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<document id="3BFF76BB95333B551AA336DA446799D3" ID-DOI="10.1649/0010-065x-69.4.739" ID-ISSN="1938-4394" ID-Zenodo-Dep="10106512" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" checkinTime="1630178374591" checkinUser="diego" docAuthor="Bares, Hannah M. &amp; Ivie, Michael A." docDate="2015" docId="03C087F9FF64A72A06D553F814577D31" docLanguage="en" docName="0010-065X-69.4.739.pdf.imf" docOrigin="The Coleopterists Bulletin 69 (4)" docSource="http://dx.doi.org/10.1649/0010-065x-69.4.739" docStyle="DocumentStyle:0305A7CAC51C960DDA50A7292A4D3620.8:TheColeopBull.2010-2016.journal_article.0cover" docStyleId="0305A7CAC51C960DDA50A7292A4D3620" docStyleName="TheColeopBull.2010-2016.journal_article.0cover" docStyleVersion="8" docTitle="Carinodula chiapanensis Bares and Ivie, new species" docType="treatment" docVersion="4" lastPageNumber="742" masterDocId="FFF9FF81FF66A7290432567F102B7E0B" masterDocTitle="A Second Species ofCarinodulaGordon, Pakaluk, and Ślipiński (Coleoptera: Coccinellidae: Microweiseinae: Carinodulini)" masterLastPageNumber="743" masterPageNumber="739" pageNumber="741" updateTime="1699637797967" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="27CE0F6778A3C9A9F089B89C6498ED38" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="3503E81D6A90DC175AFE19D5E090BFD7">
<mods:title id="A874FED55263E6EF4DFB877B43D2B087">A Second Species ofCarinodulaGordon, Pakaluk, and Ślipiński (Coleoptera: Coccinellidae: Microweiseinae: Carinodulini)</mods:title>
</mods:titleInfo>
<mods:name id="63F0D40BFDDC3281A33CD5A08B2035BC" type="personal">
<mods:role id="D05CD32779F51D732847EF495247FDBD">
<mods:roleTerm id="282CDED37A8F755E6DE337C9DDE82FD2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3CF5B1D85E6E58DB1DE8BDF79600CA2C">Bares, Hannah M.</mods:namePart>
</mods:name>
<mods:name id="A9237E63BD65916A96872EA9AA1B479A" type="personal">
<mods:role id="BD0AD6C396B8F6E88C6AE92B045D4F4E">
<mods:roleTerm id="5CDB7401261139697F4D75BC64DF7042">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7342DFE71D8B61B1DEFDE33A7B0AD101">Ivie, Michael A.</mods:namePart>
</mods:name>
<mods:typeOfResource id="92A4615483BA73B6D8CD3E8316E24BE1">text</mods:typeOfResource>
<mods:relatedItem id="DD49CAF7BCA9E1C6E7F06B3E1878A6F7" type="host">
<mods:titleInfo id="52F3F5C45539EA4B6D5C230697EF7655">
<mods:title id="309AC1C9005E8D51C609C539DCD0BE52">The Coleopterists Bulletin</mods:title>
</mods:titleInfo>
<mods:part id="255D712ED61A823290BDEEDC1F8B9AEB">
<mods:date id="BDE7128D4C44792416C48750BC750884">2015</mods:date>
<mods:detail id="8DB878E69166FF75152C0BD6739469A4" type="pubDate">
<mods:number id="4C34F0E0EA5190C4767669198D068C51">2015-12-31</mods:number>
</mods:detail>
<mods:detail id="515C7C19732ABA7CE6D59F353DE4C142" type="volume">
<mods:number id="9C86DCB52D19BA5C8F58C1FC75DF0E10">69</mods:number>
</mods:detail>
<mods:detail id="D58E247691C8ABDEAEB1A47151FBA2FD" type="issue">
<mods:number id="8742CC04FE6A2BAF386CA3AACD28841F">4</mods:number>
</mods:detail>
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<mods:start id="64277462731914F76550BDA58A60CC87">739</mods:start>
<mods:end id="82BF526278EB412AC1A259672FB6D82F">743</mods:end>
</mods:extent>
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</mods:relatedItem>
<mods:location id="ADF351593D1E06BAE7F076DFB76D58D7">
<mods:url id="5B8776A19653400083B1C27D77AAD7B5">http://dx.doi.org/10.1649/0010-065x-69.4.739</mods:url>
</mods:location>
<mods:classification id="12A744A214EEB795451DEA6C8934C534">journal article</mods:classification>
<mods:identifier id="513A051343516FDC335484CF316EF4D1" type="DOI">10.1649/0010-065x-69.4.739</mods:identifier>
<mods:identifier id="6173ED2D731E0EF81D09B44808B48F9A" type="ISSN">1938-4394</mods:identifier>
<mods:identifier id="86C1C3B66B51CB87BCD8962C5C16F671" type="Zenodo-Dep">10106512</mods:identifier>
</mods:mods>
<treatment id="03C087F9FF64A72A06D553F814577D31" ID-DOI="http://doi.org/10.5281/zenodo.5415264" ID-Zenodo-Dep="5415264" LSID="urn:lsid:plazi:treatment:03C087F9FF64A72A06D553F814577D31" httpUri="http://treatment.plazi.org/id/03C087F9FF64A72A06D553F814577D31" lastPageId="3" lastPageNumber="742" pageId="2" pageNumber="741">
<subSubSection id="C3736564FF64A72B06D553F813DB7BB3" pageId="2" pageNumber="741" type="nomenclature">
<paragraph id="8BD636EFFF64A72B06D553F813DB7BB3" blockId="2.[743,1157,1415,1464]" pageId="2" pageNumber="741">
<heading id="D09E8183FF64A72B06D553F813DB7BB3" centered="true" fontSize="10" level="2" pageId="2" pageNumber="741" reason="2">
<emphasis id="B91DEAFDFF64A72B06D553F813DB7BB3" bold="true" pageId="2" pageNumber="741">
<taxonomicName id="4C694D6CFF64A72B06D553F814AB7B96" authority="Bares and Ivie" authorityName="Bares and Ivie" box="[743,1152,1415,1437]" class="Insecta" family="Coccinellidae" genus="Carinodula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="741" phylum="Arthropoda" rank="species" species="chiapanensis" status="sp. nov.">
<emphasis id="B91DEAFDFF64A72B06D553F813F47B96" bold="true" box="[743,991,1415,1437]" italics="true" pageId="2" pageNumber="741">Carinodula chiapanensis</emphasis>
Bares and Ivie
</taxonomicName>
,
<taxonomicNameLabel id="A22E5786FF64A72B074953DC13DB7BB3" box="[891,1008,1443,1464]" pageId="2" pageNumber="741" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3736564FF64A72A06E953A7113D7A55" lastPageId="3" lastPageNumber="742" pageId="2" pageNumber="741" type="materials_examined">
<paragraph id="8BD636EFFF64A72A06E953A7116F7D5F" blockId="2.[707,1194,1495,1729]" lastBlockId="3.[151,638,725,1730]" lastPageId="3" lastPageNumber="742" pageId="2" pageNumber="741">
<emphasis id="B91DEAFDFF64A72B06E953A713AF7BE6" bold="true" box="[731,900,1496,1517]" pageId="2" pageNumber="741">Type Specimens.</emphasis>
<materialsCitation id="3B013CB2FF64A72A07A053A7116F7D5F" collectingDate="2008-05-17" collectionCode="LLAMA" country="Mexico" elevation="1520" lastPageId="3" lastPageNumber="742" latitude="15.71604" location="Custepec" longLatPrecision="1" longitude="-92.93835" pageId="2" pageNumber="741" specimenCount="16" specimenCount-female="7" specimenCount-male="8" stateProvince="Chiapas" typeStatus="holotype">
<typeStatus id="54D2884DFF64A72B07A053A713C17BE6" box="[914,1002,1496,1517]" pageId="2" pageNumber="741" type="holotype">Holotype</typeStatus>
<specimenCount id="9D6FFD66FF64A72B07C053A8142F7BE6" box="[1010,1028,1495,1517]" pageId="2" pageNumber="741" type="male">♂</specimenCount>
,
<typeStatus id="54D2884DFF64A72B002053A714757BE6" box="[1042,1118,1496,1517]" pageId="2" pageNumber="741" type="allotype">allotype</typeStatus>
<specimenCount id="9D6FFD66FF64A72B005453A814537BE6" box="[1126,1144,1495,1517]" pageId="2" pageNumber="741" type="female">♀</specimenCount>
, and
<specimenCount id="9D6FFD66FF64A72B06F1538D1377780C" box="[707,860,1522,1543]" pageId="2" pageNumber="741" type="generic" typeStatus="paratypes">seven paratypes</specimenCount>
(
<specimenCount id="9D6FFD66FF64A72B075E538D138E7803" box="[876,933,1522,1544]" pageId="2" pageNumber="741" type="female">4 ♀♀</specimenCount>
,
<specimenCount id="9D6FFD66FF64A72B0786538D13CF7803" box="[948,996,1522,1544]" pageId="2" pageNumber="741" type="male">3♂♂</specimenCount>
) labeled:
<collectingCountry id="F37E767FFF64A72B0074538D148D780C" box="[1094,1190,1522,1543]" name="Mexico" pageId="2" pageNumber="741">MEXICO</collectingCountry>
:
<collectingRegion id="49ADF80DFF64A72B06F15072133C7829" box="[707,791,1549,1570]" country="Mexico" name="Chiapas" pageId="2" pageNumber="741">Chiapas</collectingRegion>
/
<quantity id="4C919B0AFF64A72B072D507213657829" box="[799,846,1549,1570]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="741" unit="km" value="2.0">2km</quantity>
SE
<location id="8EB66034FF64A72B07B2507213CE7829" LSID="urn:lsid:plazi:treatment:03C087F9FF64A72A06D553F814577D31:8EB66034FF64A72B07B2507213CE7829" box="[896,997,1549,1570]" country="Mexico" latitude="15.71604" longLatPrecision="1" longitude="-92.93835" name="Custepec" pageId="2" pageNumber="741" stateProvince="Chiapas">Custepec</location>
,
<quantity id="4C919B0AFF64A72B07DB507214187829" box="[1001,1075,1549,1570]" metricMagnitude="3" metricUnit="m" metricValue="1.52" pageId="2" pageNumber="741" unit="m" value="1520.0">
<elevation id="0044D1DCFF64A72B07DB507214187829" box="[1001,1075,1549,1570]" metricMagnitude="3" metricUnit="m" metricValue="1.52" pageId="2" pageNumber="741" unit="m" value="1520.0">1520m</elevation>
</quantity>
/
<geoCoordinate id="EE5D5028FF64A72B0009507212FF7836" degrees="15.72099" direction="north" orientation="latitude" pageId="2" pageNumber="741" precision="1" value="15.72099">15.72099° N</geoCoordinate>
,
<geoCoordinate id="EE5D5028FF64A72B06E5505713677836" box="[727,844,1576,1597]" degrees="92.95054" direction="west" orientation="longitude" pageId="2" pageNumber="741" precision="1" value="-92.95054">92.95054°W</geoCoordinate>
/
<date id="FFD7102FFF64A72B0766505713E77836" box="[852,972,1576,1597]" pageId="2" pageNumber="741" value="2008-05-17">
<collectingDate id="EF93E9C7FF64A72B0766505713E77836" box="[852,972,1576,1597]" pageId="2" pageNumber="741" value="2008-05-17">17MAY2008</collectingDate>
</date>
;
<collectionCode id="ED78AE2AFF64A72B07E4505714017836" box="[982,1066,1576,1597]" pageId="2" pageNumber="741">LLAMA</collectionCode>
#Wm-A-02-1/ sifted mesophyll litter. Six
<typeStatus id="54D2884DFF64A72B07F3503D1431785C" box="[961,1050,1602,1623]" pageId="2" pageNumber="741" type="paratype">paratypes</typeStatus>
(
<specimenCount id="9D6FFD66FF64A72B0015503D14757853" box="[1063,1118,1602,1624]" pageId="2" pageNumber="741" type="female">2 ♀♀</specimenCount>
,
<specimenCount id="9D6FFD66FF64A72B0059503D148A7853" box="[1131,1185,1602,1624]" pageId="2" pageNumber="741" type="male">4 ♂♂</specimenCount>
) labeled:
<collectingCountry id="F37E767FFF64A72B073E502213427879" box="[780,873,1629,1650]" name="Mexico" pageId="2" pageNumber="741">MEXICO</collectingCountry>
:
<collectingRegion id="49ADF80DFF64A72B07425022139C7879" box="[880,951,1629,1650]" country="Mexico" name="Chiapas" pageId="2" pageNumber="741">Chiapas</collectingRegion>
/
<locationDeviation id="9A642FCDFF64A72B078C502214767879" box="[958,1117,1629,1650]" location="Custepec" pageId="2" pageNumber="741">
<quantity id="4C919B0AFF64A72B078C502213CD7879" box="[958,998,1629,1650]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="741" unit="km" value="3.0">3km</quantity>
SE Custepec
</locationDeviation>
,
<quantity id="4C919B0AFF64A72B0056502214897879" box="[1124,1186,1629,1650]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="2" pageNumber="741" unit="m" value="1700.0">
<elevation id="0044D1DCFF64A72B0056502214897879" box="[1124,1186,1629,1650]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="2" pageNumber="741" unit="m" value="1700.0">1700m</elevation>
</quantity>
/
<geoCoordinate id="EE5D5028FF64A72B06F1500813127887" box="[707,825,1655,1676]" degrees="15.71604" direction="north" orientation="latitude" pageId="2" pageNumber="741" precision="1" value="15.71604">15.71604°N</geoCoordinate>
<geoCoordinate id="EE5D5028FF64A72B0773500813907887" box="[833,955,1655,1676]" degrees="92.93835" direction="west" orientation="longitude" pageId="2" pageNumber="741" precision="1" value="-92.93835">92.93835°W</geoCoordinate>
/
<date id="FFD7102FFF64A72B07F05008146A7887" box="[962,1089,1655,1676]" pageId="2" pageNumber="741" value="2008-05-17">
<collectingDate id="EF93E9C7FF64A72B07F05008146A7887" box="[962,1089,1655,1676]" pageId="2" pageNumber="741" value="2008-05-17">17MAY2008</collectingDate>
</date>
;
<collectionCode id="ED78AE2AFF64A72B0076500814B77887" box="[1092,1180,1655,1676]" pageId="2" pageNumber="741">LLAMA</collectionCode>
# Wm-A-02-2/sifted mesophyll litter. One partial specimen from the type locality is not included in the type series. [In the
<collectionCode id="ED78AE2AFF65A72A057E54AA11B47CE1" box="[332,415,725,746]" pageId="3" pageNumber="742">LLAMA</collectionCode>
codes, “Wm” indicates Maxi-Winkler extraction; “A” indicates Mexico; the two-digit number indicates site number and the final digit the transect number (Longino and Anderson 2014).]
</materialsCitation>
</paragraph>
<caption id="DF166667FF65A72A049D541110F77C93" ID-DOI="http://doi.org/10.5281/zenodo.10106518" ID-Zenodo-Dep="10106518" httpUri="https://zenodo.org/record/10106518/files/figure.png" pageId="3" pageNumber="742" startId="3.[175,207,622,641]" targetBox="[156,633,194,601]" targetPageId="3">
<paragraph id="8BD636EFFF65A72A049D541112547C8A" blockId="3.[151,639,622,664]" box="[175,639,622,641]" pageId="3" pageNumber="742">
<emphasis id="B91DEAFDFF65A72A049D541110DF7C8A" bold="true" box="[175,244,622,641]" pageId="3" pageNumber="742">Fig. 10.</emphasis>
Distribution of
<taxonomicName id="4C694D6CFF65A72A05B65411120B7C8A" box="[388,544,622,641]" class="Insecta" family="Coccinellidae" genus="Carinodula" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="742" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B91DEAFDFF65A72A05B6541111C97C8B" box="[388,482,622,640]" italics="true" pageId="3" pageNumber="742">Carinodula</emphasis>
species
</taxonomicName>
in Chiapas,
</paragraph>
<paragraph id="8BD636EFFF65A72A04A554FA10F77C93" blockId="3.[151,639,622,664]" box="[151,220,645,664]" pageId="3" pageNumber="742">Mexico.</paragraph>
</caption>
<paragraph id="8BD636EFFF65A72A049D5525113D7A55" blockId="3.[151,638,725,1730]" pageId="3" pageNumber="742">
<materialsCitation id="3B013CB2FF65A72A049D552511A17DAF" country="Mexico" location="Universidad Nacional Autonoma" pageId="3" pageNumber="742" specimenCount="2" typeStatus="holotype">
<typeStatus id="54D2884DFF65A72A049D5525112D7D64" box="[175,262,858,879]" pageId="3" pageNumber="742" type="holotype">Holotype</typeStatus>
male and
<typeStatus id="54D2884DFF65A72A055D552511917D64" box="[367,442,858,879]" pageId="3" pageNumber="742" type="allotype">allotype</typeStatus>
female deposited in the collection of the
<location id="8EB66034FF65A72A0562550A12567D81" LSID="urn:lsid:plazi:treatment:03C087F9FF64A72A06D553F814577D31:8EB66034FF65A72A0562550A12567D81" box="[336,637,885,906]" country="Mexico" name="Universidad Nacional Autonoma" pageId="3" pageNumber="742">Universidad Nacional Autónoma</location>
de
<collectingCountry id="F37E767FFF65A72A048A55F0112F7DAF" box="[184,260,911,932]" name="Mexico" pageId="3" pageNumber="742">México</collectingCountry>
, DF,
<collectingCountry id="F37E767FFF65A72A050F55F011A17DAF" box="[317,394,911,932]" name="Mexico" pageId="3" pageNumber="742">Mexico</collectingCountry>
</materialsCitation>
.
<materialsCitation id="3B013CB2FF65A72A05A555F0114E7DD2" collectionCode="MT" country="United States of America" location="University Entomology Collection" municipality="Bozeman" pageId="3" pageNumber="742" specimenCount="1" stateProvince="Montana" typeStatus="paratype">
<typeStatus id="54D2884DFF65A72A05A555F011DE7DAF" box="[407,501,911,932]" pageId="3" pageNumber="742" type="paratype">Paratypes</typeStatus>
deposited in:
<collectingRegion id="49ADF80DFF65A72A04A555D511087DB4" box="[151,291,938,959]" country="United States of America" name="Montana" pageId="3" pageNumber="742">Montana State</collectingRegion>
<location id="8EB66034FF65A72A051E55D512507DB4" LSID="urn:lsid:plazi:treatment:03C087F9FF64A72A06D553F814577D31:8EB66034FF65A72A051E55D512507DB4" box="[300,635,938,959]" country="United States of America" municipality="Bozeman" name="University Entomology Collection" pageId="3" pageNumber="742" stateProvince="Montana">University Entomology Collection</location>
,
<collectingMunicipality id="6BB2AC95FF65A72A04A555BB10DF7DD2" box="[151,244,964,985]" pageId="3" pageNumber="742">Bozeman</collectingMunicipality>
,
<collectionCode id="ED78AE2AFF65A72A053355BB110D7DD2" box="[257,294,964,985]" country="Russia" name="Mus. Tinro, Vladyvostok" pageId="3" pageNumber="742">MT</collectionCode>
,
<collectingCountry id="F37E767FFF65A72A050055BB114E7DD2" box="[306,357,964,985]" name="United States of America" pageId="3" pageNumber="742">USA</collectingCountry>
</materialsCitation>
;
<materialsCitation id="3B013CB2FF65A72A054055BB12297DFF" collectionCode="DC" country="United States of America" location="National Museum of Natural History" pageId="3" pageNumber="742" specimenCount="1" stateProvince="Washington" typeStatus="paratype">
<location id="8EB66034FF65A72A054055BB112C7DFF" LSID="urn:lsid:plazi:treatment:03C087F9FF64A72A06D553F814577D31:8EB66034FF65A72A054055BB112C7DFF" country="United States of America" name="National Museum of Natural History" pageId="3" pageNumber="742" stateProvince="Washington">National Museum of Natural History</location>
,
<collectingRegion id="49ADF80DFF65A72A052755A011A57DFF" box="[277,398,991,1012]" country="United States of America" name="Washington" pageId="3" pageNumber="742">Washington</collectingRegion>
,
<collectionCode id="ED78AE2AFF65A72A05A955A011947DFF" box="[411,447,991,1012]" pageId="3" pageNumber="742">DC</collectionCode>
,
<collectingCountry id="F37E767FFF65A72A05FC55A012297DFF" box="[462,514,991,1012]" name="United States of America" pageId="3" pageNumber="742">USA</collectingCountry>
</materialsCitation>
;
<materialsCitation id="3B013CB2FF65A72A062255A011247A22" collectorName="El Colegio de la Frontera Sur" country="Switzerland" location="San Cristobal" municipality="Casas" pageId="3" pageNumber="742" specimenCount="1" typeStatus="paratype">
<collectorName id="269C5339FF65A72A062255A0117F7A04" pageId="3" pageNumber="742">El Colegio de la Frontera Sur</collectorName>
,
<location id="8EB66034FF65A72A0550558511C67A04" LSID="urn:lsid:plazi:treatment:03C087F9FF64A72A06D553F814577D31:8EB66034FF65A72A0550558511C67A04" box="[354,493,1018,1039]" country="Switzerland" municipality="Casas" name="San Cristobal" pageId="3" pageNumber="742">San Cristóbal</location>
de las
<collectingMunicipality id="6BB2AC95FF65A72A060F558512517A04" box="[573,634,1018,1039]" pageId="3" pageNumber="742">Casas</collectingMunicipality>
,
<collectingCountry id="F37E767FFF65A72A04A5526B10917A22" box="[151,186,1044,1065]" name="Switzerland" pageId="3" pageNumber="742">CH</collectingCountry>
, Mexico
</materialsCitation>
;
<materialsCitation id="3B013CB2FF65A72A0528526B116D7A4F" country="Australia" location="Canberra" pageId="3" pageNumber="742" specimenCount="1" typeStatus="paratype">
Australian National Insect Collection,
<location id="8EB66034FF65A72A04A5525010C07A4F" LSID="urn:lsid:plazi:treatment:03C087F9FF64A72A06D553F814577D31:8EB66034FF65A72A04A5525010C07A4F" box="[151,235,1071,1092]" country="Australia" name="Canberra" pageId="3" pageNumber="742">Canberra</location>
,
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</materialsCitation>
;
<materialsCitation id="3B013CB2FF65A72A057F525011397A55" country="United Kingdom" location="London" pageId="3" pageNumber="742" specimenCount="1" typeStatus="paratype">
and The Natural History Museum,
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,
<collectingCountry id="F37E767FFF65A72A04DF523611397A55" box="[237,274,1097,1118]" name="United Kingdom" pageId="3" pageNumber="742">UK</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3736564FF65A72A049D521B11127B89" pageId="3" pageNumber="742" type="diagnosis">
<paragraph id="8BD636EFFF65A72A049D521B11127B89" blockId="3.[151,638,725,1730]" pageId="3" pageNumber="742">
<emphasis id="B91DEAFDFF65A72A049D521B113F7A72" bold="true" box="[175,276,1124,1145]" pageId="3" pageNumber="742">Diagnosis.</emphasis>
<taxonomicName id="4C694D6CFF65A72A0513521B12237A72" box="[289,520,1124,1145]" class="Insecta" family="Coccinellidae" genus="Carinodula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="742" phylum="Arthropoda" rank="species" species="chiapanensis">
<emphasis id="B91DEAFDFF65A72A0513521B12237A72" box="[289,520,1124,1145]" italics="true" pageId="3" pageNumber="742">Carinodula chiapanensis</emphasis>
</taxonomicName>
is characterized by the defined anterior face on the prosternum (
<figureCitation id="13522A6AFF65A72A04AD52E610CA7AA5" box="[159,225,1177,1198]" captionStart="Figs" captionStartId="2.[197,237,1302,1321]" captionTargetBox="[190,1174,192,1279]" captionTargetId="figure-161@2.[173,1195,187,1302]" captionTargetPageId="2" captionText="Figs. 3–9. Carinodula chiapanensis. 3) Head, dorsal view; 4) Prothorax, ventral view; 5) Spermathecal capsule;" figureDoi="http://doi.org/10.5281/zenodo.10106516" httpUri="https://zenodo.org/record/10106516/files/figure.png" pageId="3" pageNumber="742">Fig. 4</figureCitation>
), truncate postcoxal lines on ventrite I (
<figureCitation id="13522A6AFF65A72A04AD52CC10F57AC3" box="[159,222,1203,1224]" captionStart="Figs" captionStartId="2.[197,237,1302,1321]" captionTargetBox="[190,1174,192,1279]" captionTargetId="figure-161@2.[173,1195,187,1302]" captionTargetPageId="2" captionText="Figs. 3–9. Carinodula chiapanensis. 3) Head, dorsal view; 4) Prothorax, ventral view; 5) Spermathecal capsule;" figureDoi="http://doi.org/10.5281/zenodo.10106516" httpUri="https://zenodo.org/record/10106516/files/figure.png" pageId="3" pageNumber="742">Fig. 9</figureCitation>
), and the shape of the aedeagus (
<figureCitation id="13522A6AFF65A72A060A52CC125D7AC3" box="[568,630,1203,1224]" captionStart="Figs" captionStartId="2.[197,237,1302,1321]" captionTargetBox="[190,1174,192,1279]" captionTargetId="figure-161@2.[173,1195,187,1302]" captionTargetPageId="2" captionText="Figs. 3–9. Carinodula chiapanensis. 3) Head, dorsal view; 4) Prothorax, ventral view; 5) Spermathecal capsule;" figureDoi="http://doi.org/10.5281/zenodo.10106516" httpUri="https://zenodo.org/record/10106516/files/figure.png" pageId="3" pageNumber="742">Fig. 6</figureCitation>
) and spermathecal capsule (
<figureCitation id="13522A6AFF65A72A059652B111CA7AE8" box="[420,481,1230,1251]" captionStart="Figs" captionStartId="2.[197,237,1302,1321]" captionTargetBox="[190,1174,192,1279]" captionTargetId="figure-161@2.[173,1195,187,1302]" captionTargetPageId="2" captionText="Figs. 3–9. Carinodula chiapanensis. 3) Head, dorsal view; 4) Prothorax, ventral view; 5) Spermathecal capsule;" figureDoi="http://doi.org/10.5281/zenodo.10106516" httpUri="https://zenodo.org/record/10106516/files/figure.png" pageId="3" pageNumber="742">Fig. 5</figureCitation>
). The new species has an 11-segmented antenna, lacks pronotal pits, and has four tarsomeres, placing it in
<taxonomicName id="4C694D6CFF65A72A063C537C125C7B13" box="[526,631,1283,1304]" class="Insecta" family="Coccinellidae" genus="Carinodula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="742" phylum="Arthropoda" rank="genus">
<emphasis id="B91DEAFDFF65A72A063C537C125C7B13" box="[526,631,1283,1304]" italics="true" pageId="3" pageNumber="742">Carinodula</emphasis>
</taxonomicName>
, the latter two characters separating it from the other genera of the tribe. The 11-segmented antenna is shared only with the Asian
<taxonomicName id="4C694D6CFF65A72A0597532C11127B89" authority="Slipinski and Jadwiszczak." authorityName="Slipinski and Jadwiszczak." class="Insecta" family="Coccinellidae" genus="Carinodulina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="742" phylum="Arthropoda" rank="genus">
<emphasis id="B91DEAFDFF65A72A0597532C12087B63" box="[421,547,1363,1384]" italics="true" pageId="3" pageNumber="742">Carinodulina</emphasis>
Ślipiński and Jadwiszczak.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C3736564FF65A72A049D53F7135A7FCD" pageId="3" pageNumber="742" type="description">
<paragraph id="8BD636EFFF65A72A049D53F7135A7FCD" blockId="3.[151,638,725,1730]" lastBlockId="3.[685,1172,193,826]" pageId="3" pageNumber="742">
<emphasis id="B91DEAFDFF65A72A049D53F7111B7B96" bold="true" box="[175,304,1416,1437]" pageId="3" pageNumber="742">Description.</emphasis>
With the characters of the tribe and genus. Length
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. Body about 1.84–2.00X longer than wide, widest at middle (
<figureCitation id="13522A6AFF65A72A04AC53A710D17BE6" box="[158,250,1495,1517]" captionStart="Figs" captionStartId="1.[175,215,1710,1729]" captionTargetBox="[166,1159,888,1689]" captionTargetId="figure-322@1.[151,1173,873,1708]" captionTargetPageId="1" captionText="Figs. 1–2. Carinodula chiapanensis. 1) Male, dorsal view; 2) Female, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10106514" httpUri="https://zenodo.org/record/10106514/files/figure.png" pageId="3" pageNumber="742">Figs. 1–2</figureCitation>
). Color yellowish brown to light brown, appendages lighter in color. Vestiture of semidecumbent, white hairs; hairs relatively long, distinctly visible at 10X magnification. Punctures on head smaller than on pronotum, separated by more than diameter of puncture; punctures on elytron slightly larger than on pronotum, separated by diameter of puncture or slightly more. Pronotum
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long,
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wide, about 0.68–0.72X longer than wide. Elytra
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long, about 2.22–2.40X longer than pronotum. Pronotum with lateral edge moderately crenulate. Prosternum with defined anterior face and intercoxal process relatively narrow, widened apically, procoxal cavities widely open (
<figureCitation id="13522A6AFF65A72A0773573813AF7F57" box="[833,900,327,348]" captionStart="Figs" captionStartId="2.[197,237,1302,1321]" captionTargetBox="[190,1174,192,1279]" captionTargetId="figure-161@2.[173,1195,187,1302]" captionTargetPageId="2" captionText="Figs. 3–9. Carinodula chiapanensis. 3) Head, dorsal view; 4) Prothorax, ventral view; 5) Spermathecal capsule;" figureDoi="http://doi.org/10.5281/zenodo.10106516" httpUri="https://zenodo.org/record/10106516/files/figure.png" pageId="3" pageNumber="742">Fig. 4</figureCitation>
). Ventrite I with truncate postcoxal lines (
<figureCitation id="13522A6AFF65A72A077D571E13A67F7D" box="[847,909,353,374]" captionStart="Figs" captionStartId="2.[197,237,1302,1321]" captionTargetBox="[190,1174,192,1279]" captionTargetId="figure-161@2.[173,1195,187,1302]" captionTargetPageId="2" captionText="Figs. 3–9. Carinodula chiapanensis. 3) Head, dorsal view; 4) Prothorax, ventral view; 5) Spermathecal capsule;" figureDoi="http://doi.org/10.5281/zenodo.10106516" httpUri="https://zenodo.org/record/10106516/files/figure.png" pageId="3" pageNumber="742">Fig. 9</figureCitation>
). Male with median patch of setae on ventrite I (
<figureCitation id="13522A6AFF65A72A07A2570313E47F9A" box="[912,975,380,401]" captionStart="Figs" captionStartId="2.[197,237,1302,1321]" captionTargetBox="[190,1174,192,1279]" captionTargetId="figure-161@2.[173,1195,187,1302]" captionTargetPageId="2" captionText="Figs. 3–9. Carinodula chiapanensis. 3) Head, dorsal view; 4) Prothorax, ventral view; 5) Spermathecal capsule;" figureDoi="http://doi.org/10.5281/zenodo.10106516" httpUri="https://zenodo.org/record/10106516/files/figure.png" pageId="3" pageNumber="742">Fig. 9</figureCitation>
). Male aedeagus as in
<figureCitation id="13522A6AFF65A72A06F557E813297FA7" box="[711,770,407,428]" captionStart="Figs" captionStartId="2.[197,237,1302,1321]" captionTargetBox="[190,1174,192,1279]" captionTargetId="figure-161@2.[173,1195,187,1302]" captionTargetPageId="2" captionText="Figs. 3–9. Carinodula chiapanensis. 3) Head, dorsal view; 4) Prothorax, ventral view; 5) Spermathecal capsule;" figureDoi="http://doi.org/10.5281/zenodo.10106516" httpUri="https://zenodo.org/record/10106516/files/figure.png" pageId="3" pageNumber="742">Fig. 6</figureCitation>
, siphon (penis) as in
<figureCitation id="13522A6AFF65A72A07E257E814217FA7" box="[976,1034,407,428]" captionStart="Figs" captionStartId="2.[197,237,1302,1321]" captionTargetBox="[190,1174,192,1279]" captionTargetId="figure-161@2.[173,1195,187,1302]" captionTargetPageId="2" captionText="Figs. 3–9. Carinodula chiapanensis. 3) Head, dorsal view; 4) Prothorax, ventral view; 5) Spermathecal capsule;" figureDoi="http://doi.org/10.5281/zenodo.10106516" httpUri="https://zenodo.org/record/10106516/files/figure.png" pageId="3" pageNumber="742">Fig. 8</figureCitation>
. Spermathecal capsule as in
<figureCitation id="13522A6AFF65A72A070057CE13467FCD" box="[818,877,433,454]" captionStart="Figs" captionStartId="2.[197,237,1302,1321]" captionTargetBox="[190,1174,192,1279]" captionTargetId="figure-161@2.[173,1195,187,1302]" captionTargetPageId="2" captionText="Figs. 3–9. Carinodula chiapanensis. 3) Head, dorsal view; 4) Prothorax, ventral view; 5) Spermathecal capsule;" figureDoi="http://doi.org/10.5281/zenodo.10106516" httpUri="https://zenodo.org/record/10106516/files/figure.png" pageId="3" pageNumber="742">Fig. 5</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3736564FF65A72A06F757B314577D31" pageId="3" pageNumber="742" type="etymology">
<paragraph id="8BD636EFFF65A72A06F757B314AA7C1D" blockId="3.[685,1172,193,826]" pageId="3" pageNumber="742">
<emphasis id="B91DEAFDFF65A72A06F757B313137FEA" bold="true" box="[709,824,460,481]" pageId="3" pageNumber="742">Etymology.</emphasis>
The species name
<taxonomicName id="4C694D6CFF65A72A07DF57B3144F7FEA" box="[1005,1124,460,481]" class="Insecta" family="Coccinellidae" genus="Carinodula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="742" phylum="Arthropoda" rank="species" species="chiapanensis">
<emphasis id="B91DEAFDFF65A72A07DF57B3144F7FEA" box="[1005,1124,460,481]" italics="true" pageId="3" pageNumber="742">chiapanensis</emphasis>
</taxonomicName>
is an adjective derived from the name of the home mountains of the species, the Sierra Madre de
<collectingRegion id="49ADF80DFF65A72A001D547E14567C1D" box="[1071,1149,513,534]" country="Mexico" name="Chiapas" pageId="3" pageNumber="742">Chiapas</collectingRegion>
.
</paragraph>
<paragraph id="8BD636EFFF65A72A06F7546414577D31" blockId="3.[685,1172,193,826]" pageId="3" pageNumber="742">
<emphasis id="B91DEAFDFF65A72A06F7546413EC7C3B" bold="true" box="[709,967,539,560]" pageId="3" pageNumber="742">Habitat and Distribution.</emphasis>
All
<taxonomicName id="4C694D6CFF65A72A07C8546314BF7C3A" box="[1018,1172,540,561]" class="Insecta" family="Coccinellidae" genus="Carinodula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="742" phylum="Arthropoda" rank="species" species="chiapanensis">
<emphasis id="B91DEAFDFF65A72A07C8546314BF7C3A" box="[1018,1172,540,561]" italics="true" pageId="3" pageNumber="742">C. chiapanensis</emphasis>
</taxonomicName>
specimens were collected with Maxi-Winklers from sifted leaf litter at two sites located about 2–3 kilometers southeast of the small town of Custepec in the Sierra Madre de
<collectingRegion id="49ADF80DFF65A72A075854F9139E7C90" box="[874,949,646,667]" country="Mexico" name="Chiapas" pageId="3" pageNumber="742">Chiapas</collectingRegion>
(
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). The localities are at mid-elevations (
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m
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) in a combination of old-growth and second-growth wet evergreen mesophyll forest (Longino and Anderson 2014).
<taxonomicName id="4C694D6CFF65A72A06C6548F13EC7D0E" box="[756,967,752,773]" class="Insecta" family="Coccinellidae" genus="Carinodula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="742" phylum="Arthropoda" rank="species" species="campbelli">
<emphasis id="B91DEAFDFF65A72A06C6548F13EC7D0E" box="[756,967,752,773]" italics="true" pageId="3" pageNumber="742">Carinodula campbelli</emphasis>
</taxonomicName>
was collected in the Central
<collectingRegion id="49ADF80DFF65A72A06CF557513617D14" box="[765,842,778,799]" country="Mexico" name="Chiapas" pageId="3" pageNumber="742">Chiapas</collectingRegion>
Highlands (
<figureCitation id="13522A6AFF65A72A07F7557514267D14" box="[965,1037,778,799]" captionStart="Fig" captionStartId="3.[175,207,622,641]" captionTargetBox="[156,633,194,601]" captionTargetId="figure-698@3.[151,638,189,606]" captionTargetPageId="3" captionText="Fig. 10. Distribution of Carinodula species in Chiapas, Mexico." figureDoi="http://doi.org/10.5281/zenodo.10106518" httpUri="https://zenodo.org/record/10106518/files/figure.png" pageId="3" pageNumber="742">Fig. 10</figureCitation>
) north of the Valle Central, which separates the two localities.
</paragraph>
</subSubSection>
</treatment>
</document>
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