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<document id="5AE94C8ED50C3E76E032232C3999C4AC" ID-DOI="10.5281/zenodo.184062" ID-GBIF-Dataset="89a3460a-6650-4941-9757-7ef48a8e77c9" ID-ISSN="1175-5326" ID-Zenodo-Dep="184062" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459940242068" checkinUser="plazi" docAuthor="Mathis, Wayne N." docDate="2008" docId="902D87820360FFC5FF19E3F6D53903FD" docLanguage="en" docName="zt01874p015.pdf" docOrigin="Zootaxa 1874" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Paraephydra Mathis, 2008, gen. nov." docType="treatment" docVersion="10" lastPageNumber="5" masterDocId="6C14FFFA0363FFC1FF8EE661D459007F" masterDocTitle="Two new neotropical genera of the shore-fly tribe Ephydrini Zetterstedt (Diptera: Ephydridae)" masterLastPageNumber="15" masterPageNumber="1" pageNumber="4" updateTime="1698230402356" updateUser="plazi">
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<mods:title id="9AE38E56BE067546A84B265BB3F2D789">Two new neotropical genera of the shore-fly tribe Ephydrini Zetterstedt (Diptera: Ephydridae)</mods:title>
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<mods:name id="505D3E6E6FCE790F861857365E25BA61" type="personal">
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<mods:namePart id="9C239C3C34CB94182EB524258B966E3A">Mathis, Wayne N.</mods:namePart>
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<mods:part id="75A63D92D7D1C402B41353B353A301CC">
<mods:date id="EB72B24709283725FABF2456A88F34C6">2008</mods:date>
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<mods:number id="E71F9AA21ADBA0F08B35C2FAF0874221">1874</mods:number>
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<mods:identifier id="2A9722B3D643C3C0C2C2702DE9FB2379" type="DOI">10.5281/zenodo.184062</mods:identifier>
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<treatment id="902D87820360FFC5FF19E3F6D53903FD" ID-DOI="http://doi.org/10.5281/zenodo.3509656" ID-GBIF-Taxon="119375858" ID-Zenodo-Dep="3509656" LSID="urn:lsid:plazi:treatment:902D87820360FFC5FF19E3F6D53903FD" httpUri="http://treatment.plazi.org/id/902D87820360FFC5FF19E3F6D53903FD" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
<subSubSection id="509E651F0360FFC2FF19E3F6D5F605CE" box="[151,431,1431,1457]" pageId="3" pageNumber="4" type="nomenclature">
<paragraph id="183B36940360FFC2FF19E3F6D5F605CE" blockId="3.[151,431,1431,1457]" box="[151,431,1431,1457]" pageId="3" pageNumber="4">
<heading id="437381F80360FFC2FF19E3F6D5F605CE" bold="true" box="[151,431,1431,1457]" fontSize="11" level="1" pageId="3" pageNumber="4" reason="1">
<emphasis id="2AF0EA860360FFC2FF19E3F6D5F605CE" bold="true" box="[151,431,1431,1457]" pageId="3" pageNumber="4">
<emphasis id="2AF0EA860360FFC2FF19E3F6D56305CE" bold="true" box="[151,314,1431,1457]" italics="true" pageId="3" pageNumber="4">
<taxonomicName id="DF844D170360FFC2FF19E3F6D56C05CE" ID-CoL="63JYV" box="[151,309,1431,1457]" class="Insecta" family="Ephydridae" genus="Paraephydra" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">Paraephydra</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="31C357FD0360FFC2FECCE3F6D5F605CE" box="[322,431,1431,1457]" pageId="3" pageNumber="4" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="509E651F0360FFC2FF19E381D7D90588" box="[151,896,1504,1527]" pageId="3" pageNumber="4" type="materials_examined">
<paragraph id="183B36940360FFC2FF19E381D7D90588" blockId="3.[151,896,1504,1527]" box="[151,896,1504,1527]" pageId="3" pageNumber="4">
<typeStatus id="C73F88360360FFC2FF19E381D4950588" box="[151,204,1504,1527]" pageId="3" pageNumber="4">Type</typeStatus>
species:
<taxonomicName id="DF844D170360FFC2FEA3E381D6240588" authority="Oliveira" authorityName="Oliveira" box="[301,637,1504,1527]" class="Insecta" family="Ephydridae" genus="Paraephydra" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="freitasi">
<emphasis id="2AF0EA860360FFC2FEA3E381D6520588" box="[301,523,1504,1527]" italics="true" pageId="3" pageNumber="4">Paraephydra freitasi</emphasis>
(Oliveira)
</taxonomicName>
, by present designation.
</paragraph>
</subSubSection>
<subSubSection id="509E651F0360FFC2FF19E043D71E0703" pageId="3" pageNumber="4" type="diagnosis">
<paragraph id="183B36940360FFC2FF19E043D6D106F3" blockId="3.[151,1437,1570,2036]" pageId="3" pageNumber="4">
<emphasis id="2AF0EA860360FFC2FF19E043D54E0643" bold="true" box="[151,279,1570,1596]" pageId="3" pageNumber="4">Diagnosis.</emphasis>
<taxonomicName id="DF844D170360FFC2FEAFE042D5E50643" box="[289,444,1571,1596]" class="Insecta" family="Ephydridae" genus="Paraephydra" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="2AF0EA860360FFC2FEAFE042D5E50643" box="[289,444,1571,1596]" italics="true" pageId="3" pageNumber="4">Paraephydra</emphasis>
</taxonomicName>
is distinguished from other genera of
<taxonomicName id="DF844D170360FFC2FC1DE043D0510643" box="[915,1032,1570,1596]" class="Insecta" family="Ephydridae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="tribe" tribe="Ephydrini">Ephydrini</taxonomicName>
by the following combination of characters: Moderately small to medium-sized shore flies, body length
<quantity id="DF7C9B710360FFC2FB86E02BD0E0061B" box="[1032,1209,1610,1636]" metricMagnitude="-3" metricUnit="m" metricValue="3.0999999999999996" metricValueMax="3.8" metricValueMin="2.4" pageId="3" pageNumber="4" unit="mm" value="3.0999999999999996" valueMax="3.8" valueMin="2.4">2.403.80 mm</quantity>
; setation normally developed, not generally appearing pilose.
</paragraph>
<paragraph id="183B36940360FFC2FF48E0FAD71E0703" blockId="3.[151,1437,1570,2036]" pageId="3" pageNumber="4">
<emphasis id="2AF0EA860360FFC2FF48E0FAD55606CB" box="[198,271,1691,1716]" italics="true" pageId="3" pageNumber="4">Head:</emphasis>
Mesofrons shiny, with metallic luster, differentiated from microtomentose parafrons; cruciate intrafrontal setae l; lateroclinate, fronto-orbital setae 2; antennal groove distinct but not deeply impressed; basal flagellomere lacking large seta inserted on lateral surface; arista as long or slightly longer than combined length of 1st 3 antennal segments, gradually tapered from base to apex, with subpectinate, dorsally branching rays on basal 2/3; postocular setae normally developed, not conspicuous; larger facial setae extended from interfoveal hump with 12, distinctly porrect to anaclinate.
</paragraph>
</subSubSection>
<subSubSection id="509E651F0360FFC5FF48E1EAD6CC01DD" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="description">
<paragraph id="183B36940360FFC5FF48E1EAD5DE00A5" blockId="3.[151,1437,1570,2036]" lastBlockId="4.[151,1437,152,898]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
<emphasis id="2AF0EA860360FFC2FF48E1EAD57C07DB" box="[198,293,1931,1956]" italics="true" pageId="3" pageNumber="4">Thorax:</emphasis>
Females with 1 prescutellar, acrostichal setae; dorsocentral setae 4 (1+3), all well developed; supra-alar seta present; posthumeral seta lacking; intrapostalar seta either weakly developed or lacking; disc of scutellum concolorous with posterior portion of scutum; females lacking dense patch of setae between posterior 2 dorsocentral setae. Hindtibia with apical, anteroventral seta, length equal to or larger than width of tibia at widest point.
</paragraph>
<paragraph id="183B36940367FFC5FF48E689D6CC01DD" blockId="4.[151,1437,152,898]" pageId="4" pageNumber="5">
<emphasis id="2AF0EA860367FFC5FF48E689D567017E" box="[198,318,232,257]" italics="true" pageId="4" pageNumber="5">Abdomen:</emphasis>
Female ventral receptacle with operculum flat, disclike. Male terminalia: symmetrical; epandrium longer than wide, narrowed ventrally, fused almost imperceptibly with base of united surstyli; surstyli fused medially except near apex; posterior surstylar process only slightly longer than lateral process; both processes apical; gonite, hypandrium, and apparently aedeagus fused to form 1 compact structure, curved anteriorly, wide basally, tapered to rounded apex.
</paragraph>
</subSubSection>
<subSubSection id="509E651F0367FFC5FF48E7CED0A60215" pageId="4" pageNumber="5" type="distribution">
<paragraph id="183B36940367FFC5FF48E7CED6D7018D" blockId="4.[151,1437,152,898]" pageId="4" pageNumber="5">
<emphasis id="2AF0EA860367FFC5FF48E7CED53E01B6" bold="true" box="[198,359,431,457]" pageId="4" pageNumber="5">Distribution.</emphasis>
<emphasis id="2AF0EA860367FFC5FEE1E7D1D65A01B6" box="[367,515,432,457]" italics="true" pageId="4" pageNumber="5">Neotropical;</emphasis>
widespread but scarce, from
<collectingCountry id="609376040367FFC5FCEEE7D1D7B501B5" box="[864,1004,432,458]" name="Puerto Rico" pageId="4" pageNumber="5">Puerto Rico</collectingCountry>
south through
<collectingCountry id="609376040367FFC5FB11E7D1D0BF01B5" box="[1183,1254,432,458]" name="Brazil" pageId="4" pageNumber="5">Brazil</collectingCountry>
to
<collectingCountry id="609376040367FFC5FA83E7D1D10801B5" box="[1293,1361,432,458]" name="Chile" pageId="4" pageNumber="5">Chile</collectingCountry>
. None of the congeners is known to be sympatric.
</paragraph>
<paragraph id="183B36940367FFC5FF48E79ED0A60215" blockId="4.[151,1437,152,898]" pageId="4" pageNumber="5">
<emphasis id="2AF0EA860367FFC5FF48E79ED5C10266" bold="true" box="[198,408,511,537]" pageId="4" pageNumber="5">Natural History.</emphasis>
Like other ephydrines,
<taxonomicName id="DF844D170367FFC5FD4EE461D7020266" box="[704,859,512,537]" class="Insecta" family="Ephydridae" genus="Paraephydra" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="2AF0EA860367FFC5FD4EE461D7020266" box="[704,859,512,537]" italics="true" pageId="4" pageNumber="5">Paraephydra</emphasis>
</taxonomicName>
occurs in wetlands. In southern
<collectingCountry id="609376040367FFC5FB7FE461D1680265" box="[1265,1329,512,538]" name="Chile" pageId="4" pageNumber="5">Chile</collectingCountry>
(Osorno Province), I collected specimens of
<taxonomicName id="DF844D170367FFC5FDC8E449D6E3023E" box="[582,698,552,577]" class="Insecta" family="Ephydridae" genus="Paraephydra" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="stauros">
<emphasis id="2AF0EA860367FFC5FDC8E449D6E3023E" box="[582,698,552,577]" italics="true" pageId="4" pageNumber="5">P. stauros</emphasis>
</taxonomicName>
in a sedge meadow near the margins of small but apparently permanent ponds. Nothing is known about the immature stages or the microhabitat of the genus.
</paragraph>
</subSubSection>
<subSubSection id="509E651F0367FFC5FF48E416D53903FD" pageId="4" pageNumber="5" type="discussion">
<paragraph id="183B36940367FFC5FF48E416D6C9029D" blockId="4.[151,1437,152,898]" pageId="4" pageNumber="5">
<emphasis id="2AF0EA860367FFC5FF48E416D50B02EE" bold="true" box="[198,338,631,657]" pageId="4" pageNumber="5">Discussion.</emphasis>
<taxonomicName id="DF844D170367FFC5FED7E419D5AD02EE" box="[345,500,632,657]" class="Insecta" family="Ephydridae" genus="Paraephydra" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="2AF0EA860367FFC5FED7E419D5AD02EE" box="[345,500,632,657]" italics="true" pageId="4" pageNumber="5">Paraephydra</emphasis>
</taxonomicName>
is proposed to accommodate
<taxonomicName id="DF844D170367FFC5FCDBE419D0C102ED" authority="Oliveira" authorityName="Oliveira" box="[853,1176,632,658]" class="Insecta" family="Ephydridae" genus="Ephydrella" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="freitasi">
<emphasis id="2AF0EA860367FFC5FCDBE419D06902EE" box="[853,1072,632,657]" italics="true" pageId="4" pageNumber="5">Ephydrella freitasi</emphasis>
Oliveira
</taxonomicName>
and
<emphasis id="2AF0EA860367FFC5FB5CE419D11502EE" box="[1234,1356,632,657]" italics="true" pageId="4" pageNumber="5">
<taxonomicName id="DF844D170367FFC5FB5CE419D11102EE" box="[1234,1352,632,657]" class="Insecta" family="Ephydridae" genus="Paraephydra" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="stauros">P. stauros</taxonomicName>
,
</emphasis>
which is similar and closely related (see “Remarks” section under
<taxonomicName id="DF844D170367FFC5FCD7E4C1D79002C6" box="[857,969,672,697]" class="Insecta" family="Ephydridae" genus="Ephydrella" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="freitasi">
<emphasis id="2AF0EA860367FFC5FCD7E4C1D79002C6" box="[857,969,672,697]" italics="true" pageId="4" pageNumber="5">P. freitasi</emphasis>
</taxonomicName>
for further comments on the classificatory history and placement of that species).
</paragraph>
<paragraph id="183B36940367FFC5FF48E491D53903FD" blockId="4.[151,1437,152,898]" pageId="4" pageNumber="5">
Sexual dimorphism is evident in the chaetotaxy of
<emphasis id="2AF0EA860367FFC5FC97E491D7E50376" box="[793,956,752,777]" italics="true" pageId="4" pageNumber="5">
<taxonomicName id="DF844D170367FFC5FC97E491D7E10376" box="[793,952,752,777]" class="Insecta" family="Ephydridae" genus="Paraephydra" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">Paraephydra</taxonomicName>
.
</emphasis>
Females, unlike males, have a prescutellar acrostichal seta that is larger than other acrostichal setulae. Based on this character,
<bibRefCitation id="7C154B650367FFC5FB5BE579D1C2034D" author="Oliveira" box="[1237,1435,792,818]" pageId="4" pageNumber="15" refString="Oliveira, S. J. de. (1954 a) Contribuicao para o conhecimento do genero &quot; Dimecoenia &quot; Cresson, 1916. II. Sobre 3 especies novas do Brasil (Diptera, Ephydridae). Revista Brasileira de Biologia, 14 (3), 269 - 278." type="journal article" year="1954" yearSuffix="a">Oliveira (1954a)</bibRefCitation>
described
<emphasis id="2AF0EA860367FFC5FE9FE521D5260326" box="[273,383,832,857]" italics="true" pageId="4" pageNumber="5">P. f re i t a s i</emphasis>
in the genus
<emphasis id="2AF0EA860367FFC5FDA5E521D6ED0326" box="[555,692,832,857]" italics="true" pageId="4" pageNumber="5">
<taxonomicName id="DF844D170367FFC5FDA5E521D6F60326" box="[555,687,832,857]" class="Insecta" family="Ephydridae" genus="Ephydrella" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">Ephydrella</taxonomicName>
,
</emphasis>
as that genus, unlike
<taxonomicName id="DF844D170367FFC5FC4FE521D16F0325" authority="Neotropical" authorityName="Neotropical" box="[961,1334,832,858]" class="Insecta" family="Ephydridae" genus="Dimecoenia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="2AF0EA860367FFC5FC4FE521D00C0326" box="[961,1109,832,857]" italics="true" pageId="4" pageNumber="5">Dimecoenia,</emphasis>
sensu Neotropical
</taxonomicName>
species, lacks these setae.
</paragraph>
</subSubSection>
</treatment>
</document>