343 lines
44 KiB
XML
343 lines
44 KiB
XML
<document ID-CLB-Dataset="21464" ID-DOI="http://dx.doi.org/10.3897/zookeys.949.51269" ID-GBIF-Dataset="e4d2c425-4596-487c-8c4f-b09330b48203" ID-PMC="PMC7381721" ID-Pensoft-Pub="1313-2970-949-1" ID-Pensoft-UUID="303D64593B3B550DBB7CE01AA547409D" ID-PubMed="32765176" ID-ZooBank="93BFA448BB7343CDBC4EA86090DA63D5" ModsDocID="1313-2970-949-1" checkinTime="1594832489974" checkinUser="pensoft" docAuthor="Salata, Sebastian & Fisher, Brian L." docDate="2020" docId="C55A1B833B4A52E991961A7F8E08366C" docLanguage="en" docName="ZooKeys 949: 1-185" docOrigin="ZooKeys 949" docSource="http://dx.doi.org/10.3897/zookeys.949.51269" docTitle="Pheidole kely Salata & Fisher 2020, sp. nov." docType="treatment" docUuid="40D3BEC5-AB9B-4891-B58B-BF60B571341B" docUuidSource="ZooBank" docVersion="6" id="303D64593B3B550DBB7CE01AA547409D" lastPageNumber="1" masterDocId="303D64593B3B550DBB7CE01AA547409D" masterDocTitle="Taxonomic revision of the Pheidole sikorae species group (Hymenoptera, Formicidae) from Madagascar" masterLastPageNumber="185" masterPageNumber="1" pageNumber="1" updateTime="1732651982216" updateUser="ExternalLinkService">
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<mods:titleInfo id="C3ACF09B768467182B3E42586FEAC831">
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<mods:title id="435D6B4D19AFAE13A390370C1217AB6D">Taxonomic revision of the Pheidole sikorae species group (Hymenoptera, Formicidae) from Madagascar</mods:title>
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</mods:titleInfo>
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<mods:name id="2873BA4ACBDFACFE85EC3D61F7DD5C28" type="personal">
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<mods:roleTerm id="BDA893B84EA3CCDDF4EAA018205490CC">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="54873CD8DF61149A38821A352424CD71">Salata, Sebastian</mods:namePart>
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<mods:affiliation id="64CD1245E52C5E6876FD59487C418DC3">Department of Entomology, California Academy of Sciences, San Francisco, CA 94118, USA</mods:affiliation>
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<mods:nameIdentifier id="CFBD5093D84FABFB795F150D34455D82" type="orcid">https://orcid.org/0000-0003-0811-2309</mods:nameIdentifier>
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<mods:nameIdentifier id="8577B6502A2A1F710B137776FC06DB9D" type="email">sdsalata@gmail.com</mods:nameIdentifier>
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<mods:namePart id="6A59C4EC1BC6DD70D1CF666986C269DF">Fisher, Brian L.</mods:namePart>
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<mods:affiliation id="40299D3D4B41815EF73F647B1E4161C8">Department of Entomology, California Academy of Sciences, San Francisco, CA 94118, USA</mods:affiliation>
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<mods:nameIdentifier id="534AEEC7895FB705DB25A55EC7C7F66B" type="orcid">https://orcid.org/0000-0002-4653-3270</mods:nameIdentifier>
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<mods:title id="07FEE4E50F721C511698F04B7C21198E">ZooKeys</mods:title>
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<mods:date id="D43A5DEE52D44A7C19A9FA25407471A7">2020</mods:date>
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<mods:detail id="00F6BC7E18FD9535DDF88A00024FCCF8" type="volume">
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<mods:number id="47C7874F1B07AC633CD3532F96BF4866">949</mods:number>
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<mods:url id="BC324C28E849AA845CE4BC7F3B74CD8B">http://dx.doi.org/10.3897/zookeys.949.51269</mods:url>
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<mods:classification id="E3A610A31159D9E84BD016E8B509FF59">journal article</mods:classification>
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<mods:identifier id="BC147B5D4F6AF0FA186CC904F0987368" type="DOI">http://dx.doi.org/10.3897/zookeys.949.51269</mods:identifier>
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<treatment id="C55A1B833B4A52E991961A7F8E08366C" ID-GBIF-Taxon="165090164" LSID="urn:lsid:zoobank.org:act:40D3BEC5-AB9B-4891-B58B-BF60B571341B" httpUri="http://treatment.plazi.org/id/C55A1B833B4A52E991961A7F8E08366C" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Formicidae" scope_order="Hymenoptera">
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<subSubSection id="0CDB39557B52825338BCF549C6693AD1" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="6CF9DD408E3417E41B66051ABD9F342D" pageId="0" pageNumber="1">
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<taxonomicName id="CB658CBC768DF2B298B59F1866E857FD" ID-CoL="CMLCF" LSID="C55A1B83-3B4A-52E9-9196-1A7F8E08366C" authority="Salata & Fisher, 2020" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole kely" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="kely" status="sp. nov.">Pheidole kely</taxonomicName>
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<taxonomicNameLabel id="A222C6AA5AF410E0F63572C6A8694737" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation id="67364F3E991A0E54EED79AFF24078670" captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Pheidole kely sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0149839) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0235022)." figureDoi="10.3897/zookeys.949.51269.figure36" httpUri="https://binary.pensoft.net/fig/432205" pageId="0" pageNumber="1">Figs 36A-F</figureCitation>
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<figureCitation id="9E3C0E5A946864C45BC841BD1B3B17E2" captionStart="Figure 63" captionStartId="F63" captionText="Figure 63. Major worker, hypostomal teeth. Pheidole alina sp. nov. (A). Pheidole ambohimanga sp. nov. (B). Pheidole analavelona sp. nov. (C). Pheidole andohahela sp. nov. (D). Pheidole anomala sp. nov. (E). Pheidole anosyenne sp. nov. (F). Pheidole antranohofa sp. nov. (G). Pheidole beanka sp. nov. (H). Pheidole befotaka sp. nov. (I). Pheidole dasos sp. nov. (J). Pheidole flavominuta sp. nov. (K). Pheidole gracilis sp. nov. (L). Pheidole haboka sp. nov. (M). Pheidole havoana sp. nov. (N). Pheidole hazo sp. nov. (O). Pheidole itremo sp. nov. (P). Pheidole joffreville sp. nov. (Q). Pheidole kely sp. nov. (R). Pheidole lavasoa sp. nov. (S). Pheidole litigiosa Forel (T). Pheidole mahamavo sp. nov. (U). Pheidole mainty sp. nov. (V). Pheidole mamiratra sp. nov. (W). Pheidole manantenina sp. nov. (X)." figureDoi="10.3897/zookeys.949.51269.figure63" httpUri="https://binary.pensoft.net/fig/432232" pageId="0" pageNumber="1">, 63R</figureCitation>
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<figureCitation id="1DEC15D2D8589D13B1C1F48E7F12AE39" captionStart="Figure 65" captionStartId="F65" captionText="Figure 65. Distribution. Pheidole alina sp. nov. (A). Pheidole ambohimanga sp. nov. (B). Pheidole analavelona sp. nov. (C). Pheidole andohahela sp. nov. (D). Pheidole anomala sp. nov. (E). Pheidole anosyenne sp. nov. (F). Pheidole antranohofa sp. nov. (G). Pheidole beanka sp. nov. (H). Pheidole befotaka sp. nov. (I). Pheidole dasos sp. nov. (J). Pheidole flavominuta sp. nov. (K). Pheidole gracilis sp. nov. (L). Pheidole haboka sp. nov. (M). Pheidole havoana sp. nov. (N). Pheidole hazo sp. nov. (O). Pheidole itremo sp. nov. (P). Pheidole joffreville sp. nov. (Q). Pheidole kely sp. nov. (R). Pheidole lavasoa sp. nov. (S). Pheidole litigiosa Forel (T)." figureDoi="10.3897/zookeys.949.51269.figure65" httpUri="https://binary.pensoft.net/fig/432234" pageId="0" pageNumber="1">, 65R</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="2A7D28964753324BEE85EAA25101E399" pageId="0" pageNumber="1" type="type material">
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<paragraph id="E5C2142A343561D125B86959A9CBC957" pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph id="1014FAD605800994CB5D67FF69139F1D" pageId="0" pageNumber="1">
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<emphasis id="48588CB5EF34D0CA4C61F85999BCCEAD" bold="true" italics="true" pageId="0" pageNumber="1">Holotype.</emphasis>
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Madagascar. • 1 major worker; Toamasina; Parc National de Zahamena;
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<geoCoordinate id="EDA3A6E910E404CEB4C6AABED3AD5C33" degrees="17.73359" direction="south" orientation="latitude" precision="1" value="-17.73359">-17.73359</geoCoordinate>
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,
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<geoCoordinate id="60E5A46413F49A050D73A558F6163A14" degrees="48.72625" direction="east" orientation="longitude" precision="1" value="48.72625">48.72625</geoCoordinate>
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; alt. 950 m; 19 Feb 2009; B. L. Fisher et al. leg.; rainforest, ex rotten log; BLF22067; CASENT0235022 (CASC).
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<emphasis id="56010E8E4DD16A7DA3A210CA6340E823" bold="true" italics="true" pageId="0" pageNumber="1">Paratype.</emphasis>
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• 1w.; same data as for holotype, CASENT0149839 (CASC).
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</paragraph>
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<caption id="10BA3701D60158FE6AA98710FE775B60" doi="10.3897/zookeys.949.51269.figure36" httpUri="https://binary.pensoft.net/fig/432205" pageId="0" pageNumber="1" start="Figure 36" startId="F36">
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<paragraph id="63E53583B9E0EF14B2E42268813A4792" pageId="0" pageNumber="1">
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<emphasis id="89FB0EB789BF65871BD3D29713E3A0DC" bold="true" pageId="0" pageNumber="1">Figure 36.</emphasis>
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<taxonomicName id="39EFE1F7FDA2592E0F066031D9ACEDCD" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole kely" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="kely">
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<emphasis id="CE9A5FE93B761D48BB8F251446193A59" italics="true" pageId="0" pageNumber="1">Pheidole kely</emphasis>
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</taxonomicName>
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sp. nov., full-face view (
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<emphasis id="8DBB17E508045E4279A662F643EB5F6C" bold="true" pageId="0" pageNumber="1">A</emphasis>
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), profile (
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<emphasis id="65AB3209B24B0E82E58F9B5B25F2E13C" bold="true" pageId="0" pageNumber="1">C</emphasis>
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), and dorsal view (
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<emphasis id="24635D818648C51A88AC6969828F5DE1" bold="true" pageId="0" pageNumber="1">E</emphasis>
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) of paratype minor worker (CASENT0149839) and full-face view (
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<emphasis id="BF9D11E1645843E30E703E2FBF7F0A26" bold="true" pageId="0" pageNumber="1">B</emphasis>
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), profile (
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<emphasis id="047E2648B9B8591E66F0401A9CF0DD22" bold="true" pageId="0" pageNumber="1">D</emphasis>
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), and dorsal view (
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<emphasis id="75F047576E4D9391A57429792C1705CB" bold="true" pageId="0" pageNumber="1">F</emphasis>
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) of holotype major worker (CASENT0235022).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="1F7D2F9FF71BD0EE14B5E49DE4E4FC39" pageId="0" pageNumber="1" type="materials_examined">
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<paragraph id="19A5E8422749D101CBFA6F2EFFBB5582" pageId="0" pageNumber="1">Other material.</paragraph>
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<paragraph id="5AF078FF01C51B3D92688360246C9890" pageId="0" pageNumber="1">
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Madagascar. -
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<emphasis id="D74977EF3DCACF9C6835E2FCB3DC1175" bold="true" pageId="0" pageNumber="1">Antsiranana</emphasis>
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: • 2w., 2s.; Betaolana Forest, along Bekona River;
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<geoCoordinate id="57E1DA61E3DAB4B59889FB580727AF65" degrees="14.52996" direction="south" orientation="latitude" precision="1" value="-14.52996">-14.52996</geoCoordinate>
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,
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<geoCoordinate id="C1724D90FBF95A1D805AFE140A7A0968" degrees="49.44039" direction="east" orientation="longitude" precision="1" value="49.44039">49.44039</geoCoordinate>
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; alt. 880 m; 4 Mar 2009; B. L. Fisher et al. leg.; rainforest, ex rotten log; BLF22452 (CASC). • 2w., 2q.; Betaolana Forest, along Bekona River;
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<geoCoordinate id="7FD86FC417A40C1A85E8F514DE59ED83" degrees="14.52996" direction="south" orientation="latitude" precision="1" value="-14.52996">-14.52996</geoCoordinate>
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,
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<geoCoordinate id="CA2846F98D2684E76E752FA4F9E1908E" degrees="49.44039" direction="east" orientation="longitude" precision="1" value="49.44039">49.44039</geoCoordinate>
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; alt. 880 m; 4 Mar 2009; B. L. Fisher et al. leg.; rainforest, ex rotten log; BLF22480 (CASC). • 2w., 2s.; Betaolana Forest, along Bekona River;
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<geoCoordinate id="27393179A42608F5180C26026724AED0" degrees="14.52996" direction="south" orientation="latitude" precision="1" value="-14.52996">-14.52996</geoCoordinate>
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,
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<geoCoordinate id="DB8FAF7731BE9FD8E4A5FE2956AE7A4A" degrees="49.44039" direction="east" orientation="longitude" precision="1" value="49.44039">49.44039</geoCoordinate>
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; alt. 880 m; 4 Mar 2009; B. L. Fisher et al. leg.; rainforest, ex rotten log; BLF22559 (CASC). • 1w., 1s.; Betaolana Forest, along Bekona River;
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<geoCoordinate id="92128BBFFCC5F86DBDC98D7E26361BF2" degrees="14.52996" direction="south" orientation="latitude" precision="1" value="-14.52996">-14.52996</geoCoordinate>
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<geoCoordinate id="7D89964A6D3A57A68CD51DB878587416" degrees="49.44039" direction="east" orientation="longitude" precision="1" value="49.44039">49.44039</geoCoordinate>
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; alt. 880 m; 5 Mar 2009; B. L. Fisher et al. leg.; rainforest, ex rotten log; BLF22591 (CASC). • 1w., 1s.; Betaolana Forest, along Bekona River;
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<geoCoordinate id="2F17E4E2D1C4B49EA63AD0E8079A7D05" degrees="14.52996" direction="south" orientation="latitude" precision="1" value="-14.52996">-14.52996</geoCoordinate>
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<geoCoordinate id="E450371916AC0FEE50697D76332112AD" degrees="49.44039" direction="east" orientation="longitude" precision="1" value="49.44039">49.44039</geoCoordinate>
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; alt. 880 m; 5 Mar 2009; B. L. Fisher et al. leg.; rainforest, ex rotten log; BLF22628 (CASC). • 2w., 2q.; Makirovana forest;
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<geoCoordinate id="1D8AAC4421E42758413A4067DFD2E8DB" degrees="14.17066" direction="south" orientation="latitude" precision="1" value="-14.17066">-14.17066</geoCoordinate>
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,
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<geoCoordinate id="E38C734427DDE24AA706D14F2CCF2942" degrees="49.95409" direction="east" orientation="longitude" precision="1" value="49.95409">49.95409</geoCoordinate>
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; alt. 415 m; 29 Apr 2011; B. L. Fisher et al. leg.; rainforest, ex rotten log; BLF26641 (CASC). • 2w.; Makirovana forest;
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<geoCoordinate id="41C67D3B354E36CE14C3D8C3A6778975" degrees="14.17066" direction="south" orientation="latitude" precision="1" value="-14.17066">-14.17066</geoCoordinate>
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,
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<geoCoordinate id="2098858EDA36D1DE72C9272F176FA878" degrees="49.95409" direction="east" orientation="longitude" precision="1" value="49.95409">49.95409</geoCoordinate>
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; alt. 415 m; 29 Apr 2011; B. L. Fisher et al. leg.; rainforest, ex root mat, ground layer; BLF26670 (CASC). • 2w.; Makirovana forest;
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<geoCoordinate id="D978F2C15AB28140E830AF54D3572AA8" degrees="14.16044" direction="south" orientation="latitude" precision="1" value="-14.16044">-14.16044</geoCoordinate>
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,
|
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<geoCoordinate id="94E22D140A6A38568AD494E55B1959FE" degrees="49.95216" direction="east" orientation="longitude" precision="1" value="49.95216">49.95216</geoCoordinate>
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; alt. 550 m; 1 May 2011; B. L. Fisher et al. leg.; rainforest, under rootmat, litter on rock; BLF26911 (CASC). • 3w., 2s.; Parc National de Marojejy, Manantenina River, 27.6 km 35°NE Andapa, 9.6 km 327°NNW Manantenina; -14.435, 49.76; alt. 775 m; 15 Nov 2003; B. L. Fisher et al. leg.; rainforest, ex rotten log; BLF08904 (CASC). • 3w., 6s.; Parc National de Marojejy, Manantenina River, 27.6 km 35°NE Andapa, 9.6 km 327°NNW Manantenina; -14.435, 49.76; alt. 775 m; 15 Nov 2003; B. L. Fisher et al. leg.; rainforest, ex rotten log; BLF08928 (CASC). • 2w.; Parc National de Marojejy, Manantenina River, 27.6 km 35°NE Andapa, 9.6 km 327°NNW Manantenina; -14.435, 49.76; alt. 775 m; 16 Nov 2003; B. L. Fisher et al. leg.; rainforest, ex rotten log; BLF08943 (CASC). • 2w., 3s.; Parc National de Marojejy, Manantenina River, 28.0 km 38°NE Andapa, 8.2 km 333°NNW Manantenina;
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||
<geoCoordinate id="C06E08F57FBC8C8F86697502871E0802" degrees="14.43667" direction="south" orientation="latitude" precision="1" value="-14.43667">-14.43667</geoCoordinate>
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,
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<geoCoordinate id="B175CC9E1BE4B159554D4B86F7B30DEF" degrees="49.775" direction="east" orientation="longitude" precision="55" value="49.775">49.775</geoCoordinate>
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||
; alt. 450 m; 12 Nov 2003; B. L. Fisher et al. leg.; rainforest, ex rotten log; BLF08749 (CASC). • 3w., 1s., 1q.; Sava Region: Parc National de Marojejy, Manantenina River, 28.0 km 24.8°NE Andapa;
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||
<geoCoordinate id="D718297DFDC04ECD97E6FD5D687D0ED9" degrees="14.43461" direction="south" orientation="latitude" precision="1" value="-14.43461">-14.43461</geoCoordinate>
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,
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<geoCoordinate id="B4A53B42D4FA097111BA9D70A7C10ED9" degrees="49.76074" direction="east" orientation="longitude" precision="1" value="49.76074">49.76074</geoCoordinate>
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; alt. 780 m; 8 Feb 2018; B. L. Fisher et al. leg.; rainforest, ex rotten log; BLF40819 (CASC). • 3w., 1s., 1m.; Sava Region: Parc National de Marojejy, Manantenina River, 28.0 km 24.8°NE Andapa;
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||
<geoCoordinate id="3B9E3FA933B034BF55A5F6C35EDA0990" degrees="14.43461" direction="south" orientation="latitude" precision="1" value="-14.43461">-14.43461</geoCoordinate>
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,
|
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<geoCoordinate id="02DE4359ECE5F565B4C2C3DF8D32F180" degrees="49.76074" direction="east" orientation="longitude" precision="1" value="49.76074">49.76074</geoCoordinate>
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; alt. 780 m; 13 Feb 2018; B. L. Fisher et al. leg.; rainforest, ex rotten log; BLF41152 (CASC). • 2w., 1s., 1q.; Sava Region: Parc National de Marojejy, near Manantenina River;
|
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<geoCoordinate id="46A97179C6550B5115FA40A493806913" degrees="14.43677" direction="south" orientation="latitude" precision="1" value="-14.43677">-14.43677</geoCoordinate>
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,
|
||
<geoCoordinate id="939B0CCB4479970C764F7114F135154F" degrees="49.77541" direction="east" orientation="longitude" precision="1" value="49.77541">49.77541</geoCoordinate>
|
||
; alt. 475 m; 7 Feb 2018; B. L. Fisher et al. leg.; rainforest, ex rotten log; BLF40761 (CASC).-
|
||
<emphasis id="606EF53394466C3964D3B8C3D4D44F1C" bold="true" pageId="0" pageNumber="1">Fianarantsoa</emphasis>
|
||
: • 2w.; Parc National
|
||
<normalizedToken id="17C2F3D6821364BC105C8A8ECA50D10B" originalValue="d’Isalo">d'Isalo</normalizedToken>
|
||
, Ambovo Springs, 29.3 km 4°N Ranohira;
|
||
<geoCoordinate id="7CC75A729FCB70FFF4478CCC6253A185" degrees="22.29833" direction="south" orientation="latitude" precision="1" value="-22.29833">-22.29833</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="C37BEFAF02A5148C731D7C11E0106A86" degrees="45.35167" direction="east" orientation="longitude" precision="1" value="45.35167">45.35167</geoCoordinate>
|
||
; alt. 990 m; 9 Feb 2003; B. L. Fisher et al. leg.; Uapaca woodland, under stone; BLF07796 (CASC).-
|
||
<emphasis id="91CFF694530D429EB3AB5F4CF810724D" bold="true" pageId="0" pageNumber="1">Toamasina</emphasis>
|
||
: • 3w.; Montagne
|
||
<normalizedToken id="15CD6787A6D4E0511B5F288B48953686" originalValue="d’Akirindro">d'Akirindro</normalizedToken>
|
||
7.6 km 341°NNW Ambinanitelo;
|
||
<geoCoordinate id="E573E5440741B432149738D0230D4F24" degrees="15.28833" direction="south" orientation="latitude" precision="1" value="-15.28833">-15.28833</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="1C7C6C3C0DB6C71302A132E6A9FC5311" degrees="49.54833" direction="east" orientation="longitude" precision="1" value="49.54833">49.54833</geoCoordinate>
|
||
; alt. 600 m; 17 Mar 2003; B. L. Fisher et al. leg.; rainforest, ex rotten log; BLF08293 (CASC). • 1w., 1s.; Parc National de Zahamena;
|
||
<geoCoordinate id="78C5214B69FE4395441A8C84F3F9985B" degrees="17.73359" direction="south" orientation="latitude" precision="1" value="-17.73359">-17.73359</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="E56145109A5CC604A25F209A9202B31B" degrees="48.72625" direction="east" orientation="longitude" precision="1" value="48.72625">48.72625</geoCoordinate>
|
||
; alt. 950 m; 19 Feb 2009; B. L. Fisher et al. leg.; rainforest, ex rotten log; BLF22093 (CASC).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D43E4A7E1C651781DDD18392ACA6975A" pageId="0" pageNumber="1" type="diagnosis">
|
||
<paragraph id="AD68F3DCAB420C150505C9162167EA83" pageId="0" pageNumber="1">Diagnosis.</paragraph>
|
||
<paragraph id="DF84E4CDF59CA40AF0240B2243842EBE" pageId="0" pageNumber="1">
|
||
Minute species.
|
||
<emphasis id="D821D822B1D9BED947932C185AC7B017" bold="true" italics="true" pageId="0" pageNumber="1">Major workers.</emphasis>
|
||
HL <1.0 mm and WL <0.8 mm; head in full-face view sub-oval, widening slightly posteriorly, with anterior and posterior sides convex, in lateral view sub-oval; ventral and dorsal faces convex; body yellowish orange; sides of head with moderately dense, moderately long, suberect to erect pilosity; entire head distinctly sculptured, medial part of frons with thick, dense, longitudinal, and interrupted rugae and smooth to indistinctly rugulate interspaces; scape, when laid back, slightly exceeding the midlength of head; mesosoma with fine and sparse rugofoveolae, pronotum with rugofoveolae reduced and smooth notches on medial parts of dorsum and lateral sides; katepisternum with smooth notch; inner hypostomal teeth distinct, large, closely spaced, triangular, with rounded apex directed upward; outer hypostomal teeth lobe-like, lower than inner teeth and approximately the same weight; inner and outer hypostomal teeth moderately closely spaced and not connected by concavity; base of first gastral tergite smooth.
|
||
<emphasis id="4ECEFC7974D7DD72061C50793B1FE82C" bold="true" italics="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
|
||
HL <0.5 mm and WL <0.6 mm, scape, when laid back, surpassing the posterior head margin by one-fifth of its length; propodeal spines small and triangular; head relatively rectangular; body dark yellow; head foveolate; medial side of frons with sparse, fine, and interrupted rugulae; area posterolateral from eyes smooth; mesosoma smooth.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="707A1A473B1BEC7BAE8CD7EDCE4113C1" pageId="0" pageNumber="1" type="description">
|
||
<paragraph id="B97F84466436AF09DE56D09463D42486" pageId="0" pageNumber="1">Description.</paragraph>
|
||
<paragraph id="A2A9989DE917E4A236B0C54BA65FC8AC" pageId="0" pageNumber="1">
|
||
<emphasis id="B21C2B415FDBCDD5F44AD7AC8877FC96" bold="true" pageId="0" pageNumber="1">Major workers.</emphasis>
|
||
Measurements (
|
||
<emphasis id="508222128C2B0BDDA45BE6FDFC37E13D" italics="true" pageId="0" pageNumber="1">N</emphasis>
|
||
= 10): HL: 0.9-0.97 (0.93); HW: 0.87-0.95 (0.91); SL: 0.48-0.53 (0.5); EL: 0.1-0.13 (0.11); WL: 0.77-0.84 (0.8); PSL: 0.12-0.15 (0.14); MTL: 0.44-0.5 (0.47); PNW: 0.39-0.46 (0.43); PTW: 0.1-0.14 (0.12); PPW: 0.25-0.31 (0.28); CI: 100.2-104.6 (102.4); SI: 53.1-58.3 (55.0); PSLI: 13.2-15.7 (14.5); PPI: 38.7-48.6 (41.8); PNI: 43.0-49.2 (47.3); MTI: 49.3-54.1 (51.5).
|
||
</paragraph>
|
||
<paragraph id="BF1F35B8164BA6D86F5D00BF3C6BF440" pageId="0" pageNumber="1">
|
||
<emphasis id="56279DE9A8B8D41A1FED060C7A49148E" bold="true" italics="true" pageId="0" pageNumber="1">Head.</emphasis>
|
||
In full-face view sub-oval, widening slightly posteriorly, with anterior and posterior sides convex (Fig.
|
||
<figureCitation id="292F099384090465F23A26B57C0E8C92" captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Pheidole kely sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0149839) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0235022)." figureDoi="10.3897/zookeys.949.51269.figure36" httpUri="https://binary.pensoft.net/fig/432205" pageId="0" pageNumber="1">36B</figureCitation>
|
||
). In lateral view sub-oval; ventral and dorsal faces convex; inner hypostomal teeth visible. Sides of the head with moderately dense, moderately long, suberect to erect pilosity; whole head with dense, long, decumbent to erect pilosity. Medial part of frons with thick, dense, longitudinal, and interrupted rugae and smooth to indistinctly rugulate interspaces; lateral sides with thick, dense, and irregular rugae, interspaces shiny and distinctly rugofoveolate; rugae more longitudinal on anterolateral sides. Occipital lobes with thinner, irregular rugae and smooth to indistinctly rugofoveolate interspaces. Area posterolateral from eyes with sculpture weaker and rugoreticulate; sculpture weakening posteriorly. Gena with relatively sparse, thick, longitudinal rugae and smooth to indistinctly rugoreticulate interspaces. Centre of clypeus smooth and shiny, lateral sides with indistinct rugulae; median notch present, moderately wide, and shallow; median longitudinal carina present; lateral longitudinal carinae absent. Scape, when laid back, slightly exceeding the midlength of head; pilosity subdecumbent to erect (Fig.
|
||
<figureCitation id="560940C32AABDB2BAACD1957AF9BA6D2" captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Pheidole kely sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0149839) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0235022)." figureDoi="10.3897/zookeys.949.51269.figure36" httpUri="https://binary.pensoft.net/fig/432205" pageId="0" pageNumber="1">36B, D</figureCitation>
|
||
). Inner hypostomal teeth distinct, large, closely spaced, triangular, with rounded apex directed upward; outer hypostomal teeth lobe-like, lower than inner teeth and approximately the same weight; inner and outer hypostomal teeth moderately closely spaced and not connected by concavity (Fig.
|
||
<figureCitation id="4F3DEF83E2EBB17B0ED567A0CA314ECB" captionStart="Figure 63" captionStartId="F63" captionText="Figure 63. Major worker, hypostomal teeth. Pheidole alina sp. nov. (A). Pheidole ambohimanga sp. nov. (B). Pheidole analavelona sp. nov. (C). Pheidole andohahela sp. nov. (D). Pheidole anomala sp. nov. (E). Pheidole anosyenne sp. nov. (F). Pheidole antranohofa sp. nov. (G). Pheidole beanka sp. nov. (H). Pheidole befotaka sp. nov. (I). Pheidole dasos sp. nov. (J). Pheidole flavominuta sp. nov. (K). Pheidole gracilis sp. nov. (L). Pheidole haboka sp. nov. (M). Pheidole havoana sp. nov. (N). Pheidole hazo sp. nov. (O). Pheidole itremo sp. nov. (P). Pheidole joffreville sp. nov. (Q). Pheidole kely sp. nov. (R). Pheidole lavasoa sp. nov. (S). Pheidole litigiosa Forel (T). Pheidole mahamavo sp. nov. (U). Pheidole mainty sp. nov. (V). Pheidole mamiratra sp. nov. (W). Pheidole manantenina sp. nov. (X)." figureDoi="10.3897/zookeys.949.51269.figure63" httpUri="https://binary.pensoft.net/fig/432232" pageId="0" pageNumber="1">63R</figureCitation>
|
||
).
|
||
<emphasis id="C0645590B96EF3277FBBF8A6329B0387" bold="true" italics="true" pageId="0" pageNumber="1">Mesosoma.</emphasis>
|
||
In lateral view, promesonotum short, angular, and moderately low, posterior mesonotum moderately steep, mesonotal process very indistinct, tubercle-like or absent; promesonotal groove absent; metanotal groove absent; propodeal spines moderately long, with moderately wide base and acute apex; humeral area laterally weakly produced (Fig.
|
||
<figureCitation id="6FCCA7D6BBB09A091A8CB24EF3357991" captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Pheidole kely sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0149839) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0235022)." figureDoi="10.3897/zookeys.949.51269.figure36" httpUri="https://binary.pensoft.net/fig/432205" pageId="0" pageNumber="1">36D</figureCitation>
|
||
). Surface shiny with fine and sparse rugofoveolae; pronotum with rugofoveolae reduced and smooth notches on medial parts of dorsum and lateral sides; katepisternum with smooth notch. Pilosity moderately sparse, long, and erect (Fig.
|
||
<figureCitation id="ADE753326D6D4140A79A820AECCC3115" captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Pheidole kely sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0149839) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0235022)." figureDoi="10.3897/zookeys.949.51269.figure36" httpUri="https://binary.pensoft.net/fig/432205" pageId="0" pageNumber="1">36D, F</figureCitation>
|
||
).
|
||
<emphasis id="A8E77A0777F745AE2BA03425ECFD8213" bold="true" italics="true" pageId="0" pageNumber="1">Petiole.</emphasis>
|
||
Shiny with fine foveolae; node smooth to finely foveolate, triangular, with rounded and thick apex, in rear view node dorsoventrally straight to slightly convex; pilosity moderately sparse and erect (Fig.
|
||
<figureCitation id="F5D78E9BE77BB119F1A6838B8B7BB64B" captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Pheidole kely sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0149839) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0235022)." figureDoi="10.3897/zookeys.949.51269.figure36" httpUri="https://binary.pensoft.net/fig/432205" pageId="0" pageNumber="1">36D, F</figureCitation>
|
||
).
|
||
<emphasis id="966EDB7A4D9D8F69D203E2244463D51E" bold="true" italics="true" pageId="0" pageNumber="1">Postpetiole.</emphasis>
|
||
Shiny and foveolate; dorsum with reduced sculpture and smooth notch; in dorsal view oval, lateral margins medially with two dentate projections; pilosity long, moderately sparse, and erect (Fig.
|
||
<figureCitation id="B895FF6590D231AC7C8E6A41A2DCF403" captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Pheidole kely sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0149839) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0235022)." figureDoi="10.3897/zookeys.949.51269.figure36" httpUri="https://binary.pensoft.net/fig/432205" pageId="0" pageNumber="1">36D, F</figureCitation>
|
||
).
|
||
<emphasis id="FE8197D5A1BD700F4F6D865E280D3252" bold="true" italics="true" pageId="0" pageNumber="1">Gaster.</emphasis>
|
||
Shiny and smooth; pilosity dense, moderately long, and erect (Fig.
|
||
<figureCitation id="1EAE55C959F23732608708AB3A240C32" captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Pheidole kely sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0149839) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0235022)." figureDoi="10.3897/zookeys.949.51269.figure36" httpUri="https://binary.pensoft.net/fig/432205" pageId="0" pageNumber="1">36D, F</figureCitation>
|
||
).
|
||
<emphasis id="3F8C9F76605A4B0A3AE177C1B6C7B1F0" bold="true" italics="true" pageId="0" pageNumber="1">Colour.</emphasis>
|
||
Yellowish orange; mandibles and gaster brown (Fig.
|
||
<figureCitation id="E833B3246006FB6AC618DC6D8DB41C08" captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Pheidole kely sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0149839) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0235022)." figureDoi="10.3897/zookeys.949.51269.figure36" httpUri="https://binary.pensoft.net/fig/432205" pageId="0" pageNumber="1">36D, F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="4E2BAE70208D8D98C44795003490ADC0" pageId="0" pageNumber="1">
|
||
<emphasis id="AA649E62EE0049AEEFD22E7AFD5818EC" bold="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
|
||
Measurements (
|
||
<emphasis id="2E8A7E405E4AA8275B7579055A368C48" italics="true" pageId="0" pageNumber="1">N</emphasis>
|
||
= 10): HL: 0.47-0.52 (0.49); HW: 0.4-0.45 (0.42); SL: 0.43-0.48 (0.45); EL: 0.09-0.11 (0.1); WL: 0.53-0.61 (0.57); PSL: 0.04-0.06 (0.05); MTL: 0.33-0.41 (0.37); PNW: 0.27-0.3 (0.28); PTW: 0.05-0.08 (0.06); PPW: 0.1-0.14 (0.12); CI: 111.9-121.1 (117.0); SI: 103.9-108.0 (106.1); PSLI: 7.9-11.7 (10.3); PPI: 48.3-64.2 (55.5); PNI: 64.9-69.7 (67.0); MTI: 80.2-98.8 (85.4).
|
||
</paragraph>
|
||
<paragraph id="C9CB881D8AB9FACADEAF1E1495A53BF6" pageId="0" pageNumber="1">
|
||
<emphasis id="CFD6959AE9A41A56681C533C6DE4AF3B" bold="true" italics="true" pageId="0" pageNumber="1">Head.</emphasis>
|
||
Cephalic margin indistinctly concave to straight (Fig.
|
||
<figureCitation id="7E1CCF104246369E5FEE95DF3A7A7BBC" captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Pheidole kely sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0149839) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0235022)." figureDoi="10.3897/zookeys.949.51269.figure36" httpUri="https://binary.pensoft.net/fig/432205" pageId="0" pageNumber="1">36A</figureCitation>
|
||
). Pilosity relatively dense, moderately long, decumbent to suberect. Sculpture shiny and foveolate; medial side of frons with sparse, fine, and interrupted rugulae; area posterolateral from eyes smooth; antennal sockets with few thick, curved outward rugae and indistinctly foveolate interspaces. Clypeus with median longitudinal carina absent; two lateral longitudinal carinae absent. Scape, when laid back, surpassing the posterior head margin by one-fifth of its length; pilosity dense, subdecumbent to erect (Fig.
|
||
<figureCitation id="39747411253457F06E869AD21661E553" captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Pheidole kely sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0149839) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0235022)." figureDoi="10.3897/zookeys.949.51269.figure36" httpUri="https://binary.pensoft.net/fig/432205" pageId="0" pageNumber="1">36A, C</figureCitation>
|
||
).
|
||
<emphasis id="F385061C753952F890E9747CFCF6ED57" bold="true" italics="true" pageId="0" pageNumber="1">Mesosoma.</emphasis>
|
||
In lateral view, promesonotum moderately high and short, arched; promesonotal groove absent; metanotal groove distinct; propodeal spines small and triangular (Fig.
|
||
<figureCitation id="29B71848D2C087772CC6E45395486698" captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Pheidole kely sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0149839) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0235022)." figureDoi="10.3897/zookeys.949.51269.figure36" httpUri="https://binary.pensoft.net/fig/432205" pageId="0" pageNumber="1">36C</figureCitation>
|
||
). Sculpture shiny and smooth. Pilosity moderately sparse, moderately long, and erect (Fig.
|
||
<figureCitation id="C5CB9FB19E017D60B40B60A30A03E436" captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Pheidole kely sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0149839) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0235022)." figureDoi="10.3897/zookeys.949.51269.figure36" httpUri="https://binary.pensoft.net/fig/432205" pageId="0" pageNumber="1">36C, E</figureCitation>
|
||
).
|
||
<emphasis id="87DAB0190B757CCF5754AFCBCEA31CDA" bold="true" italics="true" pageId="0" pageNumber="1">Gaster.</emphasis>
|
||
With sparse, erect pilosity (Fig.
|
||
<figureCitation id="DE44F6E10DA42FD2B8AE89F4A859544B" captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Pheidole kely sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0149839) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0235022)." figureDoi="10.3897/zookeys.949.51269.figure36" httpUri="https://binary.pensoft.net/fig/432205" pageId="0" pageNumber="1">36C, E</figureCitation>
|
||
).
|
||
<emphasis id="3C4436030EB2B21F089065B5BC64CD73" bold="true" italics="true" pageId="0" pageNumber="1">Colour.</emphasis>
|
||
Dark yellow, gaster and vertex brown (Fig.
|
||
<figureCitation id="1BE8E05176A813C6AFAFD73A7D521CFC" captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Pheidole kely sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0149839) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0235022)." figureDoi="10.3897/zookeys.949.51269.figure36" httpUri="https://binary.pensoft.net/fig/432205" pageId="0" pageNumber="1">36C, E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="109D5308D65B1652A66BDF9A7F70A305" pageId="0" pageNumber="1" type="etymology">
|
||
<paragraph id="F26BE182DBFC6D583BA87D241A4EA2E7" pageId="0" pageNumber="1">Etymology.</paragraph>
|
||
<paragraph id="470C189DAE2CD0CC471E41590AEEEFAE" pageId="0" pageNumber="1">Malagasy for small in reference to body size.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F86290E6546E7FDF0421253414E0A670" pageId="0" pageNumber="1" type="biology_ecology">
|
||
<paragraph id="5190F0F47EA5608E96F4F5E2F85021BA" pageId="0" pageNumber="1">Biology.</paragraph>
|
||
<paragraph id="D828DE2FC6BFC70E478D30B781B164C1" pageId="0" pageNumber="1">The species was collected between 325-990 m in elevation, in rainforest and Uapaca woodland. Nests were located in rotten logs, under rootmats, and under stones.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="50C2E70C829876B73337B12ED84EC0AD" pageId="0" pageNumber="1" type="comments">
|
||
<paragraph id="2471FB252C6C396640AF70DCE520B576" pageId="0" pageNumber="1">Comments.</paragraph>
|
||
<paragraph id="CADB76C520655D763499DEA4A7ADF732" pageId="0" pageNumber="1">
|
||
<taxonomicName id="A2D5BA77C5327A3678A5A13397B34C83" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole kely" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="kely">
|
||
<emphasis id="D2FD20B4EE2085BDDE7EFF9692D10FCD" italics="true" pageId="0" pageNumber="1">Pheidole kely</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. belongs to the group of species characterised by small body size (major workers: HL <1.05 mm, WL <0.9 mm and minor workers HL <0.5 mm, WL <0.6 mm), sub-oval, slightly widening posteriorly head with anterior and posterior sides convex in major workers, and minor workers with yellow to brown body colouration and foveolate head or head predominantly smooth and relatively oval. The group includes six species:
|
||
<taxonomicName id="9B86F5C070AD703A64480CCF7CAF5655" lsidName="P. havoana" pageId="0" pageNumber="1" rank="species" species="havoana">
|
||
<emphasis id="E24F12784295490951258204827B88BE" italics="true" pageId="0" pageNumber="1">P. havoana</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.,
|
||
<taxonomicName id="65F3B43082D01BF4B98D0E354E86C394" lsidName="P. kely" pageId="0" pageNumber="1" rank="species" species="kely">
|
||
<emphasis id="8CA262BCE9FB51694DF5F21D6D34F475" italics="true" pageId="0" pageNumber="1">P. kely</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.,
|
||
<taxonomicName id="F69318D3A1C57EC5B50D895FD09A8086" lsidName="P. parvula" pageId="0" pageNumber="1" rank="species" species="parvula">
|
||
<emphasis id="15DB6CB514A564CEF992998A2B6FFD51" italics="true" pageId="0" pageNumber="1">P. parvula</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.,
|
||
<taxonomicName id="70BB52CDD26E25250833DC074BE071CE" lsidName="P. parvulogibba" pageId="0" pageNumber="1" rank="species" species="parvulogibba">
|
||
<emphasis id="FE86B2A2C370504DE02AA44126BA8572" italics="true" pageId="0" pageNumber="1">P. parvulogibba</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.,
|
||
<taxonomicName id="7520D6BCBFB2308E4DDA12A4B951F2F9" lsidName="P. volontany" pageId="0" pageNumber="1" rank="species" species="volontany">
|
||
<emphasis id="4C17EDECFED786CE3FE1D83CAFF7460A" italics="true" pageId="0" pageNumber="1">P. volontany</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., and
|
||
<taxonomicName id="D9584D0A12D69E6677F122600D5C6209" lsidName="P. midongy" pageId="0" pageNumber="1" rank="species" species="midongy">
|
||
<emphasis id="1A28343DFEFF5CAD2C2AB2335685DBA0" italics="true" pageId="0" pageNumber="1">P. midongy</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
<taxonomicName id="485DB54C5CC64BE8B61DA813245937FA" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole kely" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="kely">
|
||
<emphasis id="F4C1B197C4CE606D0A306755C079BF5F" italics="true" pageId="0" pageNumber="1">Pheidole kely</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. is distributed from Toamasina north to Sambava and has a single record from Parc National
|
||
<normalizedToken id="BADB6A89717CC6F44360ADBDFA33C9E3" originalValue="d’Isalo">d'Isalo</normalizedToken>
|
||
in Fianarantsoa. Morphologically it is most similar to
|
||
<taxonomicName id="4C0C3E9B1070370E555AAC41CFFB2E50" lsidName="P. havoana" pageId="0" pageNumber="1" rank="species" species="havoana">
|
||
<emphasis id="7B937776A3FC505EE6354C0583CFAD60" italics="true" pageId="0" pageNumber="1">P. havoana</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. and
|
||
<taxonomicName id="3FAD54EE0E117C7E189EEC0E85BFF1A1" lsidName="P. parvulogibba" pageId="0" pageNumber="1" rank="species" species="parvulogibba">
|
||
<emphasis id="CE42E0E03E242D4521F21F928099F7EF" italics="true" pageId="0" pageNumber="1">P. parvulogibba</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., described from the Anosyenne Mts. in Toliara, and parapatric
|
||
<taxonomicName id="AB6F63723F8792708A7A273C2E62E11C" lsidName="P. parvula" pageId="0" pageNumber="1" rank="species" species="parvula">
|
||
<emphasis id="419F0B876ED6EA0AFBFEBCF03C3247DB" italics="true" pageId="0" pageNumber="1">P. parvula</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., known from two localities in the vicinity of Antananarivo. Minor workers of
|
||
<taxonomicName id="88ECCBE6C5A54099C7BEF32396505B16" lsidName="P. kely" pageId="0" pageNumber="1" rank="species" species="kely">
|
||
<emphasis id="DDF2E5E0A8AB3860702AB076BD7449BE" italics="true" pageId="0" pageNumber="1">P. kely</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. distinctly differ from those of
|
||
<taxonomicName id="F0AD92F53684D519431992ADFF191693" lsidName="P. parvula" pageId="0" pageNumber="1" rank="species" species="parvula">
|
||
<emphasis id="2635523D4354F26EF0E9329469E3C843" italics="true" pageId="0" pageNumber="1">P. parvula</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. and
|
||
<taxonomicName id="B0AA41906CD5165D01BC534F51E009D1" lsidName="P. parvulogibba" pageId="0" pageNumber="1" rank="species" species="parvulogibba">
|
||
<emphasis id="A85ADEE53AADA17C79AD078C63E6B378" italics="true" pageId="0" pageNumber="1">P. parvulogibba</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. in having a predominantly foveolate head. Minor workers of
|
||
<taxonomicName id="4BCC7B4668317212D0F9EF0072CF5DCF" lsidName="P. havoana" pageId="0" pageNumber="1" rank="species" species="havoana">
|
||
<emphasis id="D68B9333DED2A19A7A4DC433E9F49510" italics="true" pageId="0" pageNumber="1">P. havoana</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. and
|
||
<taxonomicName id="E26FC9A2BE72E908F4EC733A3586EEDA" lsidName="P. kely" pageId="0" pageNumber="1" rank="species" species="kely">
|
||
<emphasis id="A96D51ACF52D624B1B63F7A136C7F1B1" italics="true" pageId="0" pageNumber="1">P. kely</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. are indistinguishable. Major workers of
|
||
<taxonomicName id="83E51C811DFA381FD0162CED7293DA15" lsidName="P. kely" pageId="0" pageNumber="1" rank="species" species="kely">
|
||
<emphasis id="8B634F19CA8A44315BB3B9F662901740" italics="true" pageId="0" pageNumber="1">P. kely</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. differ from
|
||
<taxonomicName id="68F77F6C610EBE330401197CBDDE9806" lsidName="P. havoana" pageId="0" pageNumber="1" rank="species" species="havoana">
|
||
<emphasis id="0CCF1EA3B62693B386872009C529D825" italics="true" pageId="0" pageNumber="1">P. havoana</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. and
|
||
<taxonomicName id="976983F8B2279F018AE55CB83E1D241B" lsidName="P. parvula" pageId="0" pageNumber="1" rank="species" species="parvula">
|
||
<emphasis id="E151D2ED8DC820AA8316142B5943B39A" italics="true" pageId="0" pageNumber="1">P. parvula</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. in medial part of frons with thick, dense, longitudinal, and interrupted rugae and smooth to indistinctly rugulate interspaces; from
|
||
<taxonomicName id="6E78E117413EA3144A453BD82548B13A" lsidName="P. parvulogibba" pageId="0" pageNumber="1" rank="species" species="parvulogibba">
|
||
<emphasis id="A611B42B8ACCF30B7CE513DCA09DEAF4" italics="true" pageId="0" pageNumber="1">P. parvulogibba</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. in mesosoma with fine and sparse rugofoveolae, pronotum with rugofoveolae reduced, and smooth notches on medial parts of dorsum and its lateral sides.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |