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<document id="B66DCC773892B84B70E51C7FFB0D3464" ID-DOI="10.11646/phytotaxa.266.2.4" ID-ISSN="1179-3163" ID-Zenodo-Dep="13659734" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725388660066" checkinUser="felipe" docAuthor="Jagadeesh Ram, T. A. M. &amp; Sinha, G. P." docDate="2016" docId="F3749E5F8026F94CF2C0EF1FFA362E06" docLanguage="en" docName="phytotaxa.266.2.4.pdf" docOrigin="Phytotaxa 266 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.266.2.4" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Cryptothecia albomaculans Jagadeesh &amp; G. P. Sinha 2016, sp. nov." docType="treatment" docVersion="2" lastPageNumber="105" masterDocId="0F4DE6278027F94EF248EC2BFF9E295B" masterDocTitle="A world key to Cryptothecia and Myriostigma (Arthoniaceae), with new species and new records from the Andaman and Nicobar Islands, India" masterLastPageNumber="114" masterPageNumber="103" pageNumber="104" updateTime="1725405533529" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="75D3F054E6B0A6C010736472ACEBD735">A world key to Cryptothecia and Myriostigma (Arthoniaceae), with new species and new records from the Andaman and Nicobar Islands, India</mods:title>
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<mods:namePart id="0D5AF1F92708F7C3A7E992AA8E2923F7">Jagadeesh Ram, T. A. M.</mods:namePart>
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<mods:namePart id="C4D046BCF64019C21F8282641C31BF3A">Sinha, G. P.</mods:namePart>
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<mods:title id="2DEA5E2D0C373A7D2D6FB0C9FE29F0BC">Phytotaxa</mods:title>
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<mods:date id="7B6C9415C8D21DA81834B644A2F10B3A">2016</mods:date>
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<mods:identifier id="45B026418B919E5C91D935613F328052" type="DOI">10.11646/phytotaxa.266.2.4</mods:identifier>
<mods:identifier id="846BBAF8DACE47CCB6E68B1741671441" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="4B9B96ABA595F2909D4C8D38F3B45618" type="Zenodo-Dep">13659734</mods:identifier>
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<treatment id="F3749E5F8026F94CF2C0EF1FFA362E06" LSID="urn:lsid:plazi:treatment:F3749E5F8026F94CF2C0EF1FFA362E06" httpUri="http://treatment.plazi.org/id/F3749E5F8026F94CF2C0EF1FFA362E06" lastPageId="2" lastPageNumber="105" pageId="1" pageNumber="104">
<subSubSection id="33C77CC28026F94FF2C0EF1FFE542A28" pageId="1" pageNumber="104" type="nomenclature">
<paragraph id="7B622F498026F94FF2C0EF1FFC6E2A14" blockId="1.[136,1008,820,847]" box="[136,1008,820,847]" pageId="1" pageNumber="104">
<heading id="202A98258026F94FF2C0EF1FFC6E2A14" box="[136,1008,820,847]" fontSize="11" level="2" pageId="1" pageNumber="104" reason="3">
<taxonomicName id="BCDD54CA8026F94FF2C0EF1FFD442A14" ID-CoL="6BNKZ" authority="Jagadeesh &amp; G.P. Sinha" authorityName="Jagadeesh &amp; G. P. Sinha" authorityYear="2016" box="[136,730,820,847]" class="Arthoniomycetes" family="Arthoniaceae" genus="Cryptothecia" kingdom="Fungi" order="Arthoniales" pageId="1" pageNumber="104" phylum="Ascomycota" rank="species" species="albomaculans" status="sp. nov.">
<emphasis id="49A9F35B8026F94FF2C0EF1FFE5B2A14" bold="true" box="[136,453,820,847]" italics="true" pageId="1" pageNumber="104">Cryptothecia albomaculans</emphasis>
Jagadeesh &amp; G.P. Sinha
</taxonomicName>
,
<emphasis id="49A9F35B8026F94FF0ACEF1DFCA72A14" box="[740,825,822,847]" italics="true" pageId="1" pageNumber="104">
<taxonomicNameLabel id="529A4E208026F94FF0ACEF1DFCA72A14" box="[740,825,822,847]" pageId="1" pageNumber="104" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="E3E633CC8026F94FF10FEF1DFC3F2A14" box="[839,929,822,847]" captionStart="FIGURE 1" captionStartId="2.[136,229,1654,1675]" captionTargetBox="[158,1430,190,1629]" captionTargetId="figure-19@2.[158,1430,190,1629]" captionTargetPageId="2" captionText="FIGURE 1. A. Cryptothecia albomaculans (holotype), part of thallus with white maculiform ascigerous areas. B. C. elata (holotype), part of thallus with white raised ascigerous areas. C. C. elongata (holotype), part of thallus with slightly raised ascigerous areas. D. C. superphyllinica (holotype), part of thallus with white raised ascigerous areas. E. Myriostigma nicobaricum (holotype), part of thallus with white raised ascigerous areas. F. M. nicobaricum (paratype). Scales:A–F=1 mm." figureDoi="http://doi.org/10.5281/zenodo.13659738" httpUri="https://zenodo.org/record/13659738/files/figure.png" pageId="1" pageNumber="104">Figs 1A</figureCitation>
&amp;
<figureCitation id="E3E633CC8026F94FF18DEF1DFC772A14" box="[965,1001,822,847]" captionStart="FIGURE 2" captionStartId="3.[136,229,1672,1693]" captionTargetBox="[151,1435,675,1646]" captionTargetId="figure-229@3.[151,1436,674,1646]" captionTargetPageId="3" captionText="FIGURE 2. A. Cryptothecia albomaculans (holotype), ascospores. B. C. elata (holotype), ascospores. C–D. C. elongata (holotype). C. Ascus. D. Ascospores. E. C. superphyllinica (holotype), ascospores. F. Myriostigma nicobaricum (holotype), ascospores. Scales=20 μm." figureDoi="http://doi.org/10.5281/zenodo.13659742" httpUri="https://zenodo.org/record/13659742/files/figure.png" pageId="1" pageNumber="104">2A</figureCitation>
)
</heading>
</paragraph>
<paragraph id="7B622F498026F94FF2C0EF71FE542A28" blockId="1.[136,458,858,883]" box="[136,458,858,883]" pageId="1" pageNumber="104">MycoBank No.: MB 817295</paragraph>
</subSubSection>
<subSubSection id="33C77CC28026F94FF2C0EF8FFC842A86" pageId="1" pageNumber="104" type="description">
<paragraph id="7B622F498026F94FF2C0EF8FFC842A86" blockId="1.[136,1451,932,1061]" pageId="1" pageNumber="104">
<taxonomicName id="BCDD54CA8026F94FF2C0EF8FFE9B2AE2" box="[136,261,932,953]" class="Arthoniomycetes" family="Arthoniaceae" genus="Cryptothecia" kingdom="Fungi" order="Arthoniales" pageId="1" pageNumber="104" phylum="Ascomycota" rank="genus">
<emphasis id="49A9F35B8026F94FF2C0EF8FFE9B2AE2" box="[136,261,932,953]" italics="true" pageId="1" pageNumber="104">Cryptothecia</emphasis>
</taxonomicName>
with heteromerous thallus, whitish ascigerous areas not raised above the thallus; 8-spored asci; ellipsoid, 9–11 × 3–5 septate, 47–57 × 24–30 μm ascospores and 2’-
<emphasis id="49A9F35B8026F94FF066EFE3FDA12A86" box="[558,575,968,989]" italics="true" pageId="1" pageNumber="104">O</emphasis>
-methylperlatolic acid.
</paragraph>
</subSubSection>
<subSubSection id="33C77CC28026F94FF2C0EFC7FB562EDC" pageId="1" pageNumber="104" type="materials_examined">
<paragraph id="7B622F498026F94FF2C0EFC7FC532D7E" blockId="1.[136,1451,932,1061]" pageId="1" pageNumber="104">
<emphasis id="49A9F35B8026F94FF2C0EFC7FF5C2D5A" bold="true" box="[136,194,1004,1025]" pageId="1" pageNumber="104">Type:</emphasis>
—
<collectingCountry id="03CA6FD98026F94FF293EFC7FEBF2D5A" box="[219,289,1004,1025]" name="India" pageId="1" pageNumber="104">INDIA</collectingCountry>
. Andaman Islands, South Andaman, Wandoor, Beach forest,
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,
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, alt.
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, on
<taxonomicName id="BCDD54CA8026F94FF767EFC7FEB72D7E" class="Magnoliopsida" family="Calophyllaceae" genus="Calophyllum" kingdom="Plantae" order="Malpighiales" pageId="1" pageNumber="104" phylum="Tracheophyta" rank="species" species="inophyllum">
<emphasis id="49A9F35B8026F94FF767EFC7FEB72D7E" italics="true" pageId="1" pageNumber="104">Calophyllum inophyllum</emphasis>
</taxonomicName>
,
<date id="0F6309898026F94FF37DE83BFE152D7E" box="[309,395,1040,1061]" pageId="1" pageNumber="104" value="2011-07-08">8.7.2011</date>
,
<emphasis id="49A9F35B8026F94FF3DCE83BFD022D7E" box="[404,668,1040,1061]" italics="true" pageId="1" pageNumber="104">
<collectionCode id="1DCCB78C8026F94FF3DCE83BFE3D2D7E" box="[404,419,1040,1061]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="104">T</collectionCode>
.
<collectionCode id="1DCCB78C8026F94FF3EFE83BFE292D7E" box="[423,439,1040,1061]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="1" pageNumber="104" type="Herbarium">A</collectionCode>
.
<collectionCode id="1DCCB78C8026F94FF3F2E83BFE4C2D7E" box="[442,466,1040,1061]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="1" pageNumber="104" type="Herbarium">M</collectionCode>
. Jagadeesh Ram 573
</emphasis>
(
<typeStatus id="A46691EB8026F94FF0E0E83BFC9D2D7E" box="[680,771,1040,1061]" pageId="1" pageNumber="104" type="holotype">Holotype</typeStatus>
<collectionCode id="1DCCB78C8026F94FF141E83BFCA52D7E" box="[777,827,1040,1061]" country="India" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12692" lsid="urn:lsid:biocol.org:col:12692" name="Botanical Survey of India" pageId="1" pageNumber="104" type="Herbarium">CAL</collectionCode>
,
<typeStatus id="A46691EB8026F94FF10DE83BFC122D7E" box="[837,908,1040,1061]" pageId="1" pageNumber="104" type="isotype">Isotype</typeStatus>
<collectionCode id="1DCCB78C8026F94FF1DAE83BFC5C2D7E" box="[914,962,1040,1061]" country="India" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12690" lsid="urn:lsid:biocol.org:col:12690" name="Botanical Survey of India, Andaman and Nicobar Circle" pageId="1" pageNumber="104" type="Herbarium">PBL</collectionCode>
).
</paragraph>
<paragraph id="7B622F498026F94FF2C0E87DFB6F2C8C" blockId="1.[136,1452,1110,2035]" pageId="1" pageNumber="104">
Thallus crustose, corticolous, epiperidermal, firmly attached, irregular, up to
<quantity id="BC2582AC8026F94FF1A4E87DFBD92D34" box="[1004,1095,1110,1135]" metricMagnitude="-1" metricUnit="m" metricValue="1.05" pageId="1" pageNumber="104" unit="cm" value="10.5">10.5 cm</quantity>
across, whitish to whitish grey, smooth, ± uneven, minutely felty, lacking isidia and soredia, 90–300 μm thick, heteromerous, ecorticate, with numerous 3–18 μm wide calcium oxalate crystals; prothallus ± distinct, byssoid, thin, comprising mainly of interwoven and radiating hyphae, whitish, up to
<quantity id="BC2582AC8026F94FF3BCE8E9FDAB2D80" box="[500,565,1218,1243]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="104" unit="mm" value="2.0">2 mm</quantity>
wide, forming a dark brown line while bordering different species; photobiont layer 36–65 μm thick, photobiont trentepohlioid, cells rounded to oblong, 8–12 × 5–8 μm; medulla distinct, white, 55–235 μm. Ascomata (ascigerous areas) delimited, whitish, paler than the thallus, not raised, evenly scattered, ± rounded to linear, irregular,
<quantity id="BC2582AC8026F94FF3F7E905FDAF2C1C" box="[447,561,1326,1351]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="1" pageNumber="104" unit="mm" value="0.75" valueMax="1.0" valueMin="0.5">0.5–1 mm</quantity>
diam., thinly pruinose, partly covered by photobiont layer. Paraphysoids tightly enclosing the asci,
<collectionCode id="1DCCB78C8026F94FF313E979FEFA2C30" box="[347,356,1362,1387]" country="Romania" lsid="urn:lsid:biocol.org:col:14415" name="&amp;quot;Alexandru Ioan Cuza&amp;quot; University" pageId="1" pageNumber="104" type="Herbarium">I</collectionCode>
+ pale blue, KI+ blue. Asci frequent, aggregated in whitish ascigerous areas, externally not visible, globose to subglobose, 8-spored,
<emphasis id="49A9F35B8026F94FF3B5E95DFD8E2CD4" box="[509,528,1398,1423]" italics="true" pageId="1" pageNumber="104">c.</emphasis>
85–100 × 80–90 μm. Ascospores colourless, broadly ellipsoid, straight, muriform, 9–11 × 3–5 septate, with ± equal cells, (44–)47–57 × 24–30 μm (n=26). Chemistry: Thallus and ascigerous areas
<collectionCode id="1DCCB78C8026F94FF7CAE9B1FA092CE8" box="[1410,1431,1434,1459]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="1" pageNumber="104" type="Herbarium">K</collectionCode>
–,
<collectionCode id="1DCCB78C8026F94FF2C0E995FF052C8C" box="[136,155,1470,1495]" country="Denmark" name="University of Copenhagen" pageId="1" pageNumber="104" type="Herbarium">C</collectionCode>
–, KC–,
<collectionCode id="1DCCB78C8026F94FF2B2E995FE952C8C" box="[250,267,1470,1495]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="1" pageNumber="104" type="Herbarium">P</collectionCode>
–;
<collectionCode id="1DCCB78C8026F94FF36EE995FECF2C8C" box="[294,337,1470,1495]" country="Colombia" name="Departamento de Biologia de la Universidad del Valle" pageId="1" pageNumber="104">UV</collectionCode>
–;
<collectionCode id="1DCCB78C8026F94FF323E995FEEA2C8C" box="[363,372,1470,1495]" country="Romania" lsid="urn:lsid:biocol.org:col:14415" name="&amp;quot;Alexandru Ioan Cuza&amp;quot; University" pageId="1" pageNumber="104" type="Herbarium">I</collectionCode>
+ blue and KI+ blue in patches (in section); 2’-
<emphasis id="49A9F35B8026F94FF1CBE995FC092C8C" box="[899,919,1470,1495]" italics="true" pageId="1" pageNumber="104">
<collectionCode id="1DCCB78C8026F94FF1CBE995FC092C8C" box="[899,919,1470,1495]" country="Norway" lsid="urn:lsid:biocol.org:col:13093" name="Botanical Museum - University of Oslo" pageId="1" pageNumber="104" type="Herbarium">O</collectionCode>
</emphasis>
-methylperlatolic acid (major).
</paragraph>
<paragraph id="7B622F498026F94FF2F5E9C9FB562EDC" blockId="1.[136,1452,1110,2035]" pageId="1" pageNumber="104">
<emphasis id="49A9F35B8026F94FF2F5E9C9FE972CA0" bold="true" box="[189,265,1506,1531]" pageId="1" pageNumber="104">Notes:</emphasis>
—
<taxonomicName id="BCDD54CA8026F94FF36EE9C9FDC02CA0" authorityName="Jagadeesh &amp; G. P. Sinha" authorityYear="2016" box="[294,606,1506,1531]" class="Arthoniomycetes" family="Arthoniaceae" genus="Cryptothecia" kingdom="Fungi" order="Arthoniales" pageId="1" pageNumber="104" phylum="Ascomycota" rank="species" species="albomaculans">
<emphasis id="49A9F35B8026F94FF36EE9C9FDC02CA0" box="[294,606,1506,1531]" italics="true" pageId="1" pageNumber="104">Cryptothecia albomaculans</emphasis>
</taxonomicName>
is characterized by the heteromerous thallus, whitish ascigerous areas not raised above the thallus, ellipsoid, 9–11 × 3–5 septate, 47–57 × 24–30 μm ascospores and 2’-
<emphasis id="49A9F35B8026F94FF6E8EA2DFB2A2F44" box="[1184,1204,1542,1567]" italics="true" pageId="1" pageNumber="104">
<collectionCode id="1DCCB78C8026F94FF6E8EA2DFB2A2F44" box="[1184,1204,1542,1567]" country="Norway" lsid="urn:lsid:biocol.org:col:13093" name="Botanical Museum - University of Oslo" pageId="1" pageNumber="104" type="Herbarium">O</collectionCode>
</emphasis>
-methylperlatolic acid as lichen substance. Morphologically it somewhat resembles
<taxonomicName id="BCDD54CA8026F94FF172EA01FECB2F3C" authority="Jagadeesh, G. P. Sinha &amp; Kr. P. Singh (2009: 605)" authorityName="Jagadeesh, G. P. Sinha &amp; Kr. P. Singh" authorityPageNumber="605" authorityYear="2009" class="Arthoniomycetes" family="Arthoniaceae" genus="Cryptothecia" kingdom="Fungi" order="Arthoniales" pageId="1" pageNumber="104" phylum="Ascomycota" rank="species" species="alboglauca">
<emphasis id="49A9F35B8026F94FF172EA01FBCC2F18" box="[826,1106,1578,1603]" italics="true" pageId="1" pageNumber="104">Cryptothecia alboglauca</emphasis>
Jagadeesh, G.P. Sinha &amp; Kr.P. Singh (2009: 605)
</taxonomicName>
and
<taxonomicName id="BCDD54CA8026F94FF3C5EA65FCD02F3C" authority="Aptroot &amp; Wolseley" authorityName="Aptroot &amp; Wolseley" box="[397,846,1614,1639]" class="Arthoniomycetes" family="Arthoniaceae" genus="Cryptothecia" kingdom="Fungi" order="Arthoniales" pageId="1" pageNumber="104" phylum="Ascomycota" rank="species" species="albomaculatella">
<emphasis id="49A9F35B8026F94FF3C5EA65FDFA2F3C" box="[397,612,1614,1639]" italics="true" pageId="1" pageNumber="104">C. albomaculatella</emphasis>
Aptroot &amp; Wolseley
</taxonomicName>
(in
<bibRefCitation id="1F4C52B88026F94FF133EA65FB472F3C" author="Wolseley, P. A. &amp; Aptroot, A." box="[891,1241,1614,1639]" pageId="1" pageNumber="104" pagination="411 - 422" refId="ref20140" refString="Wolseley, P. A. &amp; Aptroot, A. (2009) The lichen genera Cryptothecia and Stirtonia in northern Thailand. Bibliotheca Lichenologica 99: 411 - 422." type="journal article" year="2009">Wolseley &amp; Aptroot 2009: 413</bibRefCitation>
) in having whitish ascigerous areas usually not raised above the thallus. All of these differ in their ascospores morphology in having large ascospores and different chemistry. Chemically it shows resemblence with
<taxonomicName id="BCDD54CA8026F94FF646EABDFA1D2FF4" authority="Patw. &amp; Makhija" authorityName="Patw. &amp; Makhija" box="[1038,1411,1686,1711]" class="Arthoniomycetes" family="Arthoniaceae" genus="Cryptothecia" kingdom="Fungi" order="Arthoniales" pageId="1" pageNumber="104" phylum="Ascomycota" rank="species" species="culbersoniae">
<emphasis id="49A9F35B8026F94FF646EABDFB212FF4" box="[1038,1215,1686,1711]" italics="true" pageId="1" pageNumber="104">C. culbersoniae</emphasis>
Patw. &amp; Makhija
</taxonomicName>
[as ‘
<emphasis id="49A9F35B8026F94FF2D9EA91FE842F88" box="[145,282,1722,1747]" italics="true" pageId="1" pageNumber="104">culbersonae</emphasis>
’] (1981: 125),
<taxonomicName id="BCDD54CA8026F94FF388EA91FCFF2F88" authority="F. Seavey (2009: 52)" authorityName="F. Seavey" authorityPageNumber="52" authorityYear="2009" box="[448,865,1722,1747]" class="Arthoniomycetes" family="Arthoniaceae" genus="Cryptothecia" kingdom="Fungi" order="Arthoniales" pageId="1" pageNumber="104" phylum="Ascomycota" rank="species" species="evergladensis">
<emphasis id="49A9F35B8026F94FF388EA91FDE62F88" box="[448,632,1722,1747]" italics="true" pageId="1" pageNumber="104">C. evergladensis</emphasis>
F.
<bibRefCitation id="1F4C52B88026F94FF0DFEA91FCFF2F88" author="Seavey, F." box="[663,865,1722,1747]" pageId="1" pageNumber="104" pagination="49 - 54" refId="ref19810" refString="Seavey, F. (2009) Cryptothecia evergladensis sp. nov. (Arthoniaceae), a new lichen species from Everglades National Park, Florida. Opuscula Philolichenum 7: 49 - 54." type="journal article" year="2009">Seavey (2009: 52)</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="BCDD54CA8026F94FF1DCEA91FA9C2F88" authority="Bungartz &amp; Elix" authorityName="Bungartz &amp; Elix" box="[916,1282,1722,1747]" class="Arthoniomycetes" family="Arthoniaceae" genus="Cryptothecia" kingdom="Fungi" order="Arthoniales" pageId="1" pageNumber="104" phylum="Ascomycota" rank="species" species="galapagoana">
<emphasis id="49A9F35B8026F94FF1DCEA91FBD92F88" box="[916,1095,1722,1747]" italics="true" pageId="1" pageNumber="104">C. galapagoana</emphasis>
Bungartz &amp; Elix
</taxonomicName>
(in
<bibRefCitation id="1F4C52B88026F94FF763EA91FE832FAC" author="Bungartz, F. &amp; Dutan-Patin, V. L. &amp; Elix, J. A." pageId="1" pageNumber="104" pagination="739 - 762" refId="ref18694" refString="Bungartz, F., Dutan-Patin, V. L. &amp; Elix, J. A. (2013) The lichen genera Cryptothecia, Herpothallon and Helminthocarpon (Arthoniales) in the Galapagos Islands, Ecuador. Lichenologist 45: 739 - 762. http: // dx. doi. org / 10.1017 / S 0024282913000522" type="journal article" year="2013">
Bungartz
<emphasis id="49A9F35B8026F94FF7DFEA91FF002FAC" italics="true" pageId="1" pageNumber="104">et al</emphasis>
. 2013: 745
</bibRefCitation>
) in having 2’-
<emphasis id="49A9F35B8026F94FF3F5EAF5FE4F2FAC" box="[445,465,1758,1783]" italics="true" pageId="1" pageNumber="104">
<collectionCode id="1DCCB78C8026F94FF3F5EAF5FE4F2FAC" box="[445,465,1758,1783]" country="Norway" lsid="urn:lsid:biocol.org:col:13093" name="Botanical Museum - University of Oslo" pageId="1" pageNumber="104" type="Herbarium">O</collectionCode>
</emphasis>
-methylperlatolic acid as a major lichen substance. However,
<taxonomicName id="BCDD54CA8026F94FF6CAEAF5FA322FAC" box="[1154,1452,1758,1783]" class="Arthoniomycetes" family="Arthoniaceae" genus="Cryptothecia" kingdom="Fungi" order="Arthoniales" pageId="1" pageNumber="104" phylum="Ascomycota" rank="species" species="culbersoniae">
<emphasis id="49A9F35B8026F94FF6CAEAF5FA322FAC" box="[1154,1452,1758,1783]" italics="true" pageId="1" pageNumber="104">Cryptothecia culbersoniae</emphasis>
</taxonomicName>
additionally has gyrophoric acid and large distinctly raised ascigerous areas.
<taxonomicName id="BCDD54CA8026F94FF64CEB29FAA72E40" authorityName="F. Seavey" authorityYear="2009" box="[1028,1337,1794,1819]" class="Arthoniomycetes" family="Arthoniaceae" genus="Cryptothecia" kingdom="Fungi" order="Arthoniales" pageId="1" pageNumber="104" phylum="Ascomycota" rank="species" species="evergladensis">
<emphasis id="49A9F35B8026F94FF64CEB29FAA72E40" box="[1028,1337,1794,1819]" italics="true" pageId="1" pageNumber="104">Cryptothecia evergladensis</emphasis>
</taxonomicName>
differs in small ascospores and additionally has psoromic acid and lichexanthone.
<taxonomicName id="BCDD54CA8026F94FF199EB0DFA9F2E64" box="[977,1281,1830,1855]" class="Arthoniomycetes" family="Arthoniaceae" genus="Cryptothecia" kingdom="Fungi" order="Arthoniales" pageId="1" pageNumber="104" phylum="Ascomycota" rank="species" species="galapagoana">
<emphasis id="49A9F35B8026F94FF199EB0DFA9F2E64" box="[977,1281,1830,1855]" italics="true" pageId="1" pageNumber="104">Cryptothecia galapagoana</emphasis>
</taxonomicName>
also differs in small ascospores and additionally with confluentic and 2’-
<emphasis id="49A9F35B8026F94FF17CEB61FCD62E38" box="[820,840,1866,1891]" italics="true" pageId="1" pageNumber="104">
<collectionCode id="1DCCB78C8026F94FF17CEB61FCD62E38" box="[820,840,1866,1891]" country="Norway" lsid="urn:lsid:biocol.org:col:13093" name="Botanical Museum - University of Oslo" pageId="1" pageNumber="104" type="Herbarium">O</collectionCode>
</emphasis>
-methylmicrophyllinic acids (
<bibRefCitation id="1F4C52B88026F94FF6D4EB61FF5D2EDC" author="Makhija, U. &amp; Patwardhan, P. G." pageId="1" pageNumber="104" pagination="57 - 72" refId="ref19472" refString="Makhija, U. &amp; Patwardhan, P. G. (1994) A contribution towards a Monograph of the lichen genus Cryptothecia (Family Arthoniaceae). In: Sarma, T. A., Saini, S. S., Trivedi, M. L. &amp; Sharma, M. (Eds.) Current Researches in Plant Sciences. Bishen Singh Mahendra Pal Singh, Dehra Dun, pp. 57 - 72." type="book chapter" year="1994">Makhija &amp; Patwardhan 1994</bibRefCitation>
,
<bibRefCitation id="1F4C52B88026F94FF286EB45FE6E2EDC" author="Wolseley, P. A. &amp; Aptroot, A." box="[206,496,1902,1927]" pageId="1" pageNumber="104" pagination="411 - 422" refId="ref20140" refString="Wolseley, P. A. &amp; Aptroot, A. (2009) The lichen genera Cryptothecia and Stirtonia in northern Thailand. Bibliotheca Lichenologica 99: 411 - 422." type="journal article" year="2009">Wolseley &amp; Aptroot 2009</bibRefCitation>
,
<bibRefCitation id="1F4C52B88026F94FF3B3EB45FCB42EDC" author="Jagadeesh Ram, T. A. M. &amp; Sinha, G. P. &amp; Singh, K. P." box="[507,810,1902,1927]" pageId="1" pageNumber="104" pagination="605 - 613" refId="ref19140" refString="Jagadeesh Ram, T. A. M., Sinha, G. P. &amp; Singh, K. P. (2009) New species and new records of Cryptothecia and Herpothallon (Arthoniales) from India. Lichenologist 41: 605 - 613. http: // dx. doi. org / 10.1017 / S 0024282909008123" type="journal article" year="2009">
Jagadeesh Ram
<emphasis id="49A9F35B8026F94FF0F8EB45FD762EDC" box="[688,744,1902,1927]" italics="true" pageId="1" pageNumber="104">et al.</emphasis>
2009
</bibRefCitation>
,
<bibRefCitation id="1F4C52B88026F94FF17DEB45FC562EDC" author="Seavey, F." box="[821,968,1902,1927]" pageId="1" pageNumber="104" pagination="49 - 54" refId="ref19810" refString="Seavey, F. (2009) Cryptothecia evergladensis sp. nov. (Arthoniaceae), a new lichen species from Everglades National Park, Florida. Opuscula Philolichenum 7: 49 - 54." type="journal article" year="2009">Seavey 2009</bibRefCitation>
,
<bibRefCitation id="1F4C52B88026F94FF19BEB45FB222EDC" author="Bungartz, F. &amp; Dutan-Patin, V. L. &amp; Elix, J. A." box="[979,1212,1902,1927]" pageId="1" pageNumber="104" pagination="739 - 762" refId="ref18694" refString="Bungartz, F., Dutan-Patin, V. L. &amp; Elix, J. A. (2013) The lichen genera Cryptothecia, Herpothallon and Helminthocarpon (Arthoniales) in the Galapagos Islands, Ecuador. Lichenologist 45: 739 - 762. http: // dx. doi. org / 10.1017 / S 0024282913000522" type="journal article" year="2013">
Bungartz
<emphasis id="49A9F35B8026F94FF608EB45FBE72EDC" box="[1088,1145,1902,1927]" italics="true" pageId="1" pageNumber="104">et al.</emphasis>
2013
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="33C77CC28026F94FF2F5EBB9FBD22E94" pageId="1" pageNumber="104" type="distribution">
<paragraph id="7B622F498026F94FF2F5EBB9FBD22E94" blockId="1.[136,1452,1110,2035]" pageId="1" pageNumber="104">
<emphasis id="49A9F35B8026F94FF2F5EBB9FE742EF0" bold="true" box="[189,490,1938,1963]" pageId="1" pageNumber="104">Distribution and habitat:</emphasis>
—
<taxonomicName id="BCDD54CA8026F94FF04EEBB9FCA22EF0" authorityName="Jagadeesh &amp; G. P. Sinha" authorityYear="2016" box="[518,828,1938,1963]" class="Arthoniomycetes" family="Arthoniaceae" genus="Cryptothecia" kingdom="Fungi" order="Arthoniales" pageId="1" pageNumber="104" phylum="Ascomycota" rank="species" species="albomaculans">
<emphasis id="49A9F35B8026F94FF04EEBB9FCA22EF0" box="[518,828,1938,1963]" italics="true" pageId="1" pageNumber="104">Cryptothecia albomaculans</emphasis>
</taxonomicName>
is known only from the
<typeStatus id="A46691EB8026F94FF618EBB9FB1E2EF0" box="[1104,1152,1938,1963]" pageId="1" pageNumber="104">type</typeStatus>
locality where it grows on
<taxonomicName id="BCDD54CA8026F94FF2C0EB9DFE002E94" box="[136,414,1974,1999]" class="Magnoliopsida" family="Calophyllaceae" genus="Calophyllum" kingdom="Plantae" order="Malpighiales" pageId="1" pageNumber="104" phylum="Tracheophyta" rank="species" species="inophyllum">
<emphasis id="49A9F35B8026F94FF2C0EB9DFE002E94" box="[136,414,1974,1999]" italics="true" pageId="1" pageNumber="104">Calophyllum inophyllum</emphasis>
</taxonomicName>
in a beach forest of South Andaman in the Andaman Islands.
</paragraph>
</subSubSection>
<subSubSection id="33C77CC28026F94CF2F5EBF1FA362E06" lastPageId="2" lastPageNumber="105" pageId="1" pageNumber="104" type="etymology">
<paragraph id="7B622F498026F94FF2F5EBF1FBE42EA8" blockId="1.[136,1452,1110,2035]" box="[189,1146,2010,2035]" pageId="1" pageNumber="104">
<emphasis id="49A9F35B8026F94FF2F5EBF1FED62EA8" bold="true" box="[189,328,2010,2035]" pageId="1" pageNumber="104">Etymology:</emphasis>
—The specific epithet refers to its maculae like whitish ascigerous areas.
</paragraph>
<caption id="2FA27FC18025F94CF2C0EA5DFC0E2FAC" ID-DOI="http://doi.org/10.5281/zenodo.13659738" ID-Zenodo-Dep="13659738" httpUri="https://zenodo.org/record/13659738/files/figure.png" pageId="2" pageNumber="105" startId="2.[136,229,1654,1675]" targetBox="[158,1430,190,1629]" targetPageId="2" targetType="figure">
<paragraph id="7B622F498025F94CF2C0EA5DFC0E2FAC" blockId="2.[136,1452,1654,1783]" pageId="2" pageNumber="105">
<emphasis id="49A9F35B8025F94CF2C0EA5DFF612FD0" bold="true" box="[136,255,1654,1675]" pageId="2" pageNumber="105">FIGURE 1.</emphasis>
A.
<taxonomicName id="BCDD54CA8025F94CF36BEA5DFDB12FD0" authorityName="Jagadeesh &amp; G. P. Sinha" authorityYear="2016" box="[291,559,1654,1675]" class="Arthoniomycetes" family="Arthoniaceae" genus="Cryptothecia" kingdom="Fungi" order="Arthoniales" pageId="2" pageNumber="105" phylum="Ascomycota" rank="species" species="albomaculans">
<emphasis id="49A9F35B8025F94CF36BEA5DFDB12FD0" box="[291,559,1654,1675]" italics="true" pageId="2" pageNumber="105">Cryptothecia albomaculans</emphasis>
</taxonomicName>
(holotype), part of thallus with white maculiform ascigerous areas. B.
<taxonomicName id="BCDD54CA8025F94CF6A5EA5DFAA42FD0" authorityName="Jagadeesh &amp; G. P. Sinha" authorityYear="2016" box="[1261,1338,1654,1675]" class="Arthoniomycetes" family="Arthoniaceae" genus="Cryptothecia" kingdom="Fungi" order="Arthoniales" pageId="2" pageNumber="105" phylum="Ascomycota" rank="species" species="elata">
<emphasis id="49A9F35B8025F94CF6A5EA5DFAA42FD0" box="[1261,1338,1654,1675]" italics="true" pageId="2" pageNumber="105">C. elata</emphasis>
</taxonomicName>
(holotype), part of thallus with white raised ascigerous areas. C.
<taxonomicName id="BCDD54CA8025F94CF0D8EAB1FC9F2FF4" authorityName="Jagadeesh &amp; G. P. Sinha" authorityYear="2016" box="[656,769,1690,1711]" class="Arthoniomycetes" family="Arthoniaceae" genus="Cryptothecia" kingdom="Fungi" order="Arthoniales" pageId="2" pageNumber="105" phylum="Ascomycota" rank="species" species="elongata">
<emphasis id="49A9F35B8025F94CF0D8EAB1FC9F2FF4" box="[656,769,1690,1711]" italics="true" pageId="2" pageNumber="105">C. elongata</emphasis>
</taxonomicName>
(holotype), part of thallus with slightly raised ascigerous areas. D.
<taxonomicName id="BCDD54CA8025F94CF7DDEAB1FE832F88" authorityName="Jagadeesh &amp; G. P. Sinha" authorityYear="2016" class="Arthoniomycetes" family="Arthoniaceae" genus="Cryptothecia" kingdom="Fungi" order="Arthoniales" pageId="2" pageNumber="105" phylum="Ascomycota" rank="species" species="superphyllinica">
<emphasis id="49A9F35B8025F94CF7DDEAB1FE832F88" italics="true" pageId="2" pageNumber="105">C. superphyllinica</emphasis>
</taxonomicName>
(holotype), part of thallus with white raised ascigerous areas. E.
<taxonomicName id="BCDD54CA8025F94CF1C0EA95FB1F2F88" authorityName="Jagadeesh &amp; G. P. Sinha" authorityYear="2016" box="[904,1153,1726,1747]" class="Arthoniomycetes" family="Arthoniaceae" genus="Myriostigma" kingdom="Fungi" order="Arthoniales" pageId="2" pageNumber="105" phylum="Ascomycota" rank="species" species="nicobaricum">
<emphasis id="49A9F35B8025F94CF1C0EA95FB1F2F88" box="[904,1153,1726,1747]" italics="true" pageId="2" pageNumber="105">Myriostigma nicobaricum</emphasis>
</taxonomicName>
(holotype), part of thallus with white raised ascigerous areas. F.
<taxonomicName id="BCDD54CA8025F94CF38BEAC9FDC22FAC" authorityName="Jagadeesh &amp; G. P. Sinha" authorityYear="2016" box="[451,604,1762,1783]" class="Arthoniomycetes" family="Arthoniaceae" genus="Myriostigma" kingdom="Fungi" order="Arthoniales" pageId="2" pageNumber="105" phylum="Ascomycota" rank="species" species="nicobaricum">
<emphasis id="49A9F35B8025F94CF38BEAC9FDC22FAC" box="[451,604,1762,1783]" italics="true" pageId="2" pageNumber="105">M. nicobaricum</emphasis>
</taxonomicName>
(paratype). Scales: A–F=1 mm.
</paragraph>
</caption>
<paragraph id="7B622F498025F94CF2F5EB0BFA352E62" blockId="2.[136,1451,1824,1885]" box="[189,1451,1824,1849]" pageId="2" pageNumber="105">
<emphasis id="49A9F35B8025F94CF2F5EB0BFDD22E62" bold="true" box="[189,588,1824,1849]" pageId="2" pageNumber="105">Additional specimen examined:</emphasis>
—
<collectingCountry id="03CA6FD98025F94CF022EB0BFD222E62" box="[618,700,1824,1849]" name="India" pageId="2" pageNumber="105">INDIA</collectingCountry>
. Andaman Islands, South Andaman, Wandoor, Beach forest,
</paragraph>
<paragraph id="7B622F498025F94CF2C0EB6FFA362E06" blockId="2.[136,1451,1824,1885]" box="[136,1448,1860,1885]" pageId="2" pageNumber="105">
<geoCoordinate id="1EE9498E8025F94CF2C0EB6FFEB12E06" box="[136,303,1860,1885]" degrees="11" direction="north" minutes="37" orientation="latitude" pageId="2" pageNumber="105" precision="1" seconds="19.0" value="11.621944">11°37’19.0” N</geoCoordinate>
,
<geoCoordinate id="1EE9498E8025F94CF372EB6FFE402E06" box="[314,478,1860,1885]" degrees="92" direction="east" minutes="37" orientation="longitude" pageId="2" pageNumber="105" precision="1" seconds="14.5" value="92.6207">92°37’14.5” E</geoCoordinate>
, alt.
<quantity id="BC2582AC8025F94CF05CEB6FFDDF2E06" box="[532,577,1860,1885]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="105" unit="m" value="8.0">8 m</quantity>
, on
<taxonomicName id="BCDD54CA8025F94CF038EB6FFC182E06" box="[624,902,1860,1885]" class="Magnoliopsida" family="Calophyllaceae" genus="Calophyllum" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="105" phylum="Tracheophyta" rank="species" species="inophyllum">
<emphasis id="49A9F35B8025F94CF038EB6FFC182E06" box="[624,902,1860,1885]" italics="true" pageId="2" pageNumber="105">Calophyllum inophyllum</emphasis>
</taxonomicName>
,
<date id="0F6309898025F94CF1DCEB6FFB982E06" box="[916,1030,1860,1885]" pageId="2" pageNumber="105" value="2015-03-10">3.10.2015</date>
,
<emphasis id="49A9F35B8025F94CF65AEB6FFACD2E06" box="[1042,1363,1860,1885]" italics="true" pageId="2" pageNumber="105">T.A.M. Jagadeesh Ram 4645</emphasis>
(PBL).
</paragraph>
</subSubSection>
</treatment>
</document>
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