183 lines
17 KiB
XML
183 lines
17 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.53.4955" ID-PMC="PMC4547023" ID-Pensoft-Pub="1314-2003-53-39" ID-Pensoft-UUID="824BD167D1535A15FC27FF98FFD3A404" ID-PubMed="26312040" ID-Zenodo-Dep="576284" ModsDocID="1314-2003-53-39" checkinTime="1451251414861" checkinUser="pensoft" docAuthor="Luong, Thien Tam, Hovenkamp, Peter H. & Sosef, Marc S. M." docDate="2015" docId="6B583D3F108144C33CA4F907BBB5BE22" docLanguage="en" docName="PhytoKeys 53: 39-71" docOrigin="PhytoKeys 53" docPubDate="2015-07-21" docSource="http://dx.doi.org/10.3897/phytokeys.53.4955" docTitle="Orthiopteris trichophylla Copel." docType="treatment" docVersion="6" id="824BD167D1535A15FC27FF98FFD3A404" lastPageNumber="60" masterDocId="824BD167D1535A15FC27FF98FFD3A404" masterDocTitle="Revision of the fern genus Orthiopteris (Saccolomataceae) in Malesia and adjacent regions" masterLastPageNumber="71" masterPageNumber="39" pageNumber="59" updateTime="1668141422086" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision of the fern genus Orthiopteris (Saccolomataceae) in Malesia and adjacent regions</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Luong, Thien Tam</mods:namePart>
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<mods:affiliation>Department of Ecology - Evolutionary Biology, Viet Nam National University Ho Chi Minh city (VNUHCM) - University of Science. 227 Nguyen Van Cu, Ho Chi Minh City, Vietnam & Naturalis Biodiversity Center, section Botany. PO Box 9517, 2300 RA Leiden, The Netherlands</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hovenkamp, Peter H.</mods:namePart>
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<mods:affiliation>Naturalis Biodiversity Center, section Botany. PO Box 9517, 2300 RA Leiden, The Netherlands</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sosef, Marc S. M.</mods:namePart>
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<mods:affiliation>Botanic Garden Meise, Nieuwelaan 38, 1860 Meise, Belgium</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>PhytoKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2015</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2015-07-21</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>53</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>39</mods:start>
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<mods:end>71</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.53.4955</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.53.4955</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2003-53-39</mods:identifier>
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<mods:identifier type="Pensoft-UUID">824BD167D1535A15FC27FF98FFD3A404</mods:identifier>
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<mods:identifier type="Zenodo-Dep">576284</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152008541" LSID="urn:lsid:plazi:treatment:6B583D3F108144C33CA4F907BBB5BE22" httpUri="http://treatment.plazi.org/id/6B583D3F108144C33CA4F907BBB5BE22" lastPageId="22" lastPageNumber="60" pageId="20" pageNumber="59">
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<subSubSection pageId="20" pageNumber="59" type="nomenclature">
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<paragraph pageId="20" pageNumber="59">
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<pageBreakToken pageId="20" pageNumber="59" start="start">8</pageBreakToken>
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.
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<taxonomicName LSID="6B583D3F-1081-44C3-3CA4-F907BBB5BE22" authority="Copel." authorityName="Copel." class="Polypodiopsida" family="Saccolomataceae" genus="Orthiopteris" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Orthiopteris trichophylla" order="Polypodiales" pageId="20" pageNumber="59" phylum="Tracheophyta" rank="species" species="trichophylla">Orthiopteris trichophylla Copel.</taxonomicName>
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ultimate segments. Arrows show distance from base of the shallowest sinuses to nearest costule / vein. a Orthiopteris ferulacea, Smith 8932 (K) b Orthiopteris cicutarioides, Carr 13257 (K) c Orthiopteris samoensis, Reinecke 97 a (L) d Orthiopteris trichophylla, Brass 12239 (L) e Orthiopteris campylura var. cautada, Carr 12518 (L) f Orthiopteris campylura var. laxa Mueller s. n. (L) g Orthiopteris campylura var. kingii, Brass 12941 (MICH) h Orthiopteris henriettae, Lam & Meeuse 5892 (L) i Orthiopteris campylura var. insularis Braithwaite 4110 (L). All scale bars 1 mm." figureDoi="10.3897/phytokeys.53.4955.figure2" httpUri="https://binary.pensoft.net/fig/47604" pageId="20" pageNumber="59">Figs 2d</figureCitation>
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Sori of Orthiopteris seen from abaxial side. a, b Orthiopteris campylura var. campylura (Cuming 119, MICH) c Orthiopteris campylura var. kingii (Brass 12941, MICH) d Orthiopteris trichophylla (Brass 12239, L) e Orthiopteris tenuis (Smith 9154, L) f Orthiopteris tenuis (Smith 7274, L) g Orthiopteris campylura var. caudata (Carr 12518, L) h Orthiopteris cicutarioides (Carr 13257, K) i, j Orthiopteris campylura var. insularis (Braithwaite 4110, L) k Orthiopteris tenuis (small segment) (Degener 14648, L) l Orthiopteris ferulacea (Smith 8932, K) m, n Orthiopteris firma (Schlechter 14890, L) o, p Orthiopteris campylura var. laxa (Mueller s. n., L) q, r Orthiopteris henriettae (Lam & Meeuse 5892, L) s Orthiopteris henriettae. t Orthiopteris samoensis (Reinecke 97 a, L) u, v Orthiopteris samoensis (Vaupel 312, L). All scale bars 1 mm, oi = outer indusium, ii = inner indusium." figureDoi="10.3897/phytokeys.53.4955.figure3" httpUri="https://binary.pensoft.net/fig/47605" pageId="20" pageNumber="59">, 3d</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="21" pageNumber="60" type="reference_group">
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<paragraph pageId="21" pageNumber="60">
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<taxonomicName class="Polypodiopsida" family="Saccolomataceae" genus="Orthiopteris" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Orthiopteris trichophylla" order="Polypodiales" pageId="21" pageNumber="60" phylum="Tracheophyta" rank="species" species="trichophylla">
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<pageBreakToken pageId="21" pageNumber="60" start="start">Orthiopteris</pageBreakToken>
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trichophylla
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</taxonomicName>
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Copel., Univ. Calif. Publ. Bot. 18: 217. 1942.
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</paragraph>
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<paragraph pageId="21" pageNumber="60">
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<taxonomicName authorityName="G. B. Nair" authorityYear="1992" baseAuthorityName="Copel." class="Polypodiopsida" family="Saccolomataceae" genus="Saccoloma" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Saccoloma trichophyllum" order="Polypodiales" pageId="21" pageNumber="60" phylum="Tracheophyta" rank="species" species="trichophyllum">Saccoloma trichophyllum</taxonomicName>
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(Copel.) G.B.Nair, J. Econ. Taxon. Bot. 16(3): 643.
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<normalizedToken originalValue="“1992”">"1992"</normalizedToken>
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[1994]. Type. Based on
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<taxonomicName class="Polypodiopsida" family="Saccolomataceae" genus="Orthiopteris" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Orthiopteris trichophylla" order="Polypodiales" pageId="21" pageNumber="60" phylum="Tracheophyta" rank="species" species="trichophylla">
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<emphasis italics="true" pageId="21" pageNumber="60">Orthiopteris trichophylla</emphasis>
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</taxonomicName>
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Copel.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="21" pageNumber="60" type="type">
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<paragraph pageId="21" pageNumber="60">Type.</paragraph>
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<paragraph pageId="21" pageNumber="60">
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PAPUA NEW GUINEA. Idenburg River, 15 km southwest of Bernhard camp, alt.1800 m, 1939,
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<emphasis italics="true" pageId="21" pageNumber="60">Brass 12027</emphasis>
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, (erroneously cited as
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<emphasis italics="true" pageId="21" pageNumber="60">Brass 12057</emphasis>
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in
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<bibRefCitation author="Copeland, EB" journalOrPublisher="University of California Publications in botany" pageId="23" pageNumber="62" pagination="217 - 226" refId="B21" refString="Copeland, EB, 1942. Filicum novarum CXXVIII Diagnoses. University of California Publications in botany 18: 217 - 226" title="Filicum novarum CXXVIII Diagnoses." volume="18" year="1942">Copeland 1942</bibRefCitation>
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, holo: MICH, 1190791* [http://quod.lib.umich.edu/h/herb2ic/x-mich1190791/mich1190791.tif]; iso: BM, 001044444, BO 1510941)
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</paragraph>
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</subSubSection>
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<subSubSection pageId="21" pageNumber="60" type="description">
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<paragraph pageId="21" pageNumber="60">Description.</paragraph>
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<paragraph pageId="21" pageNumber="60">
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Rhizome erect. Rhizome scales not seen. Fronds 120-160
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<normalizedToken originalValue="×">x</normalizedToken>
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40-50 cm; stipes stout, 40-50 cm long, 0.4-0.6 cm across (at base), dark brown; lamina deltoid, widest at base, tripinnate to quadripinnate, 100-120
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<normalizedToken originalValue="×">x</normalizedToken>
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40-50 cm, herbaceous, dark brownish green when dry, glabrous or with few scattered scales; pinnae at 30-35° to rachis, largest at base, overlapping, stalk 0.5 cm, including stalk 15-25
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<normalizedToken originalValue="×">x</normalizedToken>
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6-9 cm, lanceolate, first basiscopic pinnules of lowest pinnae enlarged; ultimate segments 1.0-1.5
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<normalizedToken originalValue="×">x</normalizedToken>
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0.5 cm, sessile or very short stalked, trapezoid, apex acute, margin with weak cartilaginous border; almost completely dissected to less than 0.2 mm from veins; lobes narrowly oblong, veins in lobes unbranched or with 1 fork, dark brown and not contrasting with lamina, percurrent. Scales on rachis mostly at base of pinnae or pinnule, very few on veins, hair-like. Sori lateral or apical on lobes, symmetric, slightly reflexed, slightly concave with lamina wings slightly recurved, 1.0-1.2
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<normalizedToken originalValue="×">x</normalizedToken>
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0.6-0.7 mm, funnelform, widest at mouth, not wider than the sterile lamina below sorus; inner indusium dull brown, firm, 4/5-2/3 as long as outer indusium, apex truncate; outer indusium truncate; sporangia 7-12 per sorus, capsule globose and rounded at apex, gradually narrowed toward base, indurated annulus cells 16-20,
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<normalizedToken originalValue="±">+/-</normalizedToken>
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equal; spores in polar view not studied, in lateral view.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="21" pageNumber="60" type="distribution">
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<paragraph pageId="21" pageNumber="60">Distribution.</paragraph>
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<paragraph pageId="21" pageNumber="60">Papua New Guinea (Idenburg River).</paragraph>
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<caption ID-Zenodo-Dep="964029" doi="10.3897/phytokeys.53.4955.figure9" httpUri="https://binary.pensoft.net/fig/47611" pageId="21" pageNumber="60" start="Figure 9" startId="F9">
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<paragraph pageId="21" pageNumber="60">
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<emphasis bold="true" pageId="21" pageNumber="60">Figure 9.</emphasis>
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Distribution of
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<taxonomicName class="Polypodiopsida" family="Saccolomataceae" genus="Orthiopteris" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Orthiopteris" order="Polypodiales" pageId="21" pageNumber="60" phylum="Tracheophyta" rank="genus">
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<emphasis italics="true" pageId="21" pageNumber="60">Orthiopteris</emphasis>
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</taxonomicName>
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species in East Malesia and the Pacific.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="21" pageNumber="60" type="ecology">
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<paragraph pageId="21" pageNumber="60">Ecology.</paragraph>
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<paragraph pageId="21" pageNumber="60">Terrestrial in bottom of ravines, rain forest, at 1800 m altitude.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="22" lastPageNumber="61" pageId="21" pageNumber="60" type="discussion">
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<paragraph pageId="21" pageNumber="60">Discussion.</paragraph>
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<paragraph lastPageId="22" lastPageNumber="61" pageId="21" pageNumber="60">
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This species can be mistaken for
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<taxonomicName class="Polypodiopsida" family="Saccolomataceae" genus="Orthiopteris" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Orthiopteris cicutarioides" order="Polypodiales" pageId="21" pageNumber="60" phylum="Tracheophyta" rank="species" species="cicutarioides">
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<emphasis italics="true" pageId="21" pageNumber="60">Orthiopteris cicutarioides</emphasis>
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</taxonomicName>
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because the frond morphology is similar, with fronds very finely dissected in both species. However,
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<taxonomicName class="Polypodiopsida" family="Saccolomataceae" genus="Orthiopteris" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Orthiopteris trichophylla" order="Polypodiales" pageId="21" pageNumber="60" phylum="Tracheophyta" rank="species" species="trichophylla">
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<emphasis italics="true" pageId="21" pageNumber="60">Orthiopteris trichophylla</emphasis>
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</taxonomicName>
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is more finely dissected with a single vein per lobe (similar to
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<taxonomicName class="Polypodiopsida" family="Saccolomataceae" genus="Orthiopteris" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Orthiopteris ferulacea" order="Polypodiales" pageId="21" pageNumber="60" phylum="Tracheophyta" rank="species" species="ferulacea">
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<emphasis italics="true" pageId="21" pageNumber="60">Orthiopteris ferulacea</emphasis>
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</taxonomicName>
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), while
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<taxonomicName class="Polypodiopsida" family="Saccolomataceae" genus="Orthiopteris" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Orthiopteris cicutarioides" order="Polypodiales" pageId="21" pageNumber="60" phylum="Tracheophyta" rank="species" species="cicutarioides">
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<emphasis italics="true" pageId="21" pageNumber="60">Orthiopteris cicutarioides</emphasis>
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</taxonomicName>
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has 2 or 3 one veins per lobe. The sori and indusia of
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<taxonomicName class="Polypodiopsida" family="Saccolomataceae" genus="Orthiopteris" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Orthiopteris trichophylla" order="Polypodiales" pageId="21" pageNumber="60" phylum="Tracheophyta" rank="species" species="trichophylla">
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<emphasis italics="true" pageId="21" pageNumber="60">Orthiopteris trichophylla</emphasis>
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</taxonomicName>
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resemble those of
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<taxonomicName authorityName="P. H. Hovenkamp & T. T. Luong" authorityYear="2015" baseAuthorityName="P. H. Hovenkamp & T. T. Luong" baseAuthorityYear="2015" class="Polypodiopsida" family="Saccolomataceae" genus="Orthiopteris" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Orthiopteris campylura subsp. var. var. kingii" order="Polypodiales" pageId="21" pageNumber="60" phylum="Tracheophyta" rank="variety" species="campylura" subSpecies="var." variety="kingii">Orthiopteris campylura var. kingii</taxonomicName>
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, being funnelform and slightly reflexed, and the possibility cannot be excluded that
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<taxonomicName class="Polypodiopsida" family="Saccolomataceae" genus="Orthiopteris" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Orthiopteris trichophylla" order="Polypodiales" pageId="21" pageNumber="60" phylum="Tracheophyta" rank="species" species="trichophylla">
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<emphasis italics="true" pageId="21" pageNumber="60">Orthiopteris trichophylla</emphasis>
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</taxonomicName>
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is locally derived from
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<taxonomicName authorityName="P. H. Hovenkamp & T. T. Luong" authorityYear="2015" baseAuthorityName="P. H. Hovenkamp & T. T. Luong" baseAuthorityYear="2015" class="Polypodiopsida" family="Saccolomataceae" genus="Orthiopteris" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Orthiopteris campylura subsp. var. var. kingii" order="Polypodiales" pageId="21" pageNumber="60" phylum="Tracheophyta" rank="variety" species="campylura" subSpecies="var." variety="kingii">Orthiopteris campylura var. kingii</taxonomicName>
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. The single specimen we examined (
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<emphasis italics="true" pageId="21" pageNumber="60">Brass 12239</emphasis>
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) differs slightly from the type (studied from scanned image of
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<emphasis italics="true" pageId="21" pageNumber="60">Brass 12027</emphasis>
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) by its shorter lobes. However, the sori and indusia are identical, and they were collected on the same day and at the same locality. Unlike in
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<taxonomicName class="Polypodiopsida" family="Saccolomataceae" genus="Orthiopteris" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Orthiopteris trichophylla" order="Polypodiales" pageId="21" pageNumber="60" phylum="Tracheophyta" rank="species" species="trichophylla">
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<emphasis italics="true" pageId="21" pageNumber="60">Orthiopteris trichophylla</emphasis>
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</taxonomicName>
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, which may represent a similar case of a locally derived taxon, both specimens of
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<taxonomicName class="Polypodiopsida" family="Saccolomataceae" genus="Orthiopteris" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Orthiopteris cicutarioides" order="Polypodiales" pageId="21" pageNumber="60" phylum="Tracheophyta" rank="species" species="cicutarioides">
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<emphasis italics="true" pageId="21" pageNumber="60">Orthiopteris cicutarioides</emphasis>
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</taxonomicName>
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look very productive with many ripe sporangia spreading over the mouth of the sorus. Brass noted that the
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<pageBreakToken pageId="22" pageNumber="61" start="start">species</pageBreakToken>
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was very common and abundant at the collecting site (Idenburg river), but we have not seen any later collections than
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<normalizedToken originalValue="Brass’s">Brass's</normalizedToken>
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in 1939. Perhaps it is just abundant locally, and it may well be currently threatened.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |