146]" italics="true" pageId="2" pageNumber="1661">Z</emphasis>
and
<emphasis id="B937EA8CFFD9FFEE5402FBB4FEEEFBB3" box="[356,372,1122,1146]" italics="true" pageId="2" pageNumber="1661">S</emphasis>
rows, on the opisthonotal part, from
<quantity id="4CBB9B7BFFD9FFEE5654FBB4FC59FBB3" box="[818,963,1122,1146]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" metricValueMax="2.2" metricValueMin="1.5" pageId="2" pageNumber="1661" unit="mm" value="18.5" valueMax="22.0" valueMin="15.0">15 to 22 µm</quantity>
, consecutively:
<emphasis id="B937EA8CFFD9FFEE511FFBB4FB0EFBB3" box="[1145,1172,1122,1146]" italics="true" pageId="2" pageNumber="1661">J1</emphasis>
(1517),
<emphasis id="B937EA8CFFD9FFEE545EFB54FEC8FB53" box="[312,338,1154,1178]" italics="true" pageId="2" pageNumber="1661">J2</emphasis>
(1517),
<emphasis id="B937EA8CFFD9FFEE54D9FB54FE40FB53" box="[447,474,1154,1178]" italics="true" pageId="2" pageNumber="1661">J3</emphasis>
(1517),
<emphasis id="B937EA8CFFD9FFEE5721FB54FDFBFB53" box="[583,609,1154,1178]" italics="true" pageId="2" pageNumber="1661">J4</emphasis>
(2022),
<emphasis id="B937EA8CFFD9FFEE57A8FB54FD75FB53" box="[718,751,1154,1178]" italics="true" pageId="2" pageNumber="1661">Z1</emphasis>
(1517),
<emphasis id="B937EA8CFFD9FFEE563AFB54FCE7FB53" box="[860,893,1154,1178]" italics="true" pageId="2" pageNumber="1661">Z2</emphasis>
(1517),
<emphasis id="B937EA8CFFD9FFEE568FFB54FB90FB53" box="[1001,1034,1154,1178]" italics="true" pageId="2" pageNumber="1661">Z3</emphasis>
(1517),
<emphasis id="B937EA8CFFD9FFEE5111FB54FB09FB53" box="[1143,1171,1154,1178]" italics="true" pageId="2" pageNumber="1661">S1</emphasis>
(1722),
<emphasis id="B937EA8CFFD9FFEE5450FB74FECEFB73" box="[310,340,1186,1210]" italics="true" pageId="2" pageNumber="1661">S2</emphasis>
(1722),
<emphasis id="B937EA8CFFD9FFEE54D8FB74FE41FB73" box="[446,475,1186,1210]" italics="true" pageId="2" pageNumber="1661">S3</emphasis>
(1722). All setae in the marginal row
<emphasis id="B937EA8CFFD9FFEE56F1FB75FC5BFB73" box="[919,961,1186,1210]" italics="true" pageId="2" pageNumber="1661">rR</emphasis>
are located on the shield, none on the soft membrane. They range in length in a row
<emphasis id="B937EA8CFFD9FFEE56DEFB14FC58FB10" box="[952,962,1218,1241]" italics="true" pageId="2" pageNumber="1661">r</emphasis>
from
<quantity id="4CBB9B7BFFD9FFEE5161FB17FB15FB13" box="[1031,1167,1217,1242]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="2" pageNumber="1661" unit="mm" value="16.0" valueMax="17.0" valueMin="15.0">15 to 17 µm</quantity>
,
<emphasis id="B937EA8CFFD9FFEE55B2FB34FF71FB33" box="[212,235,1250,1274]" italics="true" pageId="2" pageNumber="1661">r2</emphasis>
(1517),
<emphasis id="B937EA8CFFD9FFEE543CFB34FEEBFB33" box="[346,369,1250,1274]" italics="true" pageId="2" pageNumber="1661">r4</emphasis>
(1517),
<emphasis id="B937EA8CFFD9FFEE5486FB34FE6DFB33" box="[480,503,1250,1274]" italics="true" pageId="2" pageNumber="1661">r5</emphasis>
(1517),
<emphasis id="B937EA8CFFD9FFEE5700FB34FDE7FB33" box="[614,637,1250,1274]" italics="true" pageId="2" pageNumber="1661">r6</emphasis>
(1517), excepting humeral setae
<emphasis id="B937EA8CFFD9FFEE5161FB34FB85FB33" box="[1031,1055,1250,1274]" italics="true" pageId="2" pageNumber="1661">r3</emphasis>
=
<quantity id="4CBB9B7BFFD9FFEE5122FB37FB15FB33" box="[1092,1167,1249,1274]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" pageId="2" pageNumber="1661" unit="mm" value="21.0">21 µm</quantity>
, and in row
<emphasis id="B937EA8CFFD9FFEE5432FAD4FEFDFAD0" box="[340,359,1282,1305]" italics="true" pageId="2" pageNumber="1661">R</emphasis>
are about
<quantity id="4CBB9B7BFFD9FFEE5487FAD7FDFDFAD3" box="[481,615,1281,1306]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" metricValueMax="2.0" metricValueMin="1.7" pageId="2" pageNumber="1661" unit="mm" value="18.5" valueMax="20.0" valueMin="17.0">17 to 20 µm</quantity>
as given:
<emphasis id="B937EA8CFFD9FFEE57B0FAD7FD6DFAD0" box="[726,759,1281,1305]" italics="true" pageId="2" pageNumber="1661">R1</emphasis>
(1720),
<emphasis id="B937EA8CFFD9FFEE5606FAD7FC1BFAD0" box="[864,897,1281,1305]" italics="true" pageId="2" pageNumber="1661">R2</emphasis>
(1720),
<emphasis id="B937EA8CFFD9FFEE568CFAD7FB90FAD0" box="[1002,1034,1281,1305]" italics="true" pageId="2" pageNumber="1661">R3</emphasis>
(1720),
<emphasis id="B937EA8CFFD9FFEE5115FAD7FB09FAD0" box="[1139,1171,1281,1305]" italics="true" pageId="2" pageNumber="1661">R4</emphasis>
(1720),
<emphasis id="B937EA8CFFD9FFEE545AFAF4FED5FAF0" box="[316,335,1314,1337]" italics="true" pageId="2" pageNumber="1661">R</emphasis>
(1720). Setae
<emphasis id="B937EA8CFFD9FFEE5768FAF7FDC9FAF0" box="[526,595,1313,1337]" italics="true" pageId="2" pageNumber="1661">r2R5</emphasis>
at clearly delineated marginal strip. Holodorsal shield is more or less conspicuously covered with a reticulate ornamentation (
<figureCitation id="13782A1BFFD9FFEE55BAFAB7FEA6FAB0" box="[220,316,1377,1401]" captionStart="Figure 1" captionStartId="3.[144,211,1640,1662]" captionTargetBox="[211,1037,168,1608]" captionTargetId="figure@3.[211,1037,168,1608]" captionTargetPageId="3" captionText="Figure 1. Orthadenella coulsoni sp. nov., female: dorsal view of idiosoma." figureDoi="http://doi.org/10.5281/zenodo.4006374" httpUri="https://zenodo.org/record/4006374/files/figure.png" pageId="2" pageNumber="1661">Figure 1</figureCitation>
).
</paragraph>
<paragraph id="8BFC369EFFD9FFE555B2FA72FE81FDF6" blockId="2.[212,1172,1444,1691]" lastBlockId="9.[144,1104,168,575]" lastPageId="9" lastPageNumber="1668" pageId="2" pageNumber="1661">
<emphasis id="B937EA8CFFD9FFEE55B2FA72FEB5FA75" box="[212,303,1444,1468]" italics="true" pageId="2" pageNumber="1661">Ventral.</emphasis>
The tritosternum with base
<quantity id="4CBB9B7BFFD9FFEE5716FA72FD45FA75" box="[624,735,1444,1468]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="2" pageNumber="1661" unit="mm" value="10.5" valueMax="11.0" valueMin="10.0">1011 µm</quantity>
wide and
<quantity id="4CBB9B7BFFD9FFEE5633FA72FC59FA75" box="[853,963,1444,1468]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="2" pageNumber="1661" unit="mm" value="14.5" valueMax="15.0" valueMin="14.0">1415 µm</quantity>
long with laciniae
<quantity id="4CBB9B7BFFD9FFEE55B2FA12FEDFFA15" box="[212,325,1476,1500]" metricMagnitude="-2" metricUnit="m" metricValue="5.95" metricValueMax="6.2" metricValueMin="5.7" pageId="2" pageNumber="1661" unit="mm" value="59.5" valueMax="62.0" valueMin="57.0">5762 µm</quantity>
long (excluding base) with a fused area, free area for about 0.7 of total length (
<figureCitation id="13782A1BFFD9FFEE544AFA32FE38FA35" box="[300,418,1508,1532]" captionStart="Figure 3" captionStartId="5.[144,211,1463,1485]" captionTargetBox="[145,1104,168,1431]" captionTargetId="figure@5.[144,1104,168,1431]" captionTargetPageId="5" captionText="Figure 3. Orthadenella coulsoni sp. nov., female: (A) tritosternum; (B) gnathosoma; (C) sperm access system; (D) epistomes; (E) chelicera; (F) dorsal of palptarsus; (G) ventral of palptarsus." figureDoi="http://doi.org/10.5281/zenodo.4006378" httpUri="https://zenodo.org/record/4006378/files/figure.png" pageId="2" pageNumber="1661">Figure 3A</figureCitation>
). Anteriorly to the sternal shield lie two pairs of small presternal platelets. Outer platelets are smaller (57 ×
<quantity id="4CBB9B7BFFD9FFEE57DAF9D5FD6CF9D2" box="[700,758,1539,1563]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="1661" unit="mm" value="3.0">3 µm</quantity>
) than the inner ones (1115 ×
<quantity id="4CBB9B7BFFD9FFEE512AF9D5FB12F9D2" box="[1100,1160,1539,1563]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="1661" unit="mm" value="5.0">5 µm</quantity>
). Sternal shield reaches 9092 μm in length at the midline, and 120130 μm in width at the widest point, that is, at a level between the first and the second coxae. It bears three pairs of simple setae totally, of lengths:
<emphasis id="B937EA8CFFD9FFEE57A9F9B5FD75F9B2" box="[719,751,1635,1659]" italics="true" pageId="2" pageNumber="1661">st1</emphasis>
(
<quantity id="4CBB9B7BFFD9FFEE579BF9B5FCF5F9B2" box="[765,879,1635,1659]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" metricValueMax="2.5" metricValueMin="2.3" pageId="2" pageNumber="1661" unit="mm" value="24.0" valueMax="25.0" valueMin="23.0">2325 µm</quantity>
),
<emphasis id="B937EA8CFFD9FFEE56E4F9B5FC7BF9B2" box="[898,993,1635,1659]" italics="true" pageId="2" pageNumber="1661">st2 = st3</emphasis>
(
<quantity id="4CBB9B7BFFD9FFEE5697F9B5FBF9F9B2" box="[1009,1123,1635,1659]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" metricValueMax="2.0" metricValueMin="1.7" pageId="2" pageNumber="1661" unit="mm" value="18.5" valueMax="20.0" valueMin="17.0">1720 µm</quantity>
). As well as pair of gland pores
<emphasis id="B937EA8CFFD9FFEE5763F955FDA8F952" box="[517,562,1667,1691]" italics="true" pageId="2" pageNumber="1661">gst1</emphasis>
at the extensions between coxae III and two pairs of lyrifissures
<emphasis id="B937EA8CFFDEFFE95469F9CEFEC5F9F9" box="[271,351,1560,1584]" italics="true" pageId="5" pageNumber="1664">iv1, iv2</emphasis>
, but the third pair of sternal lyrifissures
<emphasis id="B937EA8CFFDEFFE95648F9CEFCD4F9F9" box="[814,846,1560,1584]" italics="true" pageId="5" pageNumber="1664">iv3</emphasis>
is absent either on the sternal shield nor metasternal shields. Posteriorly to the sternal shield are small rounded metasternal shields (10 ×
<quantity id="4CBB9B7BFFDEFFE95747F981FDF7F9A6" box="[545,621,1623,1647]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="1664" unit="mm" value="10.0">10 µm</quantity>
), with a setae
<emphasis id="B937EA8CFFDEFFE95671F98EFCADF9B9" box="[791,823,1624,1648]" italics="true" pageId="5" pageNumber="1664">st4</emphasis>
(
<quantity id="4CBB9B7BFFDEFFE9562EF981FC20F9A6" box="[840,954,1623,1648]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" metricValueMax="2.0" metricValueMin="1.9" pageId="5" pageNumber="1664" unit="mm" value="19.5" valueMax="20.0" valueMin="19.0">1920 µm</quantity>
) on it. Inner to coxae IIIV are archwise endopodal shields embracing the coxae, underlying an epigynal shield and partially fused to it. Epigynal shield broad (
<quantity id="4CBB9B7BFFDDFFEA56BEFA02FBCAFA25" box="[984,1104,1492,1516]" metricMagnitude="-2" metricUnit="m" metricValue="8.5" metricValueMax="8.8" metricValueMin="8.2" pageId="6" pageNumber="1665" unit="mm" value="85.0" valueMax="88.0" valueMin="82.0">8288 µm</quantity>
), and almost of the same length (
<emphasis id="B937EA8CFFDDFFEA577EFA23FDBEF9C5" box="[536,548,1525,1548]" italics="true" pageId="6" pageNumber="1665">c</emphasis>
.
<quantity id="4CBB9B7BFFDDFFEA5741FA22FDEFF9C5" box="[551,629,1524,1548]" metricMagnitude="-2" metricUnit="m" metricValue="9.0" pageId="6" pageNumber="1665" unit="mm" value="90.0">90 µm</quantity>
at the midline), truncate, with a pair of setae
<emphasis id="B937EA8CFFDDFFEA55B2F9C2FF6EF9E5" box="[212,244,1556,1580]" italics="true" pageId="6" pageNumber="1665">st5</emphasis>
(
<quantity id="4CBB9B7BFFDDFFEA5462F9C5FEE2F9E2" box="[260,376,1555,1580]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="6" pageNumber="1665" unit="mm" value="17.5" valueMax="20.0" valueMin="15.0">1520 µm</quantity>
). Paragenital poroids
<emphasis id="B937EA8CFFDDFFEA5717F9C2FD0BF9E5" box="[625,657,1556,1580]" italics="true" pageId="6" pageNumber="1665">iv5</emphasis>
are located outside the epigynal shield. Four scanty sclerites arranged in a one line are located posterior to an epigynal shield. Heart-shaped ventrianal shield
<quantity id="4CBB9B7BFFDDFFEA5759F985FD51F9A5" box="[575,715,1619,1644]" metricMagnitude="-1" metricUnit="m" metricValue="1.475" metricValueMax="1.5" metricValueMin="1.45" pageId="6" pageNumber="1665" unit="mm" value="147.5" valueMax="150.0" valueMin="145.0">145150 µm</quantity>
length and
<quantity id="4CBB9B7BFFDDFFEA563DF985FC7DF9A5" box="[859,999,1619,1644]" metricMagnitude="-1" metricUnit="m" metricValue="1.6" metricValueMax="1.65" metricValueMin="1.55" pageId="6" pageNumber="1665" unit="mm" value="160.0" valueMax="165.0" valueMin="155.0">155165 µm</quantity>
width with 15 setae. The shortest are the para-anal setae (
<quantity id="4CBB9B7BFFDDFFEA578EF9A5FCFAF942" box="[744,864,1651,1675]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="6" pageNumber="1665" unit="mm" value="14.0" valueMax="15.0" valueMin="13.0">1315 µm</quantity>
), noticeably longer is the postanal seta (
<quantity id="4CBB9B7BFFD2FFE55454FF7EFE38FF09" box="[306,418,168,192]" metricMagnitude="-2" metricUnit="m" metricValue="2.45" metricValueMax="2.5" metricValueMin="2.4" pageId="9" pageNumber="1668" unit="mm" value="24.5" valueMax="25.0" valueMin="24.0">2425 µm</quantity>
) and ventral setae ranging from
<quantity id="4CBB9B7BFFD2FFE55672FF7EFCC1FF08" box="[788,859,168,193]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="9" pageNumber="1668" unit="mm" value="18.0">18 µm</quantity>
to
<quantity id="4CBB9B7BFFD2FFE556E7FF7EFC5DFF08" box="[897,967,168,193]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" pageId="9" pageNumber="1668" unit="mm" value="21.0">21 µm</quantity>
as given (in µm):
<emphasis id="B937EA8CFFD2FFE555B7FF1EFF65FF29" box="[209,255,200,224]" italics="true" pageId="9" pageNumber="1668">JV1</emphasis>
(1821),
<emphasis id="B937EA8CFFD2FFE55410FF1EFE39FF29" box="[374,419,200,224]" italics="true" pageId="9" pageNumber="1668">JV2</emphasis>
(1821),
<emphasis id="B937EA8CFFD2FFE5577DFF1EFDD3FF29" box="[539,585,200,224]" italics="true" pageId="9" pageNumber="1668">JV3</emphasis>
(1821),
<emphasis id="B937EA8CFFD2FFE557A6FF1EFD77FF29" box="[704,749,200,224]" italics="true" pageId="9" pageNumber="1668">JV4</emphasis>
(1821),
<emphasis id="B937EA8CFFD2FFE55603FF1EFC02FF29" box="[869,920,200,224]" italics="true" pageId="9" pageNumber="1668">ZV2</emphasis>
(1821) except shorter
<emphasis id="B937EA8CFFD2FFE55581FF3EFE80FEC9" box="[231,282,232,256]" italics="true" pageId="9" pageNumber="1668">ZV3</emphasis>
(1517). Likewise, the sternal, genital and ventrianal shields are covered with a reticulate ornamentation (
<figureCitation id="13782A1BFFD2FFE55760FEDEFDFCFEE9" box="[518,614,264,288]" captionStart="Figure 2" captionStartId="4.[212,279,1640,1662]" captionTargetBox="[263,1120,168,1608]" captionTargetId="figure@4.[263,1121,168,1608]" captionTargetPageId="4" captionText="Figure 2. Orthadenella coulsoni sp. nov., female: ventral view of idiosoma." figureDoi="http://doi.org/10.5281/zenodo.4006376" httpUri="https://zenodo.org/record/4006376/files/figure.png" pageId="9" pageNumber="1668">Figure 2</figureCitation>
). Outside the ventrianal shield remain four pairs of simple setae (including UR3 =
<quantity id="4CBB9B7BFFD2FFE5573AFEFEFD57FE89" box="[604,717,296,320]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" metricValueMax="2.0" metricValueMin="1.7" pageId="9" pageNumber="1668" unit="mm" value="18.5" valueMax="20.0" valueMin="17.0">1720 µm</quantity>
)
<emphasis id="B937EA8CFFD2FFE557B8FEFEFC91FE89" box="[734,779,296,320]" italics="true" pageId="9" pageNumber="1668">JV5</emphasis>
(2830),
<emphasis id="B937EA8CFFD2FFE5561CFEFEFC34FE89" box="[890,942,296,320]" italics="true" pageId="9" pageNumber="1668">ZV4</emphasis>
(1620),
<emphasis id="B937EA8CFFD2FFE5517AFEFEFBCAFE89" box="[1052,1104,296,320]" italics="true" pageId="9" pageNumber="1668">ZV5</emphasis>
(2021). Peritremes long, reaching above the coxae I, stigmatas at the level of coxae IV. Peritrematal shields narrow, connected with an exopodal strip alongside coxae IV. Peritrematalexopodal shields fused, with poroids
<emphasis id="B937EA8CFFD2FFE55789FE5EFCEBFE69" box="[751,881,392,416]" italics="true" pageId="9" pageNumber="1668">ip1, ip2, ip3</emphasis>
and pores
<emphasis id="B937EA8CFFD2FFE5568DFE5EFBD2FE69" box="[1003,1096,392,416]" italics="true" pageId="9" pageNumber="1668">gp2, gp3</emphasis>
,
<emphasis id="B937EA8CFFD2FFE555F6FE7EFF33FE76" box="[144,169,424,447]" italics="true" pageId="9" pageNumber="1668">gv</emphasis>
2. Posterolaterally to the coxae IV on each side arise two pairs of metapodal sclerites. Those proximal to the coxae IV are smaller (46 ×
<quantity id="4CBB9B7BFFD2FFE55633FE11FC35FE16" box="[853,943,455,479]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="9" pageNumber="1668" unit="mm" value="5.0" valueMax="6.0" valueMin="4.0">46 µm</quantity>
), than further ones (2021 ×
<quantity id="4CBB9B7BFFD2FFE55426FE31FE2CFE36" box="[320,438,487,511]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" metricValueMax="1.3" metricValueMin="1.1" pageId="9" pageNumber="1668" unit="mm" value="12.0" valueMax="13.0" valueMin="11.0">1113 µm</quantity>
). Sperm access system is that of a Laelapid
<typeStatus id="54F8883CFFD2FFE556A2FE3EFC6DFDC9" box="[964,1015,488,512]" pageId="9" pageNumber="1668">type</typeStatus>
, with a sacculus comprised of a thick and porous layer with a numerous thin filaments (
<figureCitation id="13782A1BFFD2FFE555FEFDF1FE95FDF6" box="[152,271,551,575]" captionStart="Figure 3" captionStartId="5.[144,211,1463,1485]" captionTargetBox="[145,1104,168,1431]" captionTargetId="figure@5.[144,1104,168,1431]" captionTargetPageId="5" captionText="Figure 3. Orthadenella coulsoni sp. nov., female: (A) tritosternum; (B) gnathosoma; (C) sperm access system; (D) epistomes; (E) chelicera; (F) dorsal of palptarsus; (G) ventral of palptarsus." figureDoi="http://doi.org/10.5281/zenodo.4006378" httpUri="https://zenodo.org/record/4006378/files/figure.png" pageId="9" pageNumber="1668">Figure 3C</figureCitation>
).
</paragraph>
<caption id="DF3C6616FFD8FFEF55F6F9BEFC1CF9B7" ID-DOI="http://doi.org/10.5281/zenodo.4006374" ID-Zenodo-Dep="4006374" box="[144,902,1640,1662]" httpUri="https://zenodo.org/record/4006374/files/figure.png" pageId="3" pageNumber="1662" startId="3.[144,211,1640,1662]" targetBox="[211,1037,168,1608]" targetPageId="3">
<paragraph id="8BFC369EFFD8FFEF55F6F9BEFC1CF9B7" blockId="3.[144,902,1640,1662]" box="[144,902,1640,1662]" pageId="3" pageNumber="1662">
Figure 1.
<taxonomicName id="4C434D1DFFD8FFEF559FF9BEFE4BF9B4" authority="Gwiazdowicz &amp; Marchenko &amp; Teodorowicz, 2014" authorityName="Gwiazdowicz &amp; Marchenko &amp; Teodorowicz" authorityYear="2014" box="[249,465,1640,1661]" class="Arachnida" family="Ascidae" genus="Orthadenella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="1662" phylum="Arthropoda" rank="species" species="coulsoni" status="sp. nov.">
<emphasis id="B937EA8CFFD8FFEF559FF9BEFE4BF9B4" box="[249,465,1640,1661]" italics="true" pageId="3" pageNumber="1662">Orthadenella coulsoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20457F7FFD8FFEF54BFF9BFFDB7F9B7" box="[473,557,1641,1662]" pageId="3" pageNumber="1662" rank="species">sp. nov.</taxonomicNameLabel>
, female: dorsal view of idiosoma.
</paragraph>
</caption>
<caption id="DF3C6616FFDFFFE855B2F9BEFC48F9B7" ID-DOI="http://doi.org/10.5281/zenodo.4006376" ID-Zenodo-Dep="4006376" box="[212,978,1640,1662]" httpUri="https://zenodo.org/record/4006376/files/figure.png" pageId="4" pageNumber="1663" startId="4.[212,279,1640,1662]" targetBox="[263,1120,168,1608]" targetPageId="4">
<paragraph id="8BFC369EFFDFFFE855B2F9BEFC48F9B7" blockId="4.[212,978,1640,1662]" box="[212,978,1640,1662]" pageId="4" pageNumber="1663">
Figure 2.
<taxonomicName id="4C434D1DFFDFFFE8545BF9BFFD8FF9B7" authority="Gwiazdowicz &amp; Marchenko &amp; Teodorowicz, 2014" authorityName="Gwiazdowicz &amp; Marchenko &amp; Teodorowicz" authorityYear="2014" box="[317,533,1641,1662]" class="Arachnida" family="Ascidae" genus="Orthadenella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="1663" phylum="Arthropoda" rank="species" species="coulsoni" status="sp. nov.">
<emphasis id="B937EA8CFFDFFFE8545BF9BFFD8FF9B7" box="[317,533,1641,1662]" italics="true" pageId="4" pageNumber="1663">Orthadenella coulsoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20457F7FFDFFFE8577BF9BFFDEBF9B7" box="[541,625,1641,1662]" pageId="4" pageNumber="1663" rank="species">sp. nov.</taxonomicNameLabel>
, female: ventral view of idiosoma.
</paragraph>
</caption>
<caption id="DF3C6616FFDEFFE955F6FA61FBCAFA22" ID-DOI="http://doi.org/10.5281/zenodo.4006378" ID-Zenodo-Dep="4006378" httpUri="https://zenodo.org/record/4006378/files/figure.png" pageId="5" pageNumber="1664" startId="5.[144,211,1463,1485]" targetBox="[145,1104,168,1431]" targetPageId="5">
<paragraph id="8BFC369EFFDEFFE955F6FA61FBCAFA22" blockId="5.[144,1104,1463,1515]" pageId="5" pageNumber="1664">
Figure 3.
<taxonomicName id="4C434D1DFFDEFFE9559EFA61FE4AFA05" authority="Gwiazdowicz &amp; Marchenko &amp; Teodorowicz, 2014" authorityName="Gwiazdowicz &amp; Marchenko &amp; Teodorowicz" authorityYear="2014" box="[248,464,1463,1484]" class="Arachnida" family="Ascidae" genus="Orthadenella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="1664" phylum="Arthropoda" rank="species" species="coulsoni" status="sp. nov.">
<emphasis id="B937EA8CFFDEFFE9559EFA61FE4AFA05" box="[248,464,1463,1484]" italics="true" pageId="5" pageNumber="1664">Orthadenella coulsoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20457F7FFDEFFE954BEFA6EFDB0FA04" box="[472,554,1464,1485]" pageId="5" pageNumber="1664" rank="species">sp. nov.</taxonomicNameLabel>
, female: (A) tritosternum; (B)
<taxonomicName id="4C434D1DFFDEFFE95607FA61FC7AFA04" authorityName="Krefft" authorityYear="1871" box="[865,992,1463,1485]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="5" pageNumber="1669" phylum="Nematoda" rank="genus">gnathosoma</taxonomicName>
; (C) sperm access system; (D) epistomes; (E) chelicera; (F) dorsal of palptarsus; (G) ventral of palptarsus.
</paragraph>
</caption>
<caption id="DF3C6616FFDDFFEA55B2FA57FB1DFA5E" ID-DOI="http://doi.org/10.5281/zenodo.4006380" ID-Zenodo-Dep="4006380" box="[212,1159,1409,1431]" httpUri="https://zenodo.org/record/4006380/files/figure.png" pageId="6" pageNumber="1665" startId="6.[212,279,1409,1431]" targetBox="[292,1092,168,1377]" targetPageId="6">
<paragraph id="8BFC369EFFDDFFEA55B2FA57FB1DFA5E" blockId="6.[212,1159,1409,1431]" box="[212,1159,1409,1431]" pageId="6" pageNumber="1665">
Figure 4.
<taxonomicName id="4C434D1DFFDDFFEA545BFA57FD8FFA5F" authority="Gwiazdowicz &amp; Marchenko &amp; Teodorowicz, 2014" authorityName="Gwiazdowicz &amp; Marchenko &amp; Teodorowicz" authorityYear="2014" box="[317,533,1409,1430]" class="Arachnida" family="Ascidae" genus="Orthadenella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="1665" phylum="Arthropoda" rank="species" species="coulsoni" status="sp. nov.">
<emphasis id="B937EA8CFFDDFFEA545BFA57FD8FFA5F" box="[317,533,1409,1430]" italics="true" pageId="6" pageNumber="1665">Orthadenella coulsoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20457F7FFDDFFEA577BFA54FDEBFA5E" box="[541,625,1410,1431]" pageId="6" pageNumber="1665" rank="species">sp. nov.</taxonomicNameLabel>
, female: (A) leg I; (B) leg II; (C) leg III; (D) leg IV.
</paragraph>
</caption>
<caption id="DF3C6616FFDCFFEB55F6F9BEFCE6F9B7" ID-DOI="http://doi.org/10.5281/zenodo.4006384" ID-Zenodo-Dep="4006384" box="[144,892,1640,1662]" httpUri="https://zenodo.org/record/4006384/files/figure.png" pageId="7" pageNumber="1666" startId="7.[144,211,1640,1662]" targetBox="[196,1050,168,1608]" targetPageId="7">
<paragraph id="8BFC369EFFDCFFEB55F6F9BEFCE6F9B7" blockId="7.[144,892,1640,1662]" box="[144,892,1640,1662]" pageId="7" pageNumber="1666">
Figure 5.
<taxonomicName id="4C434D1DFFDCFFEB559FF9BFFE4BF9B7" authority="Gwiazdowicz &amp; Marchenko &amp; Teodorowicz, 2014" authorityName="Gwiazdowicz &amp; Marchenko &amp; Teodorowicz" authorityYear="2014" box="[249,465,1641,1662]" class="Arachnida" family="Ascidae" genus="Orthadenella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="1666" phylum="Arthropoda" rank="species" species="coulsoni" status="sp. nov.">
<emphasis id="B937EA8CFFDCFFEB559FF9BFFE4BF9B7" box="[249,465,1641,1662]" italics="true" pageId="7" pageNumber="1666">Orthadenella coulsoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20457F7FFDCFFEB54BFF9BFFDB7F9B7" box="[473,557,1641,1662]" pageId="7" pageNumber="1666" rank="species">sp. nov.</taxonomicNameLabel>
, male: ventral view of idiosoma.
</paragraph>
</caption>
<caption id="DF3C6616FFD3FFE455B2F9FBFEFDF9A9" ID-DOI="http://doi.org/10.5281/zenodo.4006386" ID-Zenodo-Dep="4006386" httpUri="https://zenodo.org/record/4006386/files/figure.png" pageId="8" pageNumber="1667" startId="8.[212,279,1581,1603]" targetBox="[212,1172,168,1549]" targetPageId="8">
<paragraph id="8BFC369EFFD3FFE455B2F9FBFEFDF9A9" blockId="8.[212,1172,1581,1632]" pageId="8" pageNumber="1667">
Figure 6.
<taxonomicName id="4C434D1DFFD3FFE4545DF9FBFD8AF98B" authority="Gwiazdowicz &amp; Marchenko &amp; Teodorowicz, 2014" authorityName="Gwiazdowicz &amp; Marchenko &amp; Teodorowicz" authorityYear="2014" box="[315,528,1581,1602]" class="Arachnida" family="Ascidae" genus="Orthadenella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="1667" phylum="Arthropoda" rank="species" species="coulsoni" status="sp. nov.">
<emphasis id="B937EA8CFFD3FFE4545DF9FBFD8AF98B" box="[315,528,1581,1602]" italics="true" pageId="8" pageNumber="1667">Orthadenella coulsoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20457F7FFD3FFE45773F9FBFDFCF98B" box="[533,614,1581,1602]" pageId="8" pageNumber="1667" rank="species">sp. nov.</taxonomicNameLabel>
, male: (A) tritosternum; (B)
<taxonomicName id="4C434D1DFFD3FFE4561BF9FBFC66F98A" authorityName="Krefft" authorityYear="1871" box="[893,1020,1581,1603]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="8" pageNumber="1669" phylum="Nematoda" rank="genus">gnathosoma</taxonomicName>
; (C) chelicerae; (D) epistomes.
</paragraph>
</caption>
<paragraph id="8BFC369EFFD2FFE555F6FDB1FD7FFC74" blockId="9.[144,1104,615,957]" pageId="9" pageNumber="1668">
<emphasis id="B937EA8CFFD2FFE555F6FDB1FEB8FDB6" box="[144,290,615,639]" italics="true" pageId="9" pageNumber="1668">
<taxonomicName id="4C434D1DFFD2FFE555F6FDB1FE84FDB6" authorityName="Krefft" authorityYear="1871" box="[144,286,615,639]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="9" pageNumber="1668" phylum="Nematoda" rank="genus">Gnathosoma</taxonomicName>
.
</emphasis>
Corniculi are elongated, 2630 μm long and
<quantity id="4CBB9B7BFFD2FFE55642FDB1FCF1FDB6" box="[804,875,615,639]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="1668" unit="mm" value="10.0">10 µm</quantity>
wide; seven rows of denticles are located in the hypostomatal groove (26 denticles per row); hypostomatal setae are simple of variable length:
<emphasis id="B937EA8CFFD2FFE55758FD71FDC3FD76" box="[574,601,679,703]" italics="true" pageId="9" pageNumber="1668">h1</emphasis>
2021 μm,
<emphasis id="B937EA8CFFD2FFE55787FD71FD66FD76" box="[737,764,679,703]" italics="true" pageId="9" pageNumber="1668">h2</emphasis>
2122 μm,
<emphasis id="B937EA8CFFD2FFE556E3FD71FC05FD76" box="[901,927,679,703]" italics="true" pageId="9" pageNumber="1668">h3</emphasis>
1416 μm,
<emphasis id="B937EA8CFFD2FFE5514EFD71FBD9FD76" box="[1064,1091,679,703]" italics="true" pageId="9" pageNumber="1668">h4</emphasis>
2224 μm (
<figureCitation id="13782A1BFFD2FFE55469FD10FE18FD17" box="[271,386,710,734]" captionStart="Figure 3" captionStartId="5.[144,211,1463,1485]" captionTargetBox="[145,1104,168,1431]" captionTargetId="figure@5.[144,1104,168,1431]" captionTargetPageId="5" captionText="Figure 3. Orthadenella coulsoni sp. nov., female: (A) tritosternum; (B) gnathosoma; (C) sperm access system; (D) epistomes; (E) chelicera; (F) dorsal of palptarsus; (G) ventral of palptarsus." figureDoi="http://doi.org/10.5281/zenodo.4006378" httpUri="https://zenodo.org/record/4006378/files/figure.png" pageId="9" pageNumber="1668">Figure 3B</figureCitation>
). Internal malae as long as corniculi, with a fringed laterobasal margins. Epistome with anterior margin irregularly convex, finely denticulate (
<figureCitation id="13782A1BFFD2FFE55163FD30FF2FFCD7" captionStart="Figure 3" captionStartId="5.[144,211,1463,1485]" captionTargetBox="[145,1104,168,1431]" captionTargetId="figure@5.[144,1104,168,1431]" captionTargetPageId="5" captionText="Figure 3. Orthadenella coulsoni sp. nov., female: (A) tritosternum; (B) gnathosoma; (C) sperm access system; (D) epistomes; (E) chelicera; (F) dorsal of palptarsus; (G) ventral of palptarsus." figureDoi="http://doi.org/10.5281/zenodo.4006378" httpUri="https://zenodo.org/record/4006378/files/figure.png" pageId="9" pageNumber="1668">Figure 3D</figureCitation>
). Cheliceral fixed digit
<quantity id="4CBB9B7BFFD2FFE554A9FCD0FDDBFCD6" box="[463,577,774,799]" metricMagnitude="-2" metricUnit="m" metricValue="3.9" metricValueMax="4.0" metricValueMin="3.8" pageId="9" pageNumber="1668" unit="mm" value="39.0" valueMax="40.0" valueMin="38.0">3840 µm</quantity>
long with a stout pilus dentilis; masticatory surface with a row of 12 teeth and two subapical teeth in addition to apical tooth. Cheliceral movable digit (3739) tridentate in addition to apical hook; with a transverse-diagonal groove, which appears on the basal one-third of ventral side of chelicera. Dorsal cheliceral seta, dorsal and lateral (antiaxial) lyrifissure distinct (
<figureCitation id="13782A1BFFD2FFE555FEFC73FE94FC74" box="[152,270,933,957]" captionStart="Figure 3" captionStartId="5.[144,211,1463,1485]" captionTargetBox="[145,1104,168,1431]" captionTargetId="figure@5.[144,1104,168,1431]" captionTargetPageId="5" captionText="Figure 3. Orthadenella coulsoni sp. nov., female: (A) tritosternum; (B) gnathosoma; (C) sperm access system; (D) epistomes; (E) chelicera; (F) dorsal of palptarsus; (G) ventral of palptarsus." figureDoi="http://doi.org/10.5281/zenodo.4006378" httpUri="https://zenodo.org/record/4006378/files/figure.png" pageId="9" pageNumber="1668">Figure 3E</figureCitation>
). Palps 126128 μm long (
<figureCitation id="13782A1BFFD2FFE55758FC73FD42FC74" box="[574,728,933,957]" captionStart="Figure 3" captionStartId="5.[144,211,1463,1485]" captionTargetBox="[145,1104,168,1431]" captionTargetId="figure@5.[144,1104,168,1431]" captionTargetPageId="5" captionText="Figure 3. Orthadenella coulsoni sp. nov., female: (A) tritosternum; (B) gnathosoma; (C) sperm access system; (D) epistomes; (E) chelicera; (F) dorsal of palptarsus; (G) ventral of palptarsus." figureDoi="http://doi.org/10.5281/zenodo.4006378" httpUri="https://zenodo.org/record/4006378/files/figure.png" pageId="9" pageNumber="1668">Figure 3F, G</figureCitation>
).
</paragraph>
<paragraph id="8BFC369EFFD2FFE555F6FC30FD49FB94" blockId="9.[144,1103,997,1117]" pageId="9" pageNumber="1668">
<emphasis id="B937EA8CFFD2FFE555F6FC30FF56FC34" box="[144,204,998,1021]" italics="true" pageId="9" pageNumber="1668">Legs.</emphasis>
Variable in length: I 320340 μm, II 250260 μm, III 240250 μm, IV 315325 μm. Chaetotaxy of legs is peculiar for genus
<taxonomicName id="4C434D1DFFD2FFE55644FBD0FC2BFBD7" authorityName="Athias-Henriot" authorityYear="1973" box="[802,945,1030,1054]" class="Arachnida" family="Ascidae" genus="Orthadenella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="1668" phylum="Arthropoda" rank="genus">
<emphasis id="B937EA8CFFD2FFE55644FBD0FC2BFBD7" box="[802,945,1030,1054]" italics="true" pageId="9" pageNumber="1668">Orthadenella</emphasis>
</taxonomicName>
: leg I: coxa, trochanter, femur, genu, tibia (2, 6, 12, 13, 13), leg II (2, 5, 11, 11, 10), leg III (2, 5, 7, 9, 8), leg IV (1, 4, 6, 9, 9) (
<figureCitation id="13782A1BFFD2FFE55482FB93FD5CFB94" box="[484,710,1093,1117]" captionStart="Figure 4" captionStartId="6.[212,279,1409,1431]" captionTargetBox="[292,1092,168,1377]" captionTargetId="figure@6.[292,1092,168,1377]" captionTargetPageId="6" captionText="Figure 4. Orthadenella coulsoni sp. nov., female: (A) leg I; (B) leg II; (C) leg III; (D) leg IV." figureDoi="http://doi.org/10.5281/zenodo.4006380" httpUri="https://zenodo.org/record/4006380/files/figure.png" pageId="9" pageNumber="1668">Figure 4A, B, C, D</figureCitation>
).
</paragraph>
<paragraph id="8BFC369EFFD2FFE555F4FBAEFB91FB46" blockId="9.[146,1035,1143,1168]" box="[146,1035,1143,1168]" pageId="9" pageNumber="1668">
<heading id="D0B481F2FFD2FFE555F4FBAEFB91FB46" box="[146,1035,1143,1168]" fontSize="10" level="3" pageId="9" pageNumber="1668" reason="5">
<emphasis id="B937EA8CFFD2FFE555F4FBAEFF56FB59" box="[146,204,1144,1168]" italics="true" pageId="9" pageNumber="1668">Male</emphasis>
(
<emphasis id="B937EA8CFFD2FFE555B8FBAEFF71FB46" box="[222,235,1144,1167]" italics="true" pageId="9" pageNumber="1668">n</emphasis>
= 8) Idiosoma oval,
<quantity id="4CBB9B7BFFD2FFE55486FBA1FDF0FB59" box="[480,618,1143,1168]" metricMagnitude="-1" metricUnit="m" metricValue="3.425" metricValueMax="3.55" metricValueMin="3.3" pageId="9" pageNumber="1668" unit="mm" value="342.5" valueMax="355.0" valueMin="330.0">330355 µm</quantity>
in length and
<quantity id="4CBB9B7BFFD2FFE55674FBA1FC06FB59" box="[786,924,1143,1168]" metricMagnitude="-1" metricUnit="m" metricValue="2.17" metricValueMax="2.2199999999999998" metricValueMin="2.12" pageId="9" pageNumber="1668" unit="mm" value="217.0" valueMax="222.0" valueMin="212.0">212222 µm</quantity>
in width.
</heading>
</paragraph>
<paragraph id="8BFC369EFFD2FFE555F6FB73FD86FA15" blockId="9.[144,1104,1189,1500]" pageId="9" pageNumber="1668">
<emphasis id="B937EA8CFFD2FFE555F6FB73FF78FB74" box="[144,226,1189,1213]" italics="true" pageId="9" pageNumber="1668">Dorsal.</emphasis>
Holodorsal shield bearing 43 pairs of simple setae, including 23 podonotal pairs (
<emphasis id="B937EA8CFFD2FFE555BEFB13FE79FB14" box="[216,483,1221,1245]" italics="true" pageId="9" pageNumber="1668">j1-j6, z1-z6, s1-s6, r2-r6</emphasis>
) and 20 opisthonotal pairs (
<emphasis id="B937EA8CFFD2FFE5564AFB13FF2AFB34" italics="true" pageId="9" pageNumber="1668">J1-J5, Z1-Z5, S1-S5, R1- R5</emphasis>
). Dorsal shield lightly reticulate. Measurements of podonotal setae precisely (in µm):
<emphasis id="B937EA8CFFD2FFE555ADFAD3FF45FAD4" box="[203,223,1285,1309]" italics="true" pageId="9" pageNumber="1668">j1</emphasis>
(1215),
<emphasis id="B937EA8CFFD2FFE5542FFAD3FEC4FAD4" box="[329,350,1285,1309]" italics="true" pageId="9" pageNumber="1668">j2</emphasis>
(1517),
<emphasis id="B937EA8CFFD2FFE554AEFAD3FE46FAD4" box="[456,476,1285,1309]" italics="true" pageId="9" pageNumber="1668">j3</emphasis>
(1215),
<emphasis id="B937EA8CFFD2FFE55720FAD3FDC0FAD4" box="[582,602,1285,1309]" italics="true" pageId="9" pageNumber="1668">j4</emphasis>
(1215),
<emphasis id="B937EA8CFFD2FFE557A2FAD3FD43FAD4" box="[708,729,1285,1309]" italics="true" pageId="9" pageNumber="1668">j5</emphasis>
(1215),
<emphasis id="B937EA8CFFD2FFE55625FAD3FCCDFAD4" box="[835,855,1285,1309]" italics="true" pageId="9" pageNumber="1668">j6</emphasis>
(1215),
<emphasis id="B937EA8CFFD2FFE556A7FAD3FC40FAD4" box="[961,986,1285,1309]" italics="true" pageId="9" pageNumber="1668">z1</emphasis>
(810),
<emphasis id="B937EA8CFFD2FFE55151FAD3FBD5FAD4" box="[1079,1103,1285,1309]" italics="true" pageId="9" pageNumber="1668">z2</emphasis>
(1517),
<emphasis id="B937EA8CFFD2FFE55594FAF2FE91FAF5" box="[242,267,1316,1340]" italics="true" pageId="9" pageNumber="1668">z3</emphasis>
(1215),
<emphasis id="B937EA8CFFD2FFE55413FAF2FE14FAF5" box="[373,398,1316,1340]" italics="true" pageId="9" pageNumber="1668">z4</emphasis>
(1215),
<emphasis id="B937EA8CFFD2FFE5549EFAF2FD8BFAF5" box="[504,529,1316,1340]" italics="true" pageId="9" pageNumber="1668">z5</emphasis>
(1215),
<emphasis id="B937EA8CFFD2FFE5571DFAF2FD0EFAF5" box="[635,660,1316,1340]" italics="true" pageId="9" pageNumber="1668">z6</emphasis>
(1215),
<emphasis id="B937EA8CFFD2FFE55798FAF2FC8FFAF5" box="[766,789,1316,1340]" italics="true" pageId="9" pageNumber="1668">s1</emphasis>
(1215),
<emphasis id="B937EA8CFFD2FFE55619FAF2FC0CFAF5" box="[895,918,1316,1340]" italics="true" pageId="9" pageNumber="1668">s2</emphasis>
(1517),
<emphasis id="B937EA8CFFD2FFE55166FAF2FB82FAF5" box="[1024,1048,1316,1340]" italics="true" pageId="9" pageNumber="1668">s3</emphasis>
(12 15),
<emphasis id="B937EA8CFFD2FFE555A5FA92FF40FA95" box="[195,218,1348,1372]" italics="true" pageId="9" pageNumber="1668">s4</emphasis>
(1517), s5 (1215),
<emphasis id="B937EA8CFFD2FFE554AAFA92FE7EFA95" box="[460,484,1348,1372]" italics="true" pageId="9" pageNumber="1668">s6</emphasis>
(1215),
<emphasis id="B937EA8CFFD2FFE55737FA92FD1DFA95" box="[593,647,1348,1372]" italics="true" pageId="9" pageNumber="1668">r2-r6</emphasis>
(1215). Measurements of opisthonotal setae as follows (in µm):
<emphasis id="B937EA8CFFD2FFE554CDFAB2FE5FFAB5" box="[427,453,1380,1404]" italics="true" pageId="9" pageNumber="1668">J1</emphasis>
(1012),
<emphasis id="B937EA8CFFD2FFE55757FAB2FDD1FAB5" box="[561,587,1380,1404]" italics="true" pageId="9" pageNumber="1668">J2</emphasis>
(1012),
<emphasis id="B937EA8CFFD2FFE557D1FAB2FD48FAB5" box="[695,722,1380,1404]" italics="true" pageId="9" pageNumber="1668">J3</emphasis>
(1215),
<emphasis id="B937EA8CFFD2FFE5565BFAB2FCC2FAB5" box="[829,856,1380,1404]" italics="true" pageId="9" pageNumber="1668">J4</emphasis>
(1517),
<emphasis id="B937EA8CFFD2FFE556A2FAB2FC44FAB5" box="[964,990,1380,1404]" italics="true" pageId="9" pageNumber="1668">J5</emphasis>
(56),
<emphasis id="B937EA8CFFD2FFE55149FAB2FBD5FAB5" box="[1071,1103,1380,1404]" italics="true" pageId="9" pageNumber="1668">Z1</emphasis>
(1215),
<emphasis id="B937EA8CFFD2FFE55595FA52FE8EFA55" box="[243,276,1412,1436]" italics="true" pageId="9" pageNumber="1668">Z2</emphasis>
(1215),
<emphasis id="B937EA8CFFD2FFE55418FA52FE05FA55" box="[382,415,1412,1436]" italics="true" pageId="9" pageNumber="1668">Z3</emphasis>
(1215),
<emphasis id="B937EA8CFFD2FFE5576CFA52FDB0FA55" box="[522,554,1412,1436]" italics="true" pageId="9" pageNumber="1668">Z4</emphasis>
(1720),
<emphasis id="B937EA8CFFD2FFE557F0FA52FD2CFA55" box="[662,694,1412,1436]" italics="true" pageId="9" pageNumber="1668">Z5</emphasis>
(2830),
<emphasis id="B937EA8CFFD2FFE55644FA52FCA4FA55" box="[802,830,1412,1436]" italics="true" pageId="9" pageNumber="1668">S1</emphasis>
(1215),
<emphasis id="B937EA8CFFD2FFE556CCFA52FC5DFA55" box="[938,967,1412,1436]" italics="true" pageId="9" pageNumber="1668">S2</emphasis>
(1215),
<emphasis id="B937EA8CFFD2FFE55155FA52FBCAFA55" box="[1075,1104,1412,1436]" italics="true" pageId="9" pageNumber="1668">S3</emphasis>
(1215),
<emphasis id="B937EA8CFFD2FFE55590FA72FE89FA75" box="[246,275,1444,1468]" italics="true" pageId="9" pageNumber="1668">S4</emphasis>
(1517),
<emphasis id="B937EA8CFFD2FFE554E3FA72FE3BFA75" box="[389,417,1444,1468]" italics="true" pageId="9" pageNumber="1668">S5</emphasis>
(1720),
<emphasis id="B937EA8CFFD2FFE55772FA72FDC7FA75" box="[532,605,1444,1468]" italics="true" pageId="9" pageNumber="1668">R1-R5</emphasis>
(1215). Setae
<emphasis id="B937EA8CFFD2FFE55670FA72FCC0FA75" box="[790,858,1444,1468]" italics="true" pageId="9" pageNumber="1668">r2R5</emphasis>
at clearly delineated marginal strip, likewise for female.
</paragraph>
<paragraph id="8BFC369EFFD2FFE655F6F9D2FE3AFDD6" blockId="9.[144,1104,1539,1692]" lastBlockId="10.[212,1172,168,543]" lastPageId="10" lastPageNumber="1669" pageId="9" pageNumber="1668">
<emphasis id="B937EA8CFFD2FFE555F6F9D2FF71F9D5" box="[144,235,1540,1564]" italics="true" pageId="9" pageNumber="1668">Ventral.</emphasis>
Tritosternum with base
<quantity id="4CBB9B7BFFD2FFE55762F9D2FDFCF9D5" box="[516,614,1539,1564]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="10.0" metricValueMin="7.0" pageId="9" pageNumber="1668" unit="mm" value="8.5" valueMax="10.0" valueMin="7.0">710 µm</quantity>
wide and
<quantity id="4CBB9B7BFFD2FFE557B8F9D5FCD7F9D5" box="[734,845,1539,1564]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="9" pageNumber="1668" unit="mm" value="13.5" valueMax="15.0" valueMin="12.0">1215 µm</quantity>
long with laciniae
<quantity id="4CBB9B7BFFD2FFE5514EF9D5FF4CF9F5" metricMagnitude="-2" metricUnit="m" metricValue="4.45" metricValueMax="4.7" metricValueMin="4.2" pageId="9" pageNumber="1668" unit="mm" value="44.5" valueMax="47.0" valueMin="42.0">42 47 µm</quantity>
long (excluding base) with a fused area, free area for about 0.7 of total length (
<figureCitation id="13782A1BFFD2FFE555FEF995FE8AF992" box="[152,272,1603,1627]" captionStart="Figure 6" captionStartId="8.[212,279,1581,1603]" captionTargetBox="[212,1172,168,1549]" captionTargetId="figure@8.[212,1172,168,1549]" captionTargetPageId="8" captionText="Figure 6. Orthadenella coulsoni sp. nov., male: (A) tritosternum; (B) gnathosoma; (C) chelicerae; (D) epistomes." figureDoi="http://doi.org/10.5281/zenodo.4006386" httpUri="https://zenodo.org/record/4006386/files/figure.png" pageId="9" pageNumber="1668">Figure 6A</figureCitation>
). Presternal area with a pair platelets. Peritrematal shields and peritremes as in female (poroids
<emphasis id="B937EA8CFFD2FFE554E5F9B5FD9CF9B2" box="[387,518,1635,1659]" italics="true" pageId="9" pageNumber="1668">ip1, ip2, ip3</emphasis>
and pores
<emphasis id="B937EA8CFFD2FFE557E2F9B5FD78F9B2" box="[644,738,1635,1659]" italics="true" pageId="9" pageNumber="1668">gp2, gp3</emphasis>
,
<emphasis id="B937EA8CFFD2FFE55797F9B2FC90F9B2" box="[753,778,1636,1659]" italics="true" pageId="9" pageNumber="1668">gv</emphasis>
2 are present). Sternitigenital shield
<quantity id="4CBB9B7BFFD2FFE555BDF955FEFCF955" box="[219,358,1667,1692]" metricMagnitude="-1" metricUnit="m" metricValue="1.41" metricValueMax="1.45" metricValueMin="1.3699999999999999" pageId="9" pageNumber="1668" unit="mm" value="141.0" valueMax="145.0" valueMin="137.0">137145 µm</quantity>
long and
<quantity id="4CBB9B7BFFD2FFE55486F955FDF1F955" box="[480,619,1667,1692]" metricMagnitude="-1" metricUnit="m" metricValue="1.065" metricValueMax="1.13" metricValueMin="1.0" pageId="9" pageNumber="1668" unit="mm" value="106.5" valueMax="113.0" valueMin="100.0">100113 µm</quantity>
wide at level between coxae IIIII; finely ornamented anteriorly between setae
<emphasis id="B937EA8CFFD1FFE6571EFF7EFD02FF09" box="[632,664,168,192]" italics="true" pageId="10" pageNumber="1669">st1</emphasis>
and
<emphasis id="B937EA8CFFD1FFE657B7FF7EFD6AFF09" box="[721,752,168,192]" italics="true" pageId="10" pageNumber="1669">st3</emphasis>
, posteriorly between
<emphasis id="B937EA8CFFD1FFE65684FF7EFB99FF09" box="[994,1027,168,192]" italics="true" pageId="10" pageNumber="1669">st4</emphasis>
and
<emphasis id="B937EA8CFFD1FFE6515DFF7EFBC1FF09" box="[1083,1115,168,192]" italics="true" pageId="10" pageNumber="1669">st5</emphasis>
, and lineate along lateral margins. Sternal shield with a pair of gland pores
<emphasis id="B937EA8CFFD1FFE65124FF1EFBF5FF29" box="[1090,1135,200,224]" italics="true" pageId="10" pageNumber="1669">gst1</emphasis>
at extensions between coxae III and with two pairs of lyrifissures
<emphasis id="B937EA8CFFD1FFE656BAFF3EFBAAFEC9" box="[988,1072,232,256]" italics="true" pageId="10" pageNumber="1669">iv1, iv2</emphasis>
. Shields with five pairs of setae. Measurements of sternal setae as given (in µm):
<emphasis id="B937EA8CFFD1FFE65176FEDEFBAAFEE9" box="[1040,1072,264,288]" italics="true" pageId="10" pageNumber="1669">st1</emphasis>
(2021),
<emphasis id="B937EA8CFFD1FFE655B2FEFEFF69FE89" box="[212,243,296,320]" italics="true" pageId="10" pageNumber="1669">st2</emphasis>
(1820),
<emphasis id="B937EA8CFFD1FFE65439FEFEFEE4FE89" box="[351,382,296,320]" italics="true" pageId="10" pageNumber="1669">st3</emphasis>
(1618),
<emphasis id="B937EA8CFFD1FFE6548CFEFEFD90FE89" box="[490,522,296,320]" italics="true" pageId="10" pageNumber="1669">st4</emphasis>
(1516),
<emphasis id="B937EA8CFFD1FFE65710FEFEFD0CFE89" box="[630,662,296,320]" italics="true" pageId="10" pageNumber="1669">st5</emphasis>
(1415). Ventrianal shield ornamented, midlength (
<quantity id="4CBB9B7BFFD1FFE6544DFE9EFE20FEA9" box="[299,442,328,352]" metricMagnitude="-1" metricUnit="m" metricValue="1.375" metricValueMax="1.45" metricValueMin="1.3" pageId="10" pageNumber="1669" unit="mm" value="137.5" valueMax="145.0" valueMin="130.0">130145 µm</quantity>
), greatest at midlateral width (
<quantity id="4CBB9B7BFFD1FFE65646FE9EFC37FEA9" box="[800,941,328,352]" metricMagnitude="-1" metricUnit="m" metricValue="1.625" metricValueMax="1.7" metricValueMin="1.55" pageId="10" pageNumber="1669" unit="mm" value="162.5" valueMax="170.0" valueMin="155.0">155170 µm</quantity>
) at the level of seta
<emphasis id="B937EA8CFFD1FFE655B2FEBEFE98FE49" box="[212,258,360,384]" italics="true" pageId="10" pageNumber="1669">JV1</emphasis>
, with a regularly convex lateral margins, bearing five opisthogastric setae
<emphasis id="B937EA8CFFD1FFE65100FEBEFB0EFE49" box="[1126,1172,360,384]" italics="true" pageId="10" pageNumber="1669">JV1</emphasis>
(1415),
<emphasis id="B937EA8CFFD1FFE6545FFE5EFEFCFE69" box="[313,358,392,416]" italics="true" pageId="10" pageNumber="1669">JV2</emphasis>
(1517),
<emphasis id="B937EA8CFFD1FFE654B1FE5EFD9FFE69" box="[471,517,392,416]" italics="true" pageId="10" pageNumber="1669">JV3</emphasis>
(1415),
<emphasis id="B937EA8CFFD1FFE65713FE5EFD38FE69" box="[629,674,392,416]" italics="true" pageId="10" pageNumber="1669">JV4</emphasis>
(
<quantity id="4CBB9B7BFFD1FFE657D0FE51FCB0FE56" box="[694,810,391,415]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="10" pageNumber="1669" unit="mm" value="14.5" valueMax="15.0" valueMin="14.0">1415 µm</quantity>
),
<emphasis id="B937EA8CFFD1FFE65627FE5EFCEFFE69" box="[833,885,392,416]" italics="true" pageId="10" pageNumber="1669">ZV2</emphasis>
(1415), pair of circumanal setae (1415) and post-anal seta
<quantity id="4CBB9B7BFFD1FFE6571CFE71FD59FE09" box="[634,707,423,448]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="1669" unit="mm" value="20.0">20 µm</quantity>
; bearing two pairs of poroids and pair of pores
<emphasis id="B937EA8CFFD1FFE6547DFE11FED8FE16" box="[283,322,455,479]" italics="true" pageId="10" pageNumber="1669">gv3</emphasis>
; soft opisthogastric cuticle stays with three pairs of setae
<emphasis id="B937EA8CFFD1FFE6568CFE11FB8DFE16" box="[1002,1047,455,479]" italics="true" pageId="10" pageNumber="1669">JV5</emphasis>
(20),
<emphasis id="B937EA8CFFD1FFE65106FE11FB0EFE16" box="[1120,1172,455,479]" italics="true" pageId="10" pageNumber="1669">ZV4</emphasis>
(1314),
<emphasis id="B937EA8CFFD1FFE6545DFE31FEF5FE36" box="[315,367,487,511]" italics="true" pageId="10" pageNumber="1669">ZV5</emphasis>
(1415). Opisthogastric setae
<emphasis id="B937EA8CFFD1FFE657B3FE31FC93FE36" box="[725,777,487,511]" italics="true" pageId="10" pageNumber="1669">ZV3</emphasis>
and
<emphasis id="B937EA8CFFD1FFE6562AFE31FC1AFE36" box="[844,896,487,511]" italics="true" pageId="10" pageNumber="1669">UR3</emphasis>
are absent contrary to female (
<figureCitation id="13782A1BFFD1FFE65449FDD1FE0AFDD6" box="[303,400,519,543]" captionStart="Figure 5" captionStartId="7.[144,211,1640,1662]" captionTargetBox="[196,1050,168,1608]" captionTargetId="figure@7.[196,1051,168,1608]" captionTargetPageId="7" captionText="Figure 5. Orthadenella coulsoni sp. nov., male: ventral view of idiosoma." figureDoi="http://doi.org/10.5281/zenodo.4006384" httpUri="https://zenodo.org/record/4006384/files/figure.png" pageId="10" pageNumber="1669">Figure 5</figureCitation>
).
</paragraph>
<paragraph id="8BFC369EFFD1FFE655B2FD91FD64FB94" blockId="10.[212,1172,583,1117]" pageId="10" pageNumber="1669">
<emphasis id="B937EA8CFFD1FFE655B2FD91FEFCFD96" box="[212,358,583,607]" italics="true" pageId="10" pageNumber="1669">
<taxonomicName id="4C434D1DFFD1FFE655B2FD91FEF8FD96" authorityName="Krefft" authorityYear="1871" box="[212,354,583,607]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="10" pageNumber="1669" phylum="Nematoda" rank="genus">Gnathosoma</taxonomicName>
.
</emphasis>
Deutosternum with seven rows of denticles; margins of deutosternal groove delineated laterally except posteriormost row. Number of denticles in each row varies individually in specimens: the first posteriormost row (59 denticles), the second row (812), the third row (56), the fourth row (45), 57 rows (
<quantity id="4CBB9B7BFFD1FFE65176FD70FBCDFD77" box="[1040,1111,678,702]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" metricValueMax="10.16" metricValueMin="5.08" pageId="10" pageNumber="1669" unit="in" value="3.0" valueMax="4.0" valueMin="2.0">24 in</quantity>
each row). Subcapitulum with a hypostomatic setae
<emphasis id="B937EA8CFFD1FFE6578EFD10FC99FD17" box="[744,771,710,734]" italics="true" pageId="10" pageNumber="1669">h1</emphasis>
(2022),
<emphasis id="B937EA8CFFD1FFE65608FD10FC12FD17" box="[878,904,710,734]" italics="true" pageId="10" pageNumber="1669">h2</emphasis>
(1517),
<emphasis id="B937EA8CFFD1FFE65695FD10FB94FD17" box="[1011,1038,710,734]" italics="true" pageId="10" pageNumber="1669">h3</emphasis>
(2022),
<emphasis id="B937EA8CFFD1FFE6511FFD10FB0EFD17" box="[1145,1172,710,734]" italics="true" pageId="10" pageNumber="1669">h4</emphasis>
(2022); with three pockmarked delineated areas between
<emphasis id="B937EA8CFFD1FFE65631FD30FC0EFD37" box="[855,916,742,766]" italics="true" pageId="10" pageNumber="1669">h2-h3</emphasis>
and palpcoxal seta
<emphasis id="B937EA8CFFD1FFE65114FD30FB16FD37" box="[1138,1164,742,766]" italics="true" pageId="10" pageNumber="1669">h4</emphasis>
. Form of corniculi as in female,
<quantity id="4CBB9B7BFFD1FFE6575BFCD0FD1EFCD6" box="[573,644,774,799]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="10" pageNumber="1669" unit="mm" value="22.0">22 µm</quantity>
long and
<quantity id="4CBB9B7BFFD1FFE6579FFCD1FCD7FCD6" box="[761,845,774,799]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="10" pageNumber="1669" unit="mm" value="7.5" valueMax="8.0" valueMin="7.0">78 µm</quantity>
width; internal malae longer than corniculi, with fringed lateral margins basally (
<figureCitation id="13782A1BFFD1FFE65646FCF0FC0EFCF7" box="[800,916,806,830]" captionStart="Figure 6" captionStartId="8.[212,279,1581,1603]" captionTargetBox="[212,1172,168,1549]" captionTargetId="figure@8.[212,1172,168,1549]" captionTargetPageId="8" captionText="Figure 6. Orthadenella coulsoni sp. nov., male: (A) tritosternum; (B) gnathosoma; (C) chelicerae; (D) epistomes." figureDoi="http://doi.org/10.5281/zenodo.4006386" httpUri="https://zenodo.org/record/4006386/files/figure.png" pageId="10" pageNumber="1669">Figure 6B</figureCitation>
). Cheliceral fixed digit
<quantity id="4CBB9B7BFFD1FFE655B2FC90FEDAFC96" box="[212,320,838,863]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" metricValueMax="3.2" metricValueMin="3.0" pageId="10" pageNumber="1669" unit="mm" value="31.0" valueMax="32.0" valueMin="30.0">3032 µm</quantity>
long with a stout pilus dentilis and usually with six teeth in addition to apical tooth: two large basal teeth and two smaller medium-sized teeth at a masticatory surface and two subapical teeth. Cheliceral fixed digit of one sample (eighth examined sample) with nine teeth: three large basal teeth, four medium-sized teeth at masticatory surface and two subapical teeth in addition to apical tooth. Movable digit (
<quantity id="4CBB9B7BFFD1FFE6517BFC13FB16FC14" box="[1053,1164,965,989]" metricMagnitude="-2" metricUnit="m" metricValue="2.9" metricValueMax="3.0" metricValueMin="2.8" pageId="10" pageNumber="1669" unit="mm" value="29.0" valueMax="30.0" valueMin="28.0">2830 µm</quantity>
) with one tooth in addition to an apical tooth. Spermatodactyl
<quantity id="4CBB9B7BFFD1FFE656FDFC33FB90FC37" box="[923,1034,997,1022]" metricMagnitude="-2" metricUnit="m" metricValue="4.4" metricValueMax="4.5" metricValueMin="4.3" pageId="10" pageNumber="1669" unit="mm" value="44.0" valueMax="45.0" valueMin="43.0">4345 µm</quantity>
long with a hyaline ridge above internal canal along its entire length (
<figureCitation id="13782A1BFFD1FFE65634FBD3FC5EFBD4" box="[850,964,1029,1053]" captionStart="Figure 6" captionStartId="8.[212,279,1581,1603]" captionTargetBox="[212,1172,168,1549]" captionTargetId="figure@8.[212,1172,168,1549]" captionTargetPageId="8" captionText="Figure 6. Orthadenella coulsoni sp. nov., male: (A) tritosternum; (B) gnathosoma; (C) chelicerae; (D) epistomes." figureDoi="http://doi.org/10.5281/zenodo.4006386" httpUri="https://zenodo.org/record/4006386/files/figure.png" pageId="10" pageNumber="1669">Figure 6C</figureCitation>
). Dorsal cheliceral seta, dorsal and lateral (antiaxial) lyrifissure distinct. Epistome with an anterior margin irregularly convex, finely denticulate (
<figureCitation id="13782A1BFFD1FFE6571DFB93FD6BFB94" box="[635,753,1093,1117]" captionStart="Figure 6" captionStartId="8.[212,279,1581,1603]" captionTargetBox="[212,1172,168,1549]" captionTargetId="figure@8.[212,1172,168,1549]" captionTargetPageId="8" captionText="Figure 6. Orthadenella coulsoni sp. nov., male: (A) tritosternum; (B) gnathosoma; (C) chelicerae; (D) epistomes." figureDoi="http://doi.org/10.5281/zenodo.4006386" httpUri="https://zenodo.org/record/4006386/files/figure.png" pageId="10" pageNumber="1669">Figure 6D</figureCitation>
).
</paragraph>
<paragraph id="8BFC369EFFD1FFE655B2FB50FCABFB74" blockId="10.[212,1171,1157,1213]" pageId="10" pageNumber="1669">
<emphasis id="B937EA8CFFD1FFE655B2FB50FE8AFB54" box="[212,272,1158,1181]" italics="true" pageId="10" pageNumber="1669">Legs.</emphasis>
Variable in length: I
<quantity id="4CBB9B7BFFD1FFE65746FB53FD36FB54" box="[544,684,1157,1181]" metricMagnitude="-1" metricUnit="m" metricValue="2.75" metricValueMax="2.9" metricValueMin="2.6" pageId="10" pageNumber="1669" unit="mm" value="275.0" valueMax="290.0" valueMin="260.0">260290 µm</quantity>
, II
<quantity id="4CBB9B7BFFD1FFE65788FB53FCE0FB54" box="[750,890,1157,1181]" metricMagnitude="-1" metricUnit="m" metricValue="2.2" metricValueMax="2.25" metricValueMin="2.15" pageId="10" pageNumber="1669" unit="mm" value="220.0" valueMax="225.0" valueMin="215.0">215225 µm</quantity>
, III
<quantity id="4CBB9B7BFFD1FFE656A0FB53FBC8FB54" box="[966,1106,1157,1181]" metricMagnitude="-1" metricUnit="m" metricValue="2.05" metricValueMax="2.1" metricValueMin="2.0" pageId="10" pageNumber="1669" unit="mm" value="205.0" valueMax="210.0" valueMin="200.0">200210 µm</quantity>
, IV
<quantity id="4CBB9B7BFFD1FFE655B2FB73FEFAFB74" box="[212,352,1189,1213]" metricMagnitude="-1" metricUnit="m" metricValue="2.75" metricValueMax="2.9" metricValueMin="2.6" pageId="10" pageNumber="1669" unit="mm" value="275.0" valueMax="290.0" valueMin="260.0">260290 µm</quantity>
. Leg structure and setation as in female.
</paragraph>
</subSubSection>
<subSubSection id="C3596515FFD1FFE655B2FB2BFE73FAB4" pageId="10" pageNumber="1669" type="etymology">
<paragraph id="8BFC369EFFD1FFE655B2FB2BFED7FADC" blockId="10.[212,1172,1277,1405]" box="[212,333,1277,1301]" pageId="10" pageNumber="1669">
<heading id="D0B481F2FFD1FFE655B2FB2BFED7FADC" box="[212,333,1277,1301]" fontSize="10" level="3" pageId="10" pageNumber="1669" reason="5">
<emphasis id="B937EA8CFFD1FFE655B2FB2BFED7FADC" box="[212,333,1277,1301]" italics="true" pageId="10" pageNumber="1669">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BFC369EFFD1FFE655B2FAF2FE73FAB4" blockId="10.[212,1172,1277,1405]" pageId="10" pageNumber="1669">
The species is dedicated to our friend, a scientist exploring the invertebrate fauna of the High Arctic, Prof. Dr Stephen J. Coulson from University Centre in
<collectingCountry id="F354760EFFD1FFE6514EFA92FB0AFA95" box="[1064,1168,1348,1372]" name="Svalbard and Jan Mayen" pageId="10" pageNumber="1669">Svalbard</collectingCountry>
. Longyearbyen,
<collectingCountry id="F354760EFFD1FFE654E0FAB3FE7CFAB4" box="[390,486,1381,1405]" name="Norway" pageId="10" pageNumber="1669">Norway</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3596515FFD1FFE755B2FA6AFEB5FC97" lastPageId="11" lastPageNumber="1670" pageId="10" pageNumber="1669" type="diagnosis">
<paragraph id="8BFC369EFFD1FFE655B2FA6AFE5BFA1D" blockId="10.[212,1172,1468,1691]" box="[212,449,1468,1492]" pageId="10" pageNumber="1669">
<heading id="D0B481F2FFD1FFE655B2FA6AFE5BFA1D" box="[212,449,1468,1492]" fontSize="10" level="3" pageId="10" pageNumber="1669" reason="5">
<emphasis id="B937EA8CFFD1FFE655B2FA6AFE5BFA1D" box="[212,449,1468,1492]" italics="true" pageId="10" pageNumber="1669">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BFC369EFFD1FFE755B2FA32FEB5FC97" blockId="10.[212,1172,1468,1691]" lastBlockId="11.[144,1104,168,862]" lastPageId="11" lastPageNumber="1670" pageId="10" pageNumber="1669">
The morphometric analysis of
<taxonomicName id="4C434D1DFFD1FFE6574BFA32FD30FA35" authorityName="Gwiazdowicz &amp; Marchenko &amp; Teodorowicz" authorityYear="2014" box="[557,682,1508,1532]" class="Arachnida" family="Ascidae" genus="Orthadenella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="10" pageNumber="1669" phylum="Arthropoda" rank="species" species="coulsoni">
<emphasis id="B937EA8CFFD1FFE6574BFA32FD30FA35" box="[557,682,1508,1532]" italics="true" pageId="10" pageNumber="1669">O. coulsoni</emphasis>
</taxonomicName>
shows many different diagnostic characters from the other two species of
<taxonomicName id="4C434D1DFFD1FFE65776F9D2FD06F9D5" authorityName="Athias-Henriot" authorityYear="1973" box="[528,668,1540,1564]" class="Arachnida" family="Ascidae" genus="Orthadenella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="10" pageNumber="1669" phylum="Arthropoda" rank="genus">
<emphasis id="B937EA8CFFD1FFE65776F9D2FD06F9D5" box="[528,668,1540,1564]" italics="true" pageId="10" pageNumber="1669">Orthadenella</emphasis>
</taxonomicName>
. Even a simple analysis of setae measurements gives both similarities (the same lengths of setae
<emphasis id="B937EA8CFFD1FFE65784F9F2FC98F9F5" box="[738,770,1572,1596]" italics="true" pageId="10" pageNumber="1669">Z4</emphasis>
,
<emphasis id="B937EA8CFFD1FFE65669F9F2FCB5F9F5" box="[783,815,1572,1596]" italics="true" pageId="10" pageNumber="1669">Z5</emphasis>
, and setae
<emphasis id="B937EA8CFFD1FFE656CEF9F2FC59F9F5" box="[936,963,1572,1596]" italics="true" pageId="10" pageNumber="1669">J1</emphasis>
,
<emphasis id="B937EA8CFFD1FFE656B6F9F2FC70F9F5" box="[976,1002,1572,1596]" italics="true" pageId="10" pageNumber="1669">J2</emphasis>
,
<emphasis id="B937EA8CFFD1FFE65691F9F2FB8BF9F5" box="[1015,1041,1572,1596]" italics="true" pageId="10" pageNumber="1669">J3</emphasis>
,
<emphasis id="B937EA8CFFD1FFE65179F9F2FBA3F9F5" box="[1055,1081,1572,1596]" italics="true" pageId="10" pageNumber="1669">J4</emphasis>
,
<emphasis id="B937EA8CFFD1FFE65120F9F2FBFCF9F5" box="[1094,1126,1572,1596]" italics="true" pageId="10" pageNumber="1669">Z1</emphasis>
,
<emphasis id="B937EA8CFFD1FFE65115F9F2FB0EF9F5" box="[1139,1172,1572,1596]" italics="true" pageId="10" pageNumber="1669">Z2</emphasis>
longer by 67 μm, and vertical
<emphasis id="B937EA8CFFD1FFE65741F992FDA1F995" box="[551,571,1604,1628]" italics="true" pageId="10" pageNumber="1669">j1</emphasis>
shorter by 4 μm) and dissimilarities when compared to
<emphasis id="B937EA8CFFD1FFE655B2F9B2FE10F9B2" box="[212,394,1636,1660]" italics="true" pageId="10" pageNumber="1669">
<taxonomicName id="4C434D1DFFD1FFE655B2F9B2FE1CF9B2" baseAuthorityName="De Leon" baseAuthorityYear="1963" box="[212,390,1636,1660]" class="Arachnida" family="Ascidae" genus="Orthadenella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="10" pageNumber="1669" phylum="Arthropoda" rank="species" species="tennesseensis">O. tennesseensis</taxonomicName>
.
