180 lines
21 KiB
XML
180 lines
21 KiB
XML
<document id="A767C265C9F86FED71222B296B1FF3FB" ID-DOI="10.15298/rusentj.30.3.05" ID-ISSN="0132-8069" ID-Zenodo-Dep="13179717" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722596800136" checkinUser="felipe" docAuthor="Lyubarsky, G. Yu. & Perkovsky, E. E." docDate="2021" docId="FA7A878A2543FFD31359F8601FCBFCA6" docLanguage="en" docName="RussEntomolJ.30.3.282-287.pdf" docOrigin="Russian Entomological Journal 30 (3)" docSource="http://dx.doi.org/10.15298/rusentj.30.3.05" docStyle="DocumentStyle:0D5BAC5AA0C130B3809AF578A601D837.3:RussEntomolJ.2019-.journal_article" docStyleId="0D5BAC5AA0C130B3809AF578A601D837" docStyleName="RussEntomolJ.2019-.journal_article" docStyleVersion="3" docTitle="Atomaria groehni Perkovsky et Lyubarsky 2014" docType="treatment" docVersion="2" lastPageNumber="286" masterDocId="0643FFF22542FFD71028FFD41D09FFB1" masterDocTitle="New findings of Cryptophagidae (Coleoptera: Clavicornia) from Baltic amber in the unbiased collection of the Paleontological Institute of RAS" masterLastPageNumber="287" masterPageNumber="282" pageNumber="283" updateTime="1722637898107" updateUser="ExternalLinkService">
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<mods:title id="8CA4CAFE1E732DA720E28FCEFF26F9F5">New findings of Cryptophagidae (Coleoptera: Clavicornia) from Baltic amber in the unbiased collection of the Paleontological Institute of RAS</mods:title>
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<mods:namePart id="42DAA46F50BCBCEC03F359204CA2CABB">Lyubarsky, G. Yu.</mods:namePart>
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<mods:namePart id="FA8603C9C21ADD07698FE25771815EB5">Perkovsky, E. E.</mods:namePart>
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<mods:title id="B867176A3B5AE5A68D8488D4077C62EA">Russian Entomological Journal</mods:title>
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<mods:part id="2528F6509AEBDD8BE69984F5961891D1">
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<mods:date id="4FD91C77024A1943DCDBF6A7E5867721">2021</mods:date>
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<mods:number id="8D1F763DE2C7DB258160E6CC6BA20ACE">2021-09-30</mods:number>
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<mods:number id="18EDE15EEEE8B1D6BFB10B9D02C18CEB">30</mods:number>
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<mods:url id="BB3E0E2899BC5D21DAD662BEC36C9D38">http://dx.doi.org/10.15298/rusentj.30.3.05</mods:url>
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<mods:identifier id="4690A37577D8A494A3C078262169C10C" type="DOI">10.15298/rusentj.30.3.05</mods:identifier>
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<mods:identifier id="F5F72E0BBFDE26700CA777F454A7C4D9" type="ISSN">0132-8069</mods:identifier>
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<treatment id="FA7A878A2543FFD31359F8601FCBFCA6" ID-DOI="http://doi.org/10.5281/zenodo.13179725" ID-Zenodo-Dep="13179725" LSID="urn:lsid:plazi:treatment:FA7A878A2543FFD31359F8601FCBFCA6" httpUri="http://treatment.plazi.org/id/FA7A878A2543FFD31359F8601FCBFCA6" lastPageId="4" lastPageNumber="286" pageId="1" pageNumber="283">
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<subSubSection id="3AC965172543FFD61359F860198DF859" pageId="1" pageNumber="283" type="nomenclature">
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<paragraph id="726C369C2543FFD61359F860198DF859" blockId="1.[850,1398,1971,2057]" pageId="1" pageNumber="283">
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<heading id="292481F02543FFD61359F860198DF859" centered="true" fontSize="10" level="2" pageId="1" pageNumber="283" reason="8">
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<taxonomicName id="B5D34D1F2543FFD61359F860183BF87A" authority="Perkovsky et Lyubarsky, 2014" authorityName="Perkovsky et Lyubarsky" authorityYear="2014" box="[881,1330,1971,1995]" class="Insecta" family="Cryptophagidae" genus="Atomaria" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="283" phylum="Arthropoda" rank="species" species="groehni">
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<emphasis id="40A7EA8E2543FFD61359F8601EEBF87A" box="[881,994,1971,1995]" italics="true" pageId="1" pageNumber="283">A. groehni</emphasis>
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Perkovsky et
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<bibRefCitation id="16424B6D2543FFD61453F867183BF87A" author="Lyubarsky G. Yu. & Perkovsky E. E." box="[1147,1330,1971,1995]" pageId="1" pageNumber="283" pagination="191 - 194" refId="ref4132" refString="Lyubarsky G. Yu., Perkovsky E. E. 2014. New species of the genus Nganasania from Upper Cretaceous of Taymyr (Coleoptera: Cryptophagidae) // Russian Entomol. J. Vol. 23. No. 3. P. 191 - 194." type="journal article" year="2014">Lyubarsky, 2014</bibRefCitation>
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</taxonomicName>
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<figureCitation id="EAE82A192543FFD61437F8051989F859" box="[1055,1152,2000,2025]" captionStart="Figs 5–6" captionStartId="3.[226,263,2014,2033]" captionTargetBox="[189,1399,474,2000]" captionTargetId="figure-133@3.[189,1399,474,2000]" captionTargetPageId="3" captionText="Figs 5–6. Photo of a new specimen of A. groehni from Baltic amber: 5 — dorsal view; 6 — dorsolateral view. Рис. 5–6. ВнеШний вид A. groehni иЗ балтийского ЯнтарЯ: 5 — сверху; 6 — сверху и сбоку." figureDoi="http://doi.org/10.5281/zenodo.13179723" httpUri="https://zenodo.org/record/13179723/files/figure.png" pageId="1" pageNumber="283">Figs 5–6</figureCitation>
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.
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="3AC965172543FFD3137AF8221FC3FE36" lastPageId="4" lastPageNumber="286" pageId="1" pageNumber="283" type="materials_examined">
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<paragraph id="726C369C2543FFD5137AF8221C1CFED7" blockId="1.[850,1398,1971,2057]" lastBlockId="2.[189,776,315,599]" lastPageId="2" lastPageNumber="284" pageId="1" pageNumber="283">
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MATERIAL.
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<collectionCode id="14C2AE592543FFD613E5F8221EF9F7B8" box="[973,1008,2038,2057]" country="Russia" httpUri="http://grbio.org/cool/kjjc-ziy5" name="Paleontological Institute, Russian Academy of Sciences" pageId="1" pageNumber="283">PIN</collectionCode>
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964/1325, Baltic amber, Late Eocene. Specimen is deposited in the amber collection of the
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<collectionCode id="14C2AE592540FFD51254FEE81F96FEFE" box="[636,671,316,335]" country="Russia" httpUri="http://grbio.org/cool/kjjc-ziy5" name="Paleontological Institute, Russian Academy of Sciences" pageId="2" pageNumber="284">PIN</collectionCode>
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RAS. Sex: unknown.
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</paragraph>
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<paragraph id="726C369C2540FFD510CAFEBF1F16FE60" blockId="2.[189,776,315,599]" pageId="2" pageNumber="284">
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RE-DESCRIPTION. Length of body 1.0–
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<quantity id="B52B9B792540FFD51257FEBF1FC2FE30" box="[639,715,363,385]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="2" pageNumber="284" unit="mm" value="1.1">1.1 mm</quantity>
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, body elongate (
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<figureCitation id="EAE82A192540FFD51133FE521C7BFE2D" box="[283,370,390,412]" captionStart="Figs 5–6" captionStartId="3.[226,263,2014,2033]" captionTargetBox="[189,1399,474,2000]" captionTargetId="figure-133@3.[189,1399,474,2000]" captionTargetPageId="3" captionText="Figs 5–6. Photo of a new specimen of A. groehni from Baltic amber: 5 — dorsal view; 6 — dorsolateral view. Рис. 5–6. ВнеШний вид A. groehni иЗ балтийского ЯнтарЯ: 5 — сверху; 6 — сверху и сбоку." figureDoi="http://doi.org/10.5281/zenodo.13179723" httpUri="https://zenodo.org/record/13179723/files/figure.png" pageId="2" pageNumber="284">Figs 5–6</figureCitation>
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), moderately arched, head, pronotum, and elytra light-brown.Elytra slightly convex, covered with slightly curved long elevated pubescence.
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</paragraph>
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<paragraph id="726C369C2540FFD510CAFE0219ECFE0A" blockId="2.[189,776,315,599]" lastBlockId="2.[812,1399,315,604]" pageId="2" pageNumber="284">
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Head transverse, of normal size, with hemispherical facetted eyes, strongly punctured. Antennae moderately long, with club reaching beyond base of pronotum, flagellomeres slightly elongate, antennomeres 1
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<superScript id="85A69BD42540FFD511E7FDF11CD1FD80" attach="none" box="[463,472,549,561]" fontSize="5" pageId="2" pageNumber="284">st</superScript>
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, 2
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<superScript id="85A69BD42540FFD511DAFDF11F09FD80" attach="left" box="[498,512,549,561]" fontSize="5" pageId="2" pageNumber="284">nd</superScript>
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about 1.5 times as long as broad, 3
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<superScript id="85A69BD42540FFD51125FD941C10FDFD" attach="left" box="[269,281,576,588]" fontSize="5" pageId="2" pageNumber="284">rd</superScript>
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slightly elongate, 4
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<superScript id="85A69BD42540FFD511F3FD941CEFFDFD" attach="left" box="[475,486,576,588]" fontSize="5" pageId="2" pageNumber="284">th</superScript>
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and 6
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<superScript id="85A69BD42540FFD51209FD941F25FDFD" attach="left" box="[545,556,576,588]" fontSize="5" pageId="2" pageNumber="284">th</superScript>
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antennomeres as long as broad, 5
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<superScript id="85A69BD42540FFD513BBFEEF1E97FEF6" attach="left" box="[915,926,315,327]" fontSize="5" pageId="2" pageNumber="284">th</superScript>
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1.5 times as long as broad, 7
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<superScript id="85A69BD42540FFD51486FEEF19B0FEF6" attach="none" box="[1198,1209,315,327]" fontSize="5" pageId="2" pageNumber="284">th</superScript>
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, 8
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<superScript id="85A69BD42540FFD514F8FEEF19D2FEF6" attach="left" box="[1232,1243,315,327]" fontSize="5" pageId="2" pageNumber="284">th</superScript>
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subquadrate, 9
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<superScript id="85A69BD42540FFD51543FEEF187FFEF6" attach="none" box="[1387,1398,315,327]" fontSize="5" pageId="2" pageNumber="284">th</superScript>
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slightly transverse, 10
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<superScript id="85A69BD42540FFD51423FE81191FFED0" attach="left" box="[1035,1046,341,353]" fontSize="5" pageId="2" pageNumber="284">th</superScript>
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strongly transverse, 11
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<superScript id="85A69BD42540FFD5152DFE811819FED0" attach="left" box="[1285,1296,341,353]" fontSize="5" pageId="2" pageNumber="284">th</superScript>
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obliquely oval, antennomeres 9–11 equal in width. Antennae inserted under the lateral margin of the forehead, widely separated basally, antennal insertions close to the eyes.
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</paragraph>
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<paragraph id="726C369C2540FFD4137AFE151C50FE0D" blockId="2.[812,1399,315,604]" lastBlockId="3.[189,775,315,444]" lastPageId="3" lastPageNumber="285" pageId="2" pageNumber="284">Pronotum convex, not narrowed basally, slightly transverse, 1.2–1.5 times broader than long, broadest before the middle, moderately strongly and very densely punctured (distance between punctures equal to their diameter), a single puncture smaller than the diameter of the eye facet. Pubescence long and elevated. Sides finely margined, anterior edge weakly sinuate, without excision. Pronotum broadest before middle of its length. Side borders visible from above along the entire length. Lateral margin without callosity. Posterior corners right angled, basal edge lobed, strongly sinuate, basal groove shallow.</paragraph>
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<caption id="26AC66142540FFD510CAF87A1F3AF7BC" ID-DOI="http://doi.org/10.5281/zenodo.13179721" ID-Zenodo-Dep="13179721" httpUri="https://zenodo.org/record/13179721/files/figure.png" pageId="2" pageNumber="284" startId="2.[226,263,1966,1985]" targetBox="[189,1399,634,1954]" targetPageId="2" targetType="figure">
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<paragraph id="726C369C2540FFD510CAF87A1CE1F868" blockId="2.[189,1399,1966,2061]" pageId="2" pageNumber="284">
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Figs 1–4. Photo of a new specimen of
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<taxonomicName id="B5D34D1F2540FFD51206F87A1F92F871" authorityName="Lyubarsky et Perkovsky" authorityYear="2010" box="[558,667,1966,1984]" class="Insecta" family="Cryptophagidae" genus="Micrambe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="284" phylum="Arthropoda" rank="species" species="sarnensis">
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<emphasis id="40A7EA8E2540FFD51206F87A1F92F871" box="[558,667,1966,1984]" italics="true" pageId="2" pageNumber="284">M. sarnensis</emphasis>
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</taxonomicName>
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from Baltic amber: 1 — dorsal view; 2 — lateral view; 3 — ventral view; 4 — head and pronotum, ventrolateral view.
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</paragraph>
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<paragraph id="726C369C2540FFD510CAF8361F3AF7BC" blockId="2.[189,1399,1966,2061]" pageId="2" pageNumber="284">
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Рис. 1–4. ВнеШний вид
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<taxonomicName id="B5D34D1F2540FFD51196F8361F24F845" authorityName="Lyubarsky et Perkovsky" authorityYear="2010" box="[446,557,2018,2036]" class="Insecta" family="Cryptophagidae" genus="Micrambe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="284" phylum="Arthropoda" rank="species" species="sarnensis">
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<emphasis id="40A7EA8E2540FFD51196F8361F24F845" box="[446,557,2018,2036]" italics="true" pageId="2" pageNumber="284">M. sarnensis</emphasis>
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</taxonomicName>
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иЗ балтийского ЯнтарЯ: 1 — габитус, сверху; 2 — габитус, сбоку; 3 — габитус,сниЗу; 4 — голова и переднеспинка, сниЗу и сбоку.
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</paragraph>
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</caption>
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<paragraph id="726C369C2541FFD3137AFEEF1FC3FE36" blockId="3.[812,1398,315,444]" lastBlockId="4.[189,776,315,791]" lastPageId="4" lastPageNumber="286" pageId="3" pageNumber="285">Scutellum small, transverse. Elytra short-oval, moderately convex, humeral corners rounded, maximum breadth of elytra in first third of their length, 1.6–1.7 times as long as wide and 2.5–2.7 times as long as thorax, punctuation slightly stronger and sparser than that on pronotum. Punctures in the basal part slightly stronger than those on the pronotal disk, and approximately 1.5–2 diameters apart from their lateral neighbours on an average; humeri not toothed.</paragraph>
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</subSubSection>
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<caption id="26AC66142541FFD410CAF80A193AF7BC" ID-DOI="http://doi.org/10.5281/zenodo.13179723" ID-Zenodo-Dep="13179723" httpUri="https://zenodo.org/record/13179723/files/figure.png" pageId="3" pageNumber="285" startId="3.[226,263,2014,2033]" targetBox="[189,1399,474,2000]" targetPageId="3" targetType="figure">
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<paragraph id="726C369C2541FFD410CAF80A19A2F840" blockId="3.[226,1195,2013,2061]" box="[226,1195,2013,2033]" pageId="3" pageNumber="285">
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Figs 5–6. Photo of a new specimen of
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<taxonomicName id="B5D34D1F2541FFD4121DF80A1F99F840" authorityName="Perkovsky et Lyubarsky" authorityYear="2014" box="[565,656,2014,2033]" class="Insecta" family="Cryptophagidae" genus="Atomaria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="285" phylum="Arthropoda" rank="species" species="groehni">
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<emphasis id="40A7EA8E2541FFD4121DF80A1F99F840" box="[565,656,2014,2033]" italics="true" pageId="3" pageNumber="285">A. groehni</emphasis>
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</taxonomicName>
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from Baltic amber: 5 — dorsal view; 6 — dorsolateral view.
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</paragraph>
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<paragraph id="726C369C2541FFD410CAF82E193AF7BC" blockId="3.[226,1195,2013,2061]" box="[226,1075,2041,2061]" pageId="3" pageNumber="285">
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Рис. 5–6. ВнеШний вид
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<taxonomicName id="B5D34D1F2541FFD41197F82E1F12F7BD" authorityName="Perkovsky et Lyubarsky" authorityYear="2014" box="[447,539,2041,2061]" class="Insecta" family="Cryptophagidae" genus="Atomaria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="285" phylum="Arthropoda" rank="species" species="groehni">
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<emphasis id="40A7EA8E2541FFD41197F82E1F12F7BD" box="[447,539,2041,2061]" italics="true" pageId="3" pageNumber="285">A. groehni</emphasis>
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</taxonomicName>
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иЗ балтийского ЯнтарЯ: 5 — сверху; 6 — сверху и сбоку.
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</paragraph>
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</caption>
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<subSubSection id="3AC965172546FFD310CAFE5F1FCBFCA6" pageId="4" pageNumber="286" type="discussion">
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<paragraph id="726C369C2546FFD310CAFE5F1F2AFE67" blockId="4.[189,776,315,791]" pageId="4" pageNumber="286">
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REMARKS. Representatives of
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<taxonomicName id="B5D34D1F2546FFD3120EFE581F88FE10" authorityName="Stephens" authorityYear="1829" box="[550,641,396,417]" class="Insecta" family="Cryptophagidae" genus="Atomaria" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="286" phylum="Arthropoda" rank="genus">
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<emphasis id="40A7EA8E2546FFD3120EFE581F88FE10" box="[550,641,396,417]" italics="true" pageId="4" pageNumber="286">Atomaria</emphasis>
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</taxonomicName>
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are found in all biogeographic realms including 127 species in the Palaearctic Region [
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<bibRefCitation id="16424B6D2546FFD31164FE151F11FE67" author="Johnson C. & Otero J. C. & Leschen R. A. B." box="[332,536,449,471]" pageId="4" pageNumber="286" pagination="513 - 531" refId="ref3479" refString="Johnson C., Otero J. C., Leschen R. A. B. 2007. Family Cryptophagidae // Lobl I., Smetana A. (eds.). A catalogue of Palaearctic Coleoptera. Vol. 4. Steenstrup: Apollo Books. P. 513 - 531." type="book chapter" year="2007">Johnson et al., 2007</bibRefCitation>
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].
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</paragraph>
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<paragraph id="726C369C2546FFD310CAFE081E01FD1D" blockId="4.[189,776,315,791]" pageId="4" pageNumber="286">
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<taxonomicName id="B5D34D1F2546FFD310CAFE081C84FE41" authorityName="Perkovsky et Lyubarsky" authorityYear="2014" box="[226,397,475,497]" class="Insecta" family="Cryptophagidae" genus="Atomaria" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="286" phylum="Arthropoda" rank="species" species="groehni">
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<emphasis id="40A7EA8E2546FFD310CAFE081C84FE41" box="[226,397,475,497]" italics="true" pageId="4" pageNumber="286">Atomaria groehni</emphasis>
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</taxonomicName>
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differs from similar species with lateral margin of pronotum visible from above, such as
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<taxonomicName id="B5D34D1F2546FFD31287FE221C40FD97" authority="Kolenati, 1846" authorityName="Kolenati" authorityYear="1846" class="Insecta" family="Cryptophagidae" genus="Atomaria" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="286" phylum="Arthropoda" rank="species" species="morio">
|
||
<emphasis id="40A7EA8E2546FFD31287FE221E0EFDBD" box="[687,775,502,524]" italics="true" pageId="4" pageNumber="286">A. morio</emphasis>
|
||
Kolenati, 1846
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="B5D34D1F2546FFD3115DFDC51F8DFD97" authority="Erichson, 1846" authorityName="Erichson" authorityYear="1846" box="[373,644,528,551]" class="Insecta" family="Cryptophagidae" genus="Atomaria" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="286" phylum="Arthropoda" rank="species" species="versicolor">
|
||
<emphasis id="40A7EA8E2546FFD3115DFDC51CE7FD97" box="[373,494,529,550]" italics="true" pageId="4" pageNumber="286">A. versicolor</emphasis>
|
||
Erichson, 1846
|
||
</taxonomicName>
|
||
, by small size and long elevated pubescence.
|
||
<taxonomicName id="B5D34D1F2546FFD311CAFDF81F4FFDF1" authorityName="Perkovsky et Lyubarsky" authorityYear="2014" box="[482,582,555,577]" class="Insecta" family="Cryptophagidae" genus="Atomaria" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="286" phylum="Arthropoda" rank="species" species="groehni">
|
||
<emphasis id="40A7EA8E2546FFD311CAFDF81F4FFDF1" box="[482,582,555,577]" italics="true" pageId="4" pageNumber="286">A. groehni</emphasis>
|
||
</taxonomicName>
|
||
differs from species similar to
|
||
<taxonomicName id="B5D34D1F2546FFD3110DFD921F3BFDEA" authority="Stephens, 1830" authorityName="Stephens" authorityYear="1830" box="[293,562,582,604]" class="Insecta" family="Cryptophagidae" genus="Atomaria" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="286" phylum="Arthropoda" rank="species" species="testacea">
|
||
<emphasis id="40A7EA8E2546FFD3110DFD921C98FDED" box="[293,401,582,604]" italics="true" pageId="4" pageNumber="286">A. testacea</emphasis>
|
||
Stephens,
|
||
<quantity id="B52B9B792546FFD3122AFD921F3BFDEA" box="[514,562,582,603]" metricMagnitude="1" metricUnit="m" metricValue="4.6482" pageId="4" pageNumber="286" unit="in" value="1830.0">1830</quantity>
|
||
</taxonomicName>
|
||
in pronotum slightly narrowed basally and small size.
|
||
<taxonomicName id="B5D34D1F2546FFD311DFFDB51F55FDC7" authorityName="Perkovsky et Lyubarsky" authorityYear="2014" box="[503,604,609,630]" class="Insecta" family="Cryptophagidae" genus="Atomaria" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="286" phylum="Arthropoda" rank="species" species="groehni">
|
||
<emphasis id="40A7EA8E2546FFD311DFFDB51F55FDC7" box="[503,604,609,630]" italics="true" pageId="4" pageNumber="286">A. groehni</emphasis>
|
||
</taxonomicName>
|
||
differs from most species of the genus in the structure of the antenna: the segments of the flagellum are subquadrate or slightly elongate.
|
||
</paragraph>
|
||
<paragraph id="726C369C2546FFD310CAFD651FCBFCA6" blockId="4.[189,776,315,791]" pageId="4" pageNumber="286">Among the species of amber, it differs in a combination of characters: pronotum broadest before the middle of its length; elytra covered by long elevated pubescence; side borders of pronotum visible from above along the entire length.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |