treatments-xml/data/03/C1/1D/03C11D43D31CFFECFF7F44F99B535655.xml
2024-06-21 12:22:17 +02:00

317 lines
42 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="A6E03FC8901FAD5F35DED59E2A0654D3" ID-DOI="10.5281/zenodo.196223" ID-GBIF-Dataset="648631c9-8b76-4f0f-85a4-d808b6fbfe70" ID-ISSN="1175-5326" ID-Zenodo-Dep="196223" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460240568931" checkinUser="plazi" docAuthor="Whittington, Ian D." docDate="2010" docId="03C11D43D31CFFECFF7F44F99B535655" docLanguage="en" docName="zt02519p030.pdf" docOrigin="Zootaxa 2519" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Benedeniella Johnston 1929" docType="treatment" docVersion="8" lastPageNumber="6" masterDocId="FFF8653BD318FFE9FFE847349A135202" masterDocTitle="Revision of Benedeniella Johnston, 1929 (Monogenea: Capsalidae), its assignment to Entobdellinae Bychowsky, 1957 and comments on subfamilial composition" masterLastPageNumber="30" masterPageNumber="1" pageNumber="5" updateTime="1698240612236" updateUser="plazi">
<mods:mods id="C8A2D6179E92094129A7167B819F4FD0" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="8235E296509732E5E7AA3F3A79AC4549">
<mods:title id="9171C39C819A995DE48F545516ACF323">Revision of Benedeniella Johnston, 1929 (Monogenea: Capsalidae), its assignment to Entobdellinae Bychowsky, 1957 and comments on subfamilial composition</mods:title>
</mods:titleInfo>
<mods:name id="64CA6B49614525613412DAEF3F8F1296" type="personal">
<mods:role id="20AD92B69E88BC27B6FD08CA8DBA1319">
<mods:roleTerm id="4D38CC776FB27046543EDF5E2EEF2CA8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3933CB3141055EC7FDA1C2248F09479D">Whittington, Ian D.</mods:namePart>
</mods:name>
<mods:typeOfResource id="205D3976D65718C915E7642976F0FEA4">text</mods:typeOfResource>
<mods:relatedItem id="0C655015D7CA645C0B2E7D68ED324D8F" type="host">
<mods:titleInfo id="3935F08B14AC11689DEA94A06C2B5609">
<mods:title id="7E54695B42FB99DA59E24E97F0AD0163">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="E08D325FA528465A3BF41FEDFB792D0A">
<mods:date id="F82FDF66CB0B9B4C5C92F30ADF197D5F">2010</mods:date>
<mods:detail id="497A3FB69638E3E3BC3B4FC49AE4E8F9" type="volume">
<mods:number id="06162138C50ACB7AFC5AD6BB132B8C86">2519</mods:number>
</mods:detail>
<mods:extent id="EFB6766E8B29178CE4586E99ADE25DE4" unit="page">
<mods:start id="3D33AE772962BEEF0FD155A33A56E31C">1</mods:start>
<mods:end id="74C4E6BF0A01CD3A0824665C91A92851">30</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="272D3E29A218BB6858DE42148D2CCDDC">journal article</mods:classification>
<mods:identifier id="58F0F9BF3B572931E3B7C0B4C6D42E1D" type="DOI">10.5281/zenodo.196223</mods:identifier>
<mods:identifier id="2FF25B41D56F2CA5D61AF918176948C1" type="GBIF-Dataset">648631c9-8b76-4f0f-85a4-d808b6fbfe70</mods:identifier>
<mods:identifier id="2D1C022A8111BFF7277571C3ED1F577E" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="D2D26FE6F20D2F9D74C88DD364BAAE85" type="Zenodo-Dep">196223</mods:identifier>
</mods:mods>
<treatment id="03C11D43D31CFFECFF7F44F99B535655" ID-DOI="http://doi.org/10.5281/zenodo.6207318" ID-GBIF-Taxon="119389646" ID-Zenodo-Dep="6207318" LSID="urn:lsid:plazi:treatment:03C11D43D31CFFECFF7F44F99B535655" httpUri="http://treatment.plazi.org/id/03C11D43D31CFFECFF7F44F99B535655" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
<subSubSection id="C372FFDED31CFFEDFF7F44F99F485438" pageId="4" pageNumber="5" type="nomenclature">
<paragraph id="8BD7AC55D31CFFEDFF7F44F99BE451E5" blockId="4.[151,503,973,999]" box="[151,503,973,999]" pageId="4" pageNumber="5">
<heading id="D09F1B39D31CFFEDFF7F44F99BE451E5" bold="true" box="[151,503,973,999]" fontSize="11" level="1" pageId="4" pageNumber="5" reason="1">
<taxonomicName id="4C68D7D6D31CFFEDFF7F44F99BE451E5" ID-CoL="7NQN7" authority="Johnston, 1929" authorityName="Johnston" authorityYear="1929" box="[151,503,973,999]" class="Monogenea" family="Capsalidae" genus="Benedeniella" kingdom="Animalia" order="Capsalidea" pageId="4" pageNumber="5" phylum="Platyhelminthes" rank="genus">
<emphasis id="B91C7047D31CFFEDFF7F44F99BE451E5" bold="true" box="[151,503,973,999]" pageId="4" pageNumber="5">
<emphasis id="B91C7047D31CFFEDFF7F44F99B2651E5" bold="true" box="[151,309,973,999]" italics="true" pageId="4" pageNumber="5">Benedeniella</emphasis>
<bibRefCitation id="EFF9D1A4D31CFFEDFED444F99BE451E5" author="Johnston" box="[316,503,973,999]" pageId="4" pageNumber="28" refString="Johnston, T. H. (1929) Remarks on the synonymy of certain tristomatid trematode genera. Transactions of the Royal Society of South Australia, 53, 71 - 78." type="journal article" year="1929">Johnston, 1929</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BD7AC55D31CFFEDFF7F43219F485438" blockId="4.[151,1437,1045,2004]" pageId="4" pageNumber="5">
<emphasis id="B91C7047D31CFFEDFF7F43219816562D" bold="true" box="[151,517,1045,1071]" pageId="4" pageNumber="5">Amended generic diagnosis:</emphasis>
With features of the
<taxonomicName id="4C68D7D6D31CFFEDFCE443219EA5562D" authority="Yamaguti (1963)" authorityName="Yamaguti" authorityYear="1963" box="[780,1206,1045,1071]" class="Monogenea" family="Capsalidae" kingdom="Animalia" order="Capsalidea" pageId="4" pageNumber="5" phylum="Platyhelminthes" rank="family">
Capsalidae
<emphasis id="B91C7047D31CFFEDFC73432199CC562C" box="[923,991,1045,1070]" italics="true" pageId="4" pageNumber="5">sensu</emphasis>
<bibRefCitation id="EFF9D1A4D31CFFEDFC0E43219EA5562D" author="Yamaguti" box="[998,1206,1045,1071]" pageId="4" pageNumber="30" refString="Yamaguti, S. (1963) Systema Helminthum. Volume IV. Monogenea and Aspidocotylea. InterScience Publishers, New York, USA, 699 pp." type="book" year="1963">Yamaguti (1963)</bibRefCitation>
</taxonomicName>
and Entobdellinae (amended above). Median haptoral sclerites large. Ventral haptor surface apapillate. Marginal valve conspicuous, wider anteriorly. Tendons entering haptor from body musculature conspicuous; passing through notch at proximal end of accessory sclerites before dividing, with part of tendon attaching to proximal ends of anterior hamuli and part attaching to ventral haptor tissue. Anterior attachment organs comprising ventral pair of circular, saucer-like discs, one on each ventrolateral border, bearing series of narrow complete and some incomplete grooves anterolaterally. Dorsal protuberances present, in anterior to posterior succession, in form of horn-like papillae near junction of anterior attachment organs with body, excretory papillae and conical structures at peduncle where body joins haptor. Glands of Goto absent.
<collectingCountry id="F37FECC5D31CFFEDFC0242749E355758" box="[1002,1062,1344,1370]" name="Malta" pageId="4" pageNumber="5">Male</collectingCountry>
copulatory organ a long cirrus. Cirrus sac elongate with muscular walls containing single male accessory gland reservoir. Long terminal male genital duct containing cirrus separate from long uterus for most of its length, uniting close to common genital pore, opening dorsally near left body margin at level of posterior region of left anterior attachment organ. Vagina long; vaginal pore opening dorsally near left body margin at different levels (anteriorly in
<taxonomicName id="4C68D7D6D31CFFEDFA6A42E19B395416" class="Monogenea" family="Capsalidae" genus="Benedeniella" kingdom="Animalia" order="Capsalidea" pageId="4" pageNumber="5" phylum="Platyhelminthes" rank="species" species="macrocolpa">
<emphasis id="B91C7047D31CFFEDFA6A42E19B395416" italics="true" pageId="4" pageNumber="5">B. macrocolpa</emphasis>
</taxonomicName>
; posteriorly in
<taxonomicName id="4C68D7D6D31CFFEDFE0642CF98A75416" box="[494,692,1531,1556]" class="Monogenea" family="Capsalidae" genus="Benedeniella" kingdom="Animalia" order="Capsalidea" pageId="4" pageNumber="5" phylum="Platyhelminthes" rank="species" species="posterocolpa">
<emphasis id="B91C7047D31CFFEDFE0642CF98A75416" box="[494,692,1531,1556]" italics="true" pageId="4" pageNumber="5">B. posterocolpa</emphasis>
</taxonomicName>
). Proximal region of vagina unspecialised, communicating directly with dorsal part of vitelline reservoir. Parasites of ventral skin of myliobatid elasmobranch rays.
</paragraph>
</subSubSection>
<subSubSection id="C372FFDED31CFFEDFF2D41719BCD54F6" pageId="4" pageNumber="5" type="materials_examined">
<paragraph id="8BD7AC55D31CFFEDFF2D41719EEC54A8" blockId="4.[151,1437,1045,2004]" pageId="4" pageNumber="5">
<emphasis id="B91C7047D31CFFEDFF2D41719B7A545D" bold="true" box="[197,361,1605,1631]" pageId="4" pageNumber="5">
<typeStatus id="54D312F7D31CFFEDFF2D41719B11545D" box="[197,258,1605,1631]" pageId="4" pageNumber="5">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="4C68D7D6D31CFFEDFE98417199EE545D" authority="(Luhe, 1906) Yamaguti, 1963" authorityName="(Luhe, 1906) Yamaguti" authorityYear="1963" box="[368,1021,1605,1631]" class="Monogenea" family="Capsalidae" genus="Benedeniella" kingdom="Animalia" order="Capsalidea" pageId="4" pageNumber="5" phylum="Platyhelminthes" rank="species" species="macrocolpa">
<emphasis id="B91C7047D31CFFEDFE984171988F545C" box="[368,668,1605,1630]" italics="true" pageId="4" pageNumber="5">Benedeniella macrocolpa</emphasis>
(
<bibRefCitation id="EFF9D1A4D31CFFEDFD4341719920545D" author="Luhe" box="[683,819,1605,1631]" pageId="4" pageNumber="29" refString="Luhe, M. (1906) Report on the trematode parasites from the marine fishes of Ceylon. Report to the Government of Ceylon on the Pearl Oyster Fisheries of the Gulf of Manaar, Supplementary Report, 5, 97 - 108." type="journal article" year="1906">Lühe, 1906</bibRefCitation>
)
<bibRefCitation id="EFF9D1A4D31CFFEDFCAB417199EE545D" author="Yamaguti" box="[835,1021,1605,1631]" pageId="4" pageNumber="30" refString="Yamaguti, S. (1963) Systema Helminthum. Volume IV. Monogenea and Aspidocotylea. InterScience Publishers, New York, USA, 699 pp." type="book" year="1963">Yamaguti, 1963</bibRefCitation>
</taxonomicName>
[synonyms:
<taxonomicName id="4C68D7D6D31CFFEDFB7C41719BAA5486" authority="Luhe, 1906" authorityName="Luhe" authorityYear="1906" class="Monogenea" family="Capsalidae" genus="Epibdella" higherTaxonomySource="CoL" kingdom="Animalia" order="Capsalidea" pageId="4" pageNumber="5" phylum="Platyhelminthes" rank="species" species="macrocolpa" subGenus="Benedenia">
<emphasis id="B91C7047D31CFFEDFB7C41719F15545C" box="[1172,1286,1605,1630]" italics="true" pageId="4" pageNumber="5">Epibdella</emphasis>
(
<emphasis id="B91C7047D31CFFEDFAFE41719F81545C" box="[1302,1426,1605,1630]" italics="true" pageId="4" pageNumber="5">Benedenia</emphasis>
)
<emphasis id="B91C7047D31CFFEDFF7F415F9B375486" box="[151,292,1643,1668]" italics="true" pageId="4" pageNumber="5">macrocolpa</emphasis>
<bibRefCitation id="EFF9D1A4D31CFFEDFEC4415E9BAA5486" author="Luhe" box="[300,441,1642,1668]" pageId="4" pageNumber="29" refString="Luhe, M. (1906) Report on the trematode parasites from the marine fishes of Ceylon. Report to the Government of Ceylon on the Pearl Oyster Fisheries of the Gulf of Manaar, Supplementary Report, 5, 97 - 108." type="journal article" year="1906">Lühe, 1906</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C68D7D6D31CFFEDFE2E415F9E4A5486" authority="(Luhe, 1906) MacCallum, 1927" authorityName="(Luhe, 1906) MacCallum" authorityYear="1927" box="[454,1113,1642,1668]" class="Monogenea" family="Capsalidae" genus="Epibdella" higherTaxonomySource="CoL" kingdom="Animalia" order="Capsalidea" pageId="4" pageNumber="5" phylum="Platyhelminthes" rank="species" species="macrocolpa">
<emphasis id="B91C7047D31CFFEDFE2E415F98C35486" box="[454,720,1643,1668]" italics="true" pageId="4" pageNumber="5">Epibdella macrocolpa</emphasis>
(
<bibRefCitation id="EFF9D1A4D31CFFEDFD08415E997F5486" author="Luhe" box="[736,876,1642,1668]" pageId="4" pageNumber="29" refString="Luhe, M. (1906) Report on the trematode parasites from the marine fishes of Ceylon. Report to the Government of Ceylon on the Pearl Oyster Fisheries of the Gulf of Manaar, Supplementary Report, 5, 97 - 108." type="journal article" year="1906">Lühe, 1906</bibRefCitation>
)
<bibRefCitation id="EFF9D1A4D31CFFEDFC95415E9E4A5486" author="MacCallum" box="[893,1113,1642,1668]" pageId="4" pageNumber="29" refString="MacCallum, G. A. (1927) A new ectoparasitic trematode Epibdella melleni, sp. nov. Zoopathologica, 1, 291 - 300." type="journal article" year="1927">MacCallum, 1927</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C68D7D6D31CFFEDFB8E415F986D54A8" authority="(Luhe, 1906) Johnston, 1929" authorityName="(Luhe, 1906) Johnston" authorityYear="1929" class="Monogenea" family="Capsalidae" genus="Benedenia" kingdom="Animalia" order="Capsalidea" pageId="4" pageNumber="5" phylum="Platyhelminthes" rank="species" species="macrocolpa" subGenus="Benedeniella">
<emphasis id="B91C7047D31CFFEDFB8E415F9EF75486" box="[1126,1252,1643,1668]" italics="true" pageId="4" pageNumber="5">Benedenia</emphasis>
(
<emphasis id="B91C7047D31CFFEDFB1E415F9F815486" box="[1270,1426,1643,1668]" italics="true" pageId="4" pageNumber="5">Benedeniella</emphasis>
)
<emphasis id="B91C7047D31CFFEDFF7F41A49B3154AB" box="[151,290,1680,1705]" italics="true" pageId="4" pageNumber="5">macrocolpa</emphasis>
(
<bibRefCitation id="EFF9D1A4D31CFFEDFED941A49BAA54A8" author="Luhe" box="[305,441,1680,1706]" pageId="4" pageNumber="29" refString="Luhe, M. (1906) Report on the trematode parasites from the marine fishes of Ceylon. Report to the Government of Ceylon on the Pearl Oyster Fisheries of the Gulf of Manaar, Supplementary Report, 5, 97 - 108." type="journal article" year="1906">Lühe, 1906</bibRefCitation>
)
<bibRefCitation id="EFF9D1A4D31CFFEDFE2141A4986D54A8" author="Johnston" box="[457,638,1680,1706]" pageId="4" pageNumber="28" refString="Johnston, T. H. (1929) Remarks on the synonymy of certain tristomatid trematode genera. Transactions of the Royal Society of South Australia, 53, 71 - 78." type="journal article" year="1929">Johnston, 1929</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C68D7D6D31CFFEDFD6241A49EE054A8" authority="(Luhe, 1906) Meserve, 1938" authorityName="(Luhe, 1906) Meserve" authorityYear="1938" box="[650,1267,1680,1706]" class="Monogenea" family="Capsalidae" genus="Benedenia" kingdom="Animalia" order="Capsalidea" pageId="4" pageNumber="5" phylum="Platyhelminthes" rank="species" species="macrocolpa">
<emphasis id="B91C7047D31CFFEDFD6241A4998B54AB" box="[650,920,1680,1705]" italics="true" pageId="4" pageNumber="5">Benedenia macrocolpa</emphasis>
(
<bibRefCitation id="EFF9D1A4D31CFFEDFC4041A49E2354A8" author="Luhe" box="[936,1072,1680,1706]" pageId="4" pageNumber="29" refString="Luhe, M. (1906) Report on the trematode parasites from the marine fishes of Ceylon. Report to the Government of Ceylon on the Pearl Oyster Fisheries of the Gulf of Manaar, Supplementary Report, 5, 97 - 108." type="journal article" year="1906">Lühe, 1906</bibRefCitation>
)
<bibRefCitation id="EFF9D1A4D31CFFEDFBA841A49EE054A8" author="Meserve" box="[1088,1267,1680,1706]" pageId="4" pageNumber="29" refString="Meserve, F. G. (1938) Some monogenetic trematodes from the Galapagos Islands and the neighbouring Pacific. Reports of the Allan Hancock Pacific Expeditions (1932 - 1937), 2, 31 - 89." type="journal article" year="1938">Meserve, 1938</bibRefCitation>
</taxonomicName>
].
</paragraph>
<paragraph id="8BD7AC55D31CFFEDFF2D41819BCD54F6" blockId="4.[151,1437,1045,2004]" pageId="4" pageNumber="5">
<emphasis id="B91C7047D31CFFEDFF2D41819B9B54CD" bold="true" box="[197,392,1717,1743]" pageId="4" pageNumber="5">Other species:</emphasis>
<taxonomicName id="4C68D7D6D31CFFEDFE7941819E6E54CD" authority="(Hargis, 1955) Yamaguti, 1963" authorityName="(Hargis, 1955) Yamaguti" authorityYear="1963" box="[401,1149,1717,1743]" class="Monogenea" family="Capsalidae" genus="Benedeniella" kingdom="Animalia" order="Capsalidea" pageId="4" pageNumber="5" phylum="Platyhelminthes" rank="species" species="posterocolpa">
<emphasis id="B91C7047D31CFFEDFE79418198F654CC" box="[401,741,1717,1742]" italics="true" pageId="4" pageNumber="5">Benedeniella posterocolpa</emphasis>
(
<bibRefCitation id="EFF9D1A4D31CFFEDFD10418199B254CD" author="Hargis" box="[760,929,1717,1743]" pageId="4" pageNumber="28" refString="Hargis, W. J. Jr. (1955) Monogenetic trematodes of Gulf of Mexico fishes. Part V. The superfamily Capsaloidea. Transactions of the American Microscopical Society, 74, 203 - 225." type="journal article" year="1955">Hargis, 1955</bibRefCitation>
)
<bibRefCitation id="EFF9D1A4D31CFFEDFC5B41819E6E54CD" author="Yamaguti" box="[947,1149,1717,1743]" pageId="4" pageNumber="30" refString="Yamaguti, S. (1963) Systema Helminthum. Volume IV. Monogenea and Aspidocotylea. InterScience Publishers, New York, USA, 699 pp." type="book" year="1963">Yamaguti, 1963</bibRefCitation>
</taxonomicName>
[synonym:
<taxonomicName id="4C68D7D6D31CFFEDFAFF41819BC154F6" authority="Hargis, 1955" authorityName="Hargis" authorityYear="1955" class="Monogenea" family="Capsalidae" genus="Benedenia" kingdom="Animalia" order="Capsalidea" pageId="4" pageNumber="5" phylum="Platyhelminthes" rank="species" species="posterocolpa">
<emphasis id="B91C7047D31CFFEDFAFF41819B3C54F6" italics="true" pageId="4" pageNumber="5">Benedenia posterocolpa</emphasis>
<bibRefCitation id="EFF9D1A4D31CFFEDFEDE41EE9BC154F6" author="Hargis" box="[310,466,1754,1780]" pageId="4" pageNumber="28" refString="Hargis, W. J. Jr. (1955) Monogenetic trematodes of Gulf of Mexico fishes. Part V. The superfamily Capsaloidea. Transactions of the American Microscopical Society, 74, 203 - 225." type="journal article" year="1955">Hargis, 1955</bibRefCitation>
</taxonomicName>
].
</paragraph>
</subSubSection>
<subSubSection id="C372FFDED31CFFECFF2E41CB9F9D5150" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" type="discussion">
<paragraph id="8BD7AC55D31CFFECFF2E41CB9F9D5150" blockId="4.[151,1437,1045,2004]" lastBlockId="5.[151,1438,152,1111]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
<emphasis id="B91C7047D31CFFEDFF2E41CB9B51551B" bold="true" box="[198,322,1791,1817]" pageId="4" pageNumber="5">Remarks:</emphasis>
<bibRefCitation id="EFF9D1A4D31CFFEDFEB8403498075518" author="Johnston" box="[336,532,1792,1818]" pageId="4" pageNumber="28" refString="Johnston, T. H. (1929) Remarks on the synonymy of certain tristomatid trematode genera. Transactions of the Royal Society of South Australia, 53, 71 - 78." type="journal article" year="1929">Johnston (1929)</bibRefCitation>
presented a history and examined the validity of some tristomatid (= capsalid) genera and divided
<taxonomicName id="4C68D7D6D31CFFEDFE1E4011986A553C" box="[502,633,1829,1854]" class="Monogenea" family="Capsalidae" genus="Entobdella" kingdom="Animalia" order="Capsalidea" pageId="4" pageNumber="5" phylum="Platyhelminthes" rank="genus">
<emphasis id="B91C7047D31CFFEDFE1E4011986A553C" box="[502,633,1829,1854]" italics="true" pageId="4" pageNumber="5">Entobdella</emphasis>
</taxonomicName>
and
<taxonomicName id="4C68D7D6D31CFFEDFD5C40119922553C" box="[692,817,1829,1854]" class="Monogenea" family="Capsalidae" genus="Benedenia" kingdom="Animalia" order="Capsalidea" pageId="4" pageNumber="5" phylum="Platyhelminthes" rank="genus">
<emphasis id="B91C7047D31CFFEDFD5C40119922553C" box="[692,817,1829,1854]" italics="true" pageId="4" pageNumber="5">Benedenia</emphasis>
</taxonomicName>
into subgenera based on the position of the vaginal pore in relation to the common genital pore. The subgenus
<taxonomicName id="4C68D7D6D31CFFEDFC64407F9E225566" box="[908,1073,1867,1892]" class="Monogenea" family="Capsalidae" genus="Benedeniella" kingdom="Animalia" order="Capsalidea" pageId="4" pageNumber="5" phylum="Platyhelminthes" rank="genus">
<emphasis id="B91C7047D31CFFEDFC64407F9E225566" box="[908,1073,1867,1892]" italics="true" pageId="4" pageNumber="5">Benedeniella</emphasis>
</taxonomicName>
was proposed for species of
<taxonomicName id="4C68D7D6D31CFFEDFF7F40449B09558B" box="[151,282,1904,1929]" class="Monogenea" family="Capsalidae" genus="Benedenia" kingdom="Animalia" order="Capsalidea" pageId="4" pageNumber="5" phylum="Platyhelminthes" rank="genus">
<emphasis id="B91C7047D31CFFEDFF7F40449B09558B" box="[151,282,1904,1929]" italics="true" pageId="4" pageNumber="5">Benedenia</emphasis>
</taxonomicName>
in which the vagina opened beside the common genital pore but where the vagina travelled posteriorly behind the testes and then anteriorly to the vitelline reservoir.
<bibRefCitation id="EFF9D1A4D31CFFEDFC1F40A19EA655AD" author="Johnston" box="[1015,1205,1941,1967]" pageId="4" pageNumber="28" refString="Johnston, T. H. (1929) Remarks on the synonymy of certain tristomatid trematode genera. Transactions of the Royal Society of South Australia, 53, 71 - 78." type="journal article" year="1929">Johnston (1929)</bibRefCitation>
included
<taxonomicName id="4C68D7D6D31CFFEDFAC240A1984C55D6" authority="Luhe, 1906" authorityName="Luhe" authorityYear="1906" class="Monogenea" family="Capsalidae" genus="Epibdella" higherTaxonomySource="CoL" kingdom="Animalia" order="Capsalidea" pageId="4" pageNumber="5" phylum="Platyhelminthes" rank="species" species="macrocolpa" subGenus="Benedenia">
<emphasis id="B91C7047D31CFFEDFAC240A19F8F55AC" box="[1322,1436,1941,1966]" italics="true" pageId="4" pageNumber="5">Epibdella</emphasis>
(
<emphasis id="B91C7047D31CFFEDFF49408F9B3055D6" box="[161,291,1979,2004]" italics="true" pageId="4" pageNumber="5">Benedenia</emphasis>
)
<emphasis id="B91C7047D31CFFEDFEDF408F9BDA55D6" box="[311,457,1979,2004]" italics="true" pageId="4" pageNumber="5">macrocolpa</emphasis>
<bibRefCitation id="EFF9D1A4D31CFFEDFE3A408E984C55D6" author="Luhe" box="[466,607,1978,2004]" pageId="4" pageNumber="29" refString="Luhe, M. (1906) Report on the trematode parasites from the marine fishes of Ceylon. Report to the Government of Ceylon on the Pearl Oyster Fisheries of the Gulf of Manaar, Supplementary Report, 5, 97 - 108." type="journal article" year="1906">Lühe, 1906</bibRefCitation>
</taxonomicName>
as the
<typeStatus id="54D312F7D31CFFEDFD52408E98FC55D6" box="[698,751,1978,2004]" pageId="4" pageNumber="5">type</typeStatus>
and only species of his new subgenus
<taxonomicName id="4C68D7D6D31CFFEDFB30408F9F6955D6" box="[1240,1402,1979,2004]" class="Monogenea" family="Capsalidae" genus="Benedeniella" kingdom="Animalia" order="Capsalidea" pageId="4" pageNumber="5" phylum="Platyhelminthes" rank="genus">
<emphasis id="B91C7047D31CFFEDFB30408F9F6955D6" box="[1240,1402,1979,2004]" italics="true" pageId="4" pageNumber="5">Benedeniella</emphasis>
</taxonomicName>
as
<taxonomicName id="4C68D7D6D31DFFECFF7F47AC984D52B3" box="[151,606,152,177]" class="Monogenea" family="Capsalidae" genus="Benedenia" kingdom="Animalia" order="Capsalidea" pageId="5" pageNumber="6" phylum="Platyhelminthes" rank="species" species="macrocolpa" subGenus="Benedeniella">
<emphasis id="B91C7047D31DFFECFF7F47AC984D52B3" box="[151,606,152,177]" italics="true" pageId="5" pageNumber="6">Benedenia (Benedeniella) macrocolpa</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFF9D1A4D31DFFECFD8547AC992552B0" author="Yamaguti" box="[621,822,152,178]" pageId="5" pageNumber="30" refString="Yamaguti, S. (1963) Systema Helminthum. Volume IV. Monogenea and Aspidocotylea. InterScience Publishers, New York, USA, 699 pp." type="book" year="1963">Yamaguti (1963)</bibRefCitation>
raised
<taxonomicName id="4C68D7D6D31DFFECFC6447AC9E3552B3" box="[908,1062,152,177]" class="Monogenea" family="Capsalidae" genus="Benedeniella" kingdom="Animalia" order="Capsalidea" pageId="5" pageNumber="6" phylum="Platyhelminthes" rank="genus">
<emphasis id="B91C7047D31DFFECFC6447AC9E3552B3" box="[908,1062,152,177]" italics="true" pageId="5" pageNumber="6">Benedeniella</emphasis>
</taxonomicName>
to generic rank and transferred
<taxonomicName id="4C68D7D6D31DFFECFF7F4789987652D5" authority="Hargis, 1955" authorityName="Hargis" authorityYear="1955" box="[151,613,189,215]" class="Monogenea" family="Capsalidae" genus="Benedenia" kingdom="Animalia" order="Capsalidea" pageId="5" pageNumber="6" phylum="Platyhelminthes" rank="species" species="posterocolpa">
<emphasis id="B91C7047D31DFFECFF7F47899BAD52D4" box="[151,446,189,214]" italics="true" pageId="5" pageNumber="6">Benedenia posterocolpa</emphasis>
<bibRefCitation id="EFF9D1A4D31DFFECFE2F4789987652D5" author="Hargis" box="[455,613,189,215]" pageId="5" pageNumber="28" refString="Hargis, W. J. Jr. (1955) Monogenetic trematodes of Gulf of Mexico fishes. Part V. The superfamily Capsaloidea. Transactions of the American Microscopical Society, 74, 203 - 225." type="journal article" year="1955">Hargis, 1955</bibRefCitation>
</taxonomicName>
to
<taxonomicName id="4C68D7D6D31DFFECFD664789993E52D4" box="[654,813,189,214]" class="Monogenea" family="Capsalidae" genus="Benedeniella" kingdom="Animalia" order="Capsalidea" pageId="5" pageNumber="6" phylum="Platyhelminthes" rank="genus">
<emphasis id="B91C7047D31DFFECFD664789993E52D4" box="[654,813,189,214]" italics="true" pageId="5" pageNumber="6">Benedeniella</emphasis>
</taxonomicName>
because it also possessed a long vagina but which opened posteriorly some distance from the common genital pore. Yamagutis (1963) amended generic diagnosis perpetuated errors made by
<bibRefCitation id="EFF9D1A4D31DFFECFDB2463C98FE5320" author="Luhe" box="[602,749,264,290]" pageId="5" pageNumber="29" refString="Luhe, M. (1906) Report on the trematode parasites from the marine fishes of Ceylon. Report to the Government of Ceylon on the Pearl Oyster Fisheries of the Gulf of Manaar, Supplementary Report, 5, 97 - 108." type="journal article" year="1906">Lühe (1906)</bibRefCitation>
and
<bibRefCitation id="EFF9D1A4D31DFFECFCC0463C99DD5320" author="Hargis" box="[808,974,264,290]" pageId="5" pageNumber="28" refString="Hargis, W. J. Jr. (1955) Monogenetic trematodes of Gulf of Mexico fishes. Part V. The superfamily Capsaloidea. Transactions of the American Microscopical Society, 74, 203 - 225." type="journal article" year="1955">Hargis (1955)</bibRefCitation>
regarding the anatomy of the uterus in
<taxonomicName id="4C68D7D6D31DFFECFF7F46199B5A5344" box="[151,329,301,326]" class="Monogenea" family="Capsalidae" genus="Benedenia" kingdom="Animalia" order="Capsalidea" pageId="5" pageNumber="6" phylum="Platyhelminthes" rank="species" species="macrocolpa">
<emphasis id="B91C7047D31DFFECFF7F46199B5A5344" box="[151,329,301,326]" italics="true" pageId="5" pageNumber="6">B. macrocolpa</emphasis>
</taxonomicName>
and
<taxonomicName id="4C68D7D6D31DFFECFE6E461998565344" box="[390,581,301,326]" class="Monogenea" family="Capsalidae" genus="Benedenia" kingdom="Animalia" order="Capsalidea" pageId="5" pageNumber="6" phylum="Platyhelminthes" rank="species" species="posterocolpa">
<emphasis id="B91C7047D31DFFECFE6E461998565344" box="[390,581,301,326]" italics="true" pageId="5" pageNumber="6">B. posterocolpa</emphasis>
</taxonomicName>
, respectively.
<bibRefCitation id="EFF9D1A4D31DFFECFD104619999C5345" author="Luhe" box="[760,911,301,327]" pageId="5" pageNumber="29" refString="Luhe, M. (1906) Report on the trematode parasites from the marine fishes of Ceylon. Report to the Government of Ceylon on the Pearl Oyster Fisheries of the Gulf of Manaar, Supplementary Report, 5, 97 - 108." type="journal article" year="1906">Lühe (1906)</bibRefCitation>
referred to a short uterus but a common genital pore in
<taxonomicName id="4C68D7D6D31DFFECFEBE466698175369" box="[342,516,338,363]" class="Monogenea" family="Capsalidae" genus="Benedenia" kingdom="Animalia" order="Capsalidea" pageId="5" pageNumber="6" phylum="Platyhelminthes" rank="species" species="macrocolpa">
<emphasis id="B91C7047D31DFFECFEBE466698175369" box="[342,516,338,363]" italics="true" pageId="5" pageNumber="6">B. macrocolpa</emphasis>
</taxonomicName>
inferring that this species has a long common genital duct along which eggs pass but which also accommodated the male copulatory organ. For
<taxonomicName id="4C68D7D6D31DFFECFC5E464C9F375390" authority="Hargis (1955)" authorityName="Hargis" authorityYear="1955" box="[950,1316,376,402]" class="Monogenea" family="Capsalidae" genus="Benedenia" kingdom="Animalia" order="Capsalidea" pageId="5" pageNumber="6" phylum="Platyhelminthes" rank="species" species="posterocolpa">
<emphasis id="B91C7047D31DFFECFC5E464C9E7C5393" box="[950,1135,376,401]" italics="true" pageId="5" pageNumber="6">B. posterocolpa</emphasis>
,
<bibRefCitation id="EFF9D1A4D31DFFECFB97464C9F375390" author="Hargis" box="[1151,1316,376,402]" pageId="5" pageNumber="28" refString="Hargis, W. J. Jr. (1955) Monogenetic trematodes of Gulf of Mexico fishes. Part V. The superfamily Capsaloidea. Transactions of the American Microscopical Society, 74, 203 - 225." type="journal article" year="1955">Hargis (1955)</bibRefCitation>
</taxonomicName>
described the uterus as short, opening dorsally close to the bifurcation of the gut and separate from the marginal male pore, located posterolateral to the left anterior attachment organ. The arrangement of the terminal male and female genital ducts in both
<taxonomicName id="4C68D7D6D31DFFECFE0D46DC986D5003" box="[485,638,488,513]" class="Monogenea" family="Capsalidae" genus="Benedeniella" kingdom="Animalia" order="Capsalidea" pageId="5" pageNumber="6" phylum="Platyhelminthes" rank="genus">
<emphasis id="B91C7047D31DFFECFE0D46DC986D5003" box="[485,638,488,513]" italics="true" pageId="5" pageNumber="6">Benedeniella</emphasis>
</taxonomicName>
species was resolved and clarified by
<bibRefCitation id="EFF9D1A4D31DFFECFBAB46DC9F8B5000" author="Kearn" box="[1091,1432,488,514]" pageId="5" pageNumber="28" refString="Kearn, G. C. &amp; Whittington, I. D. (1992) Diversity of reproductive behaviour in platyhelminth parasites: insemination in some benedeniine (capsalid) monogeneans. Parasitology, 104, 489 - 496." type="journal article" year="1992">Kearn &amp; Whittington (1992)</bibRefCitation>
. The amended generic diagnosis presented above reflects the fact that the terminal male genital duct containing the cirrus and the uterus are each long ducts that remain parallel but separate from each other for most of their length and unite close to the common genital pore (
<bibRefCitation id="EFF9D1A4D31DFFECFD02456C9F835070" author="Kearn" box="[746,1424,600,626]" pageId="5" pageNumber="28" refString="Kearn, G. C. &amp; Whittington, I. D. (1992) Diversity of reproductive behaviour in platyhelminth parasites: insemination in some benedeniine (capsalid) monogeneans. Parasitology, 104, 489 - 496." type="journal article" year="1992">Kearn &amp; Whittington 1992; see also redescriptions below</bibRefCitation>
). Two other
<taxonomicName id="4C68D7D6D31DFFECFEFF45499BA25094" box="[279,433,637,662]" class="Monogenea" family="Capsalidae" genus="Benedeniella" kingdom="Animalia" order="Capsalidea" pageId="5" pageNumber="6" phylum="Platyhelminthes" rank="genus">
<emphasis id="B91C7047D31DFFECFEFF45499BA25094" box="[279,433,637,662]" italics="true" pageId="5" pageNumber="6">Benedeniella</emphasis>
</taxonomicName>
species have been described, but neither is consistent with the concept for the genus as revised here. The report of
<taxonomicName id="4C68D7D6D31DFFECFDED459699C350BE" authority="Yamaguti, 1958" authorityName="Yamaguti" authorityYear="1958" box="[517,976,674,700]" class="Monogenea" family="Capsalidae" genus="Benedeniella" kingdom="Animalia" order="Capsalidea" pageId="5" pageNumber="6" phylum="Platyhelminthes" rank="species" species="congeri">
<emphasis id="B91C7047D31DFFECFDED4596991450B9" box="[517,775,674,699]" italics="true" pageId="5" pageNumber="6">Benedeniella congeri</emphasis>
<bibRefCitation id="EFF9D1A4D31DFFECFCF8459699C350BE" author="Yamaguti" box="[784,976,674,700]" pageId="5" pageNumber="30" refString="Yamaguti, S. (1958) Studies on the helminth fauna of Japan. Part 53. Trematodes of fishes, XII. Publications of the Seto Marine Biology Laboratory, 7, 53 - 88." type="journal article" year="1958">Yamaguti, 1958</bibRefCitation>
</taxonomicName>
[synonyms:
<taxonomicName id="4C68D7D6D31DFFECFB854596982D50E0" authority="(Yamaguti, 1958) Yamaguti, 1963" authorityName="(Yamaguti, 1958) Yamaguti" authorityYear="1963" class="Monogenea" family="Capsalidae" genus="Neobenedeniella" higherTaxonomySource="CoL" kingdom="Animalia" order="Capsalidea" pageId="5" pageNumber="6" phylum="Platyhelminthes" rank="species" species="congeri">
<emphasis id="B91C7047D31DFFECFB8545969F8F50B9" box="[1133,1436,674,699]" italics="true" pageId="5" pageNumber="6">Neobenedeniella congeri</emphasis>
(
<bibRefCitation id="EFF9D1A4D31DFFECFF7745FC9B7550E0" author="Yamaguti" box="[159,358,712,738]" pageId="5" pageNumber="30" refString="Yamaguti, S. (1958) Studies on the helminth fauna of Japan. Part 53. Trematodes of fishes, XII. Publications of the Seto Marine Biology Laboratory, 7, 53 - 88." type="journal article" year="1958">Yamaguti, 1958</bibRefCitation>
)
<bibRefCitation id="EFF9D1A4D31DFFECFE9F45FC982D50E0" author="Yamaguti" box="[375,574,712,738]" pageId="5" pageNumber="30" refString="Yamaguti, S. (1963) Systema Helminthum. Volume IV. Monogenea and Aspidocotylea. InterScience Publishers, New York, USA, 699 pp." type="book" year="1963">Yamaguti, 1963</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C68D7D6D31DFFECFDA345FC9ED050E0" authority="(Yamaguti, 1958) Byrnes, 1986" authorityName="(Yamaguti, 1958) Byrnes" authorityYear="1986" box="[587,1219,712,738]" class="Monogenea" family="Capsalidae" genus="Benedenia" kingdom="Animalia" order="Capsalidea" pageId="5" pageNumber="6" phylum="Platyhelminthes" rank="species" species="congeri">
<emphasis id="B91C7047D31DFFECFDA345FC992050E3" box="[587,819,712,737]" italics="true" pageId="5" pageNumber="6">Benedenia congeri</emphasis>
(
<bibRefCitation id="EFF9D1A4D31DFFECFCAB45FC9E1850E0" author="Yamaguti" box="[835,1035,712,738]" pageId="5" pageNumber="30" refString="Yamaguti, S. (1958) Studies on the helminth fauna of Japan. Part 53. Trematodes of fishes, XII. Publications of the Seto Marine Biology Laboratory, 7, 53 - 88." type="journal article" year="1958">Yamaguti, 1958</bibRefCitation>
)
<bibRefCitation id="EFF9D1A4D31DFFECFBF445FC9ED050E0" author="Byrnes" box="[1052,1219,712,738]" pageId="5" pageNumber="27" refString="Byrnes, T. (1986) Five species of Monogenea from Australian bream, Acanthopagrus spp. Australian Journal of Zoology, 34, 65 - 86." type="journal article" year="1986">Byrnes, 1986</bibRefCitation>
</taxonomicName>
] from the gills of
<taxonomicName id="4C68D7D6D31DFFECFF7F45D99BFC5105" authority="Brevoort" authorityName="Brevoort" box="[151,495,749,775]" class="Actinopterygii" family="Congridae" genus="Conger" kingdom="Animalia" order="Anguilliformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="myriaster">
<emphasis id="B91C7047D31DFFECFF7F45D99B795104" box="[151,362,749,774]" italics="true" pageId="5" pageNumber="6">Conger myriaster</emphasis>
(Brevoort)
</taxonomicName>
(Teleostei:
<taxonomicName id="4C68D7D6D31DFFECFD9545D999245105" box="[637,823,749,775]" class="Actinopterygii" kingdom="Animalia" order="Anguilliformes" pageId="5" pageNumber="6" phylum="Chordata" rank="order">Anguilliformes</taxonomicName>
:
<taxonomicName id="4C68D7D6D31DFFECFCAB45D999D05105" box="[835,963,749,775]" class="Actinopterygii" family="Congridae" kingdom="Animalia" order="Anguilliformes" pageId="5" pageNumber="6" phylum="Chordata" rank="family">Congridae</taxonomicName>
;
<taxonomicName id="4C68D7D6D31DFFECFC2745D99E335105" box="[975,1056,749,775]" class="Actinopterygii" family="Congridae" genus="Conger" kingdom="Animalia" order="Anguilliformes" pageId="5" pageNumber="6" phylum="Chordata" rank="genus">conger</taxonomicName>
and garden eels), Inland Sea of
<collectingCountry id="F37FECC5D31DFFECFF7F44269AC9512E" box="[151,218,786,812]" name="Japan" pageId="5" pageNumber="6">Japan</collectingCountry>
was dealt with by
<bibRefCitation id="EFF9D1A4D31DFFECFE50442698B6512E" box="[440,677,786,812]" pageId="5" pageNumber="29" refString="Ogawa, K., Bondad-Reantaso, M. G., Wakabayashi, H. (1995) Redescription of Benedenia epinepheli (Yamaguti, 1937) Meserve, 1938 (Monogenea: Capsalidae) from cultured and aquarium marine fishes of Japan. Canadian Journal of Fisheries and Aquatic Sciences, 52, 62 - 70." type="journal article">
Ogawa
<emphasis id="B91C7047D31DFFECFDFB442698545129" box="[531,583,786,811]" italics="true" pageId="5" pageNumber="6">et al</emphasis>
. (1995)
</bibRefCitation>
when this taxon was considered a junior synonym of
<taxonomicName id="4C68D7D6D31DFFECFAC84426988D5150" authority="(Yamaguti, 1937) Meserve, 1938" authorityName="(Yamaguti, 1937) Meserve" authorityYear="1938" class="Monogenea" family="Capsalidae" genus="Benedenia" kingdom="Animalia" order="Capsalidea" pageId="5" pageNumber="6" phylum="Platyhelminthes" rank="species" species="epinepheli">
<emphasis id="B91C7047D31DFFECFAC844269B025153" italics="true" pageId="5" pageNumber="6">Benedenia epinepheli</emphasis>
(
<bibRefCitation id="EFF9D1A4D31DFFECFEC8440C9BCE5150" author="Yamaguti" box="[288,477,824,850]" pageId="5" pageNumber="30" refString="Yamaguti, S. (1937) Studies on the Helminth Fauna of Japan. Part 19. Fourteen New Ectoparasitic Trematodes of Fishes (published by the author), 28 pp." type="book" year="1937">Yamaguti, 1937</bibRefCitation>
)
<bibRefCitation id="EFF9D1A4D31DFFECFE04440C988D5150" author="Meserve" box="[492,670,824,850]" pageId="5" pageNumber="29" refString="Meserve, F. G. (1938) Some monogenetic trematodes from the Galapagos Islands and the neighbouring Pacific. Reports of the Allan Hancock Pacific Expeditions (1932 - 1937), 2, 31 - 89." type="journal article" year="1938">Meserve, 1938</bibRefCitation>
</taxonomicName>
. This action was considered sound by Whittington
<emphasis id="B91C7047D31DFFECFB1E440C9F395153" box="[1270,1322,824,849]" italics="true" pageId="5" pageNumber="6">et al</emphasis>
. (2001).
</paragraph>
</subSubSection>
<subSubSection id="C372FFDED31DFFECFF2D44699B535655" pageId="5" pageNumber="6" type="reference_group">
<paragraph id="8BD7AC55D31DFFECFF2D44699B535655" blockId="5.[151,1438,152,1111]" pageId="5" pageNumber="6">
<taxonomicName id="4C68D7D6D31DFFECFF2D4469991B5175" ID-CoL="9dc625d0-4c00-4350-bc2b-0fec9164f73d" authority="Gupta &amp; Chanana, 1976" authorityName="Gupta &amp; Chanana" authorityYear="1976" box="[197,776,861,887]" class="Monogenea" family="Capsalidae" genus="Benedeniella" kingdom="Animalia" order="Capsalidea" pageId="5" pageNumber="6" phylum="Platyhelminthes" rank="species" species="unnithani">
<emphasis id="B91C7047D31DFFECFF2D44699BC85174" box="[197,475,861,886]" italics="true" pageId="5" pageNumber="6">Benedeniella unnithani</emphasis>
<bibRefCitation id="EFF9D1A4D31DFFECFE0B4469991B5175" author="Gupta" box="[483,776,861,887]" pageId="5" pageNumber="28" refString="Gupta, N. K. &amp; Chanana, A. (1976) Studies on some monogenetic trematodes of marine fishes from Laccadive Islands, Calicut and other places in India. Part 1. Revista Iberica de Parasitologia, 36, 267 - 275." type="journal article" year="1976">Gupta &amp; Chanana, 1976</bibRefCitation>
</taxonomicName>
was reported from the gills of a
<taxonomicName id="4C68D7D6D31DFFECFB7B44699EFE5174" ID-CoL="62L73" box="[1171,1261,861,886]" class="Actinopterygii" family="Carangidae" genus="Caranx" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="6" phylum="Chordata" rank="genus">
<emphasis id="B91C7047D31DFFECFB7B44699EFE5174" box="[1171,1261,861,886]" italics="true" pageId="5" pageNumber="6">Caranx</emphasis>
</taxonomicName>
sp. (Teleostei:
<taxonomicName id="4C68D7D6D31DFFECFF7F44B69B3A519E" ID-CoL="PC" box="[151,297,898,924]" class="Actinopterygii" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="6" phylum="Chordata" rank="order">Perciformes</taxonomicName>
:
<taxonomicName id="4C68D7D6D31DFFECFEDB44B69BAF519E" ID-CoL="7Q9" box="[307,444,898,924]" class="Actinopterygii" family="Carangidae" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="6" phylum="Chordata" rank="family">Carangidae</taxonomicName>
; jacks and pompanos) at Kavaratti,
<collectingCountry id="F37FECC5D31DFFECFCB444B69E22519E" box="[860,1073,898,924]" name="India" pageId="5" pageNumber="6">Laccadive Islands</collectingCountry>
, Arabian Sea,
<collectingCountry id="F37FECC5D31DFFECFB3244B69F09519E" box="[1242,1306,898,924]" name="India" pageId="5" pageNumber="6">India</collectingCountry>
. No details about deposition of material in a museum were provided by
<bibRefCitation id="EFF9D1A4D31DFFECFC91449C9EBD51C0" author="Gupta" box="[889,1198,936,962]" pageId="5" pageNumber="28" refString="Gupta, N. K. &amp; Chanana, A. (1976) Studies on some monogenetic trematodes of marine fishes from Laccadive Islands, Calicut and other places in India. Part 1. Revista Iberica de Parasitologia, 36, 267 - 275." type="journal article" year="1976">Gupta &amp; Chanana (1976)</bibRefCitation>
and
<bibRefCitation id="EFF9D1A4D31DFFECFB02449C9B5E51E5" author="Whittington" pageId="5" pageNumber="29" refString="Whittington, I. D. &amp; Barton, D. P. (1990) A new genus of monogenean parasites (Capsalidae: Benedeniinae) from stingrays (Rajiformes: Dasyatidae) with a description of a new species from the long-tailed stingray Himantura uarnak Forsskal from Queensland, Australia. Journal of Natural History, 24, 327 - 340." type="journal article" year="1990">Whittington &amp; Barton (1990)</bibRefCitation>
attempted to locate and borrow specimens without success. I have made two further unsuccessful efforts to find their material. Based on their description, the three capsalid specimens they reported are clearly not a species of
<taxonomicName id="4C68D7D6D31DFFECFD89432C99175633" ID-CoL="7NQN7" box="[609,772,1048,1073]" class="Monogenea" family="Capsalidae" genus="Benedeniella" kingdom="Animalia" order="Capsalidea" pageId="5" pageNumber="6" phylum="Platyhelminthes" rank="genus">
<emphasis id="B91C7047D31DFFECFD89432C99175633" box="[609,772,1048,1073]" italics="true" pageId="5" pageNumber="6">Benedeniella</emphasis>
</taxonomicName>
.
<taxonomicName id="4C68D7D6D31DFFECFCFD432C9E245633" ID-CoL="9dc625d0-4c00-4350-bc2b-0fec9164f73d" box="[789,1079,1048,1073]" class="Monogenea" family="Capsalidae" genus="Benedeniella" kingdom="Animalia" order="Capsalidea" pageId="5" pageNumber="6" phylum="Platyhelminthes" rank="species" species="unnithani">
<emphasis id="B91C7047D31DFFECFCFD432C9E245633" box="[789,1079,1048,1073]" italics="true" pageId="5" pageNumber="6">Benedeniella unnithani</emphasis>
</taxonomicName>
is here considered a species
<emphasis id="B91C7047D31DFFECFF7F43099B2A5654" box="[151,313,1085,1110]" italics="true" pageId="5" pageNumber="6">incertae sedis</emphasis>
.
</paragraph>
</subSubSection>
</treatment>
</document>