</emphasis>
A shared character between
<taxonomicName id="4C434D1DFFD1FFE657ABF9B2FCD2F9B2" authorityName="Gwiazdowicz &amp; Marchenko &amp; Teodorowicz" authorityYear="2014" box="[717,840,1635,1660]" class="Arachnida" family="Ascidae" genus="Orthadenella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="10" pageNumber="1669" phylum="Arthropoda" rank="species" species="coulsoni">
<emphasis id="B937EA8CFFD1FFE657ABF9B2FCD2F9B2" box="[717,840,1635,1660]" italics="true" pageId="10" pageNumber="1669">O. coulsoni</emphasis>
</taxonomicName>
and
<taxonomicName id="4C434D1DFFD1FFE656E7F9B5FB93F9B2" authorityName="Evans" authorityYear="1958" box="[897,1033,1635,1659]" class="Arachnida" family="Ascidae" genus="Proctolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="10" pageNumber="1669" phylum="Arthropoda" rank="species" species="lawrencei">
<emphasis id="B937EA8CFFD1FFE656E7F9B5FB93F9B2" box="[897,1033,1635,1659]" italics="true" pageId="10" pageNumber="1669">O. lawrencei</emphasis>
</taxonomicName>
is a marked dorsal reticulated patterning covering almost the whole shield, while in
<taxonomicName id="4C434D1DFFD1FFE65686F955FB17F952" baseAuthorityName="De Leon" baseAuthorityYear="1963" box="[992,1165,1667,1691]" class="Arachnida" family="Ascidae" genus="Orthadenella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="10" pageNumber="1669" phylum="Arthropoda" rank="species" species="tennesseensis">
<emphasis id="B937EA8CFFD1FFE65686F955FB17F952" box="[992,1165,1667,1691]" italics="true" pageId="10" pageNumber="1669">O. tennesseensis</emphasis>
</taxonomicName>
, this is reticulated only on the posterior and anterior border. Moreover,
<taxonomicName id="4C434D1DFFD0FFE756EFFF7EFB9BFF09" authorityName="Gwiazdowicz &amp; Marchenko &amp; Teodorowicz" authorityYear="2014" box="[905,1025,168,192]" class="Arachnida" family="Ascidae" genus="Orthadenella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="1670" phylum="Arthropoda" rank="species" species="coulsoni">
<emphasis id="B937EA8CFFD0FFE756EFFF7EFB9BFF09" box="[905,1025,168,192]" italics="true" pageId="11" pageNumber="1670">O. coulsoni</emphasis>
</taxonomicName>
and
<taxonomicName id="4C434D1DFFD0FFE75152FF7EFF6CFF29" authorityName="Evans" authorityYear="1958" class="Arachnida" family="Ascidae" genus="Proctolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="1670" phylum="Arthropoda" rank="species" species="lawrencei">
<emphasis id="B937EA8CFFD0FFE75152FF7EFF6CFF29" italics="true" pageId="11" pageNumber="1670">O. lawrencei</emphasis>
</taxonomicName>
have a humeral seta
<emphasis id="B937EA8CFFD0FFE75480FF1EFE67FF29" box="[486,509,200,224]" italics="true" pageId="11" pageNumber="1670">r3</emphasis>
conspicuously longer than the remaining setae in the marginal row
<emphasis id="B937EA8CFFD0FFE7544CFF3FFECEFEC9" box="[298,340,233,256]" italics="true" pageId="11" pageNumber="1670">rR</emphasis>
, which
<taxonomicName id="4C434D1DFFD0FFE754CCFF3EFDCDFEC9" baseAuthorityName="De Leon" baseAuthorityYear="1963" box="[426,599,232,256]" class="Arachnida" family="Ascidae" genus="Orthadenella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="1670" phylum="Arthropoda" rank="species" species="tennesseensis">
<emphasis id="B937EA8CFFD0FFE754CCFF3EFDCDFEC9" box="[426,599,232,256]" italics="true" pageId="11" pageNumber="1670">O. tennesseensis</emphasis>
</taxonomicName>
does not. This character is repeated in the
<emphasis id="B937EA8CFFD0FFE7514AFF3EFF36FEE9" italics="true" pageId="11" pageNumber="1670">S4, S5</emphasis>
and
<emphasis id="B937EA8CFFD0FFE7558EFEDEFE92FEE9" box="[232,264,264,288]" italics="true" pageId="11" pageNumber="1670">Z5</emphasis>
pairs of setae. However,
<taxonomicName id="4C434D1DFFD0FFE7574CFEDEFD2EFEE9" authorityName="Evans" authorityYear="1958" box="[554,692,264,288]" class="Arachnida" family="Ascidae" genus="Proctolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="1670" phylum="Arthropoda" rank="species" species="lawrencei">
<emphasis id="B937EA8CFFD0FFE7574CFEDEFD2EFEE9" box="[554,692,264,288]" italics="true" pageId="11" pageNumber="1670">O. lawrencei</emphasis>
</taxonomicName>
setae
<emphasis id="B937EA8CFFD0FFE7579AFEDEFC87FEE9" box="[764,797,264,288]" italics="true" pageId="11" pageNumber="1670">Z4</emphasis>
are identical to the
<emphasis id="B937EA8CFFD0FFE75164FEDEFBCAFEE9" box="[1026,1104,264,288]" italics="true" pageId="11" pageNumber="1670">Z1Z3</emphasis>
setae, whereas in
<taxonomicName id="4C434D1DFFD0FFE75436FEFEFE50FE89" authorityName="Gwiazdowicz &amp; Marchenko &amp; Teodorowicz" authorityYear="2014" box="[336,458,296,320]" class="Arachnida" family="Ascidae" genus="Orthadenella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="1670" phylum="Arthropoda" rank="species" species="coulsoni">
<emphasis id="B937EA8CFFD0FFE75436FEFEFE50FE89" box="[336,458,296,320]" italics="true" pageId="11" pageNumber="1670">O. coulsoni</emphasis>
</taxonomicName>
these setae are dissimilar. In addition, the location of pore
<emphasis id="B937EA8CFFD0FFE755F6FE9EFF35FEA9" box="[144,175,328,352]" italics="true" pageId="11" pageNumber="1670">iv5</emphasis>
is a further diagnostic character. This pore lies outside the genital shield of both
<taxonomicName id="4C434D1DFFD0FFE75152FE9EFF72FE49" authorityName="Gwiazdowicz &amp; Marchenko &amp; Teodorowicz" authorityYear="2014" class="Arachnida" family="Ascidae" genus="Orthadenella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="1670" phylum="Arthropoda" rank="species" species="coulsoni">
<emphasis id="B937EA8CFFD0FFE75152FE9EFF72FE49" italics="true" pageId="11" pageNumber="1670">O. coulsoni</emphasis>
</taxonomicName>
and
<taxonomicName id="4C434D1DFFD0FFE7547AFEBEFE2AFEB6" box="[284,432,360,384]" class="Arachnida" family="Ascidae" genus="Orthadenella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="1670" phylum="Arthropoda" rank="species" species="tenessensis">
<emphasis id="B937EA8CFFD0FFE7547AFEBEFE2AFEB6" box="[284,432,360,384]" italics="true" pageId="11" pageNumber="1670">O. tenessensis</emphasis>
</taxonomicName>
but, as is more common, on the shield in
<taxonomicName id="4C434D1DFFD0FFE75614FEBEFC63FE49" authorityName="Evans" authorityYear="1958" box="[882,1017,360,384]" class="Arachnida" family="Ascidae" genus="Proctolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="1670" phylum="Arthropoda" rank="species" species="lawrencei">
<emphasis id="B937EA8CFFD0FFE75614FEBEFC63FE49" box="[882,1017,360,384]" italics="true" pageId="11" pageNumber="1670">O. lawrencei</emphasis>
</taxonomicName>
. A very fine character separating the species is the appearance of a ventrianal shield. This has a characteristic concave anterior boarder at the level between the genital and metapodal sclerites for the entire genus, but differs in shape among species. That of
<taxonomicName id="4C434D1DFFD0FFE756DAFE11FBADFE16" authorityName="Gwiazdowicz &amp; Marchenko &amp; Teodorowicz" authorityYear="2014" box="[956,1079,455,479]" class="Arachnida" family="Ascidae" genus="Orthadenella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="1670" phylum="Arthropoda" rank="species" species="coulsoni">
<emphasis id="B937EA8CFFD0FFE756DAFE11FBADFE16" box="[956,1079,455,479]" italics="true" pageId="11" pageNumber="1670">O. coulsoni</emphasis>
</taxonomicName>
is wider than long, contrary to the other species. Likewise, the difference in the number of setae located on the ventrianal shield, excluding a circum-anal setae, is another significant character, differentiating
<taxonomicName id="4C434D1DFFD0FFE754B8FDF1FD17FDF6" baseAuthorityName="De Leon" baseAuthorityYear="1963" box="[478,653,551,575]" class="Arachnida" family="Ascidae" genus="Orthadenella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="1670" phylum="Arthropoda" rank="species" species="tennesseensis">
<emphasis id="B937EA8CFFD0FFE754B8FDF1FD17FDF6" box="[478,653,551,575]" italics="true" pageId="11" pageNumber="1670">O. tennesseensis</emphasis>
</taxonomicName>
, with five pairs, from the two remaining species, each bearing six pairs. The metapodal shields in the opistogastric region clearly distinguish
<taxonomicName id="4C434D1DFFD0FFE75469FDB1FE03FDB6" authorityName="Evans" authorityYear="1958" box="[271,409,615,639]" class="Arachnida" family="Ascidae" genus="Proctolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="1670" phylum="Arthropoda" rank="species" species="lawrencei">
<emphasis id="B937EA8CFFD0FFE75469FDB1FE03FDB6" box="[271,409,615,639]" italics="true" pageId="11" pageNumber="1670">O. lawrencei</emphasis>
</taxonomicName>
which possesses only a single pair, while
<taxonomicName id="4C434D1DFFD0FFE75615FDB1FC75FDB6" authorityName="Gwiazdowicz &amp; Marchenko &amp; Teodorowicz" authorityYear="2014" box="[883,1007,615,639]" class="Arachnida" family="Ascidae" genus="Orthadenella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="1670" phylum="Arthropoda" rank="species" species="coulsoni">
<emphasis id="B937EA8CFFD0FFE75615FDB1FC75FDB6" box="[883,1007,615,639]" italics="true" pageId="11" pageNumber="1670">O. coulsoni</emphasis>
</taxonomicName>
has two pairs composed of the larger sclerite with a smaller abutting. The epistome of
<taxonomicName id="4C434D1DFFD0FFE75152FD51FE81FD77" baseAuthorityName="De Leon" baseAuthorityYear="1963" class="Arachnida" family="Ascidae" genus="Orthadenella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="1670" phylum="Arthropoda" rank="species" species="tennesseensis">
<emphasis id="B937EA8CFFD0FFE75152FD51FE81FD77" italics="true" pageId="11" pageNumber="1670">O. tennesseensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C434D1DFFD0FFE7543FFD71FE7EFD76" authorityName="Evans" authorityYear="1958" box="[345,484,679,703]" class="Arachnida" family="Ascidae" genus="Proctolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="1670" phylum="Arthropoda" rank="species" species="lawrencei">
<emphasis id="B937EA8CFFD0FFE7543FFD71FE7EFD76" box="[345,484,679,703]" italics="true" pageId="11" pageNumber="1670">O. lawrencei</emphasis>
</taxonomicName>
is a trispinate, median process broadly triangular and extending beyond the apex of the lateral processes, contrary to
<taxonomicName id="4C434D1DFFD0FFE75630FD11FC4AFD16" authorityName="Gwiazdowicz &amp; Marchenko &amp; Teodorowicz" authorityYear="2014" box="[854,976,711,735]" class="Arachnida" family="Ascidae" genus="Orthadenella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="1670" phylum="Arthropoda" rank="species" species="coulsoni">
<emphasis id="B937EA8CFFD0FFE75630FD11FC4AFD16" box="[854,976,711,735]" italics="true" pageId="11" pageNumber="1670">O. coulsoni</emphasis>
</taxonomicName>
, which has convex epistome with homogenous denticles arranged parallel to each other. The spermatheca of
<taxonomicName id="4C434D1DFFD0FFE75477FCD0FE03FCD7" authorityName="Evans" authorityYear="1958" box="[273,409,774,798]" class="Arachnida" family="Ascidae" genus="Proctolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="1670" phylum="Arthropoda" rank="species" species="lawrencei">
<emphasis id="B937EA8CFFD0FFE75477FCD0FE03FCD7" box="[273,409,774,798]" italics="true" pageId="11" pageNumber="1670">O. lawrencei</emphasis>
</taxonomicName>
is composed of a sacculus permeated with numerous pores and cylinders, while the spermatheca of
<taxonomicName id="4C434D1DFFD0FFE75778FCF0FD03FCF7" authorityName="Gwiazdowicz &amp; Marchenko &amp; Teodorowicz" authorityYear="2014" box="[542,665,806,830]" class="Arachnida" family="Ascidae" genus="Orthadenella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="1670" phylum="Arthropoda" rank="species" species="coulsoni">
<emphasis id="B937EA8CFFD0FFE75778FCF0FD03FCF7" box="[542,665,806,830]" italics="true" pageId="11" pageNumber="1670">O. coulsoni</emphasis>
</taxonomicName>
is permeated with numerous pores and thin filaments.
</paragraph>
</subSubSection>
<subSubSection id="C3596515FFD0FFE755F6FC50FF70FA96" pageId="11" pageNumber="1670" type="key">
<paragraph id="8BFC369EFFD0FFE755F6FC50FDC7FC54" blockId="11.[144,1104,901,1021]" box="[144,605,901,926]" pageId="11" pageNumber="1670">
<heading id="D0B481F2FFD0FFE755F6FC50FDC7FC54" box="[144,605,901,926]" fontSize="10" level="3" pageId="11" pageNumber="1670" reason="5">
<emphasis id="B937EA8CFFD0FFE755F6FC50FDC7FC54" bold="true" box="[144,605,901,926]" pageId="11" pageNumber="1670">
Key to the females of genus
<taxonomicName id="4C434D1DFFD0FFE754A8FC53FDC7FC54" authorityName="Athias-Henriot" authorityYear="1973" box="[462,605,901,925]" class="Arachnida" family="Ascidae" genus="Orthadenella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="1670" phylum="Arthropoda" rank="genus">
<emphasis id="B937EA8CFFD0FFE754A8FC53FDC7FC54" bold="true" box="[462,605,901,925]" italics="true" pageId="11" pageNumber="1670">Orthadenella</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BFC369EFFD0FFE755F6FC73FDE7FC34" blockId="11.[144,1104,901,1021]" pageId="11" pageNumber="1670">
Information concerning
<taxonomicName id="4C434D1DFFD0FFE754CCFC70FDADFC77" authorityName="Evans" authorityYear="1958" box="[426,567,934,958]" class="Arachnida" family="Ascidae" genus="Proctolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="1670" phylum="Arthropoda" rank="species" species="lawrencei">
<emphasis id="B937EA8CFFD0FFE754CCFC70FE24FC77" box="[426,446,934,958]" italics="true" pageId="11" pageNumber="1670">O</emphasis>
.
<emphasis id="B937EA8CFFD0FFE754B6FC70FDADFC77" box="[464,567,934,958]" italics="true" pageId="11" pageNumber="1670">lawrencei</emphasis>
</taxonomicName>
and
<taxonomicName id="4C434D1DFFD0FFE7571EFC70FCB6FC74" baseAuthorityName="De Leon" baseAuthorityYear="1963" box="[632,812,934,958]" class="Arachnida" family="Ascidae" genus="Orthadenella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="1670" phylum="Arthropoda" rank="species" species="tennesseensis">
<emphasis id="B937EA8CFFD0FFE7571EFC70FD16FC77" box="[632,652,934,958]" italics="true" pageId="11" pageNumber="1670">O</emphasis>
.
<emphasis id="B937EA8CFFD0FFE757F8FC70FCB6FC74" box="[670,812,934,957]" italics="true" pageId="11" pageNumber="1670">tennesseensis</emphasis>
</taxonomicName>
was obtained from published descriptions and illustrations (
<bibRefCitation id="EFD24B6FFFD0FFE7572CFC13FD4EFC14" author="Evans GO" box="[586,724,965,989]" pageId="11" pageNumber="1670" pagination="177 - 229" refId="ref4916" refString="Evans GO. 1958. A revision of the British Aceosejinae (Acarina: Mesostigmata). Proc Zool Soc Lond. 131: 177 - 229. doi: 10.1111 / j. 1096 - 3642.1958. tb 00685. x" type="journal article" year="1958">Evans 1958</bibRefCitation>
;
<bibRefCitation id="EFD24B6FFFD0FFE7578CFC10FC03FC14" author="De Leon D" box="[746,921,965,989]" pageId="11" pageNumber="1670" pagination="197 - 207" refId="ref4875" refString="De Leon D. 1963. A new genus and twelve new species of mites from Mexico and Southeast United States (Acarina: Blattisocidae). Fla Entomol. 46: 197 - 207. doi: 10.2307 / 3493632" type="journal article" year="1963">De Leon 1963</bibRefCitation>
;
<bibRefCitation id="EFD24B6FFFD0FFE756D6FC13FE87FC34" author="McGraw JR &amp; Farrier MH" pageId="11" pageNumber="1670" pagination="1 - 162" refId="ref5284" refString="McGraw JR, Farrier MH. 1969. Mites of the superfamily Parasitoidea (Acarina: Mesostigmata) associated with Dendroctonus and Ips (Coleoptera: Scolytidae). N C Agric Exper Stat Tech Bull. 192: 1 - 162." type="journal article" year="1969">McGraw and Farrier 1969</bibRefCitation>
;
<bibRefCitation id="EFD24B6FFFD0FFE75448FC30FDF6FC34" author="Moraza ML &amp; Lindquist EE" box="[302,620,997,1021]" pageId="11" pageNumber="1670" pagination="1 - 25" refId="ref5327" refString="Moraza ML, Lindquist EE. 2011. A new genus of fungus-inhabiting blattisociid mites (Acari: Mesostigmata: Phytoseioidea) from Middle America, with a key to genera and subgenera of the subfamily Blattisociinae. Zootaxa. 2758: 1 - 25." type="journal article" year="2011">Moraza and Lindquist 2011</bibRefCitation>
).
</paragraph>
<key id="FFD261AFFFD0FFE755E9FBF3FBCAFAC5" pageId="11" pageNumber="1670">
<keyStep id="30B72E3BFFD0FFE755E9FBF3FBD5FBB4" pageId="11" pageNumber="1670">
<paragraph id="8BFC369EFFD0FFE755E9FBF3FBCAFB94" blockId="11.[143,1108,1061,1149]" pageId="11" pageNumber="1670">
<keyLead id="30B295ABFFD0FFE755E9FBF3FBCAFB94" pageId="11" pageNumber="1670">
1. Only one metapodal plate on each side of the body behind coxae IV .................. ......................................................................................
<taxonomicName id="4C434D1DFFD0FFE75641FB93FBCAFB94" authority="(Evans 1958)" baseAuthorityName="Evans" baseAuthorityYear="1958" box="[807,1104,1093,1117]" class="Arachnida" family="Ascidae" genus="Proctolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="1670" phylum="Arthropoda" rank="species" species="lawrencei">
<emphasis id="B937EA8CFFD0FFE75641FB93FCA1FB94" box="[807,827,1093,1117]" italics="true" pageId="11" pageNumber="1670">O</emphasis>
.
<emphasis id="B937EA8CFFD0FFE7562DFB93FC29FB94" box="[843,947,1093,1117]" italics="true" pageId="11" pageNumber="1670">lawrencei</emphasis>
(
<bibRefCitation id="EFD24B6FFFD0FFE756A5FB93FBDDFB94" author="Evans GO" box="[963,1095,1093,1117]" pageId="11" pageNumber="1670" pagination="177 - 229" refId="ref4916" refString="Evans GO. 1958. A revision of the British Aceosejinae (Acarina: Mesostigmata). Proc Zool Soc Lond. 131: 177 - 229. doi: 10.1111 / j. 1096 - 3642.1958. tb 00685. x" type="journal article" year="1958">Evans 1958</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFC369EFFD0FFE755E9FBB3FBD5FBB4" blockId="11.[143,1108,1061,1149]" box="[143,1103,1125,1149]" pageId="11" pageNumber="1670">
<keyLead id="30B295ABFFD0FFE755E9FBB3FBD5FBB4" box="[143,1103,1125,1149]" pageId="11" pageNumber="1670"> Metapodal plates divided into two small plates ................................................. 2</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B72E3BFFD0FFE755E9FB42FBCAFAC5" pageId="11" pageNumber="1670">
<paragraph id="8BFC369EFFD0FFE755E9FB42FBCAFB05" blockId="11.[143,1108,1172,1292]" pageId="11" pageNumber="1670">
<keyLead id="30B295ABFFD0FFE755E9FB42FBCAFB05" pageId="11" pageNumber="1670">
2. Interscutal setae
<emphasis id="B937EA8CFFD0FFE7541FFB43FE3DFB64" box="[377,423,1173,1197]" italics="true" pageId="11" pageNumber="1670">JV4</emphasis>
outside ventrianal shield, length and width of ventrianal shield similar ......................................................
<taxonomicName id="4C434D1DFFD0FFE75786FB63FBCAFB05" authority="(De Leon 1963)" baseAuthorityName="De Leon" baseAuthorityYear="1963" box="[736,1104,1204,1229]" class="Arachnida" family="Ascidae" genus="Orthadenella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="1670" phylum="Arthropoda" rank="species" species="tennesseensis">
<emphasis id="B937EA8CFFD0FFE75786FB63FC09FB05" box="[736,915,1205,1229]" italics="true" pageId="11" pageNumber="1670">O. tennesseensis</emphasis>
(
<bibRefCitation id="EFD24B6FFFD0FFE756C2FB62FBDDFB05" author="De Leon D" box="[932,1095,1204,1228]" pageId="11" pageNumber="1670" pagination="197 - 207" refId="ref4875" refString="De Leon D. 1963. A new genus and twelve new species of mites from Mexico and Southeast United States (Acarina: Blattisocidae). Fla Entomol. 46: 197 - 207. doi: 10.2307 / 3493632" type="journal article" year="1963">De Leon 1963</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFC369EFFD0FFE755F4FB02FBCAFAC5" blockId="11.[143,1108,1172,1292]" pageId="11" pageNumber="1670">
<keyLead id="30B295ABFFD0FFE755F4FB02FBCAFAC5" pageId="11" pageNumber="1670">
Interscutal setae
<emphasis id="B937EA8CFFD0FFE75415FB02FE3AFB25" box="[371,416,1236,1260]" italics="true" pageId="11" pageNumber="1670">JV4</emphasis>
on ventrianal shield, ventrianal shield wider than long ....... ....................................................................................................
<taxonomicName id="4C434D1DFFD0FFE756EDFB22FB92FAC5" authority="Gwiazdowicz &amp; Marchenko &amp; Teodorowicz, 2014" authorityName="Gwiazdowicz &amp; Marchenko &amp; Teodorowicz" authorityYear="2014" box="[907,1032,1268,1292]" class="Arachnida" family="Ascidae" genus="Orthadenella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="1670" phylum="Arthropoda" rank="species" species="coulsoni" status="sp. nov.">
<emphasis id="B937EA8CFFD0FFE756EDFB22FB92FAC5" box="[907,1032,1268,1292]" italics="true" pageId="11" pageNumber="1670">O. coulsoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20457F7FFD0FFE75174FB23FBCAFAC5" box="[1042,1104,1269,1292]" pageId="11" pageNumber="1670" rank="species">n. sp.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
</key>
<paragraph id="8BFC369EFFD0FFE755F6FA91FF70FA96" blockId="11.[144,1103,1351,1444]" box="[144,234,1351,1375]" pageId="11" pageNumber="1670">
<emphasis id="B937EA8CFFD0FFE755F6FA91FF70FA96" bold="true" box="[144,234,1351,1375]" pageId="11" pageNumber="1670">Funding</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3596515FFD0FFE755F6FAA6FC30FA6A" pageId="11" pageNumber="1670" type="description">
<paragraph id="8BFC369EFFD0FFE755F6FAA6FC30FA6A" blockId="11.[144,1103,1351,1444]" pageId="11" pageNumber="1670">
This study and field work of one of us (I.I. Marchenko) was supported by The Federal Fundamental Scientific Research Programme for
<date id="FFFD105EFFD0FFE757EAFA5BFD62FA6A" box="[652,760,1421,1444]" pageId="11" pageNumber="1670" value="2013" valueMax="2020">20132020</date>
[VI.51.1.7. 30.4].
</paragraph>
</subSubSection>
</treatment>
</document>