218 lines
27 KiB
XML
218 lines
27 KiB
XML
<document id="2C1A11577A4B424F2114B80DC7F6C194" ID-DOI="10.5281/zenodo.282919" ID-GBIF-Dataset="09d06f43-47e0-4f67-8ed9-54c16248c2c4" ID-ISSN="1175-5326" ID-Zenodo-Dep="282919" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460556013358" checkinUser="plazi" docAuthor="Kraus, Fred" docDate="2012" docId="03C087ACFF8BFF91AB84F8A3FEE5CFF7" docLanguage="en" docName="zt03559p036.pdf" docOrigin="Zootaxa 3559" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Cophixalus pictus Kraus, 2012, sp. nov." docType="treatment" docVersion="7" lastPageNumber="12" masterDocId="FFF9FFD4FF82FF9AAB13FF9EFFF6C860" masterDocTitle="Papuan frogs of the genus Cophixalus (Anura: Microhylidae): new synonyms, new species, and a dichotomous key" masterLastPageNumber="36" masterPageNumber="1" pageNumber="10" updateTime="1698319243426" updateUser="plazi">
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<mods:title id="ADF91BF18A319F1F400DFD9C2E6DA979">Papuan frogs of the genus Cophixalus (Anura: Microhylidae): new synonyms, new species, and a dichotomous key</mods:title>
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<mods:namePart id="4F4E9A0C60ACECC9DB2EDE3437E5C687">Kraus, Fred</mods:namePart>
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<mods:title id="9C14AD85476AC30B98B569C71EF2A95D">Zootaxa</mods:title>
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<mods:part id="26D0517AA039875ED2DEC25293657710">
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<mods:date id="EE92350A7B1931F08F3543BA38DA193D">2012</mods:date>
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<mods:number id="5FB64DD1972328F823C7B9C799D5C9CC">3559</mods:number>
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<mods:identifier id="9E5F8E1C14B0B1BABFC1397CE3B742EE" type="DOI">10.5281/zenodo.282919</mods:identifier>
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<mods:identifier id="C2D669CB81F03F60A417502842FDBA01" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03C087ACFF8BFF91AB84F8A3FEE5CFF7" ID-DOI="http://doi.org/10.5281/zenodo.6180274" ID-GBIF-Taxon="119555805" ID-Zenodo-Dep="6180274" LSID="urn:lsid:plazi:treatment:03C087ACFF8BFF91AB84F8A3FEE5CFF7" httpUri="http://treatment.plazi.org/id/03C087ACFF8BFF91AB84F8A3FEE5CFF7" lastPageId="11" lastPageNumber="12" pageId="9" pageNumber="10">
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<subSubSection id="C3736531FF8BFF93AB84F8A3FF2ECF18" pageId="9" pageNumber="10" type="nomenclature">
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<paragraph id="8BD636BAFF8BFF93AB84F8A3FE2ACF37" blockId="9.[151,476,1853,1912]" box="[151,476,1853,1879]" pageId="9" pageNumber="10">
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<heading id="D09E81D6FF8BFF93AB84F8A3FE2ACF37" bold="true" box="[151,476,1853,1879]" fontSize="11" level="1" pageId="9" pageNumber="10" reason="1">
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||
<emphasis id="B91DEAA8FF8BFF93AB84F8A3FE2ACF37" bold="true" box="[151,476,1853,1879]" pageId="9" pageNumber="10">
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<taxonomicName id="4C694D39FF8BFF93AB84F8A3FE86CF37" ID-CoL="Y44Z" box="[151,368,1853,1879]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="pictus" status="sp. nov.">
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<emphasis id="B91DEAA8FF8BFF93AB84F8A3FE86CF37" bold="true" box="[151,368,1853,1879]" italics="true" pageId="9" pageNumber="10">Cophixalus pictus</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A22E57D3FF8BFF93AA6CF8A3FE2ACF37" box="[383,476,1853,1879]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BD636BAFF8BFF93AB84F8FEFF2ECF18" blockId="9.[151,476,1853,1912]" box="[151,216,1888,1912]" pageId="9" pageNumber="10">
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<figureCitation id="13522A3FFF8BFF93AB84F8FEFF2ECF18" box="[151,216,1888,1912]" captionStart="FIGURE 5" captionStartId="10.[151,250,1988,2011]" captionTargetBox="[159,1417,185,1945]" captionTargetId="figure@10.[151,1436,167,1965]" captionTargetPageId="10" captionText="FIGURE 5. (A) Dorsal, and (B) lateral views of holotype (BPBM 1187) of Cophixalus pictus sp. nov. Scale bars are 5 mm." httpUri="https://zenodo.org/record/282924/files/figure.png" pageId="9" pageNumber="10">Fig. 5</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3736531FF8BFF93AB84F837FDE4CF84" pageId="9" pageNumber="10" type="materials_examined">
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<paragraph id="8BD636BAFF8BFF93AB84F837FDE4CF84" blockId="9.[151,1435,1960,2020]" pageId="9" pageNumber="10">
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<typeStatus id="54D28818FF8BFF93AB84F837FEF2CFA1" box="[151,260,1961,1985]" pageId="9" pageNumber="10" type="holotype">
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<emphasis id="B91DEAA8FF8BFF93AB84F837FEF2CFA1" bold="true" box="[151,260,1961,1985]" pageId="9" pageNumber="10">Holotype</emphasis>
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</typeStatus>
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.
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<collectionCode id="ED78AE7FFF8BFF93AA00F837FE97CFA0" LSID="urn:lsid:biocol.org:col:34697" box="[275,353,1961,1984]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="9" pageNumber="10">BPBM</collectionCode>
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1187, collected by J.L. Gressitt, Bomberai,
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<geoCoordinate id="EE5D507DFF8BFF93A844F836FC4ECFA0" box="[855,952,1960,1984]" direction="south" orientation="latitude" pageId="9" pageNumber="10" precision="55" value="-2.766">2.766º S</geoCoordinate>
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,
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<geoCoordinate id="EE5D507DFF8BFF93A8D5F836FBB3CFA0" box="[966,1093,1960,1984]" direction="east" orientation="longitude" pageId="9" pageNumber="10" precision="55" value="132.788">132.788º E</geoCoordinate>
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, Fakfak Division, West
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<collectingCountry id="F37E762AFF8BFF93AE44F837FA6DCFA0" box="[1367,1435,1961,1984]" name="Papua New Guinea" pageId="9" pageNumber="10">Papua</collectingCountry>
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Province,
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<collectingCountry id="F37E762AFF8BFF93AA1BF852FE8ECF84" box="[264,376,1996,2020]" name="Indonesia" pageId="9" pageNumber="10">Indonesia</collectingCountry>
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,
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<date id="FFD7107AFF8BFF93AA90F852FDF8CF84" box="[387,526,1996,2020]" pageId="9" pageNumber="10" value="1959-06-03">3 June 1959</date>
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.
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</paragraph>
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</subSubSection>
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<caption id="DF166632FF88FF90AB84F85AFA8BCFBA" box="[151,1405,1988,2011]" httpUri="https://zenodo.org/record/282924/files/figure.png" pageId="10" pageNumber="11" targetBox="[159,1417,185,1945]" targetPageId="10">
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<paragraph id="8BD636BAFF88FF90AB84F85AFA8BCFBA" blockId="10.[151,1405,1988,2011]" box="[151,1405,1988,2011]" pageId="10" pageNumber="11">
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<emphasis id="B91DEAA8FF88FF90AB84F85AFEE4CFBA" bold="true" box="[151,274,1988,2011]" pageId="10" pageNumber="11">FIGURE 5.</emphasis>
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(A) Dorsal, and (B) lateral views of holotype (BPBM 1187) of
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<taxonomicName id="4C694D39FF88FF90A88BF85BFBBBCFBB" box="[920,1101,1989,2011]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="pictus" status="sp. nov.">
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<emphasis id="B91DEAA8FF88FF90A88BF85BFBBBCFBB" box="[920,1101,1989,2011]" italics="true" pageId="10" pageNumber="11">Cophixalus pictus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22E57D3FF88FF90AF47F85BFB55CFBA" box="[1108,1187,1989,2010]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
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Scale bars are 5 mm.
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</paragraph>
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</caption>
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<subSubSection id="C3736531FF89FF91ABD4FF09FD26CB1F" pageId="11" pageNumber="12" type="diagnosis">
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<paragraph id="8BD636BAFF89FF91ABD4FF09FF02C904" blockId="11.[151,1436,151,1943]" pageId="11" pageNumber="12">
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<emphasis id="B91DEAA8FF89FF91ABD4FF09FEB2C8D0" bold="true" box="[199,324,151,176]" pageId="11" pageNumber="12">Diagnosis.</emphasis>
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A species characterized by its unique combination of small size (SV =
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<quantity id="4C919B5FFF89FF91AF8DFF09FAFEC8CF" box="[1182,1288,151,175]" metricMagnitude="-2" metricUnit="m" metricValue="1.5699999999999998" pageId="11" pageNumber="12" unit="mm" value="15.7">15.7 mm</quantity>
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) with finger discs larger than toe discs (3rdF/4thT = 1.30); first finger of normal size, with disc that lacks a circum-marginal groove; moderately large finger discs (3rdF/SV = 0.046); short legs (TL/SV = 0.46); short and narrow snout (EN/SV = 0.089, SN/EY = 0.88, EN/IN = 1.08); hidden tympanum; large eye (EY/SV = 0.15); face dark brown posteriorly, tan with dark-brown flecks anteriorly; boldly blotched with dark-brown spots laterally and ventrally (
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<figureCitation id="13522A3FFF89FF91AB8CFED2FF13C904" box="[159,229,332,356]" captionStart="FIGURE 5" captionStartId="10.[151,250,1988,2011]" captionTargetBox="[159,1417,185,1945]" captionTargetId="figure@10.[151,1436,167,1965]" captionTargetPageId="10" captionText="FIGURE 5. (A) Dorsal, and (B) lateral views of holotype (BPBM 1187) of Cophixalus pictus sp. nov. Scale bars are 5 mm." httpUri="https://zenodo.org/record/282924/files/figure.png" pageId="11" pageNumber="12">Fig. 5</figureCitation>
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).
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</paragraph>
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<paragraph id="8BD636BAFF89FF91ABD4FEF1FD26CB1F" blockId="11.[151,1436,151,1943]" pageId="11" pageNumber="12">
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<emphasis id="B91DEAA8FF89FF91ABD4FEF1FD80C9E8" bold="true" box="[199,630,367,392]" pageId="11" pageNumber="12">Comparisons with other species.</emphasis>
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The new species differs from all Papuan congeners except
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<taxonomicName id="4C694D39FF89FF91AE93FEEEFE4BC9CC" authority="Zweifel, 1956" authorityName="Zweifel" authorityYear="1956" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="daymani">
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<emphasis id="B91DEAA8FF89FF91AE93FEEEFF08C9CC" italics="true" pageId="11" pageNumber="12">C. daymani</emphasis>
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<bibRefCitation id="EFF84B4BFF89FF91AA16FE0AFE4BC9CC" author="Zweifel" box="[261,445,404,428]" pageId="11" pageNumber="12" refString="Zweifel, R. G. (1956 b) Results of the Archbold Expeditions. No. 72. Microhylid frogs from New Guinea, with descriptions of new species. American Museum Novitates, 1766, 1 - 49." type="journal article" year="1956" yearSuffix="b">Zweifel, 1956b</bibRefCitation>
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</taxonomicName>
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and
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<taxonomicName id="4C694D39FF89FF91AAEBFE0BFD9FC9CC" box="[504,617,404,428]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="kethuk">
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<emphasis id="B91DEAA8FF89FF91AAEBFE0BFD9FC9CC" box="[504,617,404,428]" italics="true" pageId="11" pageNumber="12">C. kethuk</emphasis>
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</taxonomicName>
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in having the snout shorter than the eye. It differs from
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<taxonomicName id="4C694D39FF89FF91AE01FE0BFA6AC9CC" box="[1298,1436,404,428]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="daymani">
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<emphasis id="B91DEAA8FF89FF91AE01FE0BFA6AC9CC" box="[1298,1436,404,428]" italics="true" pageId="11" pageNumber="12">C. daymani</emphasis>
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</taxonomicName>
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in its narrower snout (IN/SV = 0.90–0.108, EN/IN =
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<quantity id="4C919B5FFF89FF91A805FE29FC5BC9AF" box="[790,941,439,463]" metricMagnitude="-2" metricUnit="m" metricValue="2.1082" metricValueMax="2.2352" metricValueMin="1.9812" pageId="11" pageNumber="12" unit="in" value="0.83" valueMax="0.88" valueMin="0.78">0.78–0.88 in</quantity>
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<taxonomicName id="4C694D39FF89FF91A8A6FE26FBC8C9B0" box="[949,1086,440,464]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="daymani">
|
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<emphasis id="B91DEAA8FF89FF91A8A6FE26FBC8C9B0" box="[949,1086,440,464]" italics="true" pageId="11" pageNumber="12">C. daymani</emphasis>
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</taxonomicName>
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); it differs from
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<taxonomicName id="4C694D39FF89FF91AE18FE26FA8AC9B0" box="[1291,1404,440,464]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="kethuk">
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<emphasis id="B91DEAA8FF89FF91AE18FE26FA8AC9B0" box="[1291,1404,440,464]" italics="true" pageId="11" pageNumber="12">C. kethuk</emphasis>
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</taxonomicName>
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in lacking (vs. having in
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<taxonomicName id="4C694D39FF89FF91AAA7FE43FDDCC994" box="[436,554,476,500]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="kethuk">
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||
<emphasis id="B91DEAA8FF89FF91AAA7FE43FDDCC994" box="[436,554,476,500]" italics="true" pageId="11" pageNumber="12">C. kethuk</emphasis>
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</taxonomicName>
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) basal webbing on the toes, having a normal-sized (vs. reduced in
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<taxonomicName id="4C694D39FF89FF91AE93FE43FF13CA78" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="kethuk">
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<emphasis id="B91DEAA8FF89FF91AE93FE43FF13CA78" italics="true" pageId="11" pageNumber="12">C. kethuk</emphasis>
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</taxonomicName>
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) first finger with (vs. lacking in
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<taxonomicName id="4C694D39FF89FF91A964FD9EFD1CCA78" box="[631,746,512,536]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="kethuk">
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<emphasis id="B91DEAA8FF89FF91A964FD9EFD1CCA78" box="[631,746,512,536]" italics="true" pageId="11" pageNumber="12">C. kethuk</emphasis>
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</taxonomicName>
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) a disc, and being blotched dorsally (vs. having a dorsal hourglass pattern in
|
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<taxonomicName id="4C694D39FF89FF91AA8BFDBBFDFBCA5C" box="[408,525,548,572]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="kethuk">
|
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<emphasis id="B91DEAA8FF89FF91AA8BFDBBFDFBCA5C" box="[408,525,548,572]" italics="true" pageId="11" pageNumber="12">C. kethuk</emphasis>
|
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</taxonomicName>
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).
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<taxonomicName id="4C694D39FF89FF91A93FFDBAFCF1CA5C" box="[556,775,548,572]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="pictus">
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<emphasis id="B91DEAA8FF89FF91A93FFDBAFCF1CA5C" box="[556,775,548,572]" italics="true" pageId="11" pageNumber="12">Cophixalus pictus</emphasis>
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||
</taxonomicName>
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also differs from all Papuan congeners in its boldly blotched lateral and ventral pattern.
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<taxonomicName id="4C694D39FF89FF91A972FDD6FB58CA3F" authority="Kraus and Allison, 2000" authorityName="Kraus and Allison" authorityYear="2000" box="[609,1198,583,608]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="pulchellus">
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<emphasis id="B91DEAA8FF89FF91A972FDD6FC86CA00" box="[609,880,584,608]" italics="true" pageId="11" pageNumber="12">Cophixalus pulchellus</emphasis>
|
||
<bibRefCitation id="EFF84B4BFF89FF91A86EFDD6FB58CA3F" author="Kraus" box="[893,1198,583,607]" pageId="11" pageNumber="12" refString="Kraus, F. & Allison, A. (2000) Two new species of Cophixalus from New Guinea. Journal of Herpetology, 34, 535 - 541." type="journal article" year="2000">Kraus and Allison, 2000</bibRefCitation>
|
||
</taxonomicName>
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||
is boldly blotched dorsally with black on silver-gray, but has a dark venter and disc of third finger smaller than that of fourth toe (3rdF/4thT =
|
||
<quantity id="4C919B5FFF89FF91AA78FD11FE36CAC7" box="[363,448,655,679]" metricMagnitude="-2" metricUnit="m" metricValue="2.3876" pageId="11" pageNumber="12" unit="in" value="0.94">0.94 in</quantity>
|
||
<taxonomicName id="4C694D39FF89FF91AADAFD0EFD9ECAC8" box="[457,616,656,680]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="pulchellus">
|
||
<emphasis id="B91DEAA8FF89FF91AADAFD0EFD9ECAC8" box="[457,616,656,680]" italics="true" pageId="11" pageNumber="12">C. pulchellus</emphasis>
|
||
</taxonomicName>
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||
);
|
||
<taxonomicName id="4C694D39FF89FF91A997FD0EFCCDCAC8" box="[644,827,656,680]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="bewaniensis">
|
||
<emphasis id="B91DEAA8FF89FF91A997FD0EFCCDCAC8" box="[644,827,656,680]" italics="true" pageId="11" pageNumber="12">C. bewaniensis</emphasis>
|
||
</taxonomicName>
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has a single large dark-brown blotch laterally but has the venter flecked with brown and has a reduced first finger lacking a disc. It is uncertain if the hidden tympanum of the
|
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<typeStatus id="54D28818FF89FF91AA7DFD49FE21CA8F" box="[366,471,727,751]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
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of
|
||
<taxonomicName id="4C694D39FF89FF91A911FD46FD98CA90" box="[514,622,728,752]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="pictus">
|
||
<emphasis id="B91DEAA8FF89FF91A911FD46FD98CA90" box="[514,622,728,752]" italics="true" pageId="11" pageNumber="12">C. pictus</emphasis>
|
||
</taxonomicName>
|
||
is a preservation artifact, but this seems unlikely. If it is a reliable feature, then this character also serves to distinguish
|
||
<taxonomicName id="4C694D39FF89FF91A856FD63FC41CB74" box="[837,951,765,788]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="pictus">
|
||
<emphasis id="B91DEAA8FF89FF91A856FD63FC41CB74" box="[837,951,765,788]" italics="true" pageId="11" pageNumber="12">C. pictus</emphasis>
|
||
</taxonomicName>
|
||
from all Papuan congeners except
|
||
<taxonomicName id="4C694D39FF89FF91AE93FD63FDD3CB57" authority="Zweifel, 1956" authorityName="Zweifel" authorityYear="1956" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="cryptotympanum">
|
||
<emphasis id="B91DEAA8FF89FF91AE93FD63FEA9CB58" italics="true" pageId="11" pageNumber="12">C. cryptotympanum</emphasis>
|
||
<bibRefCitation id="EFF84B4BFF89FF91AA7AFC81FDD3CB57" author="Zweifel" box="[361,549,799,823]" pageId="11" pageNumber="12" refString="Zweifel, R. G. (1956 b) Results of the Archbold Expeditions. No. 72. Microhylid frogs from New Guinea, with descriptions of new species. American Museum Novitates, 1766, 1 - 49." type="journal article" year="1956" yearSuffix="b">Zweifel, 1956b</bibRefCitation>
|
||
</taxonomicName>
|
||
, which, in addition to the characters mentioned above, differs in having longer legs (TL/SV =
|
||
<quantity id="4C919B5FFF89FF91AAA9FCDAFDAECB3C" box="[442,600,836,860]" metricMagnitude="-2" metricUnit="m" metricValue="1.2954" metricValueMax="1.3208" metricValueMin="1.27" pageId="11" pageNumber="12" unit="in" value="0.51" valueMax="0.52" valueMin="0.5">0.50–0.52 in</quantity>
|
||
<taxonomicName id="4C694D39FF89FF91A975FCDBFCAECB3C" box="[614,856,837,860]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="cryptotympanum">
|
||
<emphasis id="B91DEAA8FF89FF91A975FCDBFCAECB3C" box="[614,856,837,860]" italics="true" pageId="11" pageNumber="12">C. cryptotympanum</emphasis>
|
||
</taxonomicName>
|
||
), a broader snout (EN/IN =
|
||
<quantity id="4C919B5FFF89FF91AFC7FCDAFA84CB3C" box="[1236,1394,836,860]" metricMagnitude="-2" metricUnit="m" metricValue="2.1717" metricValueMax="2.3114" metricValueMin="2.032" pageId="11" pageNumber="12" unit="in" value="0.855" valueMax="0.91" valueMin="0.8">0.80–0.91 in</quantity>
|
||
<taxonomicName id="4C694D39FF89FF91AE93FCDBFEA9CBE0" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="cryptotympanum">
|
||
<emphasis id="B91DEAA8FF89FF91AE93FCDBFEA9CBE0" italics="true" pageId="11" pageNumber="12">C. cryptotympanum</emphasis>
|
||
</taxonomicName>
|
||
), and a light postocular stripe.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3736531FF89FF91ABD4FC15FED4CE82" pageId="11" pageNumber="12" type="description">
|
||
<paragraph id="8BD636BAFF89FF91ABD4FC15FC8FCDA0" blockId="11.[151,1436,151,1943]" pageId="11" pageNumber="12">
|
||
<emphasis id="B91DEAA8FF89FF91ABD4FC15FE07CBC4" bold="true" box="[199,497,907,932]" pageId="11" pageNumber="12">
|
||
Description of
|
||
<typeStatus id="54D28818FF89FF91AA97FC12FE07CBC4" box="[388,497,908,932]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
. Sex uncertain because of state of preservation of internal features, but likely a juvenile male; eggs and enlarged oviducts clearly absent. Head moderately wide (HW/SV = 0.43), with steeply oblique loreal region and inflated upper lip; canthus rostralis rounded, straight when viewed from above; nostrils directed laterally, closer to tip of snout than to eyes; internarial distance narrower than distance from naris to eye (EN/IN = 1.08, IN/SV = 0.083, EN/SV = 0.089); snout slightly rounded when viewed from the side, rounded when viewed from above; eyes large (EY/SV = 0.15); eyelid approximately same width as interorbital distance; tympanum hidden. Dorsal, lateral, and ventral surfaces smooth; supratympanic fold absent. Fingers unwebbed, all bearing well-developed discs; relative lengths 3>4>2>1. Discs of F3 and F4 with poorly developed circum-marginal grooves, approximately 2 times widths of penultimate phalanges (3rdF/SV = 0.046); discs of F1 and F2 lacking such grooves, that of F2 approximately 1.5 times width of penultimate phalanx, that of F1 barely wider than penultimate phalanx. Subarticular tubercles poorly developed, metacarpal tubercles not evident. Toes unwebbed, bearing discs, with terminal grooves on T2–T5; relative lengths 4>3>5>2>1. Toe discs smaller than those of fingers (3rdF/4thT = 1.30); disc of fourth toe approximately 1.5 times as wide as penultimate phalanx (4thT/SV = 0.036); disc of first toe approximately same width as penultimate phalanx. Subarticular and metatarsal tubercles not obvious.
|
||
<collectingCountry id="F37E762AFF89FF91AABBFA36FE15CDA0" box="[424,483,1448,1472]" name="India" pageId="11" pageNumber="12">Hind</collectingCountry>
|
||
legs rather short (TL/SV = 0.46).
|
||
</paragraph>
|
||
<paragraph id="8BD636BAFF89FF91ABD4FA53FF1ECEF7" blockId="11.[151,1436,151,1943]" pageId="11" pageNumber="12">
|
||
Dorsum and upper sides brown, lighter lower on sides, with few dark-brown flecks on head and shoulders; sides with numerous dark-brown blotches, larger and more rounded ventrally (
|
||
<figureCitation id="13522A3FFF89FF91AF39FA71FB84CE67" box="[1066,1138,1519,1543]" captionStart="FIGURE 5" captionStartId="10.[151,250,1988,2011]" captionTargetBox="[159,1417,185,1945]" captionTargetId="figure@10.[151,1436,167,1965]" captionTargetPageId="10" captionText="FIGURE 5. (A) Dorsal, and (B) lateral views of holotype (BPBM 1187) of Cophixalus pictus sp. nov. Scale bars are 5 mm." httpUri="https://zenodo.org/record/282924/files/figure.png" pageId="11" pageNumber="12">Fig. 5</figureCitation>
|
||
). Front and rear of thighs brown with large dark-brown blotches. Posterior of face dark brown with few tiny tan flecks; snout tan with few dark-brown flecks (
|
||
<figureCitation id="13522A3FFF89FF91AAA1F9A9FE0CCE2F" box="[434,506,1591,1615]" captionStart="FIGURE 5" captionStartId="10.[151,250,1988,2011]" captionTargetBox="[159,1417,185,1945]" captionTargetId="figure@10.[151,1436,167,1965]" captionTargetPageId="10" captionText="FIGURE 5. (A) Dorsal, and (B) lateral views of holotype (BPBM 1187) of Cophixalus pictus sp. nov. Scale bars are 5 mm." httpUri="https://zenodo.org/record/282924/files/figure.png" pageId="11" pageNumber="12">Fig. 5</figureCitation>
|
||
).
|
||
<collectingCountry id="F37E762AFF89FF91A900F9A9FDB8CE2F" box="[531,590,1591,1615]" name="China" pageId="11" pageNumber="12">Chin</collectingCountry>
|
||
, throat, and chest dark brown with few tiny tan flecks, mostly near jaw margins; symphysis tan. Abdomen tan with dark-brown blotches and minutely stippled with black. Iris dark brown.
|
||
</paragraph>
|
||
<paragraph id="8BD636BAFF89FF91ABD4F93BFED4CE82" blockId="11.[151,1436,151,1943]" pageId="11" pageNumber="12">
|
||
<emphasis id="B91DEAA8FF89FF91ABD4F93BFE38CEDC" box="[199,462,1700,1724]" italics="true" pageId="11" pageNumber="12">Measurements (in mm).</emphasis>
|
||
—SV = 15.7, TL = 7.3, HW = 6.8, IN = 1.3, EN = 1.4, SN = 2.1, EY = 2.4, 3rdF = 0.73, 4thT = 0.56.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3736531FF89FF91ABD4F971FEE5CFF7" pageId="11" pageNumber="12" type="etymology">
|
||
<paragraph id="8BD636BAFF89FF91ABD4F971FDC5CF4A" blockId="11.[151,1436,151,1943]" pageId="11" pageNumber="12">
|
||
<emphasis id="B91DEAA8FF89FF91ABD4F971FEB8CF67" bold="true" box="[199,334,1775,1799]" pageId="11" pageNumber="12">Etymology.</emphasis>
|
||
The name is a masculine Latin adjective meaning “painted” and is in reference to the boldly blotched color pattern of the species.
|
||
</paragraph>
|
||
<paragraph id="8BD636BAFF89FF91ABD4F8A9FEE5CFF7" blockId="11.[151,1436,151,1943]" pageId="11" pageNumber="12">
|
||
<emphasis id="B91DEAA8FF89FF91ABD4F8A9FEE9CF2F" bold="true" box="[199,287,1847,1871]" pageId="11" pageNumber="12">Range.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="54D28818FF89FF91A927F8A6FD9ECF30" box="[564,616,1848,1872]" pageId="11" pageNumber="12">type</typeStatus>
|
||
locality in western New
|
||
<collectingCountry id="F37E762AFF89FF91A88BF8A9FC18CF2F" box="[920,1006,1847,1871]" name="Guinea-Bissau" pageId="11" pageNumber="12">Guinea</collectingCountry>
|
||
(
|
||
<figureCitation id="13522A3FFF89FF91AF13F8A9FBBDCF2F" box="[1024,1099,1847,1871]" captionStart="FIGURE 6" captionStartId="12.[151,250,1546,1569]" captionTargetBox="[237,1343,182,1540]" captionTargetId="figure@12.[237,1350,173,1541]" captionTargetPageId="12" captionText="FIGURE 6. Map of western New Guinea and adjacent islands showing type locality for Cophixalus pictus sp. nov. along coast of Bomberai Peninsula (filled star). Elevational contours are at 1000 m (light gray) and 1500 m (dark gray)." httpUri="https://zenodo.org/record/282925/files/figure.png" pageId="11" pageNumber="12">Fig. 6</figureCitation>
|
||
). The coordinates are along the coast, and the entire peninsula is of low elevation, so presumably this species inhabits lowland rainforest.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |