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<document id="AAF31A96A10632607B9FCF1AA352DF7C" ID-CLB-Dataset="26863" ID-DOI="10.11646/zootaxa.4603.3.3" ID-GBIF-Dataset="ccbb854c-979b-40ea-bd51-64cf20307fb0" ID-ISSN="1175-5326" ID-Zenodo-Dep="2702283" ID-ZooBank="558DEADB-EF73-4EBD-9564-AA522A2D6CF1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1557468872627" checkinUser="plazi" docAuthor="Liu, H., Liu, X. &amp; Zágoršek, K." docDate="2019" docId="530A87AB3B24F477BDAC4520E24AFEC2" docLanguage="en" docName="zootaxa.4603.3.3.pdf" docOrigin="Zootaxa 4603 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Crisia guang Liu &amp; Liu &amp; Zágoršek 2019, n. sp." docType="treatment" docVersion="7" lastPageNumber="477" masterDocId="AF33FFD33B26F473BD3B403EE675FFAA" masterDocTitle="Cyclostome bryozoans from Qingdao, South Yellow Sea, China" masterLastPageNumber="500" masterPageNumber="473" pageNumber="475" updateTime="1698728432520" updateUser="ExternalLinkService">
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<mods:title id="BB6D917971BC73B3C0DAB413708C7E56">Cyclostome bryozoans from Qingdao, South Yellow Sea, China</mods:title>
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<mods:name id="8ED039B3B588DAC7F6770AF80E521B0C" type="personal">
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<mods:namePart id="CEE7F7D929B72320CCBB1A807F54A412">Liu, H.</mods:namePart>
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<mods:namePart id="93C3DD94AA0A848FF016A51D64CDBF15">Liu, X.</mods:namePart>
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<mods:namePart id="F67CC5E33C13E3726A071B106B837278">Zágoršek, K.</mods:namePart>
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<mods:title id="2B7EC72477E9CE7195C6A2430C9E56F6">Zootaxa</mods:title>
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<mods:date id="20C22A633249D8324F3ABFCB811F71DF">2019</mods:date>
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<treatment id="530A87AB3B24F477BDAC4520E24AFEC2" ID-GBIF-Taxon="157119467" LSID="urn:lsid:plazi:treatment:530A87AB3B24F477BDAC4520E24AFEC2" httpUri="http://treatment.plazi.org/id/530A87AB3B24F477BDAC4520E24AFEC2" lastPageId="4" lastPageNumber="477" pageId="2" pageNumber="475">
<subSubSection id="93B965363B24F471BDAC4520E70BFA93" box="[151,382,1310,1337]" pageId="2" pageNumber="475" type="nomenclature">
<paragraph id="DB1C36BD3B24F471BDAC4520E70BFA93" blockId="2.[151,915,1310,1408]" box="[151,382,1310,1337]" pageId="2" pageNumber="475">
<heading id="805481D13B24F471BDAC4520E70BFA93" bold="true" box="[151,382,1310,1337]" fontSize="11" level="1" pageId="2" pageNumber="475" reason="1">
<emphasis id="E9D7EAAF3B24F471BDAC4520E70BFA93" bold="true" box="[151,382,1310,1337]" pageId="2" pageNumber="475">
<taxonomicName id="1CA34D3E3B24F471BDAC4520E740FA93" authority="Liu &amp; Liu &amp; Zágoršek, 2019" authorityName="Liu &amp; Liu &amp; Zágoršek" authorityYear="2019" box="[151,309,1310,1337]" class="Stenolaemata" family="Crisiidae" genus="Crisia" kingdom="Animalia" order="Cyclostomatida" pageId="2" pageNumber="475" phylum="Bryozoa" rank="species" species="guang" status="sp. nov.">
<emphasis id="E9D7EAAF3B24F471BDAC4520E740FA93" bold="true" box="[151,309,1310,1337]" italics="true" pageId="2" pageNumber="475">Crisia guang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F2E457D43B24F471BC074521E70BFA93" box="[316,382,1311,1337]" pageId="2" pageNumber="475" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="93B965363B24F471BDAC457CE5E6FA2A" pageId="2" pageNumber="475" type="description">
<paragraph id="DB1C36BD3B24F471BDAC457CE77EFAF7" blockId="2.[151,915,1310,1408]" box="[151,267,1346,1373]" pageId="2" pageNumber="475">
(
<figureCitation id="43982A383B24F471BDA4457CE776FAF7" box="[159,259,1346,1373]" captionStart="FIGURES 36" captionStartId="3.[151,264,1461,1486]" captionTargetBox="[164,1423,476,1409]" captionTargetId="figure@3.[164,1423,476,1409]" captionTargetPageId="3" captionText="FIGURES 36. Crisia guang n. sp. 3, 4. Holotype MBM 284486 from Yangkou (Laoshan). 5, 6. Paratypes MBM 284371 and MBM 284372 from Lingshan Island (Jiaonan), showing the frontally oriented, oval ooeciopore with straight distal margins. Scale bars: 3, 4 = 1 mm; 5, 6 = 100 µm." httpUri="https://zenodo.org/record/2702287/files/figure.png" pageId="2" pageNumber="475">Figs 36</figureCitation>
)
</paragraph>
<paragraph id="DB1C36BD3B24F471BDAC455BE5E6FA2A" blockId="2.[151,915,1310,1408]" box="[151,915,1381,1408]" pageId="2" pageNumber="475">
lsid:zoobank.org:act:
<uuid id="AF050C683B24F471BCBB455BE5E6FA2A" box="[384,915,1381,1408]" pageId="2" pageNumber="475">1E1DF6A1-190C-4689-AA38-5377226CC530</uuid>
</paragraph>
</subSubSection>
<subSubSection id="93B965363B24F471BDAC4595E502F9AA" pageId="2" pageNumber="475" type="reference_group">
<paragraph id="DB1C36BD3B24F471BDAC4595E5B5FA69" blockId="2.[151,1083,1450,1536]" box="[151,960,1450,1475]" pageId="2" pageNumber="475">
?
<taxonomicName id="1CA34D3E3B24F471BD924595E4C6FA69" ID-CoL="ZHPW" authority=": Mawatari &amp; Mawatari, 1973: p. 97" authorityName=": Mawatari &amp; Mawatari" authorityPageNumber="97" authorityYear="1973" box="[169,691,1450,1475]" class="Stenolaemata" family="Crisiidae" genus="Crisia" kingdom="Animalia" order="Cyclostomatida" pageId="2" pageNumber="475" phylum="Bryozoa" rank="species" species="eburnea">
<emphasis id="E9D7EAAF3B24F471BD924595E734FA69" box="[169,321,1451,1475]" italics="true" pageId="2" pageNumber="475">Crisia eburnea</emphasis>
:
<bibRefCitation id="BF324B4C3B24F471BC744594E401FA69" author="Mawatari, S. &amp; Mawatari, S. F." box="[335,628,1450,1475]" pageId="2" pageNumber="475" pagination="95 - 104" refId="ref12988" refString="Mawatari, S. &amp; Mawatari, S. F. (1973) Notes on marine Bryozoa from Hokkaido. 1. Crisiidae (Cyclostomata). Journal of the Faculty of Science, Hokkaido University, 19 (1), 95 - 104." type="journal article" year="1973">Mawatari &amp; Mawatari, 1973</bibRefCitation>
: p. 97
</taxonomicName>
, text-fig. 3, pl. 1, figs 7, 8.
</paragraph>
<paragraph id="DB1C36BD3B24F471BDAC45F7E24EFA48" blockId="2.[151,1083,1450,1536]" box="[151,1083,1481,1506]" pageId="2" pageNumber="475">
?
<taxonomicName id="1CA34D3E3B24F471BD9245F4E55BFA48" ID-CoL="c1047237-a7e0-49ba-8174-605b29a9591a" authority=": Mawatari &amp; Mawatari, 1973: p. 96" authorityName="Mawatari &amp; Mawatari" authorityPageNumber="96" authorityYear="1973" box="[169,814,1481,1506]" class="Stenolaemata" family="Crisiidae" genus="Crisia" kingdom="Animalia" order="Cyclostomatida" pageId="2" pageNumber="475" phylum="Bryozoa" rank="species" species="eburneo-denticulata">
<emphasis id="E9D7EAAF3B24F471BD9245F4E7CEFA4B" box="[169,443,1481,1505]" italics="true" pageId="2" pageNumber="475">Crisia eburneo-denticulata</emphasis>
:
<bibRefCitation id="BF324B4C3B24F471BCF245F7E49BFA4B" author="Mawatari, S. &amp; Mawatari, S. F." box="[457,750,1481,1506]" pageId="2" pageNumber="475" pagination="95 - 104" refId="ref12988" refString="Mawatari, S. &amp; Mawatari, S. F. (1973) Notes on marine Bryozoa from Hokkaido. 1. Crisiidae (Cyclostomata). Journal of the Faculty of Science, Hokkaido University, 19 (1), 95 - 104." type="journal article" year="1973">Mawatari &amp; Mawatari, 1973</bibRefCitation>
: p. 96
</taxonomicName>
, text-fig. 2, pl. 1, figs 5, 6.
</paragraph>
<paragraph id="DB1C36BD3B24F471BDAC45D6E502F9AA" blockId="2.[151,1083,1450,1536]" box="[151,887,1511,1536]" pageId="2" pageNumber="475">
v.
<taxonomicName id="1CA34D3E3B24F471BD9445D6E4AFF9AA" ID-CoL="c1047237-a7e0-49ba-8174-605b29a9591a" authority=": Liu, Yin &amp; Ma, 2001: 379" authorityName="Liu, Yin &amp; Ma" authorityPageNumber="379" authorityYear="2001" box="[175,730,1511,1536]" class="Stenolaemata" family="Crisiidae" genus="Crisia" kingdom="Animalia" order="Cyclostomatida" pageId="2" pageNumber="475" phylum="Bryozoa" rank="species" species="eburneo-denticulata">
<emphasis id="E9D7EAAF3B24F471BD9445D6E7B7F9AA" box="[175,450,1512,1536]" italics="true" pageId="2" pageNumber="475">Crisia eburneo-denticulata</emphasis>
:
<bibRefCitation id="BF324B4C3B24F471BCF445D9E4DDF9AA" author="Liu, X. &amp; Yin, X. &amp; Ma, J." box="[463,680,1511,1536]" pageId="2" pageNumber="475" refId="ref12772" refString="Liu, X., Yin, X. &amp; Ma, J. (2001) Biology of Marine-Fouling Bryozoans in the Coastal Waters of China. Science Press, Beijing, 860 pp." type="book" year="2001">Liu, Yin &amp; Ma, 2001</bibRefCitation>
: 379
</taxonomicName>
, pl. 4, figs 35.
</paragraph>
</subSubSection>
<subSubSection id="93B965363B24F471BDAC4613E4E2F93B" pageId="2" pageNumber="475" type="diagnosis">
<paragraph id="DB1C36BD3B24F471BDAC4613E4E2F93B" blockId="2.[151,1436,1581,2041]" pageId="2" pageNumber="475">
<emphasis id="E9D7EAAF3B24F471BDAC4613E77FF9E2" bold="true" box="[151,266,1581,1608]" pageId="2" pageNumber="475">Diagnosis</emphasis>
:
<taxonomicName id="1CA34D3E3B24F471BC274611E717F9E3" authorityName="Lamouroux" authorityYear="1812" box="[284,354,1583,1609]" class="Stenolaemata" family="Crisiidae" genus="Crisia" kingdom="Animalia" order="Cyclostomatida" pageId="2" pageNumber="475" phylum="Bryozoa" rank="genus">
<emphasis id="E9D7EAAF3B24F471BC274611E717F9E3" box="[284,354,1583,1609]" italics="true" pageId="2" pageNumber="475">Crisia</emphasis>
</taxonomicName>
with gonozooid occupying almost the entire width of the branch and brood chamber as long as 23 autozooids. Ooeciopore larger than an autozooidal aperture, facing frontally, oval, with a straight distal edge, and often adjacent to an autozooidal aperture.
</paragraph>
</subSubSection>
<subSubSection id="93B965363B24F471BDFC46A5E710F827" pageId="2" pageNumber="475" type="materials_examined">
<paragraph id="DB1C36BD3B24F471BDFC46A5E58CF91E" blockId="2.[151,1436,1581,2041]" box="[199,1017,1690,1717]" pageId="2" pageNumber="475">
<materialsCitation id="6BCB3CE03B24F471BDFC46A5E58CF91E" ID-GBIF-Occurrence="2239631087" box="[199,1017,1690,1717]" collectionCode="MBM" pageId="2" pageNumber="475" specimenCode="MBM 284486" specimenCount="1" typeStatus="holotype">
<emphasis id="E9D7EAAF3B24F471BDFC46A5E74EF91F" bold="true" box="[199,315,1691,1717]" pageId="2" pageNumber="475">
<typeStatus id="0418881F3B24F471BDFC46A5E743F91F" box="[199,310,1691,1717]" pageId="2" pageNumber="475" type="holotype">Holotype</typeStatus>
.
</emphasis>
Specimen
<specimenCode id="8B059EC63B24F471BC8C46A5E422F91E" box="[439,599,1690,1717]" collectionCode="MBM" country="0" httpUri="http://grbio.org/cool/fc7c-p25f" name="San Jose State University, Museum of Birds and Mammals" pageId="2" pageNumber="475">MBM 284486</specimenCode>
from Yangkou (Laoshan) (
<figureCitation id="43982A383B24F471BEBD46A4E59EF91E" box="[902,1003,1690,1716]" captionStart="FIGURES 36" captionStartId="3.[151,264,1461,1486]" captionTargetBox="[164,1423,476,1409]" captionTargetId="figure@3.[164,1423,476,1409]" captionTargetPageId="3" captionText="FIGURES 36. Crisia guang n. sp. 3, 4. Holotype MBM 284486 from Yangkou (Laoshan). 5, 6. Paratypes MBM 284371 and MBM 284372 from Lingshan Island (Jiaonan), showing the frontally oriented, oval ooeciopore with straight distal margins. Scale bars: 3, 4 = 1 mm; 5, 6 = 100 µm." httpUri="https://zenodo.org/record/2702287/files/figure.png" pageId="2" pageNumber="475">Figs 3, 4</figureCitation>
).
</materialsCitation>
</paragraph>
<paragraph id="DB1C36BD3B24F471BDFC4681E48AF957" blockId="2.[151,1436,1581,2041]" pageId="2" pageNumber="475">
<materialsCitation id="6BCB3CE03B24F471BDFC4681E453F972" ID-GBIF-Occurrence="2239631093" box="[199,550,1726,1753]" collectionCode="MBM" pageId="2" pageNumber="475" specimenCode="MBM 284371" specimenCount="1" typeStatus="paratype">
<emphasis id="E9D7EAAF3B24F471BDFC4681E733F973" bold="true" box="[199,326,1727,1753]" pageId="2" pageNumber="475">
<typeStatus id="0418881F3B24F471BDFC4681E734F973" box="[199,321,1727,1753]" pageId="2" pageNumber="475" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCode id="8B059EC63B24F471BC6B4681E786F972" box="[336,499,1726,1753]" collectionCode="MBM" country="0" httpUri="http://grbio.org/cool/fc7c-p25f" name="San Jose State University, Museum of Birds and Mammals" pageId="2" pageNumber="475">MBM 284371</specimenCode>
and
</materialsCitation>
<materialsCitation id="6BCB3CE03B24F471BF0A4681E5DEF973" ID-GBIF-Occurrence="2239631094" box="[561,939,1726,1753]" collectionCode="MBM" location="Lingshan Island" pageId="2" pageNumber="475" specimenCode="MBM 284372" specimenCount="1" typeStatus="paratype">
<specimenCode id="8B059EC63B24F471BF0A4681E4A1F972" box="[561,724,1726,1753]" collectionCode="MBM" country="0" httpUri="http://grbio.org/cool/fc7c-p25f" name="San Jose State University, Museum of Birds and Mammals" pageId="2" pageNumber="475">MBM 284372</specimenCode>
(
<location id="DE7C60663B24F471BFDD4680E5D1F973" LSID="urn:lsid:plazi:treatment:530A87AB3B24F477BDAC4520E24AFEC2:DE7C60663B24F471BFDD4680E5D1F973" box="[742,932,1726,1753]" name="Lingshan Island" pageId="2" pageNumber="475">Lingshan Island</location>
)
</materialsCitation>
,
<materialsCitation id="6BCB3CE03B24F471BE814681E3EDF973" ID-GBIF-Occurrence="2239631089" box="[954,1432,1726,1753]" collectionCode="MBM" location="Haijiao" municipality="Lingshan Island" pageId="2" pageNumber="475" specimenCode="MBM 284479" specimenCount="1" typeStatus="paratype">
<specimenCode id="8B059EC63B24F471BE814681E228F972" box="[954,1117,1726,1753]" collectionCode="MBM" country="0" httpUri="http://grbio.org/cool/fc7c-p25f" name="San Jose State University, Museum of Birds and Mammals" pageId="2" pageNumber="475">MBM 284479</specimenCode>
(
<location id="DE7C60663B24F471B9544680E2B3F972" LSID="urn:lsid:plazi:treatment:530A87AB3B24F477BDAC4520E24AFEC2:DE7C60663B24F471B9544680E2B3F972" box="[1135,1222,1726,1752]" municipality="Lingshan Island" name="Haijiao" pageId="2" pageNumber="475">Haijiao</location>
,
<collectingMunicipality id="3B78ACC73B24F471B9EE4680E3E5F973" box="[1237,1424,1726,1753]" pageId="2" pageNumber="475">Lingshan Island</collectingMunicipality>
)
</materialsCitation>
,
<materialsCitation id="6BCB3CE03B24F471BDAC46DDE713F956" ID-GBIF-Occurrence="2239631100" box="[151,358,1761,1789]" collectionCode="MBM" pageId="2" pageNumber="475" specimenCode="MBM 284485" specimenCount="1" typeStatus="paratype">
<specimenCode id="8B059EC63B24F471BDAC46DDE742F956" box="[151,311,1761,1789]" collectionCode="MBM" country="0" httpUri="http://grbio.org/cool/fc7c-p25f" name="San Jose State University, Museum of Birds and Mammals" pageId="2" pageNumber="475">MBM 284485</specimenCode>
and
</materialsCitation>
<materialsCitation id="6BCB3CE03B24F471BC5646DDE478F956" ID-GBIF-Occurrence="2239631086" box="[365,525,1762,1789]" collectionCode="MBM" pageId="2" pageNumber="475" specimenCode="MBM 284488" specimenCount="1" typeStatus="paratype">
<specimenCode id="8B059EC63B24F471BC5646DDE478F956" box="[365,525,1762,1789]" collectionCode="MBM" country="0" httpUri="http://grbio.org/cool/fc7c-p25f" name="San Jose State University, Museum of Birds and Mammals" pageId="2" pageNumber="475">MBM 284488</specimenCode>
</materialsCitation>
(Yangkou, Laoshan).
</paragraph>
<paragraph id="DB1C36BD3B24F471BDFC473BE710F827" blockId="2.[151,1436,1581,2041]" pageId="2" pageNumber="475">
<emphasis id="E9D7EAAF3B24F471BDFC473BE7CBF88A" bold="true" box="[199,446,1797,1824]" pageId="2" pageNumber="475">Additional material.</emphasis>
More than 30 colonies, from samples MBM 0 92255, MBM 194665, MBM 194688, MBM 194759, MBM 194937 and MBM 194942 (Haijiao, Lingshan Island), MBM 0 92292, MBM 0 92315, MBM 0 92318 and MBM 188532 (
<collectingCountry id="A3B4762D3B24F471BCEB4770E418F8C3" box="[464,621,1870,1897]" name="China" pageId="2" pageNumber="475">Jiaozhou Bay</collectingCountry>
), MBM 2843975 (Yangkou, Laoshan) and MBM 284367, MBM 284364 (Lingshan Island).
</paragraph>
</subSubSection>
<subSubSection id="93B965363B24F471BDFC47A9E7C9F87F" pageId="2" pageNumber="475" type="etymology">
<paragraph id="DB1C36BD3B24F471BDFC47A9E7C9F87F" blockId="2.[151,1436,1581,2041]" pageId="2" pageNumber="475">
<emphasis id="E9D7EAAF3B24F471BDFC47A9E73BF81B" bold="true" box="[199,334,1943,1969]" pageId="2" pageNumber="475">Etymology.</emphasis>
Referring to the wide gonozooid occupying the full width of the branch; in Chinese phonetic transcription, wide is
<taxonomicName id="1CA34D3E3B24F471BC544785E7C0F87F" authorityName="Liu &amp; Liu &amp; Zágoršek" authorityYear="2019" box="[367,437,1979,2005]" class="Stenolaemata" family="Crisiidae" genus="Crisia" kingdom="Animalia" order="Cyclostomatida" pageId="2" pageNumber="475" phylum="Bryozoa" rank="species" species="guang">
<emphasis id="E9D7EAAF3B24F471BC544785E7C0F87F" box="[367,437,1979,2005]" italics="true" pageId="2" pageNumber="475">guǎng</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="93B965363B24F470BDFC47E3E560FE05" lastPageId="3" lastPageNumber="476" pageId="2" pageNumber="475" type="description">
<paragraph id="DB1C36BD3B24F470BDFC47E3E560FE05" blockId="2.[151,1436,1581,2041]" lastBlockId="3.[151,1437,152,431]" lastPageId="3" lastPageNumber="476" pageId="2" pageNumber="475">
<emphasis id="E9D7EAAF3B24F471BDFC47E3E72CF852" bold="true" box="[199,345,2013,2040]" pageId="2" pageNumber="475">Description.</emphasis>
Width of internodes approximately
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, length approximately 1.33.8 mm; approxi- mately 1520 autozooids per internode. Autozooids approximately
<quantity id="1C5B9B583B25F470BEB040A7E597FF19" box="[907,994,153,179]" metricMagnitude="-1" metricUnit="m" metricValue="2.3" pageId="3" pageNumber="476" unit="mm" value="230.0">230 µm</quantity>
long (the zooids attached to gonozooid can exceed its length, up to
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), width approximately
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. Apertures circular, approximately
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in diameter, with peristome facing frontally. Pseudopores on the frontal wall slit-like (approximately
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long by
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wide), numerous and longitudinally oriented. Gonozooid as long as two to three autozooids, extending across entire branch width. The proximal part of the gonozooid (before dilation) is wider than the equivalent part of an autozooid. Ooeciopore frontally oriented, larger than an autozooidal aperture, oval, wider (ca.
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) than long (ca.
<quantity id="1C5B9B583B25F470BDFC414EE766FE21" box="[199,275,368,395]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="476" unit="mm" value="50.0">50 µm</quantity>
), distal edge more or less straight, adjacent to the neighbouring autozooidal aperture. Pseudopores more numerous than in autozooids but similar in shape or size.
</paragraph>
</subSubSection>
<caption id="8FDC66353B25F470BDAC458BE45EF9BC" httpUri="https://zenodo.org/record/2702287/files/figure.png" pageId="3" pageNumber="476" startId="3.[151,264,1461,1486]" targetBox="[164,1423,476,1409]" targetPageId="3">
<paragraph id="DB1C36BD3B25F470BDAC458BE45EF9BC" blockId="3.[151,1437,1461,1558]" pageId="3" pageNumber="476">
<emphasis id="E9D7EAAF3B25F470BDAC458BE74EFA64" bold="true" box="[151,315,1461,1486]" pageId="3" pageNumber="476">FIGURES 36.</emphasis>
<taxonomicName id="1CA34D3E3B25F470BC7A4588E7B3FA67" authority="Liu &amp; Liu &amp; Zágoršek, 2019" authorityName="Liu &amp; Liu &amp; Zágoršek" authorityYear="2019" box="[321,454,1462,1485]" class="Stenolaemata" family="Crisiidae" genus="Crisia" kingdom="Animalia" order="Cyclostomatida" pageId="3" pageNumber="476" phylum="Bryozoa" rank="species" species="guang" status="sp. nov.">
<emphasis id="E9D7EAAF3B25F470BC7A4588E7B3FA67" box="[321,454,1462,1485]" italics="true" pageId="3" pageNumber="476">Crisia guang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F2E457D43B25F470BCF7458BE473FA67" box="[460,518,1461,1485]" pageId="3" pageNumber="476" rank="species">
<emphasis id="E9D7EAAF3B25F470BCF7458BE473FA67" bold="true" box="[460,518,1461,1485]" pageId="3" pageNumber="476">n. sp.</emphasis>
</taxonomicNameLabel>
3, 4. Holotype MBM 284486 from Yangkou (Laoshan). 5, 6. Paratypes MBM 284371 and MBM 284372 from Lingshan Island (Jiaonan), showing the frontally oriented, oval ooeciopore with straight distal margins. Scale bars: 3, 4 = 1 mm; 5, 6 = 100 µm.
</paragraph>
</caption>
<subSubSection id="93B965363B25F470BDFC467AE2F3F960" pageId="3" pageNumber="476" type="discussion">
<paragraph id="DB1C36BD3B25F470BDFC467AE2F3F960" blockId="3.[151,1437,1604,2026]" pageId="3" pageNumber="476">
<emphasis id="E9D7EAAF3B25F470BDFC467AE74EF9F4" bold="true" box="[199,315,1604,1630]" pageId="3" pageNumber="476">Remarks.</emphasis>
The most similar species is
<taxonomicName id="1CA34D3E3B25F470BF4E467AE271F9F5" authority="(Lamarck, 1816)" authorityName="Lamarck" authorityYear="1816" baseAuthorityName="Lamarck" baseAuthorityYear="1816" box="[629,1028,1604,1631]" class="Stenolaemata" family="Crisiidae" genus="Crisia" kingdom="Animalia" order="Cyclostomatida" pageId="3" pageNumber="476" phylum="Bryozoa" rank="species" species="denticulata">
<emphasis id="E9D7EAAF3B25F470BF4E467AE535F9F4" box="[629,832,1604,1630]" italics="true" pageId="3" pageNumber="476">Crisia denticulata</emphasis>
(
<bibRefCitation id="BF324B4C3B25F470BE75467AE589F9F5" author="Lamarck, J. B. P. A. de" box="[846,1020,1604,1631]" pageId="3" pageNumber="476" refId="ref12662" refString="Lamarck, J. B. P. A. de M. (1816) Histoire naturelle des Animaux sans Vertebres ... precedee d'une introduction offrant la determination des caracteres essentiels de l'animal, sa distinction du vegetal et des autres coorps naturels, enfin, exposition des principes fondamentaux de la zoologie. Vol. 2. Verdiere, Paris, 568 pp." type="book" year="1816">Lamarck, 1816</bibRefCitation>
)
</taxonomicName>
, which based on
<bibRefCitation id="BF324B4C3B25F470B9F8467AE697F928" author="Hayward, P. J. &amp; Ryland, J. S." pageId="3" pageNumber="476" pagination="1 - 147" refId="ref12323" refString="Hayward, P. J. &amp; Ryland, J. S. (1985) Cyclostome Bryozoans. In: Kermack, D. M. &amp; Barnes, R. S. K. (Eds.), Synopses of the British Fauna. New Series. Vol. 34. E. J. Brill for the Linnaean Society, London, pp. 1 - 147." type="book chapter" year="1985">Hayward &amp; Ryland (1985)</bibRefCitation>
has a similarly extended gonozooid and a similar shape and position of the ooeciopore. The new species mainly differs in having a less convex gonozooid and in the lack of characteristic black joints between internodes. Moreover,
<taxonomicName id="1CA34D3E3B25F470BC29468EE7C4F960" baseAuthorityName="Lamarck" baseAuthorityYear="1816" box="[274,433,1712,1738]" class="Stenolaemata" family="Crisiidae" genus="Crisia" kingdom="Animalia" order="Cyclostomatida" pageId="3" pageNumber="476" phylum="Bryozoa" rank="species" species="denticulata">
<emphasis id="E9D7EAAF3B25F470BC29468EE7C4F960" box="[274,433,1712,1738]" italics="true" pageId="3" pageNumber="476">C. denticulata</emphasis>
</taxonomicName>
has been reported only from Europe (
<bibRefCitation id="BF324B4C3B25F470BE61468EE20CF960" author="Hayward, P. J. &amp; Ryland, J. S." box="[858,1145,1711,1738]" pageId="3" pageNumber="476" pagination="1 - 147" refId="ref12323" refString="Hayward, P. J. &amp; Ryland, J. S. (1985) Cyclostome Bryozoans. In: Kermack, D. M. &amp; Barnes, R. S. K. (Eds.), Synopses of the British Fauna. New Series. Vol. 34. E. J. Brill for the Linnaean Society, London, pp. 1 - 147." type="book chapter" year="1985">Hayward &amp; Ryland 1985</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="93B965363B25F477BDFC46EAE69EFEB5" lastPageId="4" lastPageNumber="477" pageId="3" pageNumber="476" type="reference_group">
<paragraph id="DB1C36BD3B25F470BDFC46EAE73EF808" blockId="3.[151,1437,1604,2026]" pageId="3" pageNumber="476">
<taxonomicName id="1CA34D3E3B25F470BDFC46EAE44EF944" ID-CoL="c1047237-a7e0-49ba-8174-605b29a9591a" authority="Smitt" authorityName="Smitt" authorityYear="1875" box="[199,571,1748,1774]" class="Stenolaemata" family="Crisiidae" genus="Crisia" kingdom="Animalia" order="Cyclostomatida" pageId="3" pageNumber="476" phylum="Bryozoa" rank="species" species="eburneo-denticulata">
<emphasis id="E9D7EAAF3B25F470BDFC46EAE782F944" box="[199,503,1748,1774]" italics="true" pageId="3" pageNumber="476">Crisia eburneo-denticulata</emphasis>
Smitt
</taxonomicName>
ms in
<bibRefCitation id="BF324B4C3B25F470BFBE46EAE572F944" author="Busk, G." box="[645,775,1747,1774]" pageId="3" pageNumber="476" refId="ref11837" refString="Busk, G. (1875) Catalogue of marine Polyzoa in the collection of the British Museum. III. Cyclostomata. Trustees of the British Museum (Natural History), London, 39 pp." type="book" year="1875">Busk, 1875</bibRefCitation>
, distinctly differs in the position of the gonozooid, occupying approximately three-fourths of the fertile branch width, and shifted slightly towards the margin of the branch. One fully developed autozooid is always situated alongside the margin of the gonozooid.
<taxonomicName id="1CA34D3E3B25F470B98F4722E698F8F0" ID-CoL="c1047237-a7e0-49ba-8174-605b29a9591a" authorityName="Smitt" authorityYear="1875" class="Stenolaemata" family="Crisiidae" genus="Crisia" kingdom="Animalia" order="Cyclostomatida" pageId="3" pageNumber="476" phylum="Bryozoa" rank="species" species="eburneo-denticulata">
<emphasis id="E9D7EAAF3B25F470B98F4722E698F8F0" italics="true" pageId="3" pageNumber="476">Crisia eburneo-denticulata</emphasis>
</taxonomicName>
has been reported mainly from the northern Atlantic (
<bibRefCitation id="BF324B4C3B25F470BE74477EE257F8F0" author="Brunel, P. &amp; Bosse, L. &amp; Lamarche, G." box="[847,1058,1856,1882]" pageId="3" pageNumber="476" pagination="1 - 405" refId="ref11792" refString="Brunel, P., Bosse, L. &amp; Lamarche, G. (1998) Catalogue of the marine invertebrates of the Estuary and Gulf of Saint-Lawrence. Canadian Special Publication of Fisheries and Aquatic Sciences, 126, 1 - 405." type="journal article" year="1998">
Brunel
<emphasis id="E9D7EAAF3B25F470BE9E477FE5ABF8F0" box="[933,990,1856,1882]" italics="true" pageId="3" pageNumber="476">et al.</emphasis>
1998
</bibRefCitation>
). The other occurrences of
<taxonomicName id="1CA34D3E3B25F470B86D477EE70FF8D4" ID-CoL="c1047237-a7e0-49ba-8174-605b29a9591a" authorityName="Smitt" authorityYear="1875" class="Stenolaemata" family="Crisiidae" genus="Crisia" kingdom="Animalia" order="Cyclostomatida" pageId="3" pageNumber="476" phylum="Bryozoa" rank="species" species="eburneo-denticulata">
<emphasis id="E9D7EAAF3B25F470B86D477EE70FF8D4" italics="true" pageId="3" pageNumber="476">Crisia eburneo-denticulata</emphasis>
</taxonomicName>
in Asia are doubtful (e.g.
<bibRefCitation id="BF324B4C3B25F470BFA0475AE57DF8D4" author="Seo, J. E." box="[667,776,1891,1918]" pageId="3" pageNumber="476" refId="ref13271" refString="Seo, J. E. (2005) Illustrated encyclopedia of fauna &amp; flora of Korea. Vol. 40. Bryozoa. Daehan Printing &amp; Publishing Co, Ltd., Seoul, 596 pp." type="book" year="2005">Seo 2005</bibRefCitation>
), so it is highly probable that this species is absent from the western Pacific.
</paragraph>
<paragraph id="DB1C36BD3B25F477BDFC4792E69EFEB5" blockId="3.[151,1437,1604,2026]" lastBlockId="4.[151,1436,153,360]" lastPageId="4" lastPageNumber="477" pageId="3" pageNumber="476">
The specimens described and illustrated by
<bibRefCitation id="BF324B4C3B25F470BF8C4792E273F86C" author="Mawatari, S. &amp; Mawatari, S. F." box="[695,1030,1964,1990]" pageId="3" pageNumber="476" pagination="95 - 104" refId="ref12988" refString="Mawatari, S. &amp; Mawatari, S. F. (1973) Notes on marine Bryozoa from Hokkaido. 1. Crisiidae (Cyclostomata). Journal of the Faculty of Science, Hokkaido University, 19 (1), 95 - 104." type="journal article" year="1973">Mawatari &amp; Mawatari (1973)</bibRefCitation>
as
<taxonomicName id="1CA34D3E3B25F470B9164792E2ADF86C" ID-CoL="ZHPW" authorityName=": Mawatari &amp; Mawatari" authorityYear="1973" box="[1069,1240,1964,1990]" class="Stenolaemata" family="Crisiidae" genus="Crisia" kingdom="Animalia" order="Cyclostomatida" pageId="3" pageNumber="476" phylum="Bryozoa" rank="species" species="eburnea">
<emphasis id="E9D7EAAF3B25F470B9164792E2ADF86C" box="[1069,1240,1964,1990]" italics="true" pageId="3" pageNumber="476">Crisia eburnea</emphasis>
</taxonomicName>
as well as
<taxonomicName id="1CA34D3E3B25F470B86D4792E70FF840" ID-CoL="c1047237-a7e0-49ba-8174-605b29a9591a" authorityName="Smitt" authorityYear="1875" class="Stenolaemata" family="Crisiidae" genus="Crisia" kingdom="Animalia" order="Cyclostomatida" pageId="3" pageNumber="476" phylum="Bryozoa" rank="species" species="eburneo-denticulata">
<emphasis id="E9D7EAAF3B25F470B86D4792E70FF840" italics="true" pageId="3" pageNumber="476">Crisia eburneo-denticulata</emphasis>
</taxonomicName>
show the same extension of the gonozooid and similar position and shape of the ooeciopore. These two species differ according to the original
<bibRefCitation id="BF324B4C3B22F477BFFE40A7E257FF19" author="Mawatari, S. &amp; Mawatari, S. F." box="[709,1058,153,179]" pageId="4" pageNumber="477" pagination="95 - 104" refId="ref12988" refString="Mawatari, S. &amp; Mawatari, S. F. (1973) Notes on marine Bryozoa from Hokkaido. 1. Crisiidae (Cyclostomata). Journal of the Faculty of Science, Hokkaido University, 19 (1), 95 - 104." type="journal article" year="1973">Mawatari &amp; Mawataris (1973)</bibRefCitation>
description only in the size of the ooeciopore. This feature might be, however, just an extreme variant in a species with ooeciopores that vary greatly in size. As we do not have the original material at our disposal, the identification of these specimens remains uncertain.
</paragraph>
</subSubSection>
<subSubSection id="93B965363B22F477BDFC4117E24AFEC2" pageId="4" pageNumber="477" type="description">
<paragraph id="DB1C36BD3B22F477BDFC4117E3EEFEE9" blockId="4.[151,1436,153,360]" box="[199,1435,297,323]" pageId="4" pageNumber="477">The most distinctive feature of this species is the very large size of the ooeciopore on the extended gonozooid.</paragraph>
<paragraph id="DB1C36BD3B22F477BDFC4172E24AFEC2" blockId="4.[151,1436,153,360]" box="[199,1087,332,360]" pageId="4" pageNumber="477">
<emphasis id="E9D7EAAF3B22F477BDFC4172E72FFECD" bold="true" box="[199,346,332,359]" pageId="4" pageNumber="477">Occurrence.</emphasis>
Haijiao (Lingshan Island),
<collectingCountry id="A3B4762D3B22F477BFB54173E553FECD" box="[654,806,333,359]" name="China" pageId="4" pageNumber="477">Jiaozhou Bay</collectingCountry>
and Yangkou (Laoshan).
</paragraph>
</subSubSection>
</treatment>
</document>