148 lines
18 KiB
XML
148 lines
18 KiB
XML
<document id="37BEDA7EF8F9F73A2226AA4E711F8799" ID-CLB-Dataset="33935" ID-DOI="http://dx.doi.org/10.3897/zookeys.131.1921" ID-GBIF-Dataset="72a81528-63ef-4531-9f9c-ade551a651d8" ID-PMC="PMC3208436" ID-Pensoft-Pub="1313-2970-131-1" ID-PubMed="22140329" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-131-1" ModsDocOrigin="ZooKeys 131" ModsDocTitle="Revision of the Southeast Asian millipede genus Orthomorpha Bollman, 1893, with the proposal of a new genus (Diplopoda, Polydesmida, Paradoxosomatidae)" checkinTime="1451249839150" checkinUser="pensoft" docAuthor="Likhitrakarn, Natdanai, Golovatch, Sergei I. & Panha, Somsak" docDate="2011" docId="33A2CF661A898C626E20DCC7D45C8122" docLanguage="en" docName="ZooKeys 131: 1-161" docOrigin="ZooKeys 131" docSource="http://dx.doi.org/10.3897/zookeys.131.1921" docTitle="Orthomorpha scabra Jeekel 1964" docType="treatment" docVersion="8" lastPageNumber="49" masterDocId="FFFF0F54FF88BD1C371C965D22453928" masterDocTitle="Revision of the Southeast Asian millipede genus Orthomorpha Bollman, 1893, with the proposal of a new genus (Diplopoda, Polydesmida, Paradoxosomatidae)" masterLastPageNumber="161" masterPageNumber="1" pageNumber="47" updateTime="1732923565725" updateUser="ExternalLinkService">
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<mods:title id="491FAC87863B5C5A6F18EBF484E7958A">Revision of the Southeast Asian millipede genus Orthomorpha Bollman, 1893, with the proposal of a new genus (Diplopoda, Polydesmida, Paradoxosomatidae)</mods:title>
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<mods:namePart id="ECFCB37202127E72837A6DC8C6A99152">Likhitrakarn, Natdanai</mods:namePart>
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<mods:namePart id="E24953AF5410E30AEF37F9FD4D6A89BA">Golovatch, Sergei I.</mods:namePart>
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<mods:namePart id="E977801D1038AF4F3B1B361464F55CCD">Panha, Somsak</mods:namePart>
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<mods:typeOfResource id="4CC197299DDFAC19D7A7B8AF4469DE98">text</mods:typeOfResource>
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<mods:title id="1D66327EC1FA38686CEF9F39EC23A1FF">ZooKeys</mods:title>
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<mods:date id="37EA3BDA9740DC3CEBEF502BAC6B3E0E">2011</mods:date>
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<mods:number id="CE71AAB82758898749203BA6E7983F63">131</mods:number>
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<mods:url id="E825922C6EDC3770CE2AE13653E327BA">http://dx.doi.org/10.3897/zookeys.131.1921</mods:url>
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<mods:classification id="C8487BF191F2E356B00FE661AB883895">journal article</mods:classification>
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<mods:identifier id="54201F6334081CE0440390C5A16D1BB6" type="DOI">http://dx.doi.org/10.3897/zookeys.131.1921</mods:identifier>
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<mods:identifier id="B1E3C597F743FA31A084EBE43E392A4A" type="Pensoft-Pub">1313-2970-131-1</mods:identifier>
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<treatment id="33A2CF661A898C626E20DCC7D45C8122" ID-GBIF-Taxon="152031147" LSID="urn:lsid:plazi:treatment:33A2CF661A898C626E20DCC7D45C8122" httpUri="http://treatment.plazi.org/id/33A2CF661A898C626E20DCC7D45C8122" lastPageId="48" lastPageNumber="49" pageId="46" pageNumber="47" scope_class="Diplopoda" scope_family="Paradoxosomatidae" scope_order="Polydesmida">
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<subSubSection id="F40F6B46EC18B34869844A52E9C58F05" pageId="46" pageNumber="47" type="nomenclature">
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<paragraph id="F8102769F88E997C053BC30E9D9BDCCF" pageId="46" pageNumber="47">
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<taxonomicName id="C366CBEF1FE92B1310EAD70F0A3E8FC9" ID-CoL="6T8NQ" LSID="http://species-id.net/wiki/Orthomorpha_scabra" authority="Jeekel, 1964" authorityName="Jeekel" authorityYear="1964" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha scabra" order="Polydesmida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="scabra">
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<pageBreakToken id="789C658A02426667CEB52899AABF2767" pageId="46" pageNumber="47" start="start">Orthomorpha</pageBreakToken>
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scabra Jeekel, 1964
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</taxonomicName>
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Figs 4142
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</paragraph>
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</subSubSection>
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<subSubSection id="CF0E176812FA709FABF78568D5293FF6" pageId="46" pageNumber="47" type="reference_group">
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<paragraph id="037C1F0F402BDBC3EE922709E59D9691" pageId="46" pageNumber="47">
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<taxonomicName id="9180E66F48A101564A2206D39C5E98DF" class="Diplopoda" family="Paradoxosomatidae" genus="Pratinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pratinus granosus" order="Polydesmida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="granosus">Pratinus granosus</taxonomicName>
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<bibRefCitation id="8967BC650DFD44F2B07E27B6CF4CCE3D" author="Attems, C" journalOrPublisher="Memoires du Museum national d'Histoire naturelle, Serie A, Zoologie" pageId="114" pageNumber="115" pagination="133 - 230" title="Myriopoden von Indochina. Expedition von Dr. C. Dawydoff (1938 - 1939)." volume="5" year="1953">Attems 1953</bibRefCitation>
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: 166 (D).
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</paragraph>
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<paragraph id="2D580BCBEE3A39691D6A127A84CAA6F3" pageId="46" pageNumber="47">
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<taxonomicName id="F0D85E4FF433F09EE3751674AD49BE9A" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha granosa" order="Polydesmida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="granosa">Orthomorpha granosa</taxonomicName>
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-
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<bibRefCitation id="A441EE4D7F6837B92F4D67D0867DC153" author="Jeekel, CAW" journalOrPublisher="Tijdschrift voor Entomologie" pageId="114" pageNumber="115" pagination="205 - 283" title="Paradoxosomatidae from Borneo (Diplopoda, Polydesmida)." volume="106" year="1963">Jeekel 1963</bibRefCitation>
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: 265 (M).
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</paragraph>
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<paragraph id="B133B4C597EA7CA44687A8737D4D50D4" pageId="46" pageNumber="47">
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<taxonomicName id="8CF3206B638559E746DAA871A36C864C" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha scabra" order="Polydesmida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="scabra">Orthomorpha scabra</taxonomicName>
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<bibRefCitation id="C5C0E2BF711905CF6ACC6B100DB5EE13" author="Jeekel, CAW" journalOrPublisher="Tijdschrift voor Entomologie" pageId="114" pageNumber="115" pagination="355 - 364" title="A new species of Orthomorpha Bollman from Thailand observed in migration, with taxonomic notes on the genus (Diplopoda)." volume="107" year="1964">Jeekel 1964</bibRefCitation>
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: 361 (N, D).
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</paragraph>
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<paragraph id="501D803FCF2821EE18D8BD65B0301A37" pageId="46" pageNumber="47">
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<taxonomicName id="F91190B2229BA7F04207EB888CCD28D6" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha scabra" order="Polydesmida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="scabra">Orthomorpha scabra</taxonomicName>
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-
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<bibRefCitation id="2252DC7D32B4F0BDD4F5615249D8DB4E" author="Jeekel, CAW" journalOrPublisher="Academisch Proefschrift, Rotterdam" pageId="114" pageNumber="115" title="On the classification and geographical distribution of the family Paradoxosomatidae (Diplopoda, Polydesmida)." year="1968">Jeekel 1968</bibRefCitation>
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: 56 (M);
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<bibRefCitation id="2D71C40F9F3B12FA4A5743BA851A1B48" author="Hoffman, RL" journalOrPublisher="Revue suisse de Zoologie" pageId="114" pageNumber="115" pagination="699 - 719" title="Diplopoda from Malayan caves, collected by M. Pierre Strinati." volume="84" year="1977">Hoffman 1977</bibRefCitation>
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: 700 (M);
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<bibRefCitation id="3AEF9E6BB5DC15D28FEB4FBB78B20A88" pageId="46" pageNumber="47">Golovatch 1998</bibRefCitation>
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: 42 (M, D).
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</paragraph>
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</subSubSection>
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<subSubSection id="224CFBB69BB8B51BD6DF8BDBC6A2FAA1" pageId="46" pageNumber="47" type="lectotype">
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<paragraph id="C807F8E24030B3D00AAD99388010E972" pageId="46" pageNumber="47">Lectotype.</paragraph>
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<paragraph id="36961171CDD7F9D7F56DB134BD385F7B" pageId="46" pageNumber="47">
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♂ of
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<taxonomicName id="1F4C1C418EA9FA2156919CAF5B17CD09" class="Diplopoda" family="Paradoxosomatidae" genus="Pratinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pratinus granosus" order="Polydesmida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="granosus">Pratinus granosus</taxonomicName>
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(NHMW-3516), Vietnam, Lamdong Prov., Peak Lang Biang, 1938-1939, leg. C. Dawydoff.
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</paragraph>
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<paragraph id="A839607578661E1F712631D9B3C6C32E" pageId="46" pageNumber="47">
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Lectotype designation proposed herewith is necessary to ensure the species is based on a complete male coming from a certain locality, because (1)
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<bibRefCitation id="ABFBEF6CD62F3386D67725926689A597" author="Attems, C" journalOrPublisher="Memoires du Museum national d'Histoire naturelle, Serie A, Zoologie" pageId="114" pageNumber="115" pagination="133 - 230" title="Myriopoden von Indochina. Expedition von Dr. C. Dawydoff (1938 - 1939)." volume="5" year="1953">Attems (1953)</bibRefCitation>
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had provided no information on the number of syntypes, and (2) he stated their provenance to have been both from Xieng Kunang, Luang Prabang Province, Laos and Peak Lang Biang, Lamdong Province, Vietnam.
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</paragraph>
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</subSubSection>
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<subSubSection id="D080B0D2C44B196E72323D658497B22A" lastPageId="48" lastPageNumber="49" pageId="46" pageNumber="47" type="redescription">
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<paragraph id="19C3CDC0732729BA1CF82F13B78C7B88" pageId="46" pageNumber="47">Redescription.</paragraph>
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<paragraph id="DD7A67F4668A1BE33A65BAB3369E3350" pageId="46" pageNumber="47">
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Length 41 mm, width of midbody pro- and metazona 3.1 and 4.4 mm, respectively (vs 38 mm in length and 4.6 mm in width, as given in the original description (
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<bibRefCitation id="3CB4CA1625F95E07278BCC659F756147" author="Attems, C" journalOrPublisher="Memoires du Museum national d'Histoire naturelle, Serie A, Zoologie" pageId="114" pageNumber="115" pagination="133 - 230" title="Myriopoden von Indochina. Expedition von Dr. C. Dawydoff (1938 - 1939)." volume="5" year="1953">Attems 1953</bibRefCitation>
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)). Coloration of alcohol material upon long-term preservation rather uniformly light brown (Fig. 41) (vs black-brown with rather contrasting light brown to yellow-brown prozona, paraterga and venter, as well as red-brown legs, as given in the original description (
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<bibRefCitation id="886E7E1A135221597EE4B29907C4E44C" author="Attems, C" journalOrPublisher="Memoires du Museum national d'Histoire naturelle, Serie A, Zoologie" pageId="114" pageNumber="115" pagination="133 - 230" title="Myriopoden von Indochina. Expedition von Dr. C. Dawydoff (1938 - 1939)." volume="5" year="1953">Attems 1953</bibRefCitation>
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)).
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</paragraph>
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<paragraph id="3622F6B35C9C96DE76B2B5EFDC925A1A" lastPageId="48" lastPageNumber="49" pageId="46" pageNumber="47">
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Head usual, clypeolabral region sparsely setose, surface of vertex smooth, epicranial suture very distinct. Antennae rather long and slender (Fig. 41A, B & J), extending behind body segment 3 dorsally. Head in width <collum <segments 3 and 4 <segment 2 <segments 5-17, gently and gradually tapering thereafter. Collum with three transverse rows of setae, 4+4 anterior, 2+2 intermediate, and 2+2 posterior setae, all borne on small knobs; caudal corner of paraterga subrectangular, nearly pointed (Fig. 41A & B). Tegument dull; metaterga coriaceous, rugulose, each postcollum one with two rows of evident tubercles, caudal row being more evident: 2+2 and 3+3 on metatergum 2, 3+2 and 4+3 on metatergum 3, 2
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<normalizedToken id="40F2A553D6268854A0113768D0885A58" originalValue="–3+2–">-3+2-</normalizedToken>
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4 until metatergum 17, 1+2 and 2+3 on metaterga 18 and 19; prozona very finely shagreened, surface below par
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<pageBreakToken id="493CB8D33E061982DED89FA164324AF0" pageId="47" pageNumber="48" start="start">aterga</pageBreakToken>
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microgranulate. Tergal setae abraded. Axial line visible, starting from metatergum 2. Paraterga very strongly developed (Fig. 41A-G), mostly slightly upturned and lying above dorsum, lying below dorsum only on segments 2 and 3, thin in lateral view, like blunt blades, a little thicker only on pore-bearing segments, on postcollum segments extending increasingly beyond rear tergal margin, all pointed. Calluses on paraterga 2 delimited by a sulcus only dorsally, on following paraterga both dorsally and ventrally, rather broad, especially so on pore-bearing segments. Paraterga 2 broad, anterior edge rounded, lateral edge with two small incisions in anterior half; posterior edge evidently concave (Fig. 41A & B). Paraterga 3 and 4 subequal, like subsequent paraterga, anterior edge broadly rounded, bordered and fused to callus, lateral edge with one small incision in anterior half. Ozopores evident, lateral, not sunken inside a groove, lying at about 1/3 of metazonital length. Transverse sulcus complete on metaterga 4-18, incomplete on metatergum 3, shallow, reaching bases of paraterga, smooth at bottom, slightly sinuate anteromedially (Fig. 41A, B, C, D & F). Stricture between pro- and metazona rather narrow, shallow, slightly ribbed at bottom down to base of paraterga. Pleurosternal carinae complete crests only on segments 2-4 (Fig. 41B, D & E), each
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<pageBreakToken id="5694324A0922427A8323FB8BF7F7B5F7" pageId="48" pageNumber="49" start="start">with</pageBreakToken>
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an evident sharp denticle caudally, thereafter increasingly strongly reduced until segment 11. Epiproct (Fig. 41E-G) conical, flattened dorsoventrally, apical papillae small; tip subtruncate; pre-apical papillae small, but visible, lying close to tip. Hypoproct (Fig. 41G) subtrapeziform, setiferous knobs at a subtruncate caudal margin small and well-separated.
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</paragraph>
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<paragraph id="86CCAA73A6482502F70F22C34A97DB8E" pageId="48" pageNumber="49">Sterna densely setose, without modifications, but with two large, rounded, fully separated, setose cones between ♂ coxae 4 (Fig. 41H & I). A paramedian pair of conspicuous, high tubercles in front of gonopod aperture. Legs long and slender, midbody ones ca 1.3-1.4 as long as body height, prefemora without modifications, tarsal brushes present until ♂ legs 5.</paragraph>
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<paragraph id="3CAEF9F2853D2E6E0FF45BBF50436261" pageId="48" pageNumber="49">
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Gonopods (Fig. 42) simple. Coxa long and slender, with several setae distodorsally. Prefemur densely setose, about 2 times shorter than femorite +
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<normalizedToken id="9E02089F2B2006A95D9B73C4C2A91809" originalValue="“postfemoral”">"postfemoral"</normalizedToken>
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part. Femorite very slender, evidently curved, slightly enlarged distad,
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<normalizedToken id="D33491364C43E7349067525828E7BC92" originalValue="“postfemoral”">"postfemoral"</normalizedToken>
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part demarcated by an oblique lateral sulcus; tip of solenophore small, trifid, with two subequal denticles (one terminal, the other middle) and a broader subterminal lobule.
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</paragraph>
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<caption id="4476F69131FF678892AF9D1F9D753453" pageId="48" pageNumber="49">
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<paragraph id="B118C02102CF1449D9BFBFF5516EB042" pageId="48" pageNumber="49">
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Figure 41.
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<taxonomicName id="CC6E5B809468EEE678D62EB8E16C0D9C" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha scabra" order="Polydesmida" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="scabra">Orthomorpha scabra</taxonomicName>
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Jeekel, 1964, ♂ lectotype. A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-F posterior part of body, lateral, dorsal and ventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively.
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</paragraph>
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</caption>
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<caption id="BF6A48A4096F7B2B22D55DB759910EC5" pageId="48" pageNumber="49">
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<paragraph id="D72C00C842172F31574D9A2EE8805F56" pageId="48" pageNumber="49">
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Figure 42.
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<taxonomicName id="8C17899E80F56E619C60FCE7CAAF3B2E" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha scabra" order="Polydesmida" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="scabra">Orthomorpha scabra</taxonomicName>
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Jeekel, 1964, ♂ lectotype. A, B right gonopod, lateral and mesal views, respectively.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="AFD68AA3320B93D34A2C8D75EA683B55" pageId="48" pageNumber="49" type="remark">
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<paragraph id="6754917DC90688FB2C92E2FCC98265B8" pageId="48" pageNumber="49">Remark.</paragraph>
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<paragraph id="D5AB7272EBB0A5AB8088CF9489D2DCB0" pageId="48" pageNumber="49">
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This species was renamed to avoid homonymy (
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<bibRefCitation id="9D04F1F45D98D66C11CCE33B2945CB6A" author="Jeekel, CAW" journalOrPublisher="Tijdschrift voor Entomologie" pageId="114" pageNumber="115" pagination="355 - 364" title="A new species of Orthomorpha Bollman from Thailand observed in migration, with taxonomic notes on the genus (Diplopoda)." volume="107" year="1964">Jeekel 1964</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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||
</document> |