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<document id="B09ED20D8D63FF62B91A6B170670ADD2" ID-DOI="10.11646/zootaxa.5023.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5225372" ID-ZooBank="E759FD85-1904-45A7-9A4D-56815FB9649B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629405269492" checkinUser="plazi" docAuthor="Yamana, Yusuke, Nakaguchi, Kazumitsu, Yamaguchi, Shuhei, Katoh, Mikio, Ogawa, Akito &amp; Ohtsuka, Susumu" docDate="2021" docId="AD40CD45413DFFDCFF02FA30FBEFFDF2" docLanguage="en" docName="zootaxa.5023.1.1.pdf" docOrigin="Zootaxa 5023 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Thyone liaoi Yamana, Ogawa &amp; Ohtsuka 2021, sp. nov." docType="treatment" docUuid="312032A1-D924-4510-90DD-81749B7B1DE6" docUuidSource="ZooBank" docVersion="3" lastPageNumber="15" masterDocId="5179B53D4137FFD2FF95FFFAFFC2FFE5" masterDocTitle="Four new dendrochirotid holothurians collected from the Seto Inland Sea and the western part of the Sea of Japan, western Japan" masterLastPageNumber="43" masterPageNumber="1" pageNumber="11" updateTime="1699072022120" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="ECF0F303618B53D0777F3D5707AB3712">Four new dendrochirotid holothurians collected from the Seto Inland Sea and the western part of the Sea of Japan, western Japan</mods:title>
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Yamana, Ogawa &amp; Ohtsuka
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[Japanese name: Kehada-namako (
<bibRefCitation id="417801A2413DFFD8FD89FA15FD7EF9EC" author="Yamana, Y." box="[540,700,1518,1545]" pageId="10" pageNumber="11" pagination="6 - 8" refId="ref28119" refString="Yamana, Y. (2016) Kii-suido shuhen no namako bunnrui no genjou [The present situation of the studies on Holothurioidea in the Kii Strait], The Umiushi Tsushin (ISSN 1341 - 5972), published by the Research Institute of Marine Invertebrates, Tokyo, 92, 6 - 8. [only in Japanese]" type="journal article" year="2016">Yamana 2016</bibRefCitation>
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(
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,
<figureCitation id="BDD260D6413DFFD8FE93F9E9FEEBF9C8" box="[262,297,1555,1581]" captionStart="FIGURE 3" captionStartId="7.[151,250,1500,1524]" captionTargetBox="[152,1426,183,1471]" captionTargetId="figure-18@7.[151,1436,181,1477]" captionTargetPageId="7" captionText="FIGURE 3. Elements of calcareous rings, viewed from the body cavity, anterior upper.A, Massinium toyoshiomaruae sp. nov., (holotype, WMNH-INV-2014-186); B, Thyone liaoi sp. nov., (holotype, WMNH-INV-2014-182, and a paratype, WMNH-INV- 2014-184); C, T. kyushuensis sp. nov., (holotype, WMNH-INV-2015-2437); D, T. toyoshiomaruae sp. nov. (holotype, WMNH- INV-2015-2483). Abbreviations: md, mid-dorsal part; mv, mid-ventral part; -ir, interradial element; -r radial element." figureDoi="http://doi.org/10.5281/zenodo.5225378" httpUri="https://zenodo.org/record/5225378/files/figure.png" pageId="10" pageNumber="11">3B</figureCitation>
,
<figureCitation id="BDD260D6413DFFD8FEA0F9E8FEB9F9C8" box="[309,379,1554,1581]" captionStart="FIGURE 5" captionStartId="11.[151,250,1888,1912]" captionTargetBox="[182,1404,182,1863]" captionTargetId="figure-18@11.[181,1406,181,1864]" captionTargetPageId="11" captionText="FIGURE 5. Scanning electron microscope images of ossicles of Thyone liaoi sp. nov., (holotype, AH, WMNH-INV-2014- 182). A, ossicles of the ventrolateral tentacle; B, the ventrolateral skin of peri-oral; C, the ventrolateral skin of pharynx; D, the ventrolateral introvert skin; E, the pedicel of the abdominal side; F and G, the middle part of the bodywall on the abdominal side, and on the dorsal side, respectively; H, the ventrolateral anal papilla." figureDoi="http://doi.org/10.5281/zenodo.5225382" httpUri="https://zenodo.org/record/5225382/files/figure.png" pageId="10" pageNumber="11">5AH</figureCitation>
)
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<taxonomicName id="E2E907D0413DFFD8FF02F9ADFDA1F98A" ID-CoL="56RN8" authority=": Liao &amp; Clark 1995: 505 - 506" authorityName="Liao &amp; Clark" authorityPageNumber="505 - 506" authorityYear="1995" box="[151,611,1623,1647]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="species" species="pedata">
<emphasis id="179DA041413DFFD8FF02F9ADFEEFF98A" box="[151,301,1623,1647]" italics="true" pageId="10" pageNumber="11">Thyone pedata</emphasis>
:
<bibRefCitation id="417801A2413DFFD8FEAFF9ADFDA1F98A" author="Liao, Y. &amp; Clark, A. M." box="[314,611,1623,1647]" pageId="10" pageNumber="11" refId="ref26030" refString="Liao, Y. &amp; Clark, A. M. (1995) The Echinoderms of Southern China. Science Press Beijing, 614 pp." type="book" year="1995">Liao &amp; Clark 1995: 505506</bibRefCitation>
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, Fig. 307 a, b;
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, Fig. a.
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<emphasis id="179DA041413DFFD8FF02F98DFF7FF96B" box="[151,189,1655,1678]" italics="true" pageId="10" pageNumber="11">non</emphasis>
:
<taxonomicName id="E2E907D0413DFFD8FF5EF98CFE34F96B" ID-CoL="56RN8" authority="Semper, 1867" authorityName="Semper" authorityYear="1867" box="[203,502,1654,1678]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="species" species="pedata">
<emphasis id="179DA041413DFFD8FF5EF98CFEA2F96B" box="[203,352,1654,1678]" italics="true" pageId="10" pageNumber="11">Thyone pedata</emphasis>
<bibRefCitation id="417801A2413DFFD8FEF2F98CFE34F96B" author="Semper, C." box="[359,502,1654,1678]" pageId="10" pageNumber="11" refId="ref27545" refString="Semper, C. (1867 - 1868) Holothurien. Reisen im Archipel der Philippinen. Teil 2. Wissenschaftliche Resultate. C. W. Kreidel, Wiesbaden, 288 pp. https: // doi. org / 10.5962 / bhl. title. 11687" type="book" year="1867">Semper, 1867</bibRefCitation>
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8: p 67. (see Remarks)
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<collectionCode id="43F8E496413DFFD8FEE8F91AFE7CF91F" box="[381,446,1760,1786]" pageId="10" pageNumber="11">INSD</collectionCode>
accession number
<accessionNumber id="3ABAE1B0413DFFD8FD0FF91AFCD7F91E" box="[666,789,1760,1787]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/LC547978" pageId="10" pageNumber="11">LC547978</accessionNumber>
, 622 bps,
<figureCitation id="BDD260D6413DFFD8FC19F91AFC0DF91F" box="[908,975,1760,1786]" captionStart="FIGURE 2" captionStartId="6.[151,250,1766,1790]" captionTargetBox="[155,1435,419,1743]" captionTargetId="figure-141@6.[151,1436,417,1743]" captionTargetPageId="6" captionText="FIGURE 2. Lateral views of the specimens, dorsal upper, anterior left. A, Massinium toyoshiomaruae sp. nov., (a) holotype, WMNH-INV-2014-186, (b) paratype, WMNH-INV-2015-2497, (c) paratype, WMNH-INV-2015-2503, (d) paratype, NSMT- E-12099-1; B, Thyone liaoi sp. nov., (e) holotype, WMNH-INV-2014-182, (f) paratype, WMNH-INV-2014-183, (g) paratype, WMNH-INV-2014-184, (h) paratype, WMNH-INV-2014-188; C, Thyone kyushuensis sp. nov., (i) holotype, WMNH-INV- 2015-2437, (j) paratype, NSMT-E-12103; D, T. toyoshiomaruae sp. nov., (k) paratype, WMNH-INV-2015-2438, (l) paratype, WMNH-INV-2015-2439, (m) holotype, WMNH-INV-2015-2483, (n) paratype, WMNH-INV-2015-2484." figureDoi="http://doi.org/10.5281/zenodo.5225376" httpUri="https://zenodo.org/record/5225376/files/figure.png" pageId="10" pageNumber="11">Fig. 2</figureCitation>
B-e)
</materialsCitation>
.
<emphasis id="179DA041413DFFD8FB84F91BFB41F91F" box="[1041,1155,1761,1786]" italics="true" pageId="10" pageNumber="11">
<typeStatus id="FA52C2F1413DFFD8FB84F91BFB41F91F" box="[1041,1155,1761,1786]" pageId="10" pageNumber="11" type="paratype">Paratypes</typeStatus>
</emphasis>
: WMNH-INV-2014-183 (ST-
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2014,
<date id="51575A93413DFFD8FEDFF8FEFDDCF8FB" box="[330,542,1796,1822]" pageId="10" pageNumber="11" value="2014-11-06">6 November 2014</date>
, length
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, width
<quantity id="E211D1B6413DFFD8FCBFF8FEFCB2F8FA" box="[810,880,1796,1823]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="10" pageNumber="11" unit="mm" value="9.0">9 mm</quantity>
, eviscerated,
<figureCitation id="BDD260D6413DFFD8FB9BF8FEFB91F8FA" box="[1038,1107,1796,1823]" captionStart="FIGURE 2" captionStartId="6.[151,250,1766,1790]" captionTargetBox="[155,1435,419,1743]" captionTargetId="figure-141@6.[151,1436,417,1743]" captionTargetPageId="6" captionText="FIGURE 2. Lateral views of the specimens, dorsal upper, anterior left. A, Massinium toyoshiomaruae sp. nov., (a) holotype, WMNH-INV-2014-186, (b) paratype, WMNH-INV-2015-2497, (c) paratype, WMNH-INV-2015-2503, (d) paratype, NSMT- E-12099-1; B, Thyone liaoi sp. nov., (e) holotype, WMNH-INV-2014-182, (f) paratype, WMNH-INV-2014-183, (g) paratype, WMNH-INV-2014-184, (h) paratype, WMNH-INV-2014-188; C, Thyone kyushuensis sp. nov., (i) holotype, WMNH-INV- 2015-2437, (j) paratype, NSMT-E-12103; D, T. toyoshiomaruae sp. nov., (k) paratype, WMNH-INV-2015-2438, (l) paratype, WMNH-INV-2015-2439, (m) holotype, WMNH-INV-2015-2483, (n) paratype, WMNH-INV-2015-2484." figureDoi="http://doi.org/10.5281/zenodo.5225376" httpUri="https://zenodo.org/record/5225376/files/figure.png" pageId="10" pageNumber="11">Fig. 2</figureCitation>
B-f); WMNH-INV-2014-184 (ST-
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2014,
<date id="51575A93413DFFD8FEAAF8D2FDCEF8A7" box="[319,524,1832,1858]" pageId="10" pageNumber="11" value="2014-11-06">6 November 2014</date>
, length
<quantity id="E211D1B6413DFFD8FDF6F8D2FD76F8A6" box="[611,692,1832,1859]" metricMagnitude="-2" metricUnit="m" metricValue="3.4" pageId="10" pageNumber="11" unit="mm" value="34.0">34 mm</quantity>
, width
<quantity id="E211D1B6413DFFD8FC93F8D2FC8BF8A6" box="[774,841,1832,1859]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="10" pageNumber="11" unit="mm" value="9.0">9 mm</quantity>
,
<figureCitation id="BDD260D6413DFFD8FCC1F8D2FC57F8A7" box="[852,917,1832,1858]" captionStart="FIGURE 2" captionStartId="6.[151,250,1766,1790]" captionTargetBox="[155,1435,419,1743]" captionTargetId="figure-141@6.[151,1436,417,1743]" captionTargetPageId="6" captionText="FIGURE 2. Lateral views of the specimens, dorsal upper, anterior left. A, Massinium toyoshiomaruae sp. nov., (a) holotype, WMNH-INV-2014-186, (b) paratype, WMNH-INV-2015-2497, (c) paratype, WMNH-INV-2015-2503, (d) paratype, NSMT- E-12099-1; B, Thyone liaoi sp. nov., (e) holotype, WMNH-INV-2014-182, (f) paratype, WMNH-INV-2014-183, (g) paratype, WMNH-INV-2014-184, (h) paratype, WMNH-INV-2014-188; C, Thyone kyushuensis sp. nov., (i) holotype, WMNH-INV- 2015-2437, (j) paratype, NSMT-E-12103; D, T. toyoshiomaruae sp. nov., (k) paratype, WMNH-INV-2015-2438, (l) paratype, WMNH-INV-2015-2439, (m) holotype, WMNH-INV-2015-2483, (n) paratype, WMNH-INV-2015-2484." figureDoi="http://doi.org/10.5281/zenodo.5225376" httpUri="https://zenodo.org/record/5225376/files/figure.png" pageId="10" pageNumber="11">Fig. 2</figureCitation>
B-g); WMNH-INV-2014-188 (ST-
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2014,
<date id="51575A93413DFFD8FA1BF8D2FE8DF883" pageId="10" pageNumber="11" value="2014-11-05">5 November 2014</date>
, anterior half only, length
<quantity id="E211D1B6413DFFD8FDE3F8B6FD05F882" box="[630,711,1868,1895]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="10" pageNumber="11" unit="mm" value="13.0">13 mm</quantity>
, width
<quantity id="E211D1B6413DFFD8FC8CF8B6FC9EF882" box="[793,860,1868,1895]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="11" unit="mm" value="5.0">5 mm</quantity>
,
<figureCitation id="BDD260D6413DFFD8FCFDF8B6FC6BF882" box="[872,937,1868,1895]" captionStart="FIGURE 2" captionStartId="6.[151,250,1766,1790]" captionTargetBox="[155,1435,419,1743]" captionTargetId="figure-141@6.[151,1436,417,1743]" captionTargetPageId="6" captionText="FIGURE 2. Lateral views of the specimens, dorsal upper, anterior left. A, Massinium toyoshiomaruae sp. nov., (a) holotype, WMNH-INV-2014-186, (b) paratype, WMNH-INV-2015-2497, (c) paratype, WMNH-INV-2015-2503, (d) paratype, NSMT- E-12099-1; B, Thyone liaoi sp. nov., (e) holotype, WMNH-INV-2014-182, (f) paratype, WMNH-INV-2014-183, (g) paratype, WMNH-INV-2014-184, (h) paratype, WMNH-INV-2014-188; C, Thyone kyushuensis sp. nov., (i) holotype, WMNH-INV- 2015-2437, (j) paratype, NSMT-E-12103; D, T. toyoshiomaruae sp. nov., (k) paratype, WMNH-INV-2015-2438, (l) paratype, WMNH-INV-2015-2439, (m) holotype, WMNH-INV-2015-2483, (n) paratype, WMNH-INV-2015-2484." figureDoi="http://doi.org/10.5281/zenodo.5225376" httpUri="https://zenodo.org/record/5225376/files/figure.png" pageId="10" pageNumber="11">Fig. 2</figureCitation>
B-h).
</paragraph>
</subSubSection>
<subSubSection id="6DF32FD8413DFFD8FF52F88AFC1FF812" pageId="10" pageNumber="11" type="diagnosis">
<paragraph id="25567C53413DFFD8FF52F88AFC1FF812" blockId="10.[151,1436,1724,2039]" pageId="10" pageNumber="11">
<emphasis id="179DA041413DFFD8FF52F88AFE80F86F" bold="true" box="[199,322,1904,1930]" pageId="10" pageNumber="11">Diagnosis.</emphasis>
Bodywall table ossicles with distorted oval or rhomboidal disc and delicate spire, mostly with one pillar formed by fusion of two pillars and spire with 46 minute apical spines. Pedicel supporting table ossicles with narrow curved disc with 24 large central perforations and 28 small distal perforations; spire high, with two (occasionally three) pillars united distally, ending in 24 minute processes.
</paragraph>
</subSubSection>
<caption id="71962CDB413CFFD9FF02F89AFC92F801" ID-DOI="http://doi.org/10.5281/zenodo.5225382" ID-Zenodo-Dep="5225382" httpUri="https://zenodo.org/record/5225382/files/figure.png" pageId="11" pageNumber="12" startId="11.[151,250,1888,1912]" targetBox="[182,1404,182,1863]" targetPageId="11">
<paragraph id="25567C53413CFFD9FF02F89AFC92F801" blockId="11.[151,1436,1888,2020]" pageId="11" pageNumber="12">
<emphasis id="179DA041413CFFD9FF02F89AFED7F89D" bold="true" box="[151,277,1888,1912]" pageId="11" pageNumber="12">FIGURE 5.</emphasis>
Scanning electron microscope images of ossicles of
<taxonomicName id="E2E907D0413CFFD9FCA9F89AFC7EF89D" authority="Yamana &amp; Nakaguchi &amp; Yamaguchi &amp; Katoh &amp; Ogawa &amp; Ohtsuka, 2021" authorityName="Yamana &amp; Nakaguchi &amp; Yamaguchi &amp; Katoh &amp; Ogawa &amp; Ohtsuka" authorityYear="2021" box="[828,956,1888,1912]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="11" pageNumber="12" phylum="Echinodermata" rank="species" species="liaoi" status="sp. nov.">
<emphasis id="179DA041413CFFD9FCA9F89AFC7EF89D" box="[828,956,1888,1912]" italics="true" pageId="11" pageNumber="12">Thyone liaoi</emphasis>
</taxonomicName>
<emphasis id="179DA041413CFFD9FC50F89AFBD5F89D" bold="true" box="[965,1047,1888,1912]" pageId="11" pageNumber="12">
<taxonomicNameLabel id="0CAE1D3A413CFFD9FC50F89AFBD5F89D" box="[965,1047,1888,1912]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, (
<emphasis id="179DA041413CFFD9FBBAF89AFB44F89D" box="[1071,1158,1888,1912]" italics="true" pageId="11" pageNumber="12">holotype</emphasis>
, AH, WMNH-INV-2014- 182). A, ossicles of the ventrolateral tentacle; B, the ventrolateral skin of peri-oral; C, the ventrolateral skin of pharynx; D, the ventrolateral introvert skin; E, the pedicel of the abdominal side; F and G, the middle part of the bodywall on the abdominal side, and on the dorsal side, respectively; H, the ventrolateral anal papilla.
</paragraph>
</caption>
<caption id="71962CDB413BFFDEFF02FE1FFAF9FE1A" box="[151,1339,485,511]" pageId="12" pageNumber="13" startId="12.[151,239,485,511]">
<paragraph id="25567C53413BFFDEFF02FE1FFAF9FE1A" blockId="12.[151,1339,485,511]" box="[151,1339,485,511]" pageId="12" pageNumber="13">
<emphasis id="179DA041413BFFDEFF02FE1FFEC8FE1A" bold="true" box="[151,266,485,511]" pageId="12" pageNumber="13">TABLE 4.</emphasis>
Counts and measurements (Mean ± sd μm) of ossicles from four specimens of
<taxonomicName id="E2E907D0413BFFDEFBCCFE1FFB1DFE1A" authority="Yamana &amp; Nakaguchi &amp; Yamaguchi &amp; Katoh &amp; Ogawa &amp; Ohtsuka, 2021" authorityName="Yamana &amp; Nakaguchi &amp; Yamaguchi &amp; Katoh &amp; Ogawa &amp; Ohtsuka" authorityYear="2021" box="[1113,1247,485,511]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="12" pageNumber="13" phylum="Echinodermata" rank="species" species="liaoi" status="sp. nov.">
<emphasis id="179DA041413BFFDEFBCCFE1FFB1DFE1A" box="[1113,1247,485,511]" italics="true" pageId="12" pageNumber="13">Thyone liaoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0CAE1D3A413BFFDEFB70FE1FFAF9FE1A" box="[1253,1339,485,511]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="25567C53413BFFDEFE2AFDF7FDF3F976" pageId="12" pageNumber="13">
<table id="57E98EF3413B002DFF37FDF7F829F976" box="[162,2027,525,1683]" gridcols="19" gridrows="35" pageId="12" pageNumber="13">
<tr id="9BD97E11413B002DFF37FDF7F829FDC1" box="[162,2027,525,548]" gridrow="0" pageId="12" pageNumber="13" rowspan-0="1" rowspan-1="1" rowspan-12="1" rowspan-18="1" rowspan-4="1" rowspan-5="1" rowspan-8="1" rowspan-9="1">
<th id="D808176D413B002DFE28FDF7FDB7FDC1" box="[445,629,525,548]" colspan="2" colspanRight="1" gridcol="2" gridrow="0" pageId="12" pageNumber="13">Plate and rod</th>
<th id="D808176D413B002DFCA9FDF7FC37FDC1" box="[828,1013,525,548]" colspan="2" colspanRight="1" gridcol="6" gridrow="0" pageId="12" pageNumber="13">Endplate</th>
<th id="D808176D413B002DFB2CFDF7FA8DFDC1" box="[1209,1359,525,548]" colspan="2" colspanRight="1" gridcol="10" gridrow="0" pageId="12" pageNumber="13">Rosette</th>
<th id="D808176D413B002DFA2CFDF7F852FDC1" box="[1465,1936,525,548]" colspan="5" colspanRight="4" gridcol="13" gridrow="0" pageId="12" pageNumber="13">Table (T) and supporting table (ST)</th>
</tr>
<tr id="9BD97E11413B002DFF37FDD6F829FDA0" box="[162,2027,556,581]" gridrow="1" pageId="12" pageNumber="13" rowspan-1="1" rowspan-12="1" rowspan-4="1" rowspan-9="1">
<th id="D808176D413B002DFF37FDD6FE9BFDA0" box="[162,345,556,581]" gridcol="0" gridrow="1" pageId="12" pageNumber="13">Samplea</th>
<td id="D808176D413B002DFE28FDD6FDB7FDA0" box="[445,629,556,581]" colspan="2" colspanRight="1" gridcol="2" gridrow="1" pageId="12" pageNumber="13">Length</td>
<td id="D808176D413B002DFD46FDD6FCBCFDA0" box="[723,894,556,581]" colspan="2" colspanRight="1" gridcol="5" gridrow="1" pageId="12" pageNumber="13">Diameter</td>
<td id="D808176D413B002DFC24FDD6FB98FDA0" box="[945,1114,556,581]" colspan="2" colspanRight="1" gridcol="7" gridrow="1" pageId="12" pageNumber="13">No. plate hole</td>
<td id="D808176D413B002DFB2CFDD6FA8DFDA0" box="[1209,1359,556,581]" colspan="2" colspanRight="1" gridcol="10" gridrow="1" pageId="12" pageNumber="13">Length</td>
<td id="D808176D413B002DFA2CFDD6F9AAFDA0" box="[1465,1640,556,581]" colspan="2" colspanRight="1" gridcol="13" gridrow="1" pageId="12" pageNumber="13">Disc length</td>
<td id="D808176D413B002DF905FDD6F8EDFDA0" box="[1680,1839,556,581]" colspan="2" colspanRight="1" gridcol="15" gridrow="1" pageId="12" pageNumber="13">No. disc hole (T)</td>
<td id="D808176D413B002DF8CDFDD6F829FDA0" box="[1880,2027,556,581]" colspan="2" colspanRight="1" gridcol="17" gridrow="1" pageId="12" pageNumber="13">Spire (ST)</td>
</tr>
<tr id="9BD97E11413B002DFF37FDB5F829FD80" box="[162,2027,591,613]" gridrow="2" pageId="12" pageNumber="13" rowspan-0="1">
<td id="D808176D413B002DFE1CFDB5FE5DFD80" box="[393,415,591,613]" gridcol="1" gridrow="2" pageId="12" pageNumber="13">
<emphasis id="179DA041413BFFDEFE1BFDB5FE58FD80" box="[398,410,591,613]" italics="true" pageId="12" pageNumber="13">n</emphasis>
</td>
<td id="D808176D413B002DFE28FDB5FDD7FD80" box="[445,533,591,613]" gridcol="2" gridrow="2" pageId="12" pageNumber="13">Mean</td>
<td id="D808176D413B002DFDA4FDB5FDB7FD80" box="[561,629,591,613]" gridcol="3" gridrow="2" pageId="12" pageNumber="13">Range</td>
<td id="D808176D413B002DFD31FDB5FD72FD80" box="[676,688,591,613]" gridcol="4" gridrow="2" pageId="12" pageNumber="13">
<emphasis id="179DA041413BFFDEFD31FDB5FD72FD80" box="[676,688,591,613]" italics="true" pageId="12" pageNumber="13">n</emphasis>
</td>
<td id="D808176D413B002DFD46FDB5FCD4FD80" box="[723,790,591,613]" gridcol="5" gridrow="2" pageId="12" pageNumber="13">Mean</td>
<td id="D808176D413B002DFCA9FDB5FCBCFD80" box="[828,894,591,613]" gridcol="6" gridrow="2" pageId="12" pageNumber="13">Range</td>
<td id="D808176D413B002DFC24FDB5FC37FD80" box="[945,1013,591,613]" gridcol="7" gridrow="2" pageId="12" pageNumber="13">Mean</td>
<td id="D808176D413B002DFB8DFDB5FB98FD80" box="[1048,1114,591,613]" gridcol="8" gridrow="2" pageId="12" pageNumber="13">Range</td>
<td id="D808176D413B002DFB11FDB5FB59FD80" box="[1156,1179,591,613]" gridcol="9" gridrow="2" pageId="12" pageNumber="13">
<emphasis id="179DA041413BFFDEFB1CFDB5FB57FD80" box="[1161,1173,591,613]" italics="true" pageId="12" pageNumber="13">n</emphasis>
</td>
<td id="D808176D413B002DFB2CFDB5FB36FD80" box="[1209,1268,591,613]" gridcol="10" gridrow="2" pageId="12" pageNumber="13">Mean</td>
<td id="D808176D413B002DFA87FDB5FA8DFD80" box="[1298,1359,591,613]" gridcol="11" gridrow="2" pageId="12" pageNumber="13">Range</td>
<td id="D808176D413B002DFAEBFDB5FA56FD80" box="[1406,1428,591,613]" gridcol="12" gridrow="2" pageId="12" pageNumber="13">
<emphasis id="179DA041413BFFDEFA16FDB5FA4DFD80" box="[1411,1423,591,613]" italics="true" pageId="12" pageNumber="13">n</emphasis>
</td>
<td id="D808176D413B002DFA2CFDB5F9C6FD80" box="[1465,1540,591,613]" gridcol="13" gridrow="2" pageId="12" pageNumber="13">Mean</td>
<td id="D808176D413B002DF9B6FDB5F9AAFD80" box="[1571,1640,591,613]" gridcol="14" gridrow="2" pageId="12" pageNumber="13">Range</td>
<td id="D808176D413B002DF905FDB5F909FD80" box="[1680,1739,591,613]" gridcol="15" gridrow="2" pageId="12" pageNumber="13">Mean</td>
<td id="D808176D413B002DF978FDB5F8EDFD80" box="[1773,1839,591,613]" gridcol="16" gridrow="2" pageId="12" pageNumber="13">Range</td>
<td id="D808176D413B002DF8CDFDB5F852FD80" box="[1880,1936,591,613]" gridcol="17" gridrow="2" pageId="12" pageNumber="13">Mean</td>
<td id="D808176D413B002DF825FDB5F829FD80" box="[1968,2027,591,613]" gridcol="18" gridrow="2" pageId="12" pageNumber="13">Range</td>
</tr>
<tr id="9BD97E11413B002DFF37FD8AF829FD62" box="[162,2027,624,647]" gridrow="3" pageId="12" pageNumber="13">
<th id="D808176D413B002DFF37FD8AF829FD62" box="[162,2027,624,647]" colspan="19" colspanRight="18" gridcol="0" gridrow="3" pageId="12" pageNumber="13">WMNH-INV-2014-182</th>
</tr>
<tr id="9BD97E11413B002DFF37FD6BF829FD4D" box="[162,2027,657,680]" gridrow="4" pageId="12" pageNumber="13" rowspan-10="1" rowspan-11="1" rowspan-17="1" rowspan-18="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413B002DFF37FD6BFE9BFD4D" box="[162,345,657,680]" gridcol="0" gridrow="4" pageId="12" pageNumber="13">Tentacle</th>
<td id="D808176D413B002DFE1CFD6BFE5DFD4D" box="[393,415,657,680]" gridcol="1" gridrow="4" pageId="12" pageNumber="13">30</td>
<td id="D808176D413B002DFE28FD6BFDD7FD4D" box="[445,533,657,680]" gridcol="2" gridrow="4" pageId="12" pageNumber="13">139 ± 108</td>
<td id="D808176D413B002DFDA4FD6BFDB7FD4D" box="[561,629,657,680]" gridcol="3" gridrow="4" pageId="12" pageNumber="13">36432</td>
<td id="D808176D413B002DFD31FD6BFD72FD4D" box="[676,688,657,680]" gridcol="4" gridrow="4" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFB11FD6BFB59FD4D" box="[1156,1179,657,680]" gridcol="9" gridrow="4" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFAEBFD6BFA56FD4D" box="[1406,1428,657,680]" gridcol="12" gridrow="4" pageId="12" pageNumber="13">4</td>
<td id="D808176D413B002DFA2CFD6BF9C6FD4D" box="[1465,1540,657,680]" gridcol="13" gridrow="4" pageId="12" pageNumber="13">147 ± 100</td>
<td id="D808176D413B002DF9B6FD6BF9AAFD4D" box="[1571,1640,657,680]" gridcol="14" gridrow="4" pageId="12" pageNumber="13">77294</td>
<td id="D808176D413B002DF905FD6BF909FD4D" box="[1680,1739,657,680]" gridcol="15" gridrow="4" pageId="12" pageNumber="13">19 ± 11</td>
<td id="D808176D413B002DF978FD6BF8EDFD4D" box="[1773,1839,657,680]" gridcol="16" gridrow="4" pageId="12" pageNumber="13">1035</td>
</tr>
<tr id="9BD97E11413B002DFF37FD4EF829FD2E" box="[162,2027,692,715]" gridrow="5" pageId="12" pageNumber="13" rowspan-17="1" rowspan-18="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413B002DFF37FD4EFE9BFD2E" box="[162,345,692,715]" gridcol="0" gridrow="5" pageId="12" pageNumber="13">Peri-oral skin</th>
<td id="D808176D413B002DFE1CFD4EFE5DFD2E" box="[393,415,692,715]" gridcol="1" gridrow="5" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFD31FD4EFD72FD2E" box="[676,688,692,715]" gridcol="4" gridrow="5" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFB11FD4EFB59FD2E" box="[1156,1179,692,715]" gridcol="9" gridrow="5" pageId="12" pageNumber="13">25</td>
<td id="D808176D413B002DFB2CFD4EFB36FD2E" box="[1209,1268,692,715]" gridcol="10" gridrow="5" pageId="12" pageNumber="13">37±11</td>
<td id="D808176D413B002DFA87FD4EFA8DFD2E" box="[1298,1359,692,715]" gridcol="11" gridrow="5" pageId="12" pageNumber="13">2369</td>
<td id="D808176D413B002DFAEBFD4EFA56FD2E" box="[1406,1428,692,715]" gridcol="12" gridrow="5" pageId="12" pageNumber="13">6</td>
<td id="D808176D413B002DFA2CFD4EF9C6FD2E" box="[1465,1540,692,715]" gridcol="13" gridrow="5" pageId="12" pageNumber="13">104 ± 11</td>
<td id="D808176D413B002DF9B6FD4EF9AAFD2E" box="[1571,1640,692,715]" gridcol="14" gridrow="5" pageId="12" pageNumber="13">82112</td>
<td id="D808176D413B002DF905FD4EF909FD2E" box="[1680,1739,692,715]" gridcol="15" gridrow="5" pageId="12" pageNumber="13">20 ± 4</td>
<td id="D808176D413B002DF978FD4EF8EDFD2E" box="[1773,1839,692,715]" gridcol="16" gridrow="5" pageId="12" pageNumber="13">1524</td>
</tr>
<tr id="9BD97E11413B002DFF37FD2DF829FD0B" box="[162,2027,727,750]" gridrow="6" pageId="12" pageNumber="13" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-17="1" rowspan-18="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413B002DFF37FD2DFE9BFD0B" box="[162,345,727,750]" gridcol="0" gridrow="6" pageId="12" pageNumber="13">Oesophagus skin</th>
<td id="D808176D413B002DFE1CFD2DFE5DFD0B" box="[393,415,727,750]" gridcol="1" gridrow="6" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFD31FD2DFD72FD0B" box="[676,688,727,750]" gridcol="4" gridrow="6" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFB11FD2DFB59FD0B" box="[1156,1179,727,750]" gridcol="9" gridrow="6" pageId="12" pageNumber="13">10</td>
<td id="D808176D413B002DFB2CFD2DFB36FD0B" box="[1209,1268,727,750]" gridcol="10" gridrow="6" pageId="12" pageNumber="13">36 ± 5</td>
<td id="D808176D413B002DFA87FD2DFA8DFD0B" box="[1298,1359,727,750]" gridcol="11" gridrow="6" pageId="12" pageNumber="13">2743</td>
<td id="D808176D413B002DFAEBFD2DFA56FD0B" box="[1406,1428,727,750]" gridcol="12" gridrow="6" pageId="12" pageNumber="13">0</td>
</tr>
<tr id="9BD97E11413B002DFF37FD03F829FCF5" box="[162,2027,761,784]" gridrow="7" pageId="12" pageNumber="13" rowspan-17="1" rowspan-18="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413B002DFF37FD03FE9BFCF5" box="[162,345,761,784]" gridcol="0" gridrow="7" pageId="12" pageNumber="13">Introvert skin</th>
<td id="D808176D413B002DFE1CFD03FE5DFCF5" box="[393,415,761,784]" gridcol="1" gridrow="7" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFD31FD03FD72FCF5" box="[676,688,761,784]" gridcol="4" gridrow="7" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFB11FD03FB59FCF5" box="[1156,1179,761,784]" gridcol="9" gridrow="7" pageId="12" pageNumber="13">4</td>
<td id="D808176D413B002DFB2CFD03FB36FCF5" box="[1209,1268,761,784]" gridcol="10" gridrow="7" pageId="12" pageNumber="13">47 ± 13</td>
<td id="D808176D413B002DFA87FD03FA8DFCF5" box="[1298,1359,761,784]" gridcol="11" gridrow="7" pageId="12" pageNumber="13">3362</td>
<td id="D808176D413B002DFAEBFD03FA56FCF5" box="[1406,1428,761,784]" gridcol="12" gridrow="7" pageId="12" pageNumber="13">12</td>
<td id="D808176D413B002DFA2CFD03F9C6FCF5" box="[1465,1540,761,784]" gridcol="13" gridrow="7" pageId="12" pageNumber="13">88 ± 11</td>
<td id="D808176D413B002DF9B6FD03F9AAFCF5" box="[1571,1640,761,784]" gridcol="14" gridrow="7" pageId="12" pageNumber="13">68108</td>
<td id="D808176D413B002DF905FD03F909FCF5" box="[1680,1739,761,784]" gridcol="15" gridrow="7" pageId="12" pageNumber="13">17 ± 4</td>
<td id="D808176D413B002DF978FD03F8EDFCF5" box="[1773,1839,761,784]" gridcol="16" gridrow="7" pageId="12" pageNumber="13">1023</td>
</tr>
<tr id="9BD97E11413B002DFF37FCE0F829FCD4" box="[162,2027,794,817]" gridrow="8" pageId="12" pageNumber="13" rowspan-10="1" rowspan-11="1" rowspan-15="1" rowspan-16="1" rowspan-2="1" rowspan-3="1">
<th id="D808176D413B002DFF37FCE0FE9BFCD4" box="[162,345,794,817]" gridcol="0" gridrow="8" pageId="12" pageNumber="13">Pedicels (ventral)</th>
<td id="D808176D413B002DFE1CFCE0FE5DFCD4" box="[393,415,794,817]" gridcol="1" gridrow="8" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFD31FCE0FD72FCD4" box="[676,688,794,817]" gridcol="4" gridrow="8" pageId="12" pageNumber="13">5</td>
<td id="D808176D413B002DFD46FCE0FCD4FCD4" box="[723,790,794,817]" gridcol="5" gridrow="8" pageId="12" pageNumber="13">176 ± 3</td>
<td id="D808176D413B002DFCA9FCE0FCBCFCD4" box="[828,894,794,817]" gridcol="6" gridrow="8" pageId="12" pageNumber="13">172179</td>
<td id="D808176D413B002DFC24FCE0FC37FCD4" box="[945,1013,794,817]" gridcol="7" gridrow="8" pageId="12" pageNumber="13">193 ± 14</td>
<td id="D808176D413B002DFB8DFCE0FB98FCD4" box="[1048,1114,794,817]" gridcol="8" gridrow="8" pageId="12" pageNumber="13">171209</td>
<td id="D808176D413B002DFB11FCE0FB59FCD4" box="[1156,1179,794,817]" gridcol="9" gridrow="8" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFAEBFCE0FA56FCD4" box="[1406,1428,794,817]" gridcol="12" gridrow="8" pageId="12" pageNumber="13">24</td>
<td id="D808176D413B002DFA2CFCE0F9C6FCD4" box="[1465,1540,794,817]" gridcol="13" gridrow="8" pageId="12" pageNumber="13">90 ± 6</td>
<td id="D808176D413B002DF9B6FCE0F9AAFCD4" box="[1571,1640,794,817]" gridcol="14" gridrow="8" pageId="12" pageNumber="13">78100</td>
<td id="D808176D413B002DF8CDFCE0F852FCD4" box="[1880,1936,794,817]" gridcol="17" gridrow="8" pageId="12" pageNumber="13">37 ± 4</td>
<td id="D808176D413B002DF825FCE0F829FCD4" box="[1968,2027,794,817]" gridcol="18" gridrow="8" pageId="12" pageNumber="13">2945</td>
</tr>
<tr id="9BD97E11413B002DFF37FCC1F829FCB7" box="[162,2027,827,850]" gridrow="9" pageId="12" pageNumber="13" rowspan-10="1" rowspan-11="1" rowspan-17="1" rowspan-18="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413B002DFF37FCC1FE9BFCB7" box="[162,345,827,850]" gridcol="0" gridrow="9" pageId="12" pageNumber="13">Bodywall (ventral)</th>
<td id="D808176D413B002DFE1CFCC1FE5DFCB7" box="[393,415,827,850]" gridcol="1" gridrow="9" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFD31FCC1FD72FCB7" box="[676,688,827,850]" gridcol="4" gridrow="9" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFB11FCC1FB59FCB7" box="[1156,1179,827,850]" gridcol="9" gridrow="9" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFAEBFCC1FA56FCB7" box="[1406,1428,827,850]" gridcol="12" gridrow="9" pageId="12" pageNumber="13">16</td>
<td id="D808176D413B002DFA2CFCC1F9C6FCB7" box="[1465,1540,827,850]" gridcol="13" gridrow="9" pageId="12" pageNumber="13">86 ± 12</td>
<td id="D808176D413B002DF9B6FCC1F9AAFCB7" box="[1571,1640,827,850]" gridcol="14" gridrow="9" pageId="12" pageNumber="13">68109</td>
<td id="D808176D413B002DF905FCC1F909FCB7" box="[1680,1739,827,850]" gridcol="15" gridrow="9" pageId="12" pageNumber="13">9 ± 2</td>
<td id="D808176D413B002DF978FCC1F8EDFCB7" box="[1773,1839,827,850]" gridcol="16" gridrow="9" pageId="12" pageNumber="13">513</td>
</tr>
<tr id="9BD97E11413B002DFF37FCA7F829FC91" box="[162,2027,861,884]" gridrow="10" pageId="12" pageNumber="13" rowspan-10="1" rowspan-11="1" rowspan-17="1" rowspan-18="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413B002DFF37FCA7FE9BFC91" box="[162,345,861,884]" gridcol="0" gridrow="10" pageId="12" pageNumber="13">Anal papilla</th>
<td id="D808176D413B002DFE1CFCA7FE5DFC91" box="[393,415,861,884]" gridcol="1" gridrow="10" pageId="12" pageNumber="13">7</td>
<td id="D808176D413B002DFE28FCA7FDD7FC91" box="[445,533,861,884]" gridcol="2" gridrow="10" pageId="12" pageNumber="13">74 ± 10</td>
<td id="D808176D413B002DFDA4FCA7FDB7FC91" box="[561,629,861,884]" gridcol="3" gridrow="10" pageId="12" pageNumber="13">6291</td>
<td id="D808176D413B002DFD31FCA7FD72FC91" box="[676,688,861,884]" gridcol="4" gridrow="10" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFB11FCA7FB59FC91" box="[1156,1179,861,884]" gridcol="9" gridrow="10" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFAEBFCA7FA56FC91" box="[1406,1428,861,884]" gridcol="12" gridrow="10" pageId="12" pageNumber="13">4</td>
<td id="D808176D413B002DFA2CFCA7F9C6FC91" box="[1465,1540,861,884]" gridcol="13" gridrow="10" pageId="12" pageNumber="13">63 ± 3</td>
<td id="D808176D413B002DF9B6FCA7F9AAFC91" box="[1571,1640,861,884]" gridcol="14" gridrow="10" pageId="12" pageNumber="13">6067</td>
<td id="D808176D413B002DF905FCA7F909FC91" box="[1680,1739,861,884]" gridcol="15" gridrow="10" pageId="12" pageNumber="13">16 ± 3</td>
<td id="D808176D413B002DF978FCA7F8EDFC91" box="[1773,1839,861,884]" gridcol="16" gridrow="10" pageId="12" pageNumber="13">1320</td>
</tr>
<tr id="9BD97E11413B002DFF37FC84F829FC70" box="[162,2027,894,917]" gridrow="11" pageId="12" pageNumber="13">
<th id="D808176D413B002DFF37FC84F829FC70" box="[162,2027,894,917]" colspan="19" colspanRight="18" gridcol="0" gridrow="11" pageId="12" pageNumber="13">WMNH-INV-2014-183</th>
</tr>
<tr id="9BD97E11413B002DFF37FC5AF829FC52" box="[162,2027,928,951]" gridrow="12" pageId="12" pageNumber="13" rowspan-10="1" rowspan-11="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-17="1" rowspan-18="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413B002DFF37FC5AFE9BFC52" box="[162,345,928,951]" gridcol="0" gridrow="12" pageId="12" pageNumber="13">Tentacle</th>
<td id="D808176D413B002DFE1CFC5AFE5DFC52" box="[393,415,928,951]" gridcol="1" gridrow="12" pageId="12" pageNumber="13">29</td>
<td id="D808176D413B002DFE28FC5AFDD7FC52" box="[445,533,928,951]" gridcol="2" gridrow="12" pageId="12" pageNumber="13">103 ± 72</td>
<td id="D808176D413B002DFDA4FC5AFDB7FC52" box="[561,629,928,951]" gridcol="3" gridrow="12" pageId="12" pageNumber="13">38295</td>
<td id="D808176D413B002DFD31FC5AFD72FC52" box="[676,688,928,951]" gridcol="4" gridrow="12" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFB11FC5AFB59FC52" box="[1156,1179,928,951]" gridcol="9" gridrow="12" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFAEBFC5AFA56FC52" box="[1406,1428,928,951]" gridcol="12" gridrow="12" pageId="12" pageNumber="13">0</td>
</tr>
<tr id="9BD97E11413B002DFF37FC3BF829FC3D" box="[162,2027,961,984]" gridrow="13" pageId="12" pageNumber="13" rowspan-10="1" rowspan-11="1" rowspan-17="1" rowspan-18="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413B002DFF37FC3BFE9BFC3D" box="[162,345,961,984]" gridcol="0" gridrow="13" pageId="12" pageNumber="13">Peri-oral skin</th>
<td id="D808176D413B002DFE1CFC3BFE5DFC3D" box="[393,415,961,984]" gridcol="1" gridrow="13" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFD31FC3BFD72FC3D" box="[676,688,961,984]" gridcol="4" gridrow="13" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFB11FC3BFB59FC3D" box="[1156,1179,961,984]" gridcol="9" gridrow="13" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFAEBFC3BFA56FC3D" box="[1406,1428,961,984]" gridcol="12" gridrow="13" pageId="12" pageNumber="13">17</td>
<td id="D808176D413B002DFA2CFC3BF9C6FC3D" box="[1465,1540,961,984]" gridcol="13" gridrow="13" pageId="12" pageNumber="13">75 ± 9</td>
<td id="D808176D413B002DF9B6FC3BF9AAFC3D" box="[1571,1640,961,984]" gridcol="14" gridrow="13" pageId="12" pageNumber="13">6492</td>
<td id="D808176D413B002DF905FC3BF909FC3D" box="[1680,1739,961,984]" gridcol="15" gridrow="13" pageId="12" pageNumber="13">18 ± 5</td>
<td id="D808176D413B002DF978FC3BF8EDFC3D" box="[1773,1839,961,984]" gridcol="16" gridrow="13" pageId="12" pageNumber="13">1225</td>
</tr>
<tr id="9BD97E11413B002DFF37FC18F829FC1C" box="[162,2027,994,1017]" gridrow="14" pageId="12" pageNumber="13" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-17="1" rowspan-18="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413B002DFF37FC18FE9BFC1C" box="[162,345,994,1017]" gridcol="0" gridrow="14" pageId="12" pageNumber="13">Oesophagus skin</th>
<td id="D808176D413B002DFE1CFC18FE5DFC1C" box="[393,415,994,1017]" gridcol="1" gridrow="14" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFD31FC18FD72FC1C" box="[676,688,994,1017]" gridcol="4" gridrow="14" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFB11FC18FB59FC1C" box="[1156,1179,994,1017]" gridcol="9" gridrow="14" pageId="12" pageNumber="13">26</td>
<td id="D808176D413B002DFB2CFC18FB36FC1C" box="[1209,1268,994,1017]" gridcol="10" gridrow="14" pageId="12" pageNumber="13">27 ± 7</td>
<td id="D808176D413B002DFA87FC18FA8DFC1C" box="[1298,1359,994,1017]" gridcol="11" gridrow="14" pageId="12" pageNumber="13">1744</td>
<td id="D808176D413B002DFAEBFC18FA56FC1C" box="[1406,1428,994,1017]" gridcol="12" gridrow="14" pageId="12" pageNumber="13">0</td>
</tr>
<tr id="9BD97E11413B002DFF37FBF9F829FBFF" box="[162,2027,1027,1050]" gridrow="15" pageId="12" pageNumber="13" rowspan-17="1" rowspan-18="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413B002DFF37FBF9FE9BFBFF" box="[162,345,1027,1050]" gridcol="0" gridrow="15" pageId="12" pageNumber="13">Introvert skin</th>
<td id="D808176D413B002DFE1CFBF9FE5DFBFF" box="[393,415,1027,1050]" gridcol="1" gridrow="15" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFD31FBF9FD72FBFF" box="[676,688,1027,1050]" gridcol="4" gridrow="15" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFB11FBF9FB59FBFF" box="[1156,1179,1027,1050]" gridcol="9" gridrow="15" pageId="12" pageNumber="13">13</td>
<td id="D808176D413B002DFB2CFBF9FB36FBFF" box="[1209,1268,1027,1050]" gridcol="10" gridrow="15" pageId="12" pageNumber="13">40 ± 16</td>
<td id="D808176D413B002DFA87FBF9FA8DFBFF" box="[1298,1359,1027,1050]" gridcol="11" gridrow="15" pageId="12" pageNumber="13">2283</td>
<td id="D808176D413B002DFAEBFBF9FA56FBFF" box="[1406,1428,1027,1050]" gridcol="12" gridrow="15" pageId="12" pageNumber="13">5</td>
<td id="D808176D413B002DFA2CFBF9F9C6FBFF" box="[1465,1540,1027,1050]" gridcol="13" gridrow="15" pageId="12" pageNumber="13">74 ± 18</td>
<td id="D808176D413B002DF9B6FBF9F9AAFBFF" box="[1571,1640,1027,1050]" gridcol="14" gridrow="15" pageId="12" pageNumber="13">4895</td>
<td id="D808176D413B002DF905FBF9F909FBFF" box="[1680,1739,1027,1050]" gridcol="15" gridrow="15" pageId="12" pageNumber="13">15 ± 5</td>
<td id="D808176D413B002DF978FBF9F8EDFBFF" box="[1773,1839,1027,1050]" gridcol="16" gridrow="15" pageId="12" pageNumber="13">1122</td>
</tr>
<tr id="9BD97E11413B002DFF37FBDFF829FBD9" box="[162,2027,1061,1084]" gridrow="16" pageId="12" pageNumber="13" rowspan-10="1" rowspan-11="1" rowspan-15="1" rowspan-16="1" rowspan-2="1" rowspan-3="1">
<th id="D808176D413B002DFF37FBDFFE9BFBD9" box="[162,345,1061,1084]" gridcol="0" gridrow="16" pageId="12" pageNumber="13">Pedicels (ventral)</th>
<td id="D808176D413B002DFE1CFBDFFE5DFBD9" box="[393,415,1061,1084]" gridcol="1" gridrow="16" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFD31FBDFFD72FBD9" box="[676,688,1061,1084]" gridcol="4" gridrow="16" pageId="12" pageNumber="13">3</td>
<td id="D808176D413B002DFD46FBDFFCD4FBD9" box="[723,790,1061,1084]" gridcol="5" gridrow="16" pageId="12" pageNumber="13">179 ± 12</td>
<td id="D808176D413B002DFCA9FBDFFCBCFBD9" box="[828,894,1061,1084]" gridcol="6" gridrow="16" pageId="12" pageNumber="13">166187</td>
<td id="D808176D413B002DFC24FBDFFC37FBD9" box="[945,1013,1061,1084]" gridcol="7" gridrow="16" pageId="12" pageNumber="13">161 ± 36</td>
<td id="D808176D413B002DFB8DFBDFFB98FBD9" box="[1048,1114,1061,1084]" gridcol="8" gridrow="16" pageId="12" pageNumber="13">127198</td>
<td id="D808176D413B002DFB11FBDFFB59FBD9" box="[1156,1179,1061,1084]" gridcol="9" gridrow="16" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFAEBFBDFFA56FBD9" box="[1406,1428,1061,1084]" gridcol="12" gridrow="16" pageId="12" pageNumber="13">19</td>
<td id="D808176D413B002DFA2CFBDFF9C6FBD9" box="[1465,1540,1061,1084]" gridcol="13" gridrow="16" pageId="12" pageNumber="13">90 ± 10</td>
<td id="D808176D413B002DF9B6FBDFF9AAFBD9" box="[1571,1640,1061,1084]" gridcol="14" gridrow="16" pageId="12" pageNumber="13">74103</td>
<td id="D808176D413B002DF8CDFBDFF852FBD9" box="[1880,1936,1061,1084]" gridcol="17" gridrow="16" pageId="12" pageNumber="13">34 ± 6</td>
<td id="D808176D413B002DF825FBDFF829FBD9" box="[1968,2027,1061,1084]" gridcol="18" gridrow="16" pageId="12" pageNumber="13">1941</td>
</tr>
<tr id="9BD97E11413B002DFF37FBBCF829FBB8" box="[162,2027,1094,1117]" gridrow="17" pageId="12" pageNumber="13" rowspan-10="1" rowspan-11="1" rowspan-17="1" rowspan-18="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413B002DFF37FBBCFE9BFBB8" box="[162,345,1094,1117]" gridcol="0" gridrow="17" pageId="12" pageNumber="13">Bodywall (ventral)</th>
<td id="D808176D413B002DFE1CFBBCFE5DFBB8" box="[393,415,1094,1117]" gridcol="1" gridrow="17" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFD31FBBCFD72FBB8" box="[676,688,1094,1117]" gridcol="4" gridrow="17" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFB11FBBCFB59FBB8" box="[1156,1179,1094,1117]" gridcol="9" gridrow="17" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFAEBFBBCFA56FBB8" box="[1406,1428,1094,1117]" gridcol="12" gridrow="17" pageId="12" pageNumber="13">28</td>
<td id="D808176D413B002DFA2CFBBCF9C6FBB8" box="[1465,1540,1094,1117]" gridcol="13" gridrow="17" pageId="12" pageNumber="13">100 ± 11</td>
<td id="D808176D413B002DF9B6FBBCF9AAFBB8" box="[1571,1640,1094,1117]" gridcol="14" gridrow="17" pageId="12" pageNumber="13">81120</td>
<td id="D808176D413B002DF905FBBCF909FBB8" box="[1680,1739,1094,1117]" gridcol="15" gridrow="17" pageId="12" pageNumber="13">9 ± 2</td>
<td id="D808176D413B002DF978FBBCF8EDFBB8" box="[1773,1839,1094,1117]" gridcol="16" gridrow="17" pageId="12" pageNumber="13">514</td>
</tr>
<tr id="9BD97E11413B002DFF37FB92F829FB9A" box="[162,2027,1128,1151]" gridrow="18" pageId="12" pageNumber="13" rowspan-10="1" rowspan-11="1" rowspan-17="1" rowspan-18="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413B002DFF37FB92FE9BFB9A" box="[162,345,1128,1151]" gridcol="0" gridrow="18" pageId="12" pageNumber="13">Anal papilla</th>
<td id="D808176D413B002DFE1CFB92FE5DFB9A" box="[393,415,1128,1151]" gridcol="1" gridrow="18" pageId="12" pageNumber="13">12</td>
<td id="D808176D413B002DFE28FB92FDD7FB9A" box="[445,533,1128,1151]" gridcol="2" gridrow="18" pageId="12" pageNumber="13">67 ± 6</td>
<td id="D808176D413B002DFDA4FB92FDB7FB9A" box="[561,629,1128,1151]" gridcol="3" gridrow="18" pageId="12" pageNumber="13">5977</td>
<td id="D808176D413B002DFD31FB92FD72FB9A" box="[676,688,1128,1151]" gridcol="4" gridrow="18" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFB11FB92FB59FB9A" box="[1156,1179,1128,1151]" gridcol="9" gridrow="18" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFAEBFB92FA56FB9A" box="[1406,1428,1128,1151]" gridcol="12" gridrow="18" pageId="12" pageNumber="13">14</td>
<td id="D808176D413B002DFA2CFB92F9C6FB9A" box="[1465,1540,1128,1151]" gridcol="13" gridrow="18" pageId="12" pageNumber="13">55 ± 7</td>
<td id="D808176D413B002DF9B6FB92F9AAFB9A" box="[1571,1640,1128,1151]" gridcol="14" gridrow="18" pageId="12" pageNumber="13">4569</td>
<td id="D808176D413B002DF905FB92F909FB9A" box="[1680,1739,1128,1151]" gridcol="15" gridrow="18" pageId="12" pageNumber="13">11 ± 2</td>
<td id="D808176D413B002DF978FB92F8EDFB9A" box="[1773,1839,1128,1151]" gridcol="16" gridrow="18" pageId="12" pageNumber="13">816</td>
</tr>
<tr id="9BD97E11413B002DFF37FB73F829FB45" box="[162,2027,1161,1184]" gridrow="19" pageId="12" pageNumber="13">
<th id="D808176D413B002DFF37FB73F829FB45" box="[162,2027,1161,1184]" colspan="19" colspanRight="18" gridcol="0" gridrow="19" pageId="12" pageNumber="13">WMNH-INV-2014-184</th>
</tr>
<tr id="9BD97E11413B002DFF37FB50F829FB24" box="[162,2027,1194,1217]" gridrow="20" pageId="12" pageNumber="13" rowspan-10="1" rowspan-11="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-17="1" rowspan-18="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413B002DFF37FB50FE9BFB24" box="[162,345,1194,1217]" gridcol="0" gridrow="20" pageId="12" pageNumber="13">Tentacle</th>
<td id="D808176D413B002DFE1CFB50FE5DFB24" box="[393,415,1194,1217]" gridcol="1" gridrow="20" pageId="12" pageNumber="13">30</td>
<td id="D808176D413B002DFE28FB50FDD7FB24" box="[445,533,1194,1217]" gridcol="2" gridrow="20" pageId="12" pageNumber="13">108 ± 69</td>
<td id="D808176D413B002DFDA4FB50FDB7FB24" box="[561,629,1194,1217]" gridcol="3" gridrow="20" pageId="12" pageNumber="13">33268</td>
<td id="D808176D413B002DFD31FB50FD72FB24" box="[676,688,1194,1217]" gridcol="4" gridrow="20" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFB11FB50FB59FB24" box="[1156,1179,1194,1217]" gridcol="9" gridrow="20" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFAEBFB50FA56FB24" box="[1406,1428,1194,1217]" gridcol="12" gridrow="20" pageId="12" pageNumber="13">0</td>
</tr>
<tr id="9BD97E11413B002DFF37FB31F829FB06" box="[162,2027,1227,1251]" gridrow="21" pageId="12" pageNumber="13" rowspan-17="1" rowspan-18="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413B002DFF37FB31FE9BFB06" box="[162,345,1227,1251]" gridcol="0" gridrow="21" pageId="12" pageNumber="13">Peri-oral skin</th>
<td id="D808176D413B002DFE1CFB31FE5DFB06" box="[393,415,1227,1251]" gridcol="1" gridrow="21" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFD31FB31FD72FB06" box="[676,688,1227,1251]" gridcol="4" gridrow="21" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFB11FB31FB59FB06" box="[1156,1179,1227,1251]" gridcol="9" gridrow="21" pageId="12" pageNumber="13">27</td>
<td id="D808176D413B002DFB2CFB31FB36FB06" box="[1209,1268,1227,1251]" gridcol="10" gridrow="21" pageId="12" pageNumber="13">38 ±1 0</td>
<td id="D808176D413B002DFA87FB31FA8DFB06" box="[1298,1359,1227,1251]" gridcol="11" gridrow="21" pageId="12" pageNumber="13">2459</td>
<td id="D808176D413B002DFAEBFB31FA56FB06" box="[1406,1428,1227,1251]" gridcol="12" gridrow="21" pageId="12" pageNumber="13">8</td>
<td id="D808176D413B002DFA2CFB31F9C6FB06" box="[1465,1540,1227,1251]" gridcol="13" gridrow="21" pageId="12" pageNumber="13">89 ± 13</td>
<td id="D808176D413B002DF9B6FB31F9AAFB06" box="[1571,1640,1227,1251]" gridcol="14" gridrow="21" pageId="12" pageNumber="13">70104</td>
<td id="D808176D413B002DF905FB31F909FB06" box="[1680,1739,1227,1251]" gridcol="15" gridrow="21" pageId="12" pageNumber="13">15 ± 6</td>
<td id="D808176D413B002DF978FB31F8EDFB06" box="[1773,1839,1227,1251]" gridcol="16" gridrow="21" pageId="12" pageNumber="13">623</td>
</tr>
<tr id="9BD97E11413B002DFF37FB17F829FAE1" box="[162,2027,1261,1284]" gridrow="22" pageId="12" pageNumber="13" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-17="1" rowspan-18="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413B002DFF37FB17FE9BFAE1" box="[162,345,1261,1284]" gridcol="0" gridrow="22" pageId="12" pageNumber="13">Oesophagus skin</th>
<td id="D808176D413B002DFE1CFB17FE5DFAE1" box="[393,415,1261,1284]" gridcol="1" gridrow="22" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFD31FB17FD72FAE1" box="[676,688,1261,1284]" gridcol="4" gridrow="22" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFB11FB17FB59FAE1" box="[1156,1179,1261,1284]" gridcol="9" gridrow="22" pageId="12" pageNumber="13">26</td>
<td id="D808176D413B002DFB2CFB17FB36FAE1" box="[1209,1268,1261,1284]" gridcol="10" gridrow="22" pageId="12" pageNumber="13">36 ±10</td>
<td id="D808176D413B002DFA87FB17FA8DFAE1" box="[1298,1359,1261,1284]" gridcol="11" gridrow="22" pageId="12" pageNumber="13">1457</td>
<td id="D808176D413B002DFAEBFB17FA56FAE1" box="[1406,1428,1261,1284]" gridcol="12" gridrow="22" pageId="12" pageNumber="13">0</td>
</tr>
<tr id="9BD97E11413B002DFF37FAF4F829FAC0" box="[162,2027,1294,1317]" gridrow="23" pageId="12" pageNumber="13" rowspan-10="1" rowspan-11="1" rowspan-17="1" rowspan-18="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413B002DFF37FAF4FE9BFAC0" box="[162,345,1294,1317]" gridcol="0" gridrow="23" pageId="12" pageNumber="13">Introvert skin</th>
<td id="D808176D413B002DFE1CFAF4FE5DFAC0" box="[393,415,1294,1317]" gridcol="1" gridrow="23" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFD31FAF4FD72FAC0" box="[676,688,1294,1317]" gridcol="4" gridrow="23" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFB11FAF4FB59FAC0" box="[1156,1179,1294,1317]" gridcol="9" gridrow="23" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFAEBFAF4FA56FAC0" box="[1406,1428,1294,1317]" gridcol="12" gridrow="23" pageId="12" pageNumber="13">16</td>
<td id="D808176D413B002DFA2CFAF4F9C6FAC0" box="[1465,1540,1294,1317]" gridcol="13" gridrow="23" pageId="12" pageNumber="13">92 ±13</td>
<td id="D808176D413B002DF9B6FAF4F9AAFAC0" box="[1571,1640,1294,1317]" gridcol="14" gridrow="23" pageId="12" pageNumber="13">71125</td>
<td id="D808176D413B002DF905FAF4F909FAC0" box="[1680,1739,1294,1317]" gridcol="15" gridrow="23" pageId="12" pageNumber="13">15 ± 4</td>
<td id="D808176D413B002DF978FAF4F8EDFAC0" box="[1773,1839,1294,1317]" gridcol="16" gridrow="23" pageId="12" pageNumber="13">1022</td>
</tr>
<tr id="9BD97E11413B002DFF37FAD5F829FAA3" box="[162,2027,1327,1350]" gridrow="24" pageId="12" pageNumber="13" rowspan-10="1" rowspan-11="1" rowspan-15="1" rowspan-16="1" rowspan-2="1" rowspan-3="1">
<th id="D808176D413B002DFF37FAD5FE9BFAA3" box="[162,345,1327,1350]" gridcol="0" gridrow="24" pageId="12" pageNumber="13">Pedicels (ventral)</th>
<td id="D808176D413B002DFE1CFAD5FE5DFAA3" box="[393,415,1327,1350]" gridcol="1" gridrow="24" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFD31FAD5FD72FAA3" box="[676,688,1327,1350]" gridcol="4" gridrow="24" pageId="12" pageNumber="13">7</td>
<td id="D808176D413B002DFD46FAD5FCD4FAA3" box="[723,790,1327,1350]" gridcol="5" gridrow="24" pageId="12" pageNumber="13">176 ± 6</td>
<td id="D808176D413B002DFCA9FAD5FCBCFAA3" box="[828,894,1327,1350]" gridcol="6" gridrow="24" pageId="12" pageNumber="13">164183</td>
<td id="D808176D413B002DFC24FAD5FC37FAA3" box="[945,1013,1327,1350]" gridcol="7" gridrow="24" pageId="12" pageNumber="13">119 ± 20</td>
<td id="D808176D413B002DFB8DFAD5FB98FAA3" box="[1048,1114,1327,1350]" gridcol="8" gridrow="24" pageId="12" pageNumber="13">85141</td>
<td id="D808176D413B002DFB11FAD5FB59FAA3" box="[1156,1179,1327,1350]" gridcol="9" gridrow="24" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFAEBFAD5FA56FAA3" box="[1406,1428,1327,1350]" gridcol="12" gridrow="24" pageId="12" pageNumber="13">23</td>
<td id="D808176D413B002DFA2CFAD5F9C6FAA3" box="[1465,1540,1327,1350]" gridcol="13" gridrow="24" pageId="12" pageNumber="13">99 ± 8</td>
<td id="D808176D413B002DF9B6FAD5F9AAFAA3" box="[1571,1640,1327,1350]" gridcol="14" gridrow="24" pageId="12" pageNumber="13">79112</td>
<td id="D808176D413B002DF8CDFAD5F852FAA3" box="[1880,1936,1327,1350]" gridcol="17" gridrow="24" pageId="12" pageNumber="13">32 ± 6</td>
<td id="D808176D413B002DF825FAD5F829FAA3" box="[1968,2027,1327,1350]" gridcol="18" gridrow="24" pageId="12" pageNumber="13">2245</td>
</tr>
<tr id="9BD97E11413B002DFF37FAABF829FA8D" box="[162,2027,1361,1384]" gridrow="25" pageId="12" pageNumber="13" rowspan-10="1" rowspan-11="1" rowspan-17="1" rowspan-18="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413B002DFF37FAABFE9BFA8D" box="[162,345,1361,1384]" gridcol="0" gridrow="25" pageId="12" pageNumber="13">Bodywall (ventral)</th>
<td id="D808176D413B002DFE1CFAABFE5DFA8D" box="[393,415,1361,1384]" gridcol="1" gridrow="25" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFD31FAABFD72FA8D" box="[676,688,1361,1384]" gridcol="4" gridrow="25" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFB11FAABFB59FA8D" box="[1156,1179,1361,1384]" gridcol="9" gridrow="25" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFAEBFAABFA56FA8D" box="[1406,1428,1361,1384]" gridcol="12" gridrow="25" pageId="12" pageNumber="13">29</td>
<td id="D808176D413B002DFA2CFAABF9C6FA8D" box="[1465,1540,1361,1384]" gridcol="13" gridrow="25" pageId="12" pageNumber="13">112 ± 20</td>
<td id="D808176D413B002DF9B6FAABF9AAFA8D" box="[1571,1640,1361,1384]" gridcol="14" gridrow="25" pageId="12" pageNumber="13">81161</td>
<td id="D808176D413B002DF905FAABF909FA8D" box="[1680,1739,1361,1384]" gridcol="15" gridrow="25" pageId="12" pageNumber="13">9 ± 6</td>
<td id="D808176D413B002DF978FAABF8EDFA8D" box="[1773,1839,1361,1384]" gridcol="16" gridrow="25" pageId="12" pageNumber="13">431</td>
</tr>
<tr id="9BD97E11413B002DFF37FA88F829FA6C" box="[162,2027,1394,1417]" gridrow="26" pageId="12" pageNumber="13" rowspan-10="1" rowspan-11="1" rowspan-17="1" rowspan-18="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413B002DFF37FA88FE9BFA6C" box="[162,345,1394,1417]" gridcol="0" gridrow="26" pageId="12" pageNumber="13">Anal papilla</th>
<td id="D808176D413B002DFE1CFA88FE5DFA6C" box="[393,415,1394,1417]" gridcol="1" gridrow="26" pageId="12" pageNumber="13">19</td>
<td id="D808176D413B002DFE28FA88FDD7FA6C" box="[445,533,1394,1417]" gridcol="2" gridrow="26" pageId="12" pageNumber="13">71 ± 15</td>
<td id="D808176D413B002DFDA4FA88FDB7FA6C" box="[561,629,1394,1417]" gridcol="3" gridrow="26" pageId="12" pageNumber="13">51107</td>
<td id="D808176D413B002DFD31FA88FD72FA6C" box="[676,688,1394,1417]" gridcol="4" gridrow="26" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFB11FA88FB59FA6C" box="[1156,1179,1394,1417]" gridcol="9" gridrow="26" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFAEBFA88FA56FA6C" box="[1406,1428,1394,1417]" gridcol="12" gridrow="26" pageId="12" pageNumber="13">7</td>
<td id="D808176D413B002DFA2CFA88F9C6FA6C" box="[1465,1540,1394,1417]" gridcol="13" gridrow="26" pageId="12" pageNumber="13">76 ± 6</td>
<td id="D808176D413B002DF9B6FA88F9AAFA6C" box="[1571,1640,1394,1417]" gridcol="14" gridrow="26" pageId="12" pageNumber="13">6886</td>
<td id="D808176D413B002DF905FA88F909FA6C" box="[1680,1739,1394,1417]" gridcol="15" gridrow="26" pageId="12" pageNumber="13">6 ± 2</td>
<td id="D808176D413B002DF978FA88F8EDFA6C" box="[1773,1839,1394,1417]" gridcol="16" gridrow="26" pageId="12" pageNumber="13">48</td>
</tr>
<tr id="9BD97E11413B002DFF37FA69F829FA4F" box="[162,2027,1427,1450]" gridrow="27" pageId="12" pageNumber="13">
<th id="D808176D413B002DFF37FA69F829FA4F" box="[162,2027,1427,1450]" colspan="19" colspanRight="18" gridcol="0" gridrow="27" pageId="12" pageNumber="13">WMNH-INV-2014-188</th>
</tr>
<tr id="9BD97E11413B002DFF37FA4EF829FA2E" box="[162,2027,1460,1483]" gridrow="28" pageId="12" pageNumber="13" rowspan-10="1" rowspan-11="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-17="1" rowspan-18="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413B002DFF37FA4EFE9BFA2E" box="[162,345,1460,1483]" gridcol="0" gridrow="28" pageId="12" pageNumber="13">Tentacle</th>
<td id="D808176D413B002DFE1CFA4EFE5DFA2E" box="[393,415,1460,1483]" gridcol="1" gridrow="28" pageId="12" pageNumber="13">30</td>
<td id="D808176D413B002DFE28FA4EFDD7FA2E" box="[445,533,1460,1483]" gridcol="2" gridrow="28" pageId="12" pageNumber="13">111 ± 76</td>
<td id="D808176D413B002DFDA4FA4EFDB7FA2E" box="[561,629,1460,1483]" gridcol="3" gridrow="28" pageId="12" pageNumber="13">35321</td>
<td id="D808176D413B002DFD31FA4EFD72FA2E" box="[676,688,1460,1483]" gridcol="4" gridrow="28" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFB11FA4EFB59FA2E" box="[1156,1179,1460,1483]" gridcol="9" gridrow="28" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFAEBFA4EFA56FA2E" box="[1406,1428,1460,1483]" gridcol="12" gridrow="28" pageId="12" pageNumber="13">0</td>
</tr>
<tr id="9BD97E11413B002DFF37FA2CF829FA08" box="[162,2027,1494,1517]" gridrow="29" pageId="12" pageNumber="13" rowspan-17="1" rowspan-18="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413B002DFF37FA2CFE9BFA08" box="[162,345,1494,1517]" gridcol="0" gridrow="29" pageId="12" pageNumber="13">Peri-oral skin</th>
<td id="D808176D413B002DFE1CFA2CFE5DFA08" box="[393,415,1494,1517]" gridcol="1" gridrow="29" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFD31FA2CFD72FA08" box="[676,688,1494,1517]" gridcol="4" gridrow="29" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFB11FA2CFB59FA08" box="[1156,1179,1494,1517]" gridcol="9" gridrow="29" pageId="12" pageNumber="13">29</td>
<td id="D808176D413B002DFB2CFA2CFB36FA08" box="[1209,1268,1494,1517]" gridcol="10" gridrow="29" pageId="12" pageNumber="13">38 ± 11</td>
<td id="D808176D413B002DFA87FA2CFA8DFA08" box="[1298,1359,1494,1517]" gridcol="11" gridrow="29" pageId="12" pageNumber="13">2574</td>
<td id="D808176D413B002DFAEBFA2CFA56FA08" box="[1406,1428,1494,1517]" gridcol="12" gridrow="29" pageId="12" pageNumber="13">4</td>
<td id="D808176D413B002DFA2CFA2CF9C6FA08" box="[1465,1540,1494,1517]" gridcol="13" gridrow="29" pageId="12" pageNumber="13">86 ± 25</td>
<td id="D808176D413B002DF9B6FA2CF9AAFA08" box="[1571,1640,1494,1517]" gridcol="14" gridrow="29" pageId="12" pageNumber="13">65117</td>
<td id="D808176D413B002DF905FA2CF909FA08" box="[1680,1739,1494,1517]" gridcol="15" gridrow="29" pageId="12" pageNumber="13">21 ±1 0</td>
<td id="D808176D413B002DF978FA2CF8EDFA08" box="[1773,1839,1494,1517]" gridcol="16" gridrow="29" pageId="12" pageNumber="13">933</td>
</tr>
<tr id="9BD97E11413B002DFF37FA0DF829F9EB" box="[162,2027,1527,1550]" gridrow="30" pageId="12" pageNumber="13" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-17="1" rowspan-18="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413B002DFF37FA0DFE9BF9EB" box="[162,345,1527,1550]" gridcol="0" gridrow="30" pageId="12" pageNumber="13">Oesophagus skin</th>
<td id="D808176D413B002DFE1CFA0DFE5DF9EB" box="[393,415,1527,1550]" gridcol="1" gridrow="30" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFD31FA0DFD72F9EB" box="[676,688,1527,1550]" gridcol="4" gridrow="30" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFB11FA0DFB59F9EB" box="[1156,1179,1527,1550]" gridcol="9" gridrow="30" pageId="12" pageNumber="13">25</td>
<td id="D808176D413B002DFB2CFA0DFB36F9EB" box="[1209,1268,1527,1550]" gridcol="10" gridrow="30" pageId="12" pageNumber="13">34 ± 5</td>
<td id="D808176D413B002DFA87FA0DFA8DF9EB" box="[1298,1359,1527,1550]" gridcol="11" gridrow="30" pageId="12" pageNumber="13">2443</td>
<td id="D808176D413B002DFAEBFA0DFA56F9EB" box="[1406,1428,1527,1550]" gridcol="12" gridrow="30" pageId="12" pageNumber="13">0</td>
</tr>
<tr id="9BD97E11413B002DFF37F9E2F829F9CA" box="[162,2027,1560,1583]" gridrow="31" pageId="12" pageNumber="13" rowspan-10="1" rowspan-11="1" rowspan-17="1" rowspan-18="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413B002DFF37F9E2FE9BF9CA" box="[162,345,1560,1583]" gridcol="0" gridrow="31" pageId="12" pageNumber="13">Introvert skin</th>
<td id="D808176D413B002DFE1CF9E2FE5DF9CA" box="[393,415,1560,1583]" gridcol="1" gridrow="31" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFD31F9E2FD72F9CA" box="[676,688,1560,1583]" gridcol="4" gridrow="31" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFB11F9E2FB59F9CA" box="[1156,1179,1560,1583]" gridcol="9" gridrow="31" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFAEBF9E2FA56F9CA" box="[1406,1428,1560,1583]" gridcol="12" gridrow="31" pageId="12" pageNumber="13">24</td>
<td id="D808176D413B002DFA2CF9E2F9C6F9CA" box="[1465,1540,1560,1583]" gridcol="13" gridrow="31" pageId="12" pageNumber="13">85 ± 14</td>
<td id="D808176D413B002DF9B6F9E2F9AAF9CA" box="[1571,1640,1560,1583]" gridcol="14" gridrow="31" pageId="12" pageNumber="13">56113</td>
<td id="D808176D413B002DF905F9E2F909F9CA" box="[1680,1739,1560,1583]" gridcol="15" gridrow="31" pageId="12" pageNumber="13">19 ± 8</td>
<td id="D808176D413B002DF978F9E2F8EDF9CA" box="[1773,1839,1560,1583]" gridcol="16" gridrow="31" pageId="12" pageNumber="13">1035</td>
</tr>
<tr id="9BD97E11413B002DFF37F9C3F829F9B4" box="[162,2027,1593,1617]" gridrow="32" pageId="12" pageNumber="13" rowspan-10="1" rowspan-11="1" rowspan-15="1" rowspan-16="1" rowspan-2="1" rowspan-3="1">
<th id="D808176D413B002DFF37F9C3FE9BF9B4" box="[162,345,1593,1617]" gridcol="0" gridrow="32" pageId="12" pageNumber="13">Pedicels (ventral)</th>
<td id="D808176D413B002DFE1CF9C3FE5DF9B4" box="[393,415,1593,1617]" gridcol="1" gridrow="32" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFD31F9C3FD72F9B4" box="[676,688,1593,1617]" gridcol="4" gridrow="32" pageId="12" pageNumber="13">3</td>
<td id="D808176D413B002DFD46F9C3FCD4F9B4" box="[723,790,1593,1617]" gridcol="5" gridrow="32" pageId="12" pageNumber="13">170 ± 12</td>
<td id="D808176D413B002DFCA9F9C3FCBCF9B4" box="[828,894,1593,1617]" gridcol="6" gridrow="32" pageId="12" pageNumber="13">160184</td>
<td id="D808176D413B002DFC24F9C3FC37F9B4" box="[945,1013,1593,1617]" gridcol="7" gridrow="32" pageId="12" pageNumber="13">143 ± 42</td>
<td id="D808176D413B002DFB8DF9C3FB98F9B4" box="[1048,1114,1593,1617]" gridcol="8" gridrow="32" pageId="12" pageNumber="13">96176</td>
<td id="D808176D413B002DFB11F9C3FB59F9B4" box="[1156,1179,1593,1617]" gridcol="9" gridrow="32" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFAEBF9C3FA56F9B4" box="[1406,1428,1593,1617]" gridcol="12" gridrow="32" pageId="12" pageNumber="13">15</td>
<td id="D808176D413B002DFA2CF9C3F9C6F9B4" box="[1465,1540,1593,1617]" gridcol="13" gridrow="32" pageId="12" pageNumber="13">91 ± 8</td>
<td id="D808176D413B002DF9B6F9C3F9AAF9B4" box="[1571,1640,1593,1617]" gridcol="14" gridrow="32" pageId="12" pageNumber="13">78106</td>
<td id="D808176D413B002DF8CDF9C3F852F9B4" box="[1880,1936,1593,1617]" gridcol="17" gridrow="32" pageId="12" pageNumber="13">38 ± 7</td>
<td id="D808176D413B002DF825F9C3F829F9B4" box="[1968,2027,1593,1617]" gridcol="18" gridrow="32" pageId="12" pageNumber="13">2651</td>
</tr>
<tr id="9BD97E11413B002DFF37F9A1F829F997" box="[162,2027,1627,1650]" gridrow="33" pageId="12" pageNumber="13" rowspan-10="1" rowspan-11="1" rowspan-17="1" rowspan-18="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="D808176D413B002DFF37F9A1FE9BF997" box="[162,345,1627,1650]" gridcol="0" gridrow="33" pageId="12" pageNumber="13">Bodywall (ventral)</th>
<td id="D808176D413B002DFE1CF9A1FE5DF997" box="[393,415,1627,1650]" gridcol="1" gridrow="33" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFD31F9A1FD72F997" box="[676,688,1627,1650]" gridcol="4" gridrow="33" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFB11F9A1FB59F997" box="[1156,1179,1627,1650]" gridcol="9" gridrow="33" pageId="12" pageNumber="13">0</td>
<td id="D808176D413B002DFAEBF9A1FA56F997" box="[1406,1428,1627,1650]" gridcol="12" gridrow="33" pageId="12" pageNumber="13">30</td>
<td id="D808176D413B002DFA2CF9A1F9C6F997" box="[1465,1540,1627,1650]" gridcol="13" gridrow="33" pageId="12" pageNumber="13">107 ± 12</td>
<td id="D808176D413B002DF9B6F9A1F9AAF997" box="[1571,1640,1627,1650]" gridcol="14" gridrow="33" pageId="12" pageNumber="13">88135</td>
<td id="D808176D413B002DF905F9A1F909F997" box="[1680,1739,1627,1650]" gridcol="15" gridrow="33" pageId="12" pageNumber="13">10 ± 2</td>
<td id="D808176D413B002DF978F9A1F8EDF997" box="[1773,1839,1627,1650]" gridcol="16" gridrow="33" pageId="12" pageNumber="13">717</td>
</tr>
<tr id="9BD97E11413B002DFF37F986F829F976" box="[162,2027,1660,1683]" gridrow="34" pageId="12" pageNumber="13" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-17="1" rowspan-18="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="D808176D413B002DFF37F986FE9BF976" box="[162,345,1660,1683]" gridcol="0" gridrow="34" pageId="12" pageNumber="13">Anal papilla</th>
<td id="D808176D413B002DFE28F986FDB7F976" box="[445,629,1660,1683]" colspan="2" colspanRight="1" gridcol="2" gridrow="34" pageId="12" pageNumber="13">Amputated</td>
</tr>
</table>
</paragraph>
<paragraph id="25567C53413BFFDEFF36F966FB81F950" blockId="12.[163,1091,1692,1717]" box="[163,1091,1692,1717]" pageId="12" pageNumber="13">
<tableNote id="D80F7DDD413BFFDEFF36F966FB81F950" box="[163,1091,1692,1717]" pageId="12" pageNumber="13" targetBox="[162,2027,525,1683]" targetPageId="12">
<superScript id="D29CD11B413BFFDEFF36F966FF6BF94F" attach="left" box="[163,169,1692,1706]" fontSize="5" pageId="12" pageNumber="13">a</superScript>
WMNH-INV: Invertebrate collection of the Wakayama Prefectural Museum of Natural History.
</tableNote>
</paragraph>
<subSubSection id="6DF32FD8413AFFDFFF52FF6CFD1FFA3E" pageId="13" pageNumber="14" type="description">
<paragraph id="25567C53413AFFDFFF52FF6CFBDEFE2A" blockId="13.[151,1437,150,2036]" pageId="13" pageNumber="14">
<emphasis id="179DA041413AFFDFFF52FF6CFE9BFF55" bold="true" box="[199,345,150,176]" pageId="13" pageNumber="14">Description.</emphasis>
Body medium, up to
<quantity id="E211D1B6413AFFDFFDC0FF6DFD64FF54" box="[597,678,151,177]" metricMagnitude="-2" metricUnit="m" metricValue="3.9" pageId="13" pageNumber="14" unit="mm" value="39.0">39 mm</quantity>
long, fusiform, curved, with both ends tapered and turned slightly upwards (
<figureCitation id="BDD260D6413AFFDFFE99FF41FEA9FF30" box="[268,363,187,213]" captionStart="FIGURE 2" captionStartId="6.[151,250,1766,1790]" captionTargetBox="[155,1435,419,1743]" captionTargetId="figure-141@6.[151,1436,417,1743]" captionTargetPageId="6" captionText="FIGURE 2. Lateral views of the specimens, dorsal upper, anterior left. A, Massinium toyoshiomaruae sp. nov., (a) holotype, WMNH-INV-2014-186, (b) paratype, WMNH-INV-2015-2497, (c) paratype, WMNH-INV-2015-2503, (d) paratype, NSMT- E-12099-1; B, Thyone liaoi sp. nov., (e) holotype, WMNH-INV-2014-182, (f) paratype, WMNH-INV-2014-183, (g) paratype, WMNH-INV-2014-184, (h) paratype, WMNH-INV-2014-188; C, Thyone kyushuensis sp. nov., (i) holotype, WMNH-INV- 2015-2437, (j) paratype, NSMT-E-12103; D, T. toyoshiomaruae sp. nov., (k) paratype, WMNH-INV-2015-2438, (l) paratype, WMNH-INV-2015-2439, (m) holotype, WMNH-INV-2015-2483, (n) paratype, WMNH-INV-2015-2484." figureDoi="http://doi.org/10.5281/zenodo.5225376" httpUri="https://zenodo.org/record/5225376/files/figure.png" pageId="13" pageNumber="14">Fig. 2B</figureCitation>
); bodywall thin, soft. Color fading upon preservation. Living body color whitish orange. Tentacles 10, arranged in a single circle, two medioventral tentacles smaller than the other eight. Tentacles and introvert transparent in most parts in living animals, and with some dark brown pigmentation. Several rows of villi surrounding oral opening. Oral disk and pharynx of the preserved specimens dark grayish brown and dark purplish brown, respectively. Pedicels long, non-retractile, covering entire body at constant density, lacking in introvert region. Color of pedicels same and slightly deeper than adjacent body. Pedicels becoming gradually smaller toward anterior and posterior from middle portion of body. Ten anal papillae and five anal teeth in radii. After preservation, color denoted above turned more yellowish and whitish than that of living color.
</paragraph>
<paragraph id="25567C53413AFFDFFF52FE23FB42FD2C" blockId="13.[151,1437,150,2036]" pageId="13" pageNumber="14">
Calcareous ring long, cracking of mosaic patterns mostly caused by the damage of material; main radial plates and interradial plates not fragmented in original condition in the several well preserved specimens (
<figureCitation id="BDD260D6413AFFDFFB7BFE06FAFDFDF2" box="[1262,1343,508,535]" captionStart="FIGURE 3" captionStartId="7.[151,250,1500,1524]" captionTargetBox="[152,1426,183,1471]" captionTargetId="figure-18@7.[151,1436,181,1477]" captionTargetPageId="7" captionText="FIGURE 3. Elements of calcareous rings, viewed from the body cavity, anterior upper.A, Massinium toyoshiomaruae sp. nov., (holotype, WMNH-INV-2014-186); B, Thyone liaoi sp. nov., (holotype, WMNH-INV-2014-182, and a paratype, WMNH-INV- 2014-184); C, T. kyushuensis sp. nov., (holotype, WMNH-INV-2015-2437); D, T. toyoshiomaruae sp. nov. (holotype, WMNH- INV-2015-2483). Abbreviations: md, mid-dorsal part; mv, mid-ventral part; -ir, interradial element; -r radial element." figureDoi="http://doi.org/10.5281/zenodo.5225378" httpUri="https://zenodo.org/record/5225378/files/figure.png" pageId="13" pageNumber="14">Fig 3B</figureCitation>
). Radial elements elongated, with paired, posterior, weakly fragmented prolongations; each with an anterior projection and notch. Interradial elements also long but without posterior prolongations with slight anterior projections; shortest in dorso-lateral elements. Radial plates and interradial plates loosely adherent. One (or rarely two) Polian vesicles in medioventral, single stone canal in mediodorsal position. Well-matured gonad situated in mid-body, in two clumps, one on each side of dorsal mesentery, most tubules not branched. Gonad lacking ossicles.
</paragraph>
<paragraph id="25567C53413AFFDFFF52FD29FACFFCD1" blockId="13.[151,1437,150,2036]" pageId="13" pageNumber="14">
Bodywall ossicles comprising abundant tables (
<figureCitation id="BDD260D6413AFFDFFD4DFD29FC8EFD08" box="[728,844,723,750]" captionStart="FIGURE 5" captionStartId="11.[151,250,1888,1912]" captionTargetBox="[182,1404,182,1863]" captionTargetId="figure-18@11.[181,1406,181,1864]" captionTargetPageId="11" captionText="FIGURE 5. Scanning electron microscope images of ossicles of Thyone liaoi sp. nov., (holotype, AH, WMNH-INV-2014- 182). A, ossicles of the ventrolateral tentacle; B, the ventrolateral skin of peri-oral; C, the ventrolateral skin of pharynx; D, the ventrolateral introvert skin; E, the pedicel of the abdominal side; F and G, the middle part of the bodywall on the abdominal side, and on the dorsal side, respectively; H, the ventrolateral anal papilla." figureDoi="http://doi.org/10.5281/zenodo.5225382" httpUri="https://zenodo.org/record/5225382/files/figure.png" pageId="13" pageNumber="14">Fig. 5F, G</figureCitation>
,
<tableCitation id="686B49E8413AFFDFFCC0FD29FC6BFD08" box="[853,937,723,749]" captionStart="TABLE 4" captionStartId="12.[151,239,485,511]" captionText="TABLE 4. Counts and measurements (Mean ± sd μm) of ossicles from four specimens of Thyone liaoi sp. nov." pageId="13" pageNumber="14">Table 4</tableCitation>
), with distorted, oval or rhomboidal disc and delicate spire, mostly with two pillars fusing into one distally, ending in 46 minute apical spines. Table disc with 816 perforations, most frequently 810; central two perforations large, peripheral perforations small.
</paragraph>
<paragraph id="25567C53413AFFDFFF52FCC4FF0BFAE0" blockId="13.[151,1437,150,2036]" pageId="13" pageNumber="14">
Tentacle ossicles comprising mostly plates of varying size and shape in trunk and branches, also table ossicles in basal parts (
<figureCitation id="BDD260D6413AFFDFFEABFC98FE5BFC98" box="[318,409,866,893]" captionStart="FIGURE 5" captionStartId="11.[151,250,1888,1912]" captionTargetBox="[182,1404,182,1863]" captionTargetId="figure-18@11.[181,1406,181,1864]" captionTargetPageId="11" captionText="FIGURE 5. Scanning electron microscope images of ossicles of Thyone liaoi sp. nov., (holotype, AH, WMNH-INV-2014- 182). A, ossicles of the ventrolateral tentacle; B, the ventrolateral skin of peri-oral; C, the ventrolateral skin of pharynx; D, the ventrolateral introvert skin; E, the pedicel of the abdominal side; F and G, the middle part of the bodywall on the abdominal side, and on the dorsal side, respectively; H, the ventrolateral anal papilla." figureDoi="http://doi.org/10.5281/zenodo.5225382" httpUri="https://zenodo.org/record/5225382/files/figure.png" pageId="13" pageNumber="14">Fig. 5A</figureCitation>
,
<tableCitation id="686B49E8413AFFDFFE30FC98FE3EFC99" box="[421,508,866,892]" captionStart="TABLE 4" captionStartId="12.[151,239,485,511]" captionText="TABLE 4. Counts and measurements (Mean ± sd μm) of ossicles from four specimens of Thyone liaoi sp. nov." pageId="13" pageNumber="14">Table 4</tableCitation>
). Table ossicles with distorted oval or rhomboidal disc, and high spires with twopillars with mostly 23 crossbeams, and 48 apical spines. Some table ossicles have oval discs, possessing two larges central perforations and small distal perforations. Plates elongated, with two, large, central perforations and small distal perforations, occasionally developing a rudimentary spire and/or extra layer in central part. Peri-oral skin ossicles comprising mostly rosettes, with tables in part adjoining tentacle bases (
<figureCitation id="BDD260D6413AFFDFFBE5FC0BFB02FBE9" box="[1136,1216,1009,1036]" captionStart="FIGURE 5" captionStartId="11.[151,250,1888,1912]" captionTargetBox="[182,1404,182,1863]" captionTargetId="figure-18@11.[181,1406,181,1864]" captionTargetPageId="11" captionText="FIGURE 5. Scanning electron microscope images of ossicles of Thyone liaoi sp. nov., (holotype, AH, WMNH-INV-2014- 182). A, ossicles of the ventrolateral tentacle; B, the ventrolateral skin of peri-oral; C, the ventrolateral skin of pharynx; D, the ventrolateral introvert skin; E, the pedicel of the abdominal side; F and G, the middle part of the bodywall on the abdominal side, and on the dorsal side, respectively; H, the ventrolateral anal papilla." figureDoi="http://doi.org/10.5281/zenodo.5225382" httpUri="https://zenodo.org/record/5225382/files/figure.png" pageId="13" pageNumber="14">Fig.5B</figureCitation>
,
<tableCitation id="686B49E8413AFFDFFB5BFC0BFAE5FBEE" box="[1230,1319,1009,1035]" captionStart="TABLE 4" captionStartId="12.[151,239,485,511]" captionText="TABLE 4. Counts and measurements (Mean ± sd μm) of ossicles from four specimens of Thyone liaoi sp. nov." pageId="13" pageNumber="14">Table 4</tableCitation>
). Rosettes have granulated, mulberry-like, densely branched in more than one plane branches. Tables with distorted oval or rhomboidal discs, high spires with two-pillars, mostly with 13 crossbeams and 46 apical spines. Occasionally plates also exist, some with rudimentary spires and/or extra layer in central part. Pharyngeal villi have rosettes with granulated, mulberry-like (
<figureCitation id="BDD260D6413AFFDFFE5CFB7AFDE1FB7F" box="[457,547,1151,1178]" captionStart="FIGURE 5" captionStartId="11.[151,250,1888,1912]" captionTargetBox="[182,1404,182,1863]" captionTargetId="figure-18@11.[181,1406,181,1864]" captionTargetPageId="11" captionText="FIGURE 5. Scanning electron microscope images of ossicles of Thyone liaoi sp. nov., (holotype, AH, WMNH-INV-2014- 182). A, ossicles of the ventrolateral tentacle; B, the ventrolateral skin of peri-oral; C, the ventrolateral skin of pharynx; D, the ventrolateral introvert skin; E, the pedicel of the abdominal side; F and G, the middle part of the bodywall on the abdominal side, and on the dorsal side, respectively; H, the ventrolateral anal papilla." figureDoi="http://doi.org/10.5281/zenodo.5225382" httpUri="https://zenodo.org/record/5225382/files/figure.png" pageId="13" pageNumber="14">Fig. 5C</figureCitation>
,
<tableCitation id="686B49E8413AFFDFFDA4FB7AFD4BFB7F" box="[561,649,1152,1178]" captionStart="TABLE 4" captionStartId="12.[151,239,485,511]" captionText="TABLE 4. Counts and measurements (Mean ± sd μm) of ossicles from four specimens of Thyone liaoi sp. nov." pageId="13" pageNumber="14">Table 4</tableCitation>
). Introvert with tables and mulberry-like rosettes (
<figureCitation id="BDD260D6413AFFDFFB5BFB7AFAEBFB7F" box="[1230,1321,1151,1178]" captionStart="FIGURE 5" captionStartId="11.[151,250,1888,1912]" captionTargetBox="[182,1404,182,1863]" captionTargetId="figure-18@11.[181,1406,181,1864]" captionTargetPageId="11" captionText="FIGURE 5. Scanning electron microscope images of ossicles of Thyone liaoi sp. nov., (holotype, AH, WMNH-INV-2014- 182). A, ossicles of the ventrolateral tentacle; B, the ventrolateral skin of peri-oral; C, the ventrolateral skin of pharynx; D, the ventrolateral introvert skin; E, the pedicel of the abdominal side; F and G, the middle part of the bodywall on the abdominal side, and on the dorsal side, respectively; H, the ventrolateral anal papilla." figureDoi="http://doi.org/10.5281/zenodo.5225382" httpUri="https://zenodo.org/record/5225382/files/figure.png" pageId="13" pageNumber="14">Fig. 5D</figureCitation>
,
<tableCitation id="686B49E8413AFFDFFAA2FB7AFA4DFB7F" box="[1335,1423,1152,1178]" captionStart="TABLE 4" captionStartId="12.[151,239,485,511]" captionText="TABLE 4. Counts and measurements (Mean ± sd μm) of ossicles from four specimens of Thyone liaoi sp. nov." pageId="13" pageNumber="14">Table 4</tableCitation>
). Table ossicles with distorted, oval or rhomboidal disc, and low spires of two-pillars, with mostly 12 crossbeams, and 48 minute apical spines, occasionally with an extra layer in central part. Rosettes with granulated, mulberrylike.
</paragraph>
<paragraph id="25567C53413AFFDFFF52FAF5FD1FFA3E" blockId="13.[151,1437,150,2036]" pageId="13" pageNumber="14">
Most pedicels with endplate and supporting tables (
<figureCitation id="BDD260D6413AFFDFFC80FAF5FCAFFACC" box="[789,877,1294,1321]" captionStart="FIGURE 5" captionStartId="11.[151,250,1888,1912]" captionTargetBox="[182,1404,182,1863]" captionTargetId="figure-18@11.[181,1406,181,1864]" captionTargetPageId="11" captionText="FIGURE 5. Scanning electron microscope images of ossicles of Thyone liaoi sp. nov., (holotype, AH, WMNH-INV-2014- 182). A, ossicles of the ventrolateral tentacle; B, the ventrolateral skin of peri-oral; C, the ventrolateral skin of pharynx; D, the ventrolateral introvert skin; E, the pedicel of the abdominal side; F and G, the middle part of the bodywall on the abdominal side, and on the dorsal side, respectively; H, the ventrolateral anal papilla." figureDoi="http://doi.org/10.5281/zenodo.5225382" httpUri="https://zenodo.org/record/5225382/files/figure.png" pageId="13" pageNumber="14">Fig. 5E</figureCitation>
,
<tableCitation id="686B49E8413AFFDFFCEFFAF5FC10FACC" box="[890,978,1295,1321]" captionStart="TABLE 4" captionStartId="12.[151,239,485,511]" captionText="TABLE 4. Counts and measurements (Mean ± sd μm) of ossicles from four specimens of Thyone liaoi sp. nov." pageId="13" pageNumber="14">Table 4</tableCitation>
). Supporting tables with narrow, curved disc, with 24 large, central perforations and 28 small, distal perforations; spire high, with two (occasionally three) pillars united distally, ending in 24 minute processes (
<figureCitation id="BDD260D6413AFFDFFC84FAACFCABFA94" box="[785,873,1366,1393]" captionStart="FIGURE 5" captionStartId="11.[151,250,1888,1912]" captionTargetBox="[182,1404,182,1863]" captionTargetId="figure-18@11.[181,1406,181,1864]" captionTargetPageId="11" captionText="FIGURE 5. Scanning electron microscope images of ossicles of Thyone liaoi sp. nov., (holotype, AH, WMNH-INV-2014- 182). A, ossicles of the ventrolateral tentacle; B, the ventrolateral skin of peri-oral; C, the ventrolateral skin of pharynx; D, the ventrolateral introvert skin; E, the pedicel of the abdominal side; F and G, the middle part of the bodywall on the abdominal side, and on the dorsal side, respectively; H, the ventrolateral anal papilla." figureDoi="http://doi.org/10.5281/zenodo.5225382" httpUri="https://zenodo.org/record/5225382/files/figure.png" pageId="13" pageNumber="14">Fig. 5E</figureCitation>
,
<tableCitation id="686B49E8413AFFDFFCE3FAACFC0CFA95" box="[886,974,1366,1392]" captionStart="TABLE 4" captionStartId="12.[151,239,485,511]" captionText="TABLE 4. Counts and measurements (Mean ± sd μm) of ossicles from four specimens of Thyone liaoi sp. nov." pageId="13" pageNumber="14">Table 4</tableCitation>
). Endplates with holes arranged in three concentric areas,
<emphasis id="179DA041413AFFDFFECEFA81FE43FA71" box="[347,385,1403,1428]" italics="true" pageId="13" pageNumber="14">viz.</emphasis>
, commarginally-elongated holes near rim, radially-elongated holes in mid-region, and almost circular holes in central area. Anal papillae with small, irregular supporting plates and tables (
<figureCitation id="BDD260D6413AFFDFFB0AFA67FB35FA5D" box="[1183,1271,1437,1464]" captionStart="FIGURE 5" captionStartId="11.[151,250,1888,1912]" captionTargetBox="[182,1404,182,1863]" captionTargetId="figure-18@11.[181,1406,181,1864]" captionTargetPageId="11" captionText="FIGURE 5. Scanning electron microscope images of ossicles of Thyone liaoi sp. nov., (holotype, AH, WMNH-INV-2014- 182). A, ossicles of the ventrolateral tentacle; B, the ventrolateral skin of peri-oral; C, the ventrolateral skin of pharynx; D, the ventrolateral introvert skin; E, the pedicel of the abdominal side; F and G, the middle part of the bodywall on the abdominal side, and on the dorsal side, respectively; H, the ventrolateral anal papilla." figureDoi="http://doi.org/10.5281/zenodo.5225382" httpUri="https://zenodo.org/record/5225382/files/figure.png" pageId="13" pageNumber="14">Fig. 5H</figureCitation>
,
<tableCitation id="686B49E8413AFFDFFA94FA67FA94FA5D" box="[1281,1366,1437,1464]" captionStart="TABLE 4" captionStartId="12.[151,239,485,511]" captionText="TABLE 4. Counts and measurements (Mean ± sd μm) of ossicles from four specimens of Thyone liaoi sp. nov." pageId="13" pageNumber="14">Table 4</tableCitation>
). Skin around anus with relatively small and simple tables.
</paragraph>
</subSubSection>
<subSubSection id="6DF32FD8413AFFDCFF52FA1FFBC4FF1D" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="discussion">
<paragraph id="25567C53413AFFDCFF52FA1FFBC4FF1D" blockId="13.[151,1437,150,2036]" lastBlockId="14.[151,1437,150,535]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
<emphasis id="179DA041413AFFDFFF52FA1FFEF9FA1A" bold="true" box="[199,315,1509,1535]" pageId="13" pageNumber="14">Remarks.</emphasis>
<taxonomicName id="E2E907D0413AFFDFFEDFFA1FFE1DFA1A" authority="Yamana &amp; Nakaguchi &amp; Yamaguchi &amp; Katoh &amp; Ogawa &amp; Ohtsuka, 2021" authorityName="Yamana &amp; Nakaguchi &amp; Yamaguchi &amp; Katoh &amp; Ogawa &amp; Ohtsuka" authorityYear="2021" box="[330,479,1509,1535]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="13" pageNumber="14" phylum="Echinodermata" rank="species" species="liaoi" status="sp. nov.">
<emphasis id="179DA041413AFFDFFEDFFA1FFE1DFA1A" box="[330,479,1509,1535]" italics="true" pageId="13" pageNumber="14">Thyone liaoi</emphasis>
</taxonomicName>
<emphasis id="179DA041413AFFDFFE7DFA1FFD85FA1A" bold="true" box="[488,583,1509,1535]" pageId="13" pageNumber="14">
<taxonomicNameLabel id="0CAE1D3A413AFFDFFE7DFA1FFD85FA1A" box="[488,583,1509,1535]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was previously reported as a regional form of “
<taxonomicName id="E2E907D0413AFFDFFB10FA1FFF15F9C6" authority="Semper, 1867" authorityName="Semper" authorityYear="1867" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="13" pageNumber="14" phylum="Echinodermata" rank="species" species="pedata">
<emphasis id="179DA041413AFFDFFB10FA1FFAF6FA1A" box="[1157,1332,1509,1535]" italics="true" pageId="13" pageNumber="14">Thyone pedata</emphasis>
<bibRefCitation id="417801A2413AFFDFFAA8FA1FFF15F9C6" author="Semper, C." pageId="13" pageNumber="14" refId="ref27545" refString="Semper, C. (1867 - 1868) Holothurien. Reisen im Archipel der Philippinen. Teil 2. Wissenschaftliche Resultate. C. W. Kreidel, Wiesbaden, 288 pp. https: // doi. org / 10.5962 / bhl. title. 11687" type="book" year="1867">Semper, 1867</bibRefCitation>
</taxonomicName>
,” from the Chinese and Japanese waters (
<emphasis id="179DA041413AFFDFFD55F9F0FD28F9C6" box="[704,746,1546,1571]" italics="true" pageId="13" pageNumber="14">e.g.</emphasis>
,
<bibRefCitation id="417801A2413AFFDFFD6FF9F3FC19F9C1" author="Liao, Y. &amp; Clark, A. M." box="[762,987,1545,1572]" pageId="13" pageNumber="14" refId="ref26030" refString="Liao, Y. &amp; Clark, A. M. (1995) The Echinoderms of Southern China. Science Press Beijing, 614 pp." type="book" year="1995">Liao &amp; Clark 1995</bibRefCitation>
,
<bibRefCitation id="417801A2413AFFDFFC73F9F3FB4EF9C1" author="Yamana, Y." box="[998,1164,1545,1572]" pageId="13" pageNumber="14" pagination="6 - 8" refId="ref28119" refString="Yamana, Y. (2016) Kii-suido shuhen no namako bunnrui no genjou [The present situation of the studies on Holothurioidea in the Kii Strait], The Umiushi Tsushin (ISSN 1341 - 5972), published by the Research Institute of Marine Invertebrates, Tokyo, 92, 6 - 8. [only in Japanese]" type="journal article" year="2016">Yamana 2016</bibRefCitation>
), because the bodywall ossicle morphologies (reported in
<bibRefCitation id="417801A2413AFFDFFDA6F9D6FCEAF9A2" author="Clark, A. M. &amp; Rowe, F. W. E." box="[563,808,1580,1607]" pageId="13" pageNumber="14" refId="ref25186" refString="Clark, A. M. &amp; Rowe, F. W. E. (1971) Monograph of shallow-water Indo-West Pacific echinoderms. British Museum of Natural History, London, 238 pp." type="book" year="1971">Clark &amp; Rowe 1971</bibRefCitation>
;
<bibRefCitation id="417801A2413AFFDFFCA3F9D6FBDCF9A2" author="Liao, Y. &amp; Clark, A. M." box="[822,1054,1580,1607]" pageId="13" pageNumber="14" refId="ref26030" refString="Liao, Y. &amp; Clark, A. M. (1995) The Echinoderms of Southern China. Science Press Beijing, 614 pp." type="book" year="1995">Liao &amp; Clark 1995</bibRefCitation>
; and the present study) of these species resemble each other. However,
<bibRefCitation id="417801A2413AFFDFFDF7F9AAFCCAF98E" author="Semper, C." box="[610,776,1616,1643]" pageId="13" pageNumber="14" refId="ref27545" refString="Semper, C. (1867 - 1868) Holothurien. Reisen im Archipel der Philippinen. Teil 2. Wissenschaftliche Resultate. C. W. Kreidel, Wiesbaden, 288 pp. https: // doi. org / 10.5962 / bhl. title. 11687" type="book" year="1867">Semper (1867</bibRefCitation>
8) plainly stated that the gonadal tubules of this species have needle-shaped ossicles and that the bodywall possesses quite large ossicles. (These ossicle morphologies were not mentioned in
<bibRefCitation id="417801A2413AFFDFFE32F962FD86F957" author="Semper, C." box="[423,580,1688,1714]" pageId="13" pageNumber="14" refId="ref27545" refString="Semper, C. (1867 - 1868) Holothurien. Reisen im Archipel der Philippinen. Teil 2. Wissenschaftliche Resultate. C. W. Kreidel, Wiesbaden, 288 pp. https: // doi. org / 10.5962 / bhl. title. 11687" type="book" year="1867">Semper 1867</bibRefCitation>
8, p 67). The presence of such ossicles were not observed in the present new species. From seas worldwide, there are more than 60 nominal species and subspecies in the genus
<taxonomicName id="E2E907D0413AFFDFFAD3F941FA5EF930" authorityName="Oken" authorityYear="1815" box="[1350,1436,1723,1749]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="13" pageNumber="14" phylum="Echinodermata" rank="genus">
<emphasis id="179DA041413AFFDFFAD3F941FA5EF930" box="[1350,1436,1723,1749]" italics="true" pageId="13" pageNumber="14">Thyone</emphasis>
</taxonomicName>
(65 or 68 species/subspecies nominated in WoRMS 2021b), in which 14 species are presently classified into the Group III (comprising species with both rosettes and tables in the introvert) congeners which had been implied by
<bibRefCitation id="417801A2413AFFDFFEB4F8DDFDC6F8A4" author="Deichmann, E." box="[289,516,1831,1857]" pageId="13" pageNumber="14" pagination="43 - 236" refId="ref25222" refString="Deichmann, E. (1930) The Holothurians of the Western Part of the Atlantic Ocean. Bulletin of the Museum of Comparative Zooology at Harvard College, 71, 43 - 236." type="journal article" year="1930">Deichmann (1930)</bibRefCitation>
,
<bibRefCitation id="417801A2413AFFDFFD87F8DDFD09F8A4" author="Panning, A." box="[530,715,1831,1857]" pageId="13" pageNumber="14" pagination="404 - 470" refId="ref26870" refString="Panning, A. (1949) Versuch einer Neuordnung der Familie Cucumariidae (Holothurioidea, Dendrochirota). Zoologischen Jahrbuchern, 78, 404 - 470." type="journal article" year="1949">Panning (1949)</bibRefCitation>
and Pawson &amp; Millar (1981). The present new species also should be involved into this group as 15th species (Table 8). Among the 15 species of Group III, this species have the four identical characters: 1) granulated, mulberry-like small rosettes (
<figureCitation id="BDD260D6413AFFDFFBABF894FB7FF86C" box="[1086,1213,1902,1929]" captionStart="FIGURE 5" captionStartId="11.[151,250,1888,1912]" captionTargetBox="[182,1404,182,1863]" captionTargetId="figure-18@11.[181,1406,181,1864]" captionTargetPageId="11" captionText="FIGURE 5. Scanning electron microscope images of ossicles of Thyone liaoi sp. nov., (holotype, AH, WMNH-INV-2014- 182). A, ossicles of the ventrolateral tentacle; B, the ventrolateral skin of peri-oral; C, the ventrolateral skin of pharynx; D, the ventrolateral introvert skin; E, the pedicel of the abdominal side; F and G, the middle part of the bodywall on the abdominal side, and on the dorsal side, respectively; H, the ventrolateral anal papilla." figureDoi="http://doi.org/10.5281/zenodo.5225382" httpUri="https://zenodo.org/record/5225382/files/figure.png" pageId="13" pageNumber="14">Fig. 5BD</figureCitation>
) are detected from the peri-oral, pharynx, and introvert skin; 2) length (approx. 70160 µm) of the bodywall table disc (
<figureCitation id="BDD260D6413AFFDFFAF8F868FF1EF835" captionStart="FIGURE 5" captionStartId="11.[151,250,1888,1912]" captionTargetBox="[182,1404,182,1863]" captionTargetId="figure-18@11.[181,1406,181,1864]" captionTargetPageId="11" captionText="FIGURE 5. Scanning electron microscope images of ossicles of Thyone liaoi sp. nov., (holotype, AH, WMNH-INV-2014- 182). A, ossicles of the ventrolateral tentacle; B, the ventrolateral skin of peri-oral; C, the ventrolateral skin of pharynx; D, the ventrolateral introvert skin; E, the pedicel of the abdominal side; F and G, the middle part of the bodywall on the abdominal side, and on the dorsal side, respectively; H, the ventrolateral anal papilla." figureDoi="http://doi.org/10.5281/zenodo.5225382" httpUri="https://zenodo.org/record/5225382/files/figure.png" pageId="13" pageNumber="14">Fig. 5F, G</figureCitation>
) is exceedingly larger than that of the other 14 species (Table 8-2); 3) length (approx. 30430 µm) of the tentacle ossicles (
<figureCitation id="BDD260D6413AFFDFFE29F823FDD8F811" box="[444,538,2009,2036]" captionStart="FIGURE 5" captionStartId="11.[151,250,1888,1912]" captionTargetBox="[182,1404,182,1863]" captionTargetId="figure-18@11.[181,1406,181,1864]" captionTargetPageId="11" captionText="FIGURE 5. Scanning electron microscope images of ossicles of Thyone liaoi sp. nov., (holotype, AH, WMNH-INV-2014- 182). A, ossicles of the ventrolateral tentacle; B, the ventrolateral skin of peri-oral; C, the ventrolateral skin of pharynx; D, the ventrolateral introvert skin; E, the pedicel of the abdominal side; F and G, the middle part of the bodywall on the abdominal side, and on the dorsal side, respectively; H, the ventrolateral anal papilla." figureDoi="http://doi.org/10.5281/zenodo.5225382" httpUri="https://zenodo.org/record/5225382/files/figure.png" pageId="13" pageNumber="14">Fig. 5A</figureCitation>
) is exceedingly larger than that of the other 14 species (Table 8-2); 4) the two-pillared spire of the bodywall table is united one-pillar with 46 minute apical spines (
<figureCitation id="BDD260D64139FFDCFB74FF6DFAA2FF54" box="[1249,1376,150,177]" captionStart="FIGURE 5" captionStartId="11.[151,250,1888,1912]" captionTargetBox="[182,1404,182,1863]" captionTargetId="figure-18@11.[181,1406,181,1864]" captionTargetPageId="11" captionText="FIGURE 5. Scanning electron microscope images of ossicles of Thyone liaoi sp. nov., (holotype, AH, WMNH-INV-2014- 182). A, ossicles of the ventrolateral tentacle; B, the ventrolateral skin of peri-oral; C, the ventrolateral skin of pharynx; D, the ventrolateral introvert skin; E, the pedicel of the abdominal side; F and G, the middle part of the bodywall on the abdominal side, and on the dorsal side, respectively; H, the ventrolateral anal papilla." figureDoi="http://doi.org/10.5281/zenodo.5225382" httpUri="https://zenodo.org/record/5225382/files/figure.png" pageId="14" pageNumber="15">Fig. 5F, G</figureCitation>
), but not forming a crown. From these features, this new species is easily distinguished from the 14 species of the Group III and other 24 species of undetectable group of
<taxonomicName id="E2E907D04139FFDCFCA1FF24FC48FF1D" authorityName="Oken" authorityYear="1815" box="[820,906,222,248]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="14" pageNumber="15" phylum="Echinodermata" rank="genus">
<emphasis id="179DA0414139FFDCFCA1FF24FC48FF1D" box="[820,906,222,248]" italics="true" pageId="14" pageNumber="15">Thyone</emphasis>
</taxonomicName>
(Table 8).
</paragraph>
</subSubSection>
<subSubSection id="6DF32FD84139FFDCFF52FEF8FA52FE81" pageId="14" pageNumber="15" type="distribution">
<paragraph id="25567C534139FFDCFF52FEF8FA52FE81" blockId="14.[151,1437,150,535]" pageId="14" pageNumber="15">
<emphasis id="179DA0414139FFDCFF52FEF8FEA3FEF9" bold="true" box="[199,353,258,284]" pageId="14" pageNumber="15">Distribution.</emphasis>
Known from the Gulf of
<collectingCountry id="5DFE3CC34139FFDCFDE5FEF8FD7DFEF9" box="[624,703,258,284]" name="Vietnam" pageId="14" pageNumber="15">Tonkin</collectingCountry>
,
<collectingCountry id="5DFE3CC34139FFDCFD5CFEF8FCCEFEF9" box="[713,780,258,284]" name="China" pageId="14" pageNumber="15">China</collectingCountry>
, depth
<quantity id="E211D1B64139FFDCFCC2FEF8FC4FFEF9" box="[855,909,258,285]" metricMagnitude="1" metricUnit="m" metricValue="5.5" pageId="14" pageNumber="15" unit="m" value="55.0">55 m</quantity>
(
<bibRefCitation id="417801A24139FFDCFC0FFEF8FBABFEF8" author="Liao, Y. &amp; Clark, A. M." box="[922,1129,258,285]" pageId="14" pageNumber="15" refId="ref26030" refString="Liao, Y. &amp; Clark, A. M. (1995) The Echinoderms of Southern China. Science Press Beijing, 614 pp." type="book" year="1995">Liao &amp; Clark 1995</bibRefCitation>
), and several localities in the Seto Inland Sea,
<collectingCountry id="5DFE3CC34139FFDCFEDAFEDCFE53FEA5" box="[335,401,294,320]" name="Japan" pageId="14" pageNumber="15">Japan</collectingCountry>
: middle and eastern parts of the Seto Inland Sea, sand bottom, depth
<quantity id="E211D1B64139FFDCFBEAFEDFFB74FEA5" box="[1151,1206,293,320]" metricMagnitude="1" metricUnit="m" metricValue="2.6" pageId="14" pageNumber="15" unit="m" value="26.0">26 m</quantity>
(ST-
<quantity id="E211D1B64139FFDCFB78FEDCFAD4FEDA" box="[1261,1302,293,320]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="14" pageNumber="15" unit="in" value="4.0">4 in</quantity>
2014); mud bottom, depth
<quantity id="E211D1B64139FFDCFEA1FEB3FE51FE81" box="[308,403,329,356]" metricMagnitude="1" metricUnit="m" metricValue="2.6" metricValueMax="2.7" metricValueMin="2.5" pageId="14" pageNumber="15" unit="m" value="26.0" valueMax="27.0" valueMin="25.0">2527 m</quantity>
(ST-
<quantity id="E211D1B64139FFDCFE5FFEB3FE31FE86" box="[458,499,329,355]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="14" pageNumber="15" unit="in" value="7.0">7 in</quantity>
2014), and northen part of the Kii Strait, sand bottom, shallow water (
<bibRefCitation id="417801A24139FFDCFB7CFEB3FA46FE81" author="Yamana, Y." box="[1257,1412,329,356]" pageId="14" pageNumber="15" pagination="6 - 8" refId="ref28119" refString="Yamana, Y. (2016) Kii-suido shuhen no namako bunnrui no genjou [The present situation of the studies on Holothurioidea in the Kii Strait], The Umiushi Tsushin (ISSN 1341 - 5972), published by the Research Institute of Marine Invertebrates, Tokyo, 92, 6 - 8. [only in Japanese]" type="journal article" year="2016">Yamana 2016</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="6DF32FD84139FFDCFF52FE97FBEFFDF2" pageId="14" pageNumber="15" type="etymology">
<paragraph id="25567C534139FFDCFF52FE97FB58FE4E" blockId="14.[151,1437,150,535]" pageId="14" pageNumber="15">
<emphasis id="179DA0414139FFDCFF52FE97FE8CFE62" bold="true" box="[199,334,365,391]" pageId="14" pageNumber="15">Etymology.</emphasis>
The species named in honor of the famous researcher Dr. Yulin Liao of the Institute of Oceanology, Academia Sinica (IOAS) Qingdao,
<collectingCountry id="5DFE3CC34139FFDCFDB3FE6BFDAEFE4E" box="[550,620,401,427]" name="China" pageId="14" pageNumber="15">China</collectingCountry>
, who first reported this species as
<emphasis id="179DA0414139FFDCFC79FE6BFB58FE4E" box="[1004,1178,401,427]" italics="true" pageId="14" pageNumber="15">
<taxonomicName id="E2E907D04139FFDCFC79FE6BFB54FE4E" authorityName="Semper" authorityYear="1867" box="[1004,1174,401,427]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="14" pageNumber="15" phylum="Echinodermata" rank="species" species="pedata">Thyone pedata</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="25567C534139FFDCFF52FE4FFBEFFDF2" blockId="14.[151,1437,150,535]" pageId="14" pageNumber="15">
<emphasis id="179DA0414139FFDCFF52FE4FFDD5FE2B" bold="true" box="[199,535,436,463]" pageId="14" pageNumber="15">Molecular data comparison.</emphasis>
In BLAST searches, the obtained partial COI gene sequence is deeply divergent from all other sequences. A sequence identified as
<taxonomicName id="E2E907D04139FFDCFD48FE22FC5BFE16" box="[733,921,472,499]" class="Holothuroidea" family="Synallactidae" genus="Paelopatides" kingdom="Animalia" order="Synallactida" pageId="14" pageNumber="15" phylum="Echinodermata" rank="species" species="undetermined">
<emphasis id="179DA0414139FFDCFD48FE22FCADFE17" box="[733,879,472,498]" italics="true" pageId="14" pageNumber="15">Paelopatides</emphasis>
sp.
</taxonomicName>
(INSD accession number
<accessionNumber id="3ABAE1B04139FFDCFB5DFE22FA85FE16" box="[1224,1351,472,499]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KX874356" pageId="14" pageNumber="15">KX874356</accessionNumber>
, 80.6% similarity with 100% coverage) was closest (meaningless at generic/family level).
</paragraph>
</subSubSection>
</treatment>
</document>