treatments-xml/data/03/C0/87/03C087AC8723BF3AFF61F942FC62FE97.xml
2024-06-21 12:22:17 +02:00

463 lines
62 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="00DE365F361D604B30A330615CFEA666" ID-CLB-Dataset="5386" ID-DOI="10.1111/j.1096-3642.2008.00441.x" ID-GBIF-Dataset="7d64f10b-205f-4051-bdae-bfbc9f432201" ID-ISSN="0024-4082" ID-Zenodo-Dep="4634967" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1616618902228" checkinUser="carolina" docAuthor="Schubart, Christoph D. &amp; Ng, Peter K. L." docDate="2008" docId="03C087AC8723BF3AFF61F942FC62FE97" docLanguage="en" docName="ZoolJLinnSoc.154.211-221.pdf" docOrigin="Zoological Journal of the Linnean Society 154 (2)" docSource="https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2008.00441.x" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.2:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="2" docTitle="Sundathelphusa MOLLUSCIVORA 2008, SP. NOV." docType="treatment" docVersion="7" lastPageNumber="218" masterDocId="FFF9FFD48720BF3DFFB2FFB6FF80FFC1" masterDocTitle="A new molluscivore crab from Lake Poso confirms multiple colonization of ancient lakes in Sulawesi by freshwater crabs (Decapoda: Brachyura)" masterLastPageNumber="221" masterPageNumber="211" pageNumber="214" updateTime="1698934849678" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods id="4BF79C0CABB940A045BE1838542083DA" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="7098332BDB8C993B819F1F305A1E8186">
<mods:title id="C61D0EBC17ADB20F022B94A906663A21">A new molluscivore crab from Lake Poso confirms multiple colonization of ancient lakes in Sulawesi by freshwater crabs (Decapoda: Brachyura)</mods:title>
</mods:titleInfo>
<mods:name id="1AF05F3D477B24AF2F727A596F8A0D42" type="personal">
<mods:role id="1E6C8CF41F90C0C9E90DF9AE89EF276A">
<mods:roleTerm id="99868899023E83E7B40C0B04F7867BE9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="188EE7E8FCDD0ABA26457A5D6876EA10">Schubart, Christoph D.</mods:namePart>
</mods:name>
<mods:name id="E786D1CDC7899A7CBF4D876BF207C364" type="personal">
<mods:role id="29EAB444063061507A68792C6E372CC3">
<mods:roleTerm id="65DD3C164B02D49BA82D9F510BBC60AB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="12180D0583A024B609C3663CB36DAD1B">Ng, Peter K. L.</mods:namePart>
</mods:name>
<mods:typeOfResource id="951E47298BAA9589BAD9545ACD3E23BA">text</mods:typeOfResource>
<mods:relatedItem id="AB7DFC7298762BD932D302043C655737" type="host">
<mods:titleInfo id="5A2557A4131C876888C35F4F1F422D15">
<mods:title id="69275C3F3F2DF22B312DE563FD1EBF5A">Zoological Journal of the Linnean Society</mods:title>
</mods:titleInfo>
<mods:part id="E260D520BD9CA5399FA9A33F90EA85C6">
<mods:date id="36D53DD294D5043091E370144D828292">2008</mods:date>
<mods:detail id="7CD930E3E7FB18324C8FEF860D26FAE1" type="pubDate">
<mods:number id="B0352BA6F0BDE3CBBE517E8A1B305759">2008-10-31</mods:number>
</mods:detail>
<mods:detail id="FCEB463E205141464FA991B5EC2C2592" type="volume">
<mods:number id="E455D4DD8575DD532268882865E2D74A">154</mods:number>
</mods:detail>
<mods:detail id="7BAEC58BB68B8801A64178688DFEC8A6" type="issue">
<mods:number id="1A4DF3EC882A8669B36C06E7D3584AC8">2</mods:number>
</mods:detail>
<mods:extent id="8D1C9ACCC9CAC79ACF87E7F3EB0BECA0" unit="page">
<mods:start id="4C1781701B4BDF90DC1A495867E7FFB6">211</mods:start>
<mods:end id="3DDE80F3AF977354316001FAC7E0634E">221</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="4C9C8429798B518E13561010B090F42E">
<mods:url id="6EDA32CFD1EFCB1C821FA0EF78EC9BDF">https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2008.00441.x</mods:url>
</mods:location>
<mods:classification id="CDFA0F101E32FF8CA23C9772EBB5DE3A">journal article</mods:classification>
<mods:identifier id="636EEA92611EB9715C37486F635DF087" type="CLB-Dataset">5386</mods:identifier>
<mods:identifier id="BCF63E9DE7C727025EADEEDFFD57D999" type="DOI">10.1111/j.1096-3642.2008.00441.x</mods:identifier>
<mods:identifier id="9760210D766D338660E22B85A80BD69B" type="GBIF-Dataset">7d64f10b-205f-4051-bdae-bfbc9f432201</mods:identifier>
<mods:identifier id="A6AB46287207865B5F4DBE523BF0447D" type="ISSN">0024-4082</mods:identifier>
<mods:identifier id="3181BD7F84AE34B78EBA89D1A57B9A38" type="Zenodo-Dep">4634967</mods:identifier>
</mods:mods>
<treatment id="03C087AC8723BF3AFF61F942FC62FE97" ID-DOI="http://doi.org/10.5281/zenodo.5114357" ID-GBIF-Taxon="183631401" ID-Zenodo-Dep="5114357" LSID="urn:lsid:plazi:treatment:03C087AC8723BF3AFF61F942FC62FE97" httpUri="http://treatment.plazi.org/id/03C087AC8723BF3AFF61F942FC62FE97" lastPageId="7" lastPageNumber="218" pageId="3" pageNumber="214">
<subSubSection id="C37365318723BF3EFF61F942FD5CF8C8" box="[211,732,1780,1801]" pageId="3" pageNumber="214" type="nomenclature">
<paragraph id="8BD636BA8723BF3EFF61F942FD5CF8C8" blockId="3.[211,732,1780,1834]" box="[211,732,1780,1801]" pageId="3" pageNumber="214">
<heading id="D09E81D68723BF3EFF61F942FD5CF8C8" box="[211,732,1780,1801]" centered="true" fontSize="9" level="2" pageId="3" pageNumber="214" reason="3">
<taxonomicName id="4C694D398723BF3EFF61F942FDEDF8C9" ID-CoL="7Q3CF" authority="MOLLUSCIVORA" authorityName="MOLLUSCIVORA" authorityYear="2008" box="[211,621,1780,1801]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="214" phylum="Arthropoda" rank="genus" status="sp. nov.">SUNDATHELPHUSA MOLLUSCIVORA</taxonomicName>
<emphasis id="B91DEAA88723BF3EFDC5F941FD5CF8C8" bold="true" box="[631,732,1780,1801]" pageId="3" pageNumber="214">
<taxonomicNameLabel id="A22E57D38723BF3EFDC5F941FD5CF8C8" box="[631,732,1780,1801]" pageId="3" pageNumber="214" rank="species">SP. NOV.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37365318723BF3AFE2CF8A2FC62FE97" lastPageId="7" lastPageNumber="218" pageId="3" pageNumber="214" type="description">
<paragraph id="8BD636BA8723BF3EFE2CF8A2FD90F8EB" blockId="3.[211,732,1780,1834]" box="[414,528,1812,1834]" pageId="3" pageNumber="214">
(
<figureCitation id="13522A3F8723BF3EFE15F8A2FD88F8EB" box="[423,520,1812,1834]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStart-3="Figure 4" captionStartId-0="3.[826,905,1207,1226]" captionStartId-1="4.[144,223,1284,1303]" captionStartId-2="5.[164,243,1417,1436]" captionStartId-3="6.[144,223,1313,1332]" captionTargetBox-0="[830,1438,195,1177]" captionTargetBox-1="[148,756,196,1254]" captionTargetBox-2="[329,1277,200,1387]" captionTargetBox-3="[309,1258,204,1273]" captionTargetId-0="figure-513@3.[830,1438,195,1177]" captionTargetId-1="figure-497@4.[148,756,196,1254]" captionTargetId-2="figure-168@5.[323,1283,195,1387]" captionTargetId-3="figure-236@6.[303,1263,194,1283]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="Figure 1. Sundathelphusa molluscivora sp. nov. Paratype male (23.5 by 19.0 mm) (ZRC 2000.1703). A, overall view; B, frontal view; C, ventral view." captionText-1="Figure 2. Sundathelphusa molluscivora sp. nov. Major chelae. A, male (23.5 by 19.0 mm) (ZRC 2000.1703); B, holotype male (24.6 by 20.4 mm) (MZB 1480); C, female (24.3 by 20.0 mm) (ZRC 2000.1703)." captionText-2="Figure 3. Sundathelphusa molluscivora sp. nov. Paratype male (23.5 by 19.0 mm) (ZRC 2000.1703). A, right side of carapace; B, anterior thoracic sternum; C, left third maxilliped; D, right chelipedal carpus; E, male abdomen; F, right fourth ambulatory leg; G, frontal median triangle. Scale bars: AF, 5.0 mm; G, 1.0 mm." captionText-3="Figure 4. Sundathelphusa molluscivora sp. nov. Gonopods, paratype male (23.5 by 19.0 mm) (ZRC 2000.1703). A and C, left G1, ventral view; B and D, left G1, dorsal view; E, left G2. Scale bars: A, B, E, 1.0 mm; C, D, 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4634969" figureDoi-1="http://doi.org/10.5281/zenodo.4634971" figureDoi-2="http://doi.org/10.5281/zenodo.4634973" figureDoi-3="http://doi.org/10.5281/zenodo.4634975" httpUri-0="https://zenodo.org/record/4634969/files/figure.png" httpUri-1="https://zenodo.org/record/4634971/files/figure.png" httpUri-2="https://zenodo.org/record/4634973/files/figure.png" httpUri-3="https://zenodo.org/record/4634975/files/figure.png" pageId="3" pageNumber="214">FIG. 14</figureCitation>
)
</paragraph>
<paragraph id="8BD636BA8723BF3EFF16F88CFB74FA47" blockId="3.[164,779,1850,1903]" lastBlockId="3.[826,1442,1362,1568]" pageId="3" pageNumber="214">
<materialsCitation id="3B013CE78723BF3EFF16F88CFB74FA47" ID-GBIF-Occurrence="3325232301" collectingDate="2000-01-22" collectionCode="MZB" collectorName="C. D. Schubart &amp; Schubart, T. M &amp; D. Wowor" country="Indonesia" elevation="2" location="Lake Poso" pageId="3" pageNumber="214" specimenCode="MZB 1480" specimenCount="1" stateProvince="Sulawesi Utara" typeStatus="holotype">
<typeStatus id="54D288188723BF3EFF16F88CFE8AF891" box="[164,266,1850,1872]" pageId="3" pageNumber="214" type="holotype">Holotype</typeStatus>
male (24.6 by
<quantity id="4C919B5F8723BF3EFE4AF88CFDDBF891" box="[504,603,1850,1872]" metricMagnitude="-2" metricUnit="m" metricValue="2.04" pageId="3" pageNumber="214" unit="mm" value="20.4">20.4 mm</quantity>
) (
<specimenCode id="DBCF9EC18723BF3EFDC9F88CFD7FF891" box="[635,767,1850,1872]" collectionCode="MZB" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="3" pageNumber="214" type="Museum">MZB 1480</specimenCode>
), under rocks,
<emphasis id="B91DEAA88723BF3EFE8AF8EFFECAF8AF" box="[312,330,1881,1902]" italics="true" pageId="3" pageNumber="214">c.</emphasis>
<elevation id="0044D1898723BF3EFEE3F8EFFE46F8AE" box="[337,454,1881,1903]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="214" unit="m" value="2.0">
<quantity id="4C919B5F8723BF3EFEE3F8EFFEFBF8AF" box="[337,379,1881,1902]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="214" unit="m" value="2.0">2 m</quantity>
depth
</elevation>
,
<location id="8EB660618723BF3EFE63F8EFFDC8F8AF" LSID="urn:lsid:plazi:treatment:03C087AC8723BF3AFF61F942FC62FE97:8EB660618723BF3EFE63F8EFFDC8F8AF" box="[465,584,1881,1903]" country="Indonesia" name="Lake Poso" pageId="3" pageNumber="214" stateProvince="Sulawesi Utara">Lake Poso</location>
, west coast, near Taipa,
<collectingRegion id="49ADF8588723BF3EFC39FAE4FC74FAA9" box="[907,1012,1362,1384]" country="Indonesia" name="Sulawesi Utara" pageId="3" pageNumber="214">Sulawesi</collectingRegion>
,
<collectingCountry id="F37E762A8723BF3EFBB0FAE4FBF4FAA9" box="[1026,1140,1362,1384]" name="Indonesia" pageId="3" pageNumber="214">Indonesia</collectingCountry>
, coll.
<collectorName id="269C536C8723BF3EFB0BFAE4FAE0FAA9" box="[1209,1376,1362,1384]" pageId="3" pageNumber="214">C.D. Schubart</collectorName>
, T.M. Leong &amp;
<collectorName id="269C536C8723BF3EFC15FAC7FB9AFA47" box="[935,1050,1393,1414]" pageId="3" pageNumber="214">D. Wowor</collectorName>
,
<date id="FFD7107A8723BF3EFB95FAC7FB70FA47" box="[1063,1264,1393,1414]" pageId="3" pageNumber="214" value="2000-01-22">
<collectingDate id="EF93E9928723BF3EFB95FAC7FB70FA47" box="[1063,1264,1393,1414]" pageId="3" pageNumber="214" value="2000-01-22">22 January 2000</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="8BD636BA8723BF3EFCE1FA26FBC7F9E1" blockId="3.[826,1442,1362,1568]" pageId="3" pageNumber="214">
<materialsCitation id="3B013CE78723BF3EFCE1FA26FBC3F9E1" ID-GBIF-Occurrence="3325232302" collectingDate="2000-01-22" collectionCode="ZRC, MZB" collectorName="C. D. Schubart &amp; Schubart, T. M &amp; D. Wowor" country="Indonesia" elevation="2" location="Lake Poso" pageId="3" pageNumber="214" specimenCode="ZRC 2000.1703, MZB 1481" specimenCount="1" stateProvince="Sulawesi Utara" typeStatus="paratype">
<typeStatus id="54D288188723BF3EFCE1FA26FC46FA64" box="[851,966,1424,1445]" pageId="3" pageNumber="214" type="paratype">Paratypes</typeStatus>
eight males (smallest 11.7 by
<quantity id="4C919B5F8723BF3EFA8AFA26FA1CFA64" box="[1336,1436,1424,1445]" metricMagnitude="-2" metricUnit="m" metricValue="1.02" pageId="3" pageNumber="214" unit="mm" value="10.2">10.2 mm</quantity>
, largest 23.5 by
<quantity id="4C919B5F8723BF3EFC5BFA18FBCFFA05" box="[1001,1103,1454,1476]" metricMagnitude="-2" metricUnit="m" metricValue="2.04" pageId="3" pageNumber="214" unit="mm" value="20.4">20.4 mm</quantity>
), eight females (smallest 12.0 by 10.0 mm, largest 24.7 by
<quantity id="4C919B5F8723BF3EFB3EFA7BFB71FA23" box="[1164,1265,1485,1506]" metricMagnitude="-2" metricUnit="m" metricValue="2.03" pageId="3" pageNumber="214" unit="mm" value="20.3">20.3 mm</quantity>
), two juveniles (
<specimenCode id="DBCF9EC18723BF3EFCF3FA5DFC74F9C0" box="[833,1012,1515,1537]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="3" pageNumber="214">ZRC 2000.1703</specimenCode>
), two males, two females (
<specimenCode id="DBCF9EC18723BF3EFAA9FA5DFA14F9C0" box="[1307,1428,1515,1537]" collectionCode="MZB" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="3" pageNumber="214" type="Museum">MZB 1481</specimenCode>
), same data as holotype
</materialsCitation>
.
</paragraph>
<paragraph id="8BD636BA8723BF39FC88F9D3FD34FA1A" blockId="3.[826,1442,1636,1903]" lastBlockId="4.[144,758,1446,1499]" lastPageId="4" lastPageNumber="215" pageId="3" pageNumber="214">
<emphasis id="B91DEAA88723BF3EFC88F9D3FC31F9BB" box="[826,945,1637,1658]" italics="true" pageId="3" pageNumber="214">Diagnosis:</emphasis>
Dorsal surface of carapace convex, relatively smooth, not prominently inflated or swollen. Frontal medium triangle complete, margins completely fused. External orbital angle broadly triangular; anterolateral margin distinctly convex, with one distinct tooth. Adult chelipeds markedly heterochelous in both sexes; major chela with strong molariform teeth at the bases of the cutting edges of fingers. Male abdomen distinctly T-shaped: segment 6 longer than telson. First gonopods gently curving outwards; terminal segment
<emphasis id="B91DEAA88724BF39FED4FA73FEF8FA1B" box="[358,376,1477,1498]" italics="true" pageId="4" pageNumber="215">c.</emphasis>
0.3 times the total length.
</paragraph>
<caption id="DF1666328724BF39FF22FAB2FD92FAAE" ID-DOI="http://doi.org/10.5281/zenodo.4634971" ID-Zenodo-Dep="4634971" httpUri="https://zenodo.org/record/4634971/files/figure.png" pageId="4" pageNumber="215" startId="4.[144,223,1284,1303]" targetBox="[148,756,196,1254]" targetPageId="4">
<paragraph id="8BD636BA8724BF39FF22FAB2FD92FAAE" blockId="4.[144,759,1284,1391]" pageId="4" pageNumber="215">
<emphasis id="B91DEAA88724BF39FF22FAB2FD77FAD6" bold="true" box="[144,759,1284,1303]" pageId="4" pageNumber="215">
Figure 2.
<taxonomicName id="4C694D398724BF39FEB6FAB2FDF6FAD6" authority="Schubart &amp; Ng, 2008" authorityName="Schubart &amp; Ng" authorityYear="2008" box="[260,630,1284,1303]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="215" phylum="Arthropoda" rank="species" species="molluscivora" status="sp. nov.">
<emphasis id="B91DEAA88724BF39FEB6FAB2FDF6FAD6" bold="true" box="[260,630,1284,1303]" italics="true" pageId="4" pageNumber="215">Sundathelphusa molluscivora</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E57D38724BF39FD3CFAB2FD77FAD6" box="[654,759,1284,1303]" pageId="4" pageNumber="215" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Major chelae. A, male (23.5 by 19.0 mm) (ZRC 2000.1703); B, holotype male (24.6 by 20.4 mm) (MZB 1480); C, female (24.3 by 20.0 mm) (ZRC 2000.1703).
</paragraph>
</caption>
<paragraph id="8BD636BA8724BF39FF22F9BFFAFAFC9F" blockId="4.[144,759,1544,1903]" lastBlockId="4.[806,1422,198,1903]" pageId="4" pageNumber="215">
<emphasis id="B91DEAA88724BF39FF22F9BFFE21F9DC" box="[144,417,1544,1566]" italics="true" pageId="4" pageNumber="215">
Description of
<typeStatus id="54D288188724BF39FE8EF9BEFE1BF9DC" box="[316,411,1544,1565]" pageId="4" pageNumber="215" type="holotype">holotype</typeStatus>
:
</emphasis>
Carapace appears quadrate, widest point of carapace about one-third of the distance from front, carapace gently convex transversely; regions poorly demarcated, relatively smooth; cervical groove distinct, broad; H-shaped gastrocardiac groove distinct; shallow transverse depression between cardiac and intestinal regions. Frontal median triangle complete, lateral and dorsal margins cristate, dorsal and lateral margins fused, but suture still visible. Epigastric cristae: low, rounded, but still discernible, separated from each other by shallow cleft. Postorbital cristae: very low, rugose, almost indiscernible. Frontal margin truncate, about half the maximum carapace width, appears vaguely bilobed from dorsal view; supraorbital margin entire, subparallel with frontal margin. External orbital tooth broadly triangular, clearly directed anteriorly, outer margin uneven, straight, twice as long as inner margin, separated from anterolateral margin by distinct V-shaped cleft, tip not extending beyond level of front. Anterolateral margin distinctly convex, slightly granular, and gently curving to join posterolateral margin. Posterolateral margin gently convex, converging gradually towards more concave posterior carapace margin. Suborbital, sub-branchial, and pterygostomial regions covered with scattered oblique short striae and small granules. Basal antennal segment large, subquadrate. Posterior margin of epistome with large median triangular lobe; lateral margins sinuous. Third maxilliped with ischium rectangular, submedian oblique sulcus distinct; merus subquadrate, with gentle median depression, anteroexternal angle rounded; tip of exopod reaching to midpoint of outer margin of merus, flagellum long.
</paragraph>
<paragraph id="8BD636BA8724BF39FC8DFCD1FC25FA89" blockId="4.[806,1422,198,1903]" pageId="4" pageNumber="215">Chelipeds prominently unequal. Merus with slightly serrated margins, no distinct subdistal tooth. Carpus with outer surface rugose, with welldeveloped inner distal spine that is somewhat laterally flattened, and proximal basal part with several small sharp tubercles. Outer surface of minor palm gently rugose to almost smooth; fingers longer than palm, cutting edges lined with numerous teeth and denticles, tip pectinated. Outer surface of major palm smooth or finely punctate; fingers subequal to length of palm, proximal two-thirds of cutting edge of pollex with large molariform tooth, rest of edge with distinct teeth; proximal one-fifth of cutting edge of dactylus with relatively smaller molariform tooth, rest of edge with numerous teeth and denticles, slender, tip pectinated.</paragraph>
<paragraph id="8BD636BA8724BF39FC8DFAE7FAEBF9DF" blockId="4.[806,1422,198,1903]" pageId="4" pageNumber="215">Ambulatory legs not elongate, second leg longest. Merus with dorsal, slightly serrate margin, subdistal angle with low tooth but no spine. Carpus elongate, dorsal margin gently serrated, ventral margin smooth, and outer surface with two low ridges. Dorsal and ventral margins of propodus serrated. Dactylus slender, slightly flattened laterally, gently curved.</paragraph>
<paragraph id="8BD636BA8724BF38FC8DF991FD59F9E8" blockId="4.[806,1422,198,1903]" lastBlockId="5.[164,779,1524,1791]" lastPageId="5" lastPageNumber="216" pageId="4" pageNumber="215">First to fifth thoracic sternites smooth, with scattered punctae; all sternites fused, no sutures evident. Abdominal cavity reaching to imaginary line joining anterior edge of coxae of chelipeds. Male abdomen distinctly T-shaped. Segment 1 very narrow longitudinally, proximal and distal margins gently sinuous. Segment 2 subrectangular transversely. Segments 35 gradually more trapezoidal; lateral margins of segment 3 convex; lateral margins of segments 4 and 5, straight to gently concave. Segment 6 longitudinally rectangular, longer than telson, and lateral margins gently sinuous. Telson longitudinally triangular, lateral margins concave, and tip rounded.</paragraph>
<caption id="DF1666328725BF38FF16FA3FFBD1FA16" ID-DOI="http://doi.org/10.5281/zenodo.4634973" ID-Zenodo-Dep="4634973" httpUri="https://zenodo.org/record/4634973/files/figure.png" pageId="5" pageNumber="216" startId="5.[164,243,1417,1436]" targetBox="[329,1277,200,1387]" targetPageId="5">
<paragraph id="8BD636BA8725BF38FF16FA3FFBD1FA16" blockId="5.[164,1441,1417,1495]" pageId="5" pageNumber="216">
<emphasis id="B91DEAA88725BF38FF16FA3FFD58FA5D" bold="true" box="[164,728,1417,1436]" pageId="5" pageNumber="216">
Figure 3.
<taxonomicName id="4C694D398725BF38FEABFA3FFDFAFA5D" authority="Schubart &amp; Ng, 2008" authorityName="Schubart &amp; Ng" authorityYear="2008" box="[281,634,1417,1436]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="216" phylum="Arthropoda" rank="species" species="molluscivora" status="sp. nov.">
<emphasis id="B91DEAA88725BF38FEABFA3FFDFAFA5D" bold="true" box="[281,634,1417,1436]" italics="true" pageId="5" pageNumber="216">Sundathelphusa molluscivora</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E57D38725BF38FD33FA3FFD58FA5D" box="[641,728,1417,1436]" pageId="5" pageNumber="216" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Paratype male (23.5 by 19.0 mm) (ZRC 2000.1703). A, right side of carapace; B, anterior thoracic sternum; C, left third maxilliped; D, right chelipedal carpus; E, male abdomen; F, right fourth ambulatory leg; G, frontal median triangle. Scale bars: AF, 5.0 mm; G, 1.0 mm.
</paragraph>
</caption>
<paragraph id="8BD636BA8725BF38FF0EF984FED1F93E" blockId="5.[164,779,1524,1791]" pageId="5" pageNumber="216">
First gonopods gently curving outwards; subterminal segment with outer margin gently concave; terminal segment
<emphasis id="B91DEAA88725BF38FECFF9C6FE0FF944" box="[381,399,1648,1669]" italics="true" pageId="5" pageNumber="216">c.</emphasis>
0.3 times total gonopod length, slightly curved, subcylindrical, tip gently rounded. Second gonopods elongated, much longer than first; distal segment long,
<emphasis id="B91DEAA88725BF38FE10F97DFE34F921" box="[418,436,1739,1760]" italics="true" pageId="5" pageNumber="216">c.</emphasis>
0.7 times the length of the basal segment.
</paragraph>
<paragraph id="8BD636BA8725BF38FF16F8AAFAC6F920" blockId="5.[164,779,1820,1903]" lastBlockId="5.[826,1441,1524,1761]" pageId="5" pageNumber="216">
<emphasis id="B91DEAA88725BF38FF16F8AAFE9BF8F0" box="[164,283,1820,1841]" italics="true" pageId="5" pageNumber="216">
<typeStatus id="54D288188725BF38FF16F8AAFE95F8F0" box="[164,277,1820,1841]" pageId="5" pageNumber="216" type="paratype">Paratypes</typeStatus>
:
</emphasis>
The
<typeStatus id="54D288188725BF38FEE3F8AAFE37F8F0" box="[337,439,1820,1841]" pageId="5" pageNumber="216" type="paratype">paratype</typeStatus>
males agree with the
<typeStatus id="54D288188725BF38FD18F8AAFC8BF8F3" box="[682,779,1820,1842]" pageId="5" pageNumber="216" type="holotype">holotype</typeStatus>
male in all major aspects, although in smaller specimens, the major chela is relatively smaller. The carapaces of smaller specimens also tend to be more quadrate. Females agree with the males in most non-sexual characters, although the lateral margins in larger specimens are slightly more convex, giving them a somewhat more inflated appearance. The heterochely in females is also less prominent, with the major claw being much smaller than in males of equivalent sizes, but still showing molariform dentation.
</paragraph>
<paragraph id="8BD636BA8725BF38FC88F8AAFA81F8AE" blockId="5.[826,1441,1820,1903]" pageId="5" pageNumber="216">
<emphasis id="B91DEAA88725BF38FC88F8AAFC56F8F0" box="[826,982,1820,1841]" italics="true" pageId="5" pageNumber="216">Colour in life:</emphasis>
Pale yellow; carapace darker, somewhat olive; legs lighter; chelae homogeneously yellow, except shining white molariform teeth.
</paragraph>
<caption id="DF1666328726BF3BFF22FA97FAB1FA93" ID-DOI="http://doi.org/10.5281/zenodo.4634975" ID-Zenodo-Dep="4634975" httpUri="https://zenodo.org/record/4634975/files/figure.png" pageId="6" pageNumber="217" startId="6.[144,223,1313,1332]" targetBox="[309,1258,204,1273]" targetPageId="6">
<paragraph id="8BD636BA8726BF3BFF22FA97FAB1FA93" blockId="6.[144,1421,1313,1362]" pageId="6" pageNumber="217">
<emphasis id="B91DEAA88726BF3BFF22FA97FD40FAF5" bold="true" box="[144,704,1313,1332]" pageId="6" pageNumber="217">
Figure 4.
<taxonomicName id="4C694D398726BF3BFEB6FA97FDE4FAF5" authority="Schubart &amp; Ng, 2008" authorityName="Schubart &amp; Ng" authorityYear="2008" box="[260,612,1313,1332]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="217" phylum="Arthropoda" rank="species" species="molluscivora" status="sp. nov.">
<emphasis id="B91DEAA88726BF3BFEB6FA97FDE4FAF5" bold="true" box="[260,612,1313,1332]" italics="true" pageId="6" pageNumber="217">Sundathelphusa molluscivora</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E57D38726BF3BFDD8FA97FD40FAF5" box="[618,704,1313,1332]" pageId="6" pageNumber="217" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Gonopods, paratype male (23.5 by 19.0 mm) (ZRC 2000.1703). A and C, left G1, ventral view; B and D, left G1, dorsal view; E, left G2. Scale bars: A, B, E, 1.0 mm; C, D, 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BD636BA8726BF3BFF22FA38FEEAF9FC" blockId="6.[144,759,1422,1597]" pageId="6" pageNumber="217">
<emphasis id="B91DEAA88726BF3BFF22FA38FE8DFA62" box="[144,269,1422,1443]" italics="true" pageId="6" pageNumber="217">Etymology:</emphasis>
The name molluscivora (mollusc-eating) is used as an adjective, and is derived from the taxon name Mollusca and the Latin verb vorare (to devour). It makes reference to the apparent mollusceating habit of these crabs, as suggested by their chelar morphology.
</paragraph>
<paragraph id="8BD636BA8726BF3BFF22F9D2FE69F976" blockId="6.[144,759,1636,1719]" pageId="6" pageNumber="217">
<emphasis id="B91DEAA88726BF3BFF22F9D2FEA1F9B8" box="[144,289,1636,1657]" italics="true" pageId="6" pageNumber="217">Distribution:</emphasis>
This species has only been found from its
<typeStatus id="54D288188726BF3BFF01F935FF64F959" box="[179,228,1667,1688]" pageId="6" pageNumber="217">type</typeStatus>
locality in Lake Poso (
<collectingRegion id="49ADF8588726BF3BFE54F935FDCFF958" box="[486,591,1667,1689]" country="Indonesia" name="Sulawesi Utara" pageId="6" pageNumber="217">Sulawesi</collectingRegion>
,
<collectingCountry id="F37E762A8726BF3BFDE9F935FD4EF958" box="[603,718,1667,1689]" name="Indonesia" pageId="6" pageNumber="217">Indonesia</collectingCountry>
), to which it is probably endemic.
</paragraph>
<paragraph id="8BD636BA8726BF3BFF22F968FC6DF9FC" blockId="6.[144,759,1758,1903]" lastBlockId="6.[806,1421,1422,1903]" pageId="6" pageNumber="217">
<emphasis id="B91DEAA88726BF3BFF22F968FF7BF932" box="[144,251,1758,1779]" italics="true" pageId="6" pageNumber="217">Remarks:</emphasis>
The genus
<taxonomicName id="4C694D398726BF3BFECCF968FD37F935" authority="Bott, 1969" authorityName="Bott" authorityYear="1969" box="[382,695,1758,1780]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="217" phylum="Arthropoda" rank="genus">
<emphasis id="B91DEAA88726BF3BFECCF968FDBAF932" box="[382,570,1758,1779]" italics="true" pageId="6" pageNumber="217">Sundathelphusa</emphasis>
<bibRefCitation id="EFF84B4B8726BF3BFDF3F969FD37F935" author="Bott R" box="[577,695,1758,1780]" pageId="6" pageNumber="217" pagination="359 - 366" refId="ref5734" refString="Bott R. 1969. Die Flusskrabben aus Asien und ihre Klassifikation (Crustacea, Decapoda). Senckenbergiana biologica, Frankfurt 50: 359 - 366." type="journal article" year="1969">Bott, 1969</bibRefCitation>
</taxonomicName>
(
<typeStatus id="54D288188726BF3BFD77F969FD77F935" box="[709,759,1759,1780]" pageId="6" pageNumber="217">type</typeStatus>
species
<taxonomicName id="4C694D398726BF3BFF42F948FF49F8F0" authority="De Man, 1902" authorityName="De Man" authorityYear="1902" class="Malacostraca" family="Potamidae" genus="Potamon" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="217" phylum="Arthropoda" rank="species" species="cassiope" subGenus="Geothelphusa">
<emphasis id="B91DEAA88726BF3BFF42F948FED5F8D2" box="[240,341,1790,1811]" italics="true" pageId="6" pageNumber="217">Potamon</emphasis>
(
<emphasis id="B91DEAA88726BF3BFEDFF94BFD87F8D3" box="[365,519,1789,1810]" italics="true" pageId="6" pageNumber="217">Geothelphusa</emphasis>
)
<emphasis id="B91DEAA88726BF3BFD92F948FDFCF8D2" box="[544,636,1790,1811]" italics="true" pageId="6" pageNumber="217">cassiope</emphasis>
<bibRefCitation id="EFF84B4B8726BF3BFD3EF94BFF49F8F0" author="De Man JG" pageId="6" pageNumber="217" pagination="467 - 929" refId="ref5982" refString="De Man JG. 1902. Die von Herrn Professor Kukenthal in Indischen Archipel gesammelten Dekapoden und Stomatopoden. In: Kukenthal W, ed. Ergebnisse einer zoologischen Forschungsreise in den Molukken und Borneo. Abhandlungen der Senckenberg Naturforschenden Gesellschaft 25: 467 - 929." type="journal article" year="1902">De Man, 1902</bibRefCitation>
</taxonomicName>
, by original designation) currently contains 28 species from the
<collectingCountry id="F37E762A8726BF3BFEE1F88CFE57F891" box="[339,471,1850,1872]" name="Philippines" pageId="6" pageNumber="217">Philippines</collectingCountry>
,
<collectingRegion id="49ADF8588726BF3BFE56F88CFDCDF891" box="[484,589,1850,1872]" country="Indonesia" name="Sulawesi Utara" pageId="6" pageNumber="217">Sulawesi</collectingRegion>
,
<collectingCountry id="F37E762A8726BF3BFDE8F88CFD43F891" box="[602,707,1850,1872]" name="Indonesia" pageId="6" pageNumber="217">Moluccas</collectingCountry>
and eastern Borneo (
<bibRefCitation id="EFF84B4B8726BF3BFEF9F8EFFDBBF8AF" author="Ng PKL &amp; Stuebing R" box="[331,571,1881,1903]" pageId="6" pageNumber="217" pagination="13 - 19" refId="ref6559" refString="Ng PKL, Stuebing R. 1989. Description of a new species of montane freshwater crab of the genus Sundathelphusa Bott, 1969 (Crustacea: Brachyura: Decapoda: Brachyura: Gecarcinucoidea) from Borneo. Malayan Nature Journal 43: 13 - 19." type="journal article" year="1989">Ng &amp; Stuebing, 1989</bibRefCitation>
;
<bibRefCitation id="EFF84B4B8726BF3BFDF5F8EFFD2FF8AF" author="Ng PKL" box="[583,687,1881,1902]" pageId="6" pageNumber="217" pagination="13 - 24" refId="ref6472" refString="Ng PKL. 1991. On two species of Archipelothelphusa Bott, 1969 (Crustacea: Decapoda: Brachyura: Sundathelphusidae) from Luzon, Philippines, one of which is new. Zoologische Mededelingen, Leiden 65: 13 - 24." type="journal article" year="1991">Ng, 1991</bibRefCitation>
;
<bibRefCitation id="EFF84B4B8726BF3BFD08F8EFFC24FA65" author="Ng PKL &amp; Sket B" box="[698,932,1422,1902]" pageId="6" pageNumber="217" pagination="695 - 706" refId="ref6515" refString="Ng PKL, Sket B. 1996. The freshwater crab fauna (Crustacea: Brachyura) of the Philippines. IV. On a collection of Parathelphusidae from Bohol. Proceedings of the Biological Society of Washington 109: 695 - 706." type="journal article" year="1996">Ng &amp; Sket, 1996</bibRefCitation>
;
<bibRefCitation id="EFF84B4B8726BF3BFC00FA38FB18FA65" author="Takeda M &amp; Ng PKL" box="[946,1176,1422,1444]" pageId="6" pageNumber="217" pagination="1123 - 1127" refId="ref7127" refString="Takeda M, Ng PKL. 2001. The freshwater crab fauna (Crustacea, Brachyura) of the Philippines. VI. A new cavernicolous crab from Mindanao. Zoological Science, Tokyo 18: 1123 - 1127." type="journal article" year="2001">Takeda &amp; Ng, 2001</bibRefCitation>
). The genus is dominant in the
<collectingCountry id="F37E762A8726BF3BFC7FFA1AFBD1FA03" box="[973,1105,1452,1474]" name="Philippines" pageId="6" pageNumber="217">Philippines</collectingCountry>
, with only three species known from Sulawesi thus far, viz.
<taxonomicName id="4C694D398726BF3BFB63FA7DFBCAF9C1" authority="(De Man, 1902)" baseAuthorityName="De Man" baseAuthorityYear="1902" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="217" phylum="Arthropoda" rank="species" species="cassiope">
<emphasis id="B91DEAA88726BF3BFB63FA7DFC02FA3E" italics="true" pageId="6" pageNumber="217">Sundathelphusa cassiope</emphasis>
(
<bibRefCitation id="EFF84B4B8726BF3BFC24FA5CFBC2F9C1" author="De Man JG" box="[918,1090,1514,1536]" pageId="6" pageNumber="217" pagination="467 - 929" refId="ref5982" refString="De Man JG. 1902. Die von Herrn Professor Kukenthal in Indischen Archipel gesammelten Dekapoden und Stomatopoden. In: Kukenthal W, ed. Ergebnisse einer zoologischen Forschungsreise in den Molukken und Borneo. Abhandlungen der Senckenberg Naturforschenden Gesellschaft 25: 467 - 929." type="journal article" year="1902">De Man, 1902</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C694D398726BF3BFBE8FA5CFBA8F9DF" authority="(Schenkel, 1902)" baseAuthorityName="Schenkel" baseAuthorityYear="1902" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="217" phylum="Arthropoda" rank="species" species="minahassae">
<emphasis id="B91DEAA88726BF3BFBE8FA5CFCCAF9DF" italics="true" pageId="6" pageNumber="217">Sundathelphusa minahassae</emphasis>
(
<bibRefCitation id="EFF84B4B8726BF3BFCD7F9BEFBA1F9DF" author="Schenkel E" box="[869,1057,1544,1566]" pageId="6" pageNumber="217" pagination="485 - 618" refId="ref6853" refString="Schenkel E. 1902. Beitrag zur Kenntnis der Dekapodenfauna von Celebes. Verhandlungen der Naturforschenden Gesellschaft Basel 13: 485 - 618." type="journal article" year="1902">Schenkel, 1902</bibRefCitation>
)
</taxonomicName>
, and
<taxonomicName id="4C694D398726BF3BFBCCF9BEFC69F9FC" authority="(Schenkel, 1902)" baseAuthorityName="Schenkel" baseAuthorityYear="1902" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="217" phylum="Arthropoda" rank="species" species="rubra">
<emphasis id="B91DEAA88726BF3BFBCCF9BEFA0DF9DC" box="[1150,1421,1544,1565]" italics="true" pageId="6" pageNumber="217">Sundathelphusa rubra</emphasis>
(
<bibRefCitation id="EFF84B4B8726BF3BFC9CF991FC61F9FC" author="Schenkel E" box="[814,993,1575,1597]" pageId="6" pageNumber="217" pagination="485 - 618" refId="ref6853" refString="Schenkel E. 1902. Beitrag zur Kenntnis der Dekapodenfauna von Celebes. Verhandlungen der Naturforschenden Gesellschaft Basel 13: 485 - 618." type="journal article" year="1902">Schenkel, 1902</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="8BD636BA8726BF3AFC8DF9F3FD5CFEF9" blockId="6.[806,1421,1422,1903]" lastBlockId="7.[164,779,198,1903]" lastPageId="7" lastPageNumber="218" pageId="6" pageNumber="217">
The authors have examined photographs of the whole
<typeStatus id="54D288188726BF3BFCC7F9D2FC26F9B8" box="[885,934,1636,1657]" pageId="6" pageNumber="217">type</typeStatus>
specimens, with figures of the G1s and G2s of
<taxonomicName id="4C694D398726BF3BFCCBF935FC76F959" baseAuthorityName="De Man" baseAuthorityYear="1902" box="[889,1014,1667,1688]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="217" phylum="Arthropoda" rank="species" species="cassiope">
<emphasis id="B91DEAA88726BF3BFCCBF935FC76F959" box="[889,1014,1667,1688]" italics="true" pageId="6" pageNumber="217">S. cassiope</emphasis>
</taxonomicName>
,
<taxonomicName id="4C694D398726BF3BFBB5F935FB2EF959" baseAuthorityName="Schenkel" baseAuthorityYear="1902" box="[1031,1198,1667,1688]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="217" phylum="Arthropoda" rank="species" species="minahassae">
<emphasis id="B91DEAA88726BF3BFBB5F935FB2EF959" box="[1031,1198,1667,1688]" italics="true" pageId="6" pageNumber="217">S. minahassae</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C694D398726BF3BFB41F935FAD4F959" baseAuthorityName="Schenkel" baseAuthorityYear="1902" box="[1267,1364,1667,1688]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="217" phylum="Arthropoda" rank="species" species="rubra">
<emphasis id="B91DEAA88726BF3BFB41F935FAD4F959" box="[1267,1364,1667,1688]" italics="true" pageId="6" pageNumber="217">S. rubra</emphasis>
</taxonomicName>
that were prepared for us by Oliver Chia (National University of
<collectingCountry id="F37E762A8726BF3BFC29F976FB8FF917" box="[923,1039,1728,1750]" name="Singapore" pageId="6" pageNumber="217">Singapore</collectingCountry>
) at our request (see also
<bibRefCitation id="EFF84B4B8726BF3BFA88F976FC10F935" author="Chia OCK &amp; Ng PKL" pageId="6" pageNumber="217" pagination="381 - 428" refId="ref5860" refString="Chia OCK, Ng PKL. 2006. The freshwater crabs of Sulawesi, with descriptions of two new genera and four new species (Crustacea: Decapoda: Brachyura: Parathelphusidae). Raffles Bulletin of Zoology Singapore 54: 381 - 428." type="journal article" year="2006">Chia &amp; Ng, 2006</bibRefCitation>
). As such, we have no doubt that our present specimens from Poso belong to a new species. The most obvious character that distinguishes adults of
<taxonomicName id="4C694D398726BF3BFAC4F8AAFC38F88E" authority="Schubart &amp; Ng, 2008" authorityName="Schubart &amp; Ng" authorityYear="2008" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="217" phylum="Arthropoda" rank="species" species="molluscivora" status="sp. nov.">
<emphasis id="B91DEAA88726BF3BFAC4F8AAFC38F88E" italics="true" pageId="6" pageNumber="217">S. molluscivora</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E57D38726BF3BFC70F88DFB9AF891" box="[962,1050,1851,1872]" pageId="6" pageNumber="217" rank="species">sp. nov.</taxonomicNameLabel>
from all congeners is the large molariform tooth on the larger chela. Such a character is present in only two other freshwater crabs of the region, viz.
<taxonomicName id="4C694D398727BF3AFEC0FF53FF63FED8" authority="(Schenkel, 1902)" baseAuthorityName="Schenkel" baseAuthorityYear="1902" class="Malacostraca" family="Gecarcinucidae" genus="Syntripsa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="218" phylum="Arthropoda" rank="species" species="matannensis">
<emphasis id="B91DEAA88727BF3AFEC0FF53FD03FF3B" box="[370,643,229,250]" italics="true" pageId="7" pageNumber="218">Syntripsa matannensis</emphasis>
(
<bibRefCitation id="EFF84B4B8727BF3AFD2EFF52FF5BFED8" author="Schenkel E" pageId="7" pageNumber="218" pagination="485 - 618" refId="ref6853" refString="Schenkel E. 1902. Beitrag zur Kenntnis der Dekapodenfauna von Celebes. Verhandlungen der Naturforschenden Gesellschaft Basel 13: 485 - 618." type="journal article" year="1902">Schenkel, 1902</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C694D398727BF3AFEAFFEB5FD4CFED8" authority="Chia &amp; Ng, 2006" authorityName="Chia &amp; Ng" authorityYear="2006" box="[285,716,259,281]" class="Malacostraca" family="Gecarcinucidae" genus="Syntripsa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="218" phylum="Arthropoda" rank="species" species="flavichela">
<emphasis id="B91DEAA88727BF3AFEAFFEB5FD81FED9" box="[285,513,259,280]" italics="true" pageId="7" pageNumber="218">Syntripsa flavichela</emphasis>
<bibRefCitation id="EFF84B4B8727BF3AFDBBFEB5FD4CFED8" author="Chia OCK &amp; Ng PKL" box="[521,716,259,281]" pageId="7" pageNumber="218" pagination="381 - 428" refId="ref5860" refString="Chia OCK, Ng PKL. 2006. The freshwater crabs of Sulawesi, with descriptions of two new genera and four new species (Crustacea: Decapoda: Brachyura: Parathelphusidae). Raffles Bulletin of Zoology Singapore 54: 381 - 428." type="journal article" year="2006">Chia &amp; Ng, 2006</bibRefCitation>
</taxonomicName>
, both from the Malili lake system in central Sulawesi.
</paragraph>
<paragraph id="8BD636BA8727BF3AFF0EFEF6FEEFFCC2" blockId="7.[164,779,198,1903]" pageId="7" pageNumber="218">
The carapaces of
<taxonomicName id="4C694D398727BF3AFE35FEF7FD83FE97" baseAuthorityName="De Man" baseAuthorityYear="1902" box="[391,515,321,342]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="218" phylum="Arthropoda" rank="species" species="cassiope">
<emphasis id="B91DEAA88727BF3AFE35FEF7FD83FE97" box="[391,515,321,342]" italics="true" pageId="7" pageNumber="218">S. cassiope</emphasis>
</taxonomicName>
and
<taxonomicName id="4C694D398727BF3AFDF2FEF7FD22FE94" baseAuthorityName="Schenkel" baseAuthorityYear="1902" box="[576,674,320,342]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="218" phylum="Arthropoda" rank="species" species="rubra">
<emphasis id="B91DEAA88727BF3AFDF2FEF7FD22FE94" box="[576,674,320,342]" italics="true" pageId="7" pageNumber="218">S. rubra</emphasis>
</taxonomicName>
are both prominently swollen, with the dorsal surface strongly convex and the branchial regions swollen. In addition, their G1 terminal segments are proportionately much shorter. The same is true of
<taxonomicName id="4C694D398727BF3AFE40FE0DFE58FE2F" authority="Schenkel, 1902" authorityName="Schenkel" authorityYear="1902" class="Malacostraca" family="Potamidae" genus="Potamon" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="218" phylum="Arthropoda" rank="species" species="angustipes" subGenus="Geothelphusa">
<emphasis id="B91DEAA88727BF3AFE40FE0DFDD7FE11" box="[498,599,443,464]" italics="true" pageId="7" pageNumber="218">Potamon</emphasis>
(
<emphasis id="B91DEAA88727BF3AFDDAFE0DFC82FE11" box="[616,770,443,464]" italics="true" pageId="7" pageNumber="218">Geothelphusa</emphasis>
)
<emphasis id="B91DEAA88727BF3AFF16FE6CFE9DFE2E" box="[164,285,474,495]" italics="true" pageId="7" pageNumber="218">angustipes</emphasis>
<bibRefCitation id="EFF84B4B8727BF3AFE95FE6FFE58FE2F" author="Schenkel E" box="[295,472,473,495]" pageId="7" pageNumber="218" pagination="485 - 618" refId="ref6853" refString="Schenkel E. 1902. Beitrag zur Kenntnis der Dekapodenfauna von Celebes. Verhandlungen der Naturforschenden Gesellschaft Basel 13: 485 - 618." type="journal article" year="1902">Schenkel, 1902</bibRefCitation>
</taxonomicName>
, a species that was tentatively synonymized with
<taxonomicName id="4C694D398727BF3AFE78FE4EFDACFDCC" baseAuthorityName="Schenkel" baseAuthorityYear="1902" box="[458,556,504,525]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="218" phylum="Arthropoda" rank="species" species="rubra">
<emphasis id="B91DEAA88727BF3AFE78FE4EFDACFDCC" box="[458,556,504,525]" italics="true" pageId="7" pageNumber="218">S. rubra</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFF84B4B8727BF3AFDEEFE4EFC8BFDCF" author="Bott R" box="[604,779,504,526]" pageId="7" pageNumber="218" pagination="1 - 338" refId="ref5762" refString="Bott R. 1970. Die Susswasserkrabben von Europa, Asien, Australien und ihre Stammesgeschichte. Eine Revision der Potamoidea und Parathelphusoidea (Crustacea, Decapoda). Abhandlungen der Senckenberg Naturforschenden Gesellschaft, Frankfurt 526: 1 - 338." type="journal article" year="1970">Bott (1970: 74)</bibRefCitation>
(see also
<bibRefCitation id="EFF84B4B8727BF3AFEA1FDA0FD82FDED" author="Schenkel E" box="[275,514,534,556]" pageId="7" pageNumber="218" pagination="485 - 618" refId="ref6853" refString="Schenkel E. 1902. Beitrag zur Kenntnis der Dekapodenfauna von Celebes. Verhandlungen der Naturforschenden Gesellschaft Basel 13: 485 - 618." type="journal article" year="1902">Schenkel, 1902: 533</bibRefCitation>
, pl. 11 fig. 17;
<bibRefCitation id="EFF84B4B8727BF3AFD07FDA0FE92FD8A" author="Chia OCK &amp; Ng PKL" pageId="7" pageNumber="218" pagination="381 - 428" refId="ref5860" refString="Chia OCK, Ng PKL. 2006. The freshwater crabs of Sulawesi, with descriptions of two new genera and four new species (Crustacea: Decapoda: Brachyura: Parathelphusidae). Raffles Bulletin of Zoology Singapore 54: 381 - 428." type="journal article" year="2006">Chia &amp; Ng, 2006</bibRefCitation>
). In the form of the flat carapace and the structure of the G1,
<taxonomicName id="4C694D398727BF3AFE2AFDE2FDCCFDA8" authority="Schubart &amp; Ng, 2008" authorityName="Schubart &amp; Ng" authorityYear="2008" box="[408,588,596,617]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="218" phylum="Arthropoda" rank="species" species="molluscivora" status="sp. nov.">
<emphasis id="B91DEAA88727BF3AFE2AFDE2FDCCFDA8" box="[408,588,596,617]" italics="true" pageId="7" pageNumber="218">S. molluscivora</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E57D38727BF3AFDE4FDE2FD2EFDA8" box="[598,686,596,617]" pageId="7" pageNumber="218" rank="species">sp. nov.</taxonomicNameLabel>
is morphologically closest to
<taxonomicName id="4C694D398727BF3AFE0BFDC5FDE5FD46" baseAuthorityName="Schenkel" baseAuthorityYear="1902" box="[441,613,626,648]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="218" phylum="Arthropoda" rank="species" species="minahassae">
<emphasis id="B91DEAA88727BF3AFE0BFDC5FDE5FD46" box="[441,613,626,648]" italics="true" pageId="7" pageNumber="218">S. minahassae</emphasis>
</taxonomicName>
. However,
<taxonomicName id="4C694D398727BF3AFD41FDC5FEB6FD67" authority="Schubart &amp; Ng, 2008" authorityName="Schubart &amp; Ng" authorityYear="2008" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="218" phylum="Arthropoda" rank="species" species="molluscivora" status="sp. nov.">
<emphasis id="B91DEAA88727BF3AFD41FDC5FEB6FD67" italics="true" pageId="7" pageNumber="218">S. molluscivora</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E57D38727BF3AFE8CFD27FE15FD67" box="[318,405,657,678]" pageId="7" pageNumber="218" rank="species">sp. nov.</taxonomicNameLabel>
can easily be separated, in that its anterolateral margin is more strongly convex, and the posterolateral regions are smoother, with very low transverse striae.
</paragraph>
<paragraph id="8BD636BA8727BF3AFF0EFCBAFC8BFB6F" blockId="7.[164,779,198,1903]" pageId="7" pageNumber="218">
Two other species of
<taxonomicName id="4C694D398727BF3AFE70FCBDFDFEFCE1" authorityName="Bott" authorityYear="1969" box="[450,638,779,800]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="218" phylum="Arthropoda" rank="genus">
<emphasis id="B91DEAA88727BF3AFE70FCBDFDFEFCE1" box="[450,638,779,800]" italics="true" pageId="7" pageNumber="218">Sundathelphusa</emphasis>
</taxonomicName>
are known from near
<collectingRegion id="49ADF8588727BF3AFE93FC9CFE08FC81" box="[289,392,810,832]" country="Indonesia" name="Sulawesi Utara" pageId="7" pageNumber="218">Sulawesi</collectingRegion>
in the
<collectingCountry id="F37E762A8727BF3AFE53FC9CFDCDFC81" box="[481,589,810,832]" name="Indonesia" pageId="7" pageNumber="218">Moluccas</collectingCountry>
, viz.
<taxonomicName id="4C694D398727BF3AFD3EFC9CFE7EFC9E" authority="(Roux, 1911)" baseAuthorityName="Roux" baseAuthorityYear="1911" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="218" phylum="Arthropoda" rank="species" species="aruana">
<emphasis id="B91DEAA88727BF3AFD3EFC9CFECFFC9F" italics="true" pageId="7" pageNumber="218">Sundathelphusa aruana</emphasis>
(
<bibRefCitation id="EFF84B4B8727BF3AFED9FCFFFE76FC9E" author="Roux J" box="[363,502,841,863]" pageId="7" pageNumber="218" pagination="81 - 106" refId="ref6803" refString="Roux J. 1911. Nouvelles especes de Decapodes d'eau douce provenant de Papouasie. Notes of the Leyden Museum 33: 81 - 106." type="journal article" year="1911">Roux, 1911</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C694D398727BF3AFDFDFCFEFDB4FCBC" authority="(De Man, 1902)" baseAuthorityName="De Man" baseAuthorityYear="1902" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="218" phylum="Arthropoda" rank="species" species="halmaherensis">
<emphasis id="B91DEAA88727BF3AFDFDFCFEFECAFCBD" italics="true" pageId="7" pageNumber="218">Sundathelphusa halmaherensis</emphasis>
(
<bibRefCitation id="EFF84B4B8727BF3AFEDBFCD1FDACFCBC" author="De Man JG" box="[361,556,871,893]" pageId="7" pageNumber="218" pagination="467 - 929" refId="ref5982" refString="De Man JG. 1902. Die von Herrn Professor Kukenthal in Indischen Archipel gesammelten Dekapoden und Stomatopoden. In: Kukenthal W, ed. Ergebnisse einer zoologischen Forschungsreise in den Molukken und Borneo. Abhandlungen der Senckenberg Naturforschenden Gesellschaft 25: 467 - 929." type="journal article" year="1902">De Man, 1902</bibRefCitation>
)
</taxonomicName>
.
<taxonomicName id="4C694D398727BF3AFDFDFCD1FEB6FC5A" authority="Schubart &amp; Ng, 2008" authorityName="Schubart &amp; Ng" authorityYear="2008" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="218" phylum="Arthropoda" rank="species" species="molluscivora" status="sp. nov.">
<emphasis id="B91DEAA88727BF3AFDFDFCD1FEB6FC5A" italics="true" pageId="7" pageNumber="218">Sundathelphusa molluscivora</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E57D38727BF3AFEF1FC30FE1FFC5A" box="[323,415,902,923]" pageId="7" pageNumber="218" rank="species">sp. nov.</taxonomicNameLabel>
differs from
<taxonomicName id="4C694D398727BF3AFDF6FC30FD3CFC5A" baseAuthorityName="Roux" baseAuthorityYear="1911" box="[580,700,902,923]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="218" phylum="Arthropoda" rank="species" species="aruana">
<emphasis id="B91DEAA88727BF3AFDF6FC30FD3CFC5A" box="[580,700,902,923]" italics="true" pageId="7" pageNumber="218">S. aruana</emphasis>
</taxonomicName>
in its prominently flatter carapace, with the latter resembling
<taxonomicName id="4C694D398727BF3AFF5AFC75FEE6FC19" baseAuthorityName="De Man" baseAuthorityYear="1902" box="[232,358,963,984]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="218" phylum="Arthropoda" rank="species" species="cassiope">
<emphasis id="B91DEAA88727BF3AFF5AFC75FEE6FC19" box="[232,358,963,984]" italics="true" pageId="7" pageNumber="218">S. cassiope</emphasis>
</taxonomicName>
in carapace physiognomy (cf.
<typeStatus id="54D288188727BF3AFD7CFC75FC8BFC19" box="[718,779,963,984]" pageId="7" pageNumber="218">types</typeStatus>
examined, see also
<bibRefCitation id="EFF84B4B8727BF3AFE23FC54FDBEFC36" author="Bott R" box="[401,574,993,1015]" pageId="7" pageNumber="218" pagination="1 - 338" refId="ref5762" refString="Bott R. 1970. Die Susswasserkrabben von Europa, Asien, Australien und ihre Stammesgeschichte. Eine Revision der Potamoidea und Parathelphusoidea (Crustacea, Decapoda). Abhandlungen der Senckenberg Naturforschenden Gesellschaft, Frankfurt 526: 1 - 338." type="journal article" year="1970">Bott, 1970: 75</bibRefCitation>
, 76;
<bibRefCitation id="EFF84B4B8727BF3AFDCCFC57FF5EFBD7" author="Chia OCK &amp; Ng PKL" pageId="7" pageNumber="218" pagination="381 - 428" refId="ref5860" refString="Chia OCK, Ng PKL. 2006. The freshwater crabs of Sulawesi, with descriptions of two new genera and four new species (Crustacea: Decapoda: Brachyura: Parathelphusidae). Raffles Bulletin of Zoology Singapore 54: 381 - 428." type="journal article" year="2006">Chia &amp; Ng, 2006</bibRefCitation>
: figs 42, 43). The status of
<taxonomicName id="4C694D398727BF3AFD96FBB6FD6CFBD4" baseAuthorityName="De Man" baseAuthorityYear="1902" box="[548,748,1024,1045]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="218" phylum="Arthropoda" rank="species" species="halmaherensis">
<emphasis id="B91DEAA88727BF3AFD96FBB6FD6CFBD4" box="[548,748,1024,1045]" italics="true" pageId="7" pageNumber="218">S. halmaherensis</emphasis>
</taxonomicName>
is not clear, as it is only known from juveniles, but it is a species with a clearly more rugose carapace (see
<bibRefCitation id="EFF84B4B8727BF3AFD59FB88FEDAFBB3" author="De Man JG" pageId="7" pageNumber="218" pagination="467 - 929" refId="ref5982" refString="De Man JG. 1902. Die von Herrn Professor Kukenthal in Indischen Archipel gesammelten Dekapoden und Stomatopoden. In: Kukenthal W, ed. Ergebnisse einer zoologischen Forschungsreise in den Molukken und Borneo. Abhandlungen der Senckenberg Naturforschenden Gesellschaft 25: 467 - 929." type="journal article" year="1902">De Man, 1902: 561</bibRefCitation>
, pl. 20, fig. 17;
<bibRefCitation id="EFF84B4B8727BF3AFDBDFBEAFD35FBB0" author="Bott R" box="[527,693,1116,1137]" pageId="7" pageNumber="218" pagination="1 - 338" refId="ref5762" refString="Bott R. 1970. Die Susswasserkrabben von Europa, Asien, Australien und ihre Stammesgeschichte. Eine Revision der Potamoidea und Parathelphusoidea (Crustacea, Decapoda). Abhandlungen der Senckenberg Naturforschenden Gesellschaft, Frankfurt 526: 1 - 338." type="journal article" year="1970">Bott, 1970: 78</bibRefCitation>
, pl. 14, figs 6769;
<bibRefCitation id="EFF84B4B8727BF3AFE9AFBCCFD9EFB51" author="Cai Y &amp; Ng PKL" box="[296,542,1146,1168]" pageId="7" pageNumber="218" pagination="665 - 695" refId="ref5833" refString="Cai Y, Ng PKL. 2001. The freshwater decapod crustaceans of Halmahera, Indonesia. Journal of Crustacean Biology 21: 665 - 695." type="journal article" year="2001">Cai &amp; Ng, 2001: 686</bibRefCitation>
), and is not conspecific with the much smoother
<taxonomicName id="4C694D398727BF3AFE49FB2FFD2DFB6F" authority="Schubart &amp; Ng, 2008" authorityName="Schubart &amp; Ng" authorityYear="2008" box="[507,685,1177,1198]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="218" phylum="Arthropoda" rank="species" species="molluscivora" status="sp. nov.">
<emphasis id="B91DEAA88727BF3AFE49FB2FFD2DFB6F" box="[507,685,1177,1198]" italics="true" pageId="7" pageNumber="218">S. molluscivora</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E57D38727BF3AFD07FB2FFC8BFB6F" box="[693,779,1177,1198]" pageId="7" pageNumber="218" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BD636BA8727BF3AFF0EFB0EFC88F9FC" blockId="7.[164,779,198,1903]" pageId="7" pageNumber="218">
The molecular phylogeny places the new species
<taxonomicName id="4C694D398727BF3AFD41FB0EFEB6FB2A" authority="Schubart &amp; Ng, 2008" authorityName="Schubart &amp; Ng" authorityYear="2008" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="218" phylum="Arthropoda" rank="species" species="molluscivora" status="sp. nov.">
<emphasis id="B91DEAA88727BF3AFD41FB0EFEB6FB2A" italics="true" pageId="7" pageNumber="218">S. molluscivora</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E57D38727BF3AFE8EFB61FE11FB2D" box="[316,401,1239,1260]" pageId="7" pageNumber="218" rank="species">sp. nov.</taxonomicNameLabel>
next to an unidentified species of the genus, with high confidence values. This sister species was found in a small stream draining into Lake Poso, but could not be identified to species level, because only one juvenile individual was available, and thereby lacked diagnostic morphological characters. Also, the third representative of
<taxonomicName id="4C694D398727BF3AFDFDFA38FC8BFA62" authorityName="Bott" authorityYear="1969" box="[591,779,1422,1443]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="218" phylum="Arthropoda" rank="genus">
<emphasis id="B91DEAA88727BF3AFDFDFA38FC8BFA62" box="[591,779,1422,1443]" italics="true" pageId="7" pageNumber="218">Sundathelphusa</emphasis>
</taxonomicName>
in this study,
<taxonomicName id="4C694D398727BF3AFEFDFA1BFE79FA00" baseAuthorityName="Schenkel" baseAuthorityYear="1902" box="[335,505,1452,1474]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="218" phylum="Arthropoda" rank="species" species="minahassae">
<emphasis id="B91DEAA88727BF3AFEFDFA1BFE79FA00" box="[335,505,1452,1474]" italics="true" pageId="7" pageNumber="218">S. minahassae</emphasis>
</taxonomicName>
, belongs to this clade.
<taxonomicName id="4C694D398727BF3AFF16FA7DFE79FA21" baseAuthorityName="Schenkel" baseAuthorityYear="1902" box="[164,505,1483,1504]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="218" phylum="Arthropoda" rank="species" species="minahassae">
<emphasis id="B91DEAA88727BF3AFF16FA7DFE79FA21" box="[164,505,1483,1504]" italics="true" pageId="7" pageNumber="218">Sundathelphusa minahassae</emphasis>
</taxonomicName>
was collected in the northeastern part of
<collectingRegion id="49ADF8588727BF3AFE1EFA5CFD95F9C1" box="[428,533,1514,1536]" country="Indonesia" name="Sulawesi Utara" pageId="7" pageNumber="218">Sulawesi</collectingRegion>
, and the very long branches document pronounced genetic differences between the two species from the central highlands.
</paragraph>
<paragraph id="8BD636BA8727BF3AFF0EF9F3FC62FE97" blockId="7.[164,779,198,1903]" lastBlockId="7.[826,1441,198,342]" pageId="7" pageNumber="218">
The three species of
<taxonomicName id="4C694D398727BF3AFE17F9F3FDE1F99B" authorityName="Bott" authorityYear="1969" box="[421,609,1605,1626]" class="Malacostraca" family="Gecarcinucidae" genus="Sundathelphusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="218" phylum="Arthropoda" rank="genus">
<emphasis id="B91DEAA88727BF3AFE17F9F3FDE1F99B" box="[421,609,1605,1626]" italics="true" pageId="7" pageNumber="218">Sundathelphusa</emphasis>
</taxonomicName>
as a group are reciprocally monophyletic to all the other species of freshwater crabs from
<collectingRegion id="49ADF8588727BF3AFE74F935FDADF958" box="[454,557,1667,1689]" country="Indonesia" name="Sulawesi Utara" pageId="7" pageNumber="218">Sulawesi</collectingRegion>
belonging to the genera
<taxonomicName id="4C694D398727BF3AFEB3F917FDC6F977" authority=", Migmathelphsua" authorityName="Migmathelphsua" box="[257,582,1697,1719]" class="Malacostraca" family="Gecarcinucidae" genus="Syntripsa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="218" phylum="Arthropoda" rank="genus">
<emphasis id="B91DEAA88727BF3AFEB3F917FEF0F977" box="[257,368,1697,1718]" italics="true" pageId="7" pageNumber="218">Syntripsa</emphasis>
,
<emphasis id="B91DEAA88727BF3AFE37F917FDC6F977" box="[389,582,1697,1718]" italics="true" pageId="7" pageNumber="218">Migmathelphsua</emphasis>
</taxonomicName>
,
<taxonomicName id="4C694D398727BF3AFDEEF917FC84F977" box="[604,772,1697,1718]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="218" phylum="Arthropoda" rank="genus">
<emphasis id="B91DEAA88727BF3AFDEEF917FC84F977" box="[604,772,1697,1718]" italics="true" pageId="7" pageNumber="218">Parathelphusa</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C694D398727BF3AFF69F976FE22F914" authorityName="Balss" authorityYear="1933" box="[219,418,1728,1749]" class="Malacostraca" family="Gecarcinucidae" genus="Nautilothelphusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="218" phylum="Arthropoda" rank="genus">
<emphasis id="B91DEAA88727BF3AFF69F976FE22F914" box="[219,418,1728,1749]" italics="true" pageId="7" pageNumber="218">Nautilothelphusa</emphasis>
</taxonomicName>
. Within this latter group, the genus
<taxonomicName id="4C694D398727BF3AFF5CF969FEDDF935" box="[238,349,1759,1780]" class="Malacostraca" family="Gecarcinucidae" genus="Syntripsa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="218" phylum="Arthropoda" rank="genus">
<emphasis id="B91DEAA88727BF3AFF5CF969FEDDF935" box="[238,349,1759,1780]" italics="true" pageId="7" pageNumber="218">Syntripsa</emphasis>
</taxonomicName>
from the Malili Lakes holds the most basal position. The next split separates tributaries and the lake from the Poso system from the tributaries and lakes from the Malili system, giving the overall speciation pattern a clear hydrogeographic component. The genera
<taxonomicName id="4C694D398727BF3AFBEAFF70FB47FF1A" box="[1112,1223,198,219]" class="Malacostraca" family="Gecarcinucidae" genus="Syntripsa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="218" phylum="Arthropoda" rank="genus">
<emphasis id="B91DEAA88727BF3AFBEAFF70FB47FF1A" box="[1112,1223,198,219]" italics="true" pageId="7" pageNumber="218">Syntripsa</emphasis>
</taxonomicName>
,
<taxonomicName id="4C694D398727BF3AFB6BFF70FA1AFF1A" box="[1241,1434,198,219]" class="Malacostraca" family="Gecarcinucidae" genus="Migmathelphusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="218" phylum="Arthropoda" rank="genus">
<emphasis id="B91DEAA88727BF3AFB6BFF70FA1AFF1A" box="[1241,1434,198,219]" italics="true" pageId="7" pageNumber="218">Migmathelphusa</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C694D398727BF3AFCCCFF52FBC5FF38" authorityName="Balss" authorityYear="1933" box="[894,1093,228,249]" class="Malacostraca" family="Gecarcinucidae" genus="Nautilothelphusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="218" phylum="Arthropoda" rank="genus">
<emphasis id="B91DEAA88727BF3AFCCCFF52FBC5FF38" box="[894,1093,228,249]" italics="true" pageId="7" pageNumber="218">Nautilothelphusa</emphasis>
</taxonomicName>
exclusively comprise lake crabs, demonstrating from their phylogenetic position that the lakes must have been colonized several times independently.
</paragraph>
</subSubSection>
</treatment>
</document>