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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1087.64382" ID-GBIF-Dataset="9c9b7f37-a72e-4610-8ac5-908f601d1121" ID-PMC="PMC8888540" ID-Pensoft-Pub="1313-2970-1087-19" ID-Pensoft-UUID="ECD9B4DC2A3357AABC04DB88FB7D40B1" ID-PubMed="35437365" ID-ZooBank="94F2384E640E4A58B8B4D9D06675D2C2" ModsDocID="1313-2970-1087-19" checkinTime="1645627321867" checkinUser="pensoft" docAuthor="Nupponen 1, Kari &amp; Sihvonen, Pasi" docDate="2022" docId="271E6A3F1D8D5B2388724D971B4C6CE9" docLanguage="en" docName="ZooKeys 1087: 19-104" docOrigin="ZooKeys 1087" docPubDate="2022-02-22" docSource="http://dx.doi.org/10.3897/zookeys.1087.64382" docTitle="Landryia ankylosauroides Nupponen, sp. nov., genus combination 2022, incertae sedis" docType="treatment" docUuid="5173B006-37BB-467F-A1CF-187F5B523FC2" docUuidSource="ZooBank" docVersion="4" id="ECD9B4DC2A3357AABC04DB88FB7D40B1" lastPageNumber="19" masterDocId="ECD9B4DC2A3357AABC04DB88FB7D40B1" masterDocTitle="Revision of Neotropical Scythrididae moths and descriptions of 22 new species from Argentina, Chile, and Peru (Lepidoptera, Gelechioidea)" masterLastPageNumber="104" masterPageNumber="19" pageNumber="19" updateTime="1668151506650" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Revision of Neotropical Scythrididae moths and descriptions of 22 new species from Argentina, Chile, and Peru (Lepidoptera, Gelechioidea)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Nupponen 1, Kari</mods:namePart>
<mods:affiliation>Merenneidontie 19 D, FI- 02320 Espoo, Finland</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sihvonen, Pasi</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-2237-9325</mods:nameIdentifier>
<mods:affiliation>Finnish Museum of Natural History, P. O. Box 17, Pohjoinen Rautatiekatu 13, 00014 University of Helsinki, Finland</mods:affiliation>
<mods:nameIdentifier type="email">pasi.sihvonen@helsinki.fi</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-02-22</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1087</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>19</mods:start>
<mods:end>104</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1087.64382</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1087.64382</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1087-19</mods:identifier>
<mods:identifier type="ZooBank">94F2384E640E4A58B8B4D9D06675D2C2</mods:identifier>
<mods:identifier type="Pensoft-UUID">ECD9B4DC2A3357AABC04DB88FB7D40B1</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="193585865" LSID="urn:lsid:zoobank.org:act:5173B006-37BB-467F-A1CF-187F5B523FC2" httpUri="http://treatment.plazi.org/id/271E6A3F1D8D5B2388724D971B4C6CE9" lastPageNumber="19" pageId="0" pageNumber="19">
<subSubSection pageId="0" pageNumber="19" type="nomenclature">
<paragraph pageId="0" pageNumber="19">
<taxonomicName LSID="http://zoobank.org/5173B006-37BB-467F-A1CF-187F5B523FC2" authority="Nupponen, sp. nov., genus combination" authorityName="Nupponen, sp. nov., genus combination" authorityYear="2022" class="Insecta" family="Scythrididae" genus="Landryia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Landryia ankylosauroides" order="Lepidoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="ankylosauroides" status="incertae sedis">Landryia ankylosauroides Nupponen, sp. nov., genus combination</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="19">incertae sedis</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="19" type="description">
<paragraph pageId="0" pageNumber="19">
<figureCitation captionStart="Figures 711" captionStartId="F2" captionText="Figures 7 - 11. Scythrididae adults, genera Rhamphura and Landryia 7 R. angulisociella Nupponen, sp. nov., genus combination incertae sedis, male, holotype 8 R. curvisociella Nupponen, sp. nov., genus combination incertae sedis, male, holotype 9 R. tetrafasciella Nupponen, sp. nov., genus combination incertae sedis, female, holotype 10 A L. ankylosauroides Nupponen sp. nov., genus combination incertae sedis, male, holotype 10 B L. ankylosauroides Nupponen, sp. nov., genus combination incertae sedis, male, paratype 11 L. chilensis Nupponen, sp. nov., genus combination incertae sedis, male, holotype." figureDoi="10.3897/zookeys.1087.64382.figures7-11" httpUri="https://binary.pensoft.net/fig/649316" pageId="0" pageNumber="19">Figs 10</figureCitation>
<figureCitation captionStart="Figures 4142" captionStartId="F10" captionText="Figures 41 - 42. Male genitalia of Rhamphura and Landryia 41 R. curvisociella Nupponen sp. nov., genus combination incertae sedis, holotype, slide 1 / 12 Dec. 2019 KN 42 L. ankylosauroides Nupponen, sp. nov., genus combination incertae sedis, holotype, above (lateral view): slide 4 / 13 Dec. 2019 KN, below (ventral view): slide 2 / 13 Dec. 2019 KN (paratype)." figureDoi="10.3897/zookeys.1087.64382.figures41-42" httpUri="https://binary.pensoft.net/fig/649324" pageId="0" pageNumber="19">, 42</figureCitation>
<figureCitation captionStart="Figure 65" captionStartId="F23" captionText="Figure 65. Female genitalia of Landryia ankylosauroides Nupponen sp. nov., genus combination incertae sedis, paratype, slide 1 / 15 Dec. 2019 KN." figureDoi="10.3897/zookeys.1087.64382.figure65" httpUri="https://binary.pensoft.net/fig/649337" pageId="0" pageNumber="19">, 65</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="19" type="type material">
<paragraph pageId="0" pageNumber="19">Type material.</paragraph>
<paragraph pageId="0" pageNumber="19">
<emphasis bold="true" pageId="0" pageNumber="19">
<emphasis italics="true" pageId="0" pageNumber="19">Holotype</emphasis>
.
</emphasis>
Argentina • ♂; prov. Santiago del Estero, Pozo Honda village S, by salt lake;
<geoCoordinate degrees="27" direction="south" minutes="17.2" orientation="latitude" precision="92" value="-27.286667">27°17.2'S</geoCoordinate>
,
<geoCoordinate degrees="64" direction="west" minutes="28.0" orientation="longitude" precision="92" value="-64.46667">64°28.0'W</geoCoordinate>
; 260 m a.s.l.; 20 Sep. 2017; K. Nupponen &amp; R. Haverinen leg.; [BOLD sample ID] KN01059; [genitalia slide] K. Nupponen prep. No. 4/13 Jan. 2019; coll. NUPP (MZH).
</paragraph>
<paragraph pageId="0" pageNumber="19">
<emphasis bold="true" pageId="0" pageNumber="19">
<emphasis italics="true" pageId="0" pageNumber="19">Paratypes</emphasis>
.
</emphasis>
Argentina • 20 ♂, 9 ♀; same data as for holotype; [BOLD sample ID] KN01060; [genitalia slide] K. Nupponen prep. No. 2/13 Jan. 2019 ♂; coll. NUPP; • 21 ♂, 12 ♀; same data as for holotype except collecting date; 19 Sep. 2017; [BOLD sample IDs] KN01061, KN01062; [genitalia slide] K. Nupponen prep. No. 1/15 Dec. 2019 ♀; coll. NUPP; • 1 ♀; prov. La Rioja, valley east of Sierra de Sanogasta;
<geoCoordinate degrees="29" direction="south" minutes="51.7" orientation="latitude" precision="92" value="-29.861668">29°51.7'S</geoCoordinate>
,
<geoCoordinate degrees="67" direction="west" minutes="09.9" orientation="longitude" precision="92" value="-67.165">67°09.9'W</geoCoordinate>
; 670 m a.s.l.; 22 Sep. 2017; K. Nupponen &amp; R. Haverinen leg.; coll. NUPP.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="19" type="diagnosis">
<paragraph pageId="0" pageNumber="19">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="19">
A pale streak in forewing is diagnostic. In the male genitalia of
<taxonomicName lsidName="L. ankylosauroides" pageId="0" pageNumber="19" rank="species" species="ankylosauroides">
<emphasis italics="true" pageId="0" pageNumber="19">L. ankylosauroides</emphasis>
</taxonomicName>
, the S-shaped distal arm of gnathos is distinctive, and similar structure is found only in
<taxonomicName lsidName="L. chilensis" pageId="0" pageNumber="19" rank="species" species="chilensis">
<emphasis italics="true" pageId="0" pageNumber="19">L. chilensis</emphasis>
</taxonomicName>
. The two taxa are readily separated by several details in the male genitalia: in
<taxonomicName lsidName="L. ankylosauroides" pageId="0" pageNumber="19" rank="species" species="ankylosauroides">
<emphasis italics="true" pageId="0" pageNumber="19">L. ankylosauroides</emphasis>
</taxonomicName>
the left valva is much shorter than the right one (in
<taxonomicName lsidName="L. chilensis" pageId="0" pageNumber="19" rank="species" species="chilensis">
<emphasis italics="true" pageId="0" pageNumber="19">L. chilensis</emphasis>
</taxonomicName>
valvae ca. equal length) and the right valva is without large distal lobe (in
<taxonomicName lsidName="L. chilensis" pageId="0" pageNumber="19" rank="species" species="chilensis">
<emphasis italics="true" pageId="0" pageNumber="19">L. chilensis</emphasis>
</taxonomicName>
a large distal lobe is present), tergum VIII has narrow lateral arms with melanised spikes (in
<taxonomicName lsidName="L. chilensis" pageId="0" pageNumber="19" rank="species" species="chilensis">
<emphasis italics="true" pageId="0" pageNumber="19">L. chilensis</emphasis>
</taxonomicName>
spikes are absent and posterior margin is deeply concave). In the female genitalia of
<taxonomicName lsidName="L. ankylosauroides" pageId="0" pageNumber="19" rank="species" species="ankylosauroides">
<emphasis italics="true" pageId="0" pageNumber="19">L. ankylosauroides</emphasis>
</taxonomicName>
, sterigma is an inverted cone, which resembles that of
<taxonomicName class="Reptilia" family="Amaranthaceae" kingdom="Animalia" lsidName="R. pozohondaensis" pageId="0" pageNumber="19" phylum="Chordata" rank="species" species="pozohondaensis">
<emphasis italics="true" pageId="0" pageNumber="19">R. pozohondaensis</emphasis>
</taxonomicName>
, but differs by trapezoid posterior flap (parallel triangular flaps in
<taxonomicName class="Reptilia" family="Amaranthaceae" kingdom="Animalia" lsidName="R. pozohondaensis" pageId="0" pageNumber="19" phylum="Chordata" rank="species" species="pozohondaensis">
<emphasis italics="true" pageId="0" pageNumber="19">R. pozohondaensis</emphasis>
</taxonomicName>
) and absence of cleavage at anterior tip.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="19" type="description">
<paragraph pageId="0" pageNumber="19">Description.</paragraph>
<paragraph pageId="0" pageNumber="19">
Wingspan 10.5-12 mm. Head, collar, tegula and thorax pale fuscous; few white scales around eye, and small blotch of same colour at medioposterior margin of thorax. Neck tuft and haustellum white. Scape dorsally dark brown, ventrally dirty white, pecten longer than diameter of scape. Flagellum dark brown, 0.65
<normalizedToken originalValue="×">x</normalizedToken>
length of forewing, in male ciliate, sensillae ~ 0.75
<normalizedToken originalValue="×">x</normalizedToken>
as long as diameter of flagellum. Labial palps: palpomere I white; lower surface of posterior 1/2 of palpomere II and palpomere III dark brown, otherwise white. Legs cream, upper surfaces more or less mixed with different tones of brown. Abdomen dorsally fuscous, ventrally dirty white. Forewing grey, costal area slightly darker than dorsal one; more or less distinct white streak in forewing from base to termen, in dorsal margin edged by interrupted dark brown line; few white scales at apical area. Hindwing pale grey.
</paragraph>
<paragraph pageId="0" pageNumber="19">
<emphasis bold="true" pageId="0" pageNumber="19">
<emphasis italics="true" pageId="0" pageNumber="19">Male genitalia</emphasis>
.
</emphasis>
Uncus heavily sclerotised, subtriangular, basal part heart-shaped. Gnathos base small belt; distal arm long, strongly sigmoid (S-shaped), tip club-shaped covered by minute spines. Tegumen rectangular. Phallus 0.7
<normalizedToken originalValue="×">x</normalizedToken>
length of right valva, straight, shaped as elongated bottle. Valvae asymmetrical, fused at basal 1/2, dorsal margins setose; left valva short, oval; right valva 1.4
<normalizedToken originalValue="×">x</normalizedToken>
longer than left, of constant width, subapically with small extension, apex bent, heavily sclerotised, tip shallowly indented. Saccus as long as right valva, triangular. Sternum VIII large hexagonal plate, medioposteriorly deeply U-shaped; posterior margin with two asymmetrical and diverging extensions, longer one with numerous long and thin setae; latero-anterior corners with parallel long and narrow extensions, tips spatulate. Tergum VIII trapezoid basally, anterior margin concave; medioposteriorly with digitate extension; mediolaterally at both sides long and upwards directed extensions, distal 1/2 with ~ ten long and heavily sclerotised spiniform setae.
</paragraph>
<paragraph pageId="0" pageNumber="19">
<emphasis bold="true" pageId="0" pageNumber="19">
<emphasis italics="true" pageId="0" pageNumber="19">Female genitalia</emphasis>
.
</emphasis>
Sterigma triangular. Ostium small, situated at anterior tip of sterigma. Sternum VII trapezoid; lateroposteriorly small triangular flaps at both sides, anterior corners extended. Sternum VIII with two, suboval, sclerotised plates. Apophyses anteriores 0.35
<normalizedToken originalValue="×">x</normalizedToken>
length of apophyses posteriores.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="19" type="etymology">
<paragraph pageId="0" pageNumber="19">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="19">
Latinised adjective in the nominative singular. The species name alludes to the shape of the gnathos arm, reminiscent of the tail of
<taxonomicName authorityName="Brown" authorityYear="1908" class="Reptilia" family="Ankylosauridae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Dinosauria" pageId="0" pageNumber="19" phylum="Chordata" rank="family">Ankylosauridae</taxonomicName>
(
<taxonomicName class="Reptilia" kingdom="Animalia" lsidName="" pageId="0" pageNumber="19" phylum="Chordata" rank="class">Reptilia</taxonomicName>
:
<taxonomicName genus="Dinosauria" lsidName="" pageId="0" pageNumber="19" rank="genus">Dinosauria</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="19" type="distribution">
<paragraph pageId="0" pageNumber="19">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="19">NW Argentina.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="19" type="habitat">
<paragraph pageId="0" pageNumber="19">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="19">
The habitat at the type locality of Pozo Honda is a dry shrubby area near a salt lake shore (Fig.
<figureCitation captionStart="Figure 78" captionStartId="F36" captionText="Figure 78. Collecting site of Scythris bicoloristrigella sp. nov. and Rhamphura spiniuncus sp. nov.: Argentina, Andes Mts. (1620 m), salt lake by Cordillera del Tigre, 26 Jan. 2017." figureDoi="10.3897/zookeys.1087.64382.figure78" httpUri="https://binary.pensoft.net/fig/649350" pageId="0" pageNumber="19">78</figureCitation>
); the other collecting site is an open valley with halophytic vegetation.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="19" type="genetic data">
<paragraph pageId="0" pageNumber="19">Genetic data.</paragraph>
<paragraph pageId="0" pageNumber="19">
BIN: BOLD:ADZ2684 (
<emphasis italics="true" pageId="0" pageNumber="19">n</emphasis>
= 3 from Argentina). Genetically rather homogenous, maximum variation 0.32%. Nearest neighbour: North American
<taxonomicName baseAuthorityName="Clemens" baseAuthorityYear="1860" class="Insecta" family="Scythrididae" genus="Landryia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Landryia matutella" order="Lepidoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="matutella">
<emphasis italics="true" pageId="0" pageNumber="19">Landryia matutella</emphasis>
</taxonomicName>
(Clemens, 1860) (
<taxonomicName authorityName="Nupponen 1 &amp; Sihvonen" authorityYear="2022" family="Scythrididae" kingdom="Animalia" lsidName="" order="Lepidoptera" pageId="0" pageNumber="19" rank="family">Scythrididae</taxonomicName>
, BIN: BOLD:AAE6120, 1.25%).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="19" type="remarks">
<paragraph pageId="0" pageNumber="19">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="19">
Based on our COI maximum likelihood phylogeny, the South American taxa
<taxonomicName authorityName="Nupponen, sp. nov., genus combination" authorityYear="2022" class="Insecta" family="Scythrididae" genus="Landryia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Landryia ankylosauroides" order="Lepidoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="ankylosauroides">
<emphasis italics="true" pageId="0" pageNumber="19">Landryia ankylosauroides</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Nupponen, sp. nov., genus combination" authorityYear="2022" class="Insecta" family="Scythrididae" genus="Landryia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Landryia chilensis" order="Lepidoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="chilensis">
<emphasis italics="true" pageId="0" pageNumber="19">Landryia chilensis</emphasis>
</taxonomicName>
group inside a large clade, whose taxa are classified in
<taxonomicName authorityName="Kemal &amp; Kocak" authorityYear="2006" class="Insecta" family="Scythrididae" genus="Landryia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Landryia" order="Lepidoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="19">Landryia</emphasis>
</taxonomicName>
on BOLD (Suppl. material 2). However,
<taxonomicName authorityName="Nupponen, sp. nov., genus combination" authorityYear="2022" class="Insecta" family="Scythrididae" genus="Landryia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Landryia ankylosauroides" order="Lepidoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="ankylosauroides">
<emphasis italics="true" pageId="0" pageNumber="19">Landryia ankylosauroides</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Nupponen, sp. nov., genus combination" authorityYear="2022" class="Insecta" family="Scythrididae" genus="Landryia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Landryia chilensis" order="Lepidoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="chilensis">
<emphasis italics="true" pageId="0" pageNumber="19">Landryia chilensis</emphasis>
</taxonomicName>
do not have the diagnostic morphological characters of
<taxonomicName authorityName="Kemal &amp; Kocak" authorityYear="2006" class="Insecta" family="Scythrididae" genus="Landryia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Landryia" order="Lepidoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="19">Landryia</emphasis>
</taxonomicName>
, such as a greatly enlarged bulbus ejaculatorius (unless accidentally removed during dissection) in the male genitalia and the pincer-like projections on the caudal margin of female sternum VII (
<bibRefCitation DOI="https://doi.org/10.4039/entm123160fv" author="Landry, JF" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="0" pageNumber="19" pagination="1 - 341" refId="B22" refString="Landry, JF, 1991. Systematics of Nearctic Scythrididae (Lepidoptera: Gelechioidea): phylogeny and classification of supraspecific taxa, with a review of described species. Memoirs of the Entomological Society of Canada 160: 1 - 341, DOI: https://doi.org/10.4039/entm123160fv" title="Systematics of Nearctic Scythrididae (Lepidoptera: Gelechioidea): phylogeny and classification of supraspecific taxa, with a review of described species." url="https://doi.org/10.4039/entm123160fv" volume="160" year="1991">Landry 1991</bibRefCitation>
). Also, male sternum VIII of
<taxonomicName authorityName="Nupponen, sp. nov., genus combination" authorityYear="2022" class="Insecta" family="Scythrididae" genus="Landryia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Landryia ankylosauroides" order="Lepidoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="ankylosauroides">
<emphasis italics="true" pageId="0" pageNumber="19">Landryia ankylosauroides</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Nupponen, sp. nov., genus combination" authorityYear="2022" class="Insecta" family="Scythrididae" genus="Landryia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Landryia chilensis" order="Lepidoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="chilensis">
<emphasis italics="true" pageId="0" pageNumber="19">Landryia chilensis</emphasis>
</taxonomicName>
are distinct with their spiniform setae and long apodemes, but such are not present in North American
<taxonomicName authorityName="Kemal &amp; Kocak" authorityYear="2006" class="Insecta" family="Scythrididae" genus="Landryia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Landryia" order="Lepidoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="19">Landryia</emphasis>
</taxonomicName>
(
<bibRefCitation DOI="https://doi.org/10.4039/entm123160fv" author="Landry, JF" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="0" pageNumber="19" pagination="1 - 341" refId="B22" refString="Landry, JF, 1991. Systematics of Nearctic Scythrididae (Lepidoptera: Gelechioidea): phylogeny and classification of supraspecific taxa, with a review of described species. Memoirs of the Entomological Society of Canada 160: 1 - 341, DOI: https://doi.org/10.4039/entm123160fv" title="Systematics of Nearctic Scythrididae (Lepidoptera: Gelechioidea): phylogeny and classification of supraspecific taxa, with a review of described species." url="https://doi.org/10.4039/entm123160fv" volume="160" year="1991">Landry 1991</bibRefCitation>
). Further, North American
<taxonomicName lsidName="L. matutella" pageId="0" pageNumber="19" rank="species" species="matutella">
<emphasis italics="true" pageId="0" pageNumber="19">L. matutella</emphasis>
</taxonomicName>
, which is genetically the nearest neighbour to taxon
<taxonomicName authorityName="Nupponen, sp. nov., genus combination" authorityYear="2022" class="Insecta" family="Scythrididae" genus="Landryia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Landryia ankylosauroides" order="Lepidoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="ankylosauroides">
<emphasis italics="true" pageId="0" pageNumber="19">Landryia ankylosauroides</emphasis>
</taxonomicName>
, is morphologically different. We therefore classified these two taxa in
<taxonomicName authorityName="Kemal &amp; Kocak" authorityYear="2006" class="Insecta" family="Scythrididae" genus="Landryia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Landryia" order="Lepidoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="19">Landryia</emphasis>
</taxonomicName>
(incertae sedis), highlighting the need for further research.
</paragraph>
</subSubSection>
</treatment>
</document>