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<document ID-DOI="10.1206/352.1" ID-ISSN="0003-0090" ID-Zenodo-Dep="5405335" approvalRequired="503" approvalRequired_for_taxonomicNames="62" approvalRequired_for_textStreams="429" approvalRequired_for_treatments="12" checkinTime="1630345222680" checkinUser="tatiana" docAuthor="Nesbitt, Sterling J." docDate="2011" docId="357D771BFF9BFF91EDF6FBD4FEF8FE56" docLanguage="en" docName="B352.pdf" docOrigin="Bulletin of the American Museum of Natural History 2011 (352)" docSource="http://www.bioone.org/doi/abs/10.1206/352.1" docStyle="DocumentStyle:C5E2DA72A22EF33813C92A197453A310.3:BulAmeMusNatHis.2011-.journal_article.0cover" docStyleId="C5E2DA72A22EF33813C92A197453A310" docStyleName="BulAmeMusNatHis.2011-.journal_article.0cover" docStyleVersion="3" docTitle="Pisanosaurus mertii Casamiquela 1967" docType="treatment" docVersion="6" lastPageNumber="50" masterDocId="C9440F63FFABFFA0EF17FFF9FFBFFF8E" masterDocTitle="The Early Evolution Of Archosaurs: Relationships And The Origin Of Major Clades" masterLastPageNumber="292" masterPageNumber="1" pageNumber="49" updateTime="1631052925311" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title>The Early Evolution Of Archosaurs: Relationships And The Origin Of Major Clades</mods:title>
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<mods:namePart>Nesbitt, Sterling J.</mods:namePart>
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<mods:date>2011</mods:date>
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<mods:number>2011-04-29</mods:number>
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<mods:number>2011</mods:number>
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<mods:number>352</mods:number>
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<mods:identifier type="DOI">10.1206/352.1</mods:identifier>
<mods:identifier type="ISSN">0003-0090</mods:identifier>
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<heading box="[737,1193,1068,1090]" centered="true" fontSize="9" level="2" pageId="48" pageNumber="49" reason="2">
<taxonomicName authority="Casamiquela, 1967" authorityName="Casamiquela" authorityYear="1967" box="[737,1193,1068,1090]" class="Reptilia" family="Pisanosauridae" genus="Pisanosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="mertii">
<emphasis box="[737,962,1069,1090]" italics="true" pageId="48" pageNumber="49">Pisanosaurus mertii</emphasis>
<bibRefCitation author="Casamiquela, R. M." box="[971,1193,1068,1090]" pageId="48" pageNumber="49" pagination="47 - 64" refId="ref186758" refString="Casamiquela, R. M. 1967. Un nuevo dinosaurio ornitisquio Triasico (Pisanosaurus mertii; Ornithopoda) de la Formacion Ischigualasto, Argentina. Ameghiniana 5: 47 - 64." type="journal article" year="1967">Casamiquela, 1967</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
<paragraph blockId="48.[701,1228,1113,1759]" pageId="48" pageNumber="49">
AGE: Late Carnianearly Norian, Late Triassic (
<bibRefCitation author="Sereno, P. C. &amp; C. A. Forster &amp; R. R. Rogers &amp; A. M. Monetta" box="[815,1060,1144,1166]" pageId="48" pageNumber="49" pagination="64 - 66" refId="ref200535" refString="Sereno, P. C., C. A. Forster, R. R. Rogers, and A. M. Monetta. 1993. Primitive dinosaur skeleton from Argentina and the early evolution of Dinosauria. Nature 361: 64 - 66." type="journal article" year="1993">Rogers et al., 1993</bibRefCitation>
, adjusted for the new Triassic timescale of
<bibRefCitation author="Muttoni, G." pageId="48" pageNumber="49" pagination="1043 - 1058" refId="ref194940" refString="Muttoni, G., et al. 2004. Tethyan magnetostratigraphy from Pizzi Mondello (Sicily) and correlation to the Late Triassic Newark astrochronological polarity time scale. Geological Society of America Bulletin 116: 1043 - 1058." type="journal article" year="2004">Muttoni et al., 2004</bibRefCitation>
).
</paragraph>
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<subSubSection pageId="48" pageNumber="49" type="description">
<paragraph blockId="48.[701,1228,1113,1759]" pageId="48" pageNumber="49">
OCCURRENCE: Middle portion of the Ischigualasto Formation, Ischigualasto basin,
<collectingCountry box="[751,872,1293,1315]" name="Argentina" pageId="48" pageNumber="49">Argentina</collectingCountry>
.
</paragraph>
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<subSubSection pageId="48" pageNumber="49" type="materials_examined">
<paragraph blockId="48.[701,1228,1113,1759]" pageId="48" pageNumber="49">
<typeStatus box="[730,858,1321,1345]" pageId="48" pageNumber="49">HOLOTYPE</typeStatus>
: PVL 2577, tooth-bearing elements, vertebrae, incomplete hind limb, impression of the pelvis.
</paragraph>
</subSubSection>
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<paragraph blockId="48.[701,1228,1113,1759]" lastBlockId="49.[93,619,213,472]" lastPageId="49" lastPageNumber="50" pageId="48" pageNumber="49">
REMARKS: Since its discovery,
<taxonomicName class="Reptilia" family="Pisanosauridae" genus="Pisanosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="48" pageNumber="49" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="48" pageNumber="49">Pisanosaurus</emphasis>
</taxonomicName>
was considered the most primitive ornithischian (
<bibRefCitation author="Casamiquela, R. M." box="[867,1089,1471,1493]" pageId="48" pageNumber="49" pagination="47 - 64" refId="ref186758" refString="Casamiquela, R. M. 1967. Un nuevo dinosaurio ornitisquio Triasico (Pisanosaurus mertii; Ornithopoda) de la Formacion Ischigualasto, Argentina. Ameghiniana 5: 47 - 64." type="journal article" year="1967">Casamiquela, 1967</bibRefCitation>
;
<bibRefCitation author="Bonaparte, J. F." pageId="48" pageNumber="49" pagination="808 - 820" refId="ref184976" refString="Bonaparte, J. F. 1976. Pisanosaurus mertii Casimiquela and the origin of the Ornithischia. Journal of Paleontology 50: 808 - 820." type="journal article" year="1976">Bonaparte, 1976</bibRefCitation>
;
<bibRefCitation author="Weishampel, D. B. &amp; L. M. Witmer" box="[770,1130,1501,1523]" pageId="48" pageNumber="49" pagination="486 - 497" refId="ref202678" refString="Weishampel, D. B., and L. M. Witmer. 1990. Heterodontosauridae. In D. B. Weishampel, P. Dodson, and H. Osmolska (editors), The Dinosauria: 486 - 497. Berkeley: University of California Press." type="book chapter" year="1990">Weishampel and Witmer, 1990</bibRefCitation>
;
<bibRefCitation author="Sereno, P. C." pageId="48" pageNumber="49" pagination="168 - 197" refId="ref200069" refString="Sereno, P. C. 1991 b. Lesothosaurus, '' Fabrosaurids, '' and the early evolution of Ornithischia. Journal of Vertebrate Paleontology 11: 168 - 197." type="journal article" year="1991">Sereno, 1991b</bibRefCitation>
;
<bibRefCitation author="Irmis, R. B. &amp; W. G. Parker &amp; S. J. Nesbitt &amp; J. Liu" box="[786,1000,1530,1552]" pageId="48" pageNumber="49" pagination="3 - 22" refId="ref192454" refString="Irmis, R. B., W. G. Parker, S. J. Nesbitt, and J. Liu. 2007 a. Ornithischian dinosaurs: the Triassic record. Historical Biology 18: 3 - 22." type="journal article" year="2007">Irmis et al., 2007a</bibRefCitation>
;
<bibRefCitation author="Butler, R. J. &amp; R. M. H. Smith &amp; D. B. Norman" box="[1012,1223,1530,1552]" pageId="48" pageNumber="49" pagination="2041 - 2046" refId="ref186235" refString="Butler, R. J., R. M. H. Smith, and D. B. Norman. 2007. A primitive ornithischian dinosaur from the Late Triassic of South Africa, and the early evolution and diversification of Ornithischia. Proceedings of the Royal Society of London B Biological Sciences 274: 2041 - 2046." type="journal article" year="2007">Butler et al., 2007</bibRefCitation>
,
<bibRefCitation author="Butler, R. J. &amp; L. B. Porro &amp; D. B. Norman" box="[701,777,1560,1582]" pageId="48" pageNumber="49" pagination="702 - 711" refId="ref186334" refString="Butler, R. J., L. B. Porro, and D. B. Norman. 2008 b. A juvenile skull of the primitive ornithischian dinosaur Heterodontosaurus tucki from the ' Stormberg' of southern Africa. Journal of Vertebrate Paleontology 28: 702 - 711." type="journal article" year="2008">2008b</bibRefCitation>
). However, questions about the association of the material plagued certainty regarding the validity of the taxon. Our current understanding of the taxon can be better understood only with the discovery of a new specimen. However,
<taxonomicName box="[955,1104,1708,1729]" class="Reptilia" family="Pisanosauridae" genus="Pisanosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="48" pageNumber="49" phylum="Chordata" rank="genus">
<emphasis box="[955,1104,1708,1729]" italics="true" pageId="48" pageNumber="49">Pisanosaurus</emphasis>
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is almost always found as the basalmost member of
<taxonomicName box="[93,237,213,235]" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="49" pageNumber="50" phylum="Chordata" rank="order">Ornithischia</taxonomicName>
(
<bibRefCitation author="Langer, M. C. &amp; M. J. Benton" box="[249,539,213,235]" pageId="49" pageNumber="50" pagination="309 - 358" refId="ref193175" refString="Langer, M. C., and M. J. Benton. 2006. Early dinosaurs: a phylogenetic study. Journal of Systematic Palaeontology 4: 309 - 358." type="journal article" year="2006">Langer and Benton, 2006</bibRefCitation>
;
<bibRefCitation author="Butler, R. J. &amp; R. M. H. Smith &amp; D. B. Norman" pageId="49" pageNumber="50" pagination="2041 - 2046" refId="ref186235" refString="Butler, R. J., R. M. H. Smith, and D. B. Norman. 2007. A primitive ornithischian dinosaur from the Late Triassic of South Africa, and the early evolution and diversification of Ornithischia. Proceedings of the Royal Society of London B Biological Sciences 274: 2041 - 2046." type="journal article" year="2007">Butler et al., 2007</bibRefCitation>
,
<bibRefCitation author="Butler, R. J. &amp; L. B. Porro &amp; D. B. Norman" box="[229,301,242,264]" pageId="49" pageNumber="50" pagination="702 - 711" refId="ref186334" refString="Butler, R. J., L. B. Porro, and D. B. Norman. 2008 b. A juvenile skull of the primitive ornithischian dinosaur Heterodontosaurus tucki from the ' Stormberg' of southern Africa. Journal of Vertebrate Paleontology 28: 702 - 711." type="journal article" year="2008">2008b</bibRefCitation>
;
<bibRefCitation author="Irmis, R. B. &amp; W. G. Parker &amp; S. J. Nesbitt &amp; J. Liu" box="[311,516,242,264]" pageId="49" pageNumber="50" pagination="3 - 22" refId="ref192454" refString="Irmis, R. B., W. G. Parker, S. J. Nesbitt, and J. Liu. 2007 a. Ornithischian dinosaurs: the Triassic record. Historical Biology 18: 3 - 22." type="journal article" year="2007">Irmis et al., 2007a</bibRefCitation>
) because of the combination of ornithischian synapomorphies and archosaur plesiomorphies, such as the anteroventrally directed pubis.
</paragraph>
</subSubSection>
<subSubSection pageId="49" pageNumber="50" type="key">
<paragraph blockId="49.[93,619,213,472]" pageId="49" pageNumber="50">
KEY REFERENCES:
<bibRefCitation author="Casamiquela, R. M." box="[376,615,361,383]" pageId="49" pageNumber="50" pagination="47 - 64" refId="ref186758" refString="Casamiquela, R. M. 1967. Un nuevo dinosaurio ornitisquio Triasico (Pisanosaurus mertii; Ornithopoda) de la Formacion Ischigualasto, Argentina. Ameghiniana 5: 47 - 64." type="journal article" year="1967">Casamiquela, 1967</bibRefCitation>
;
<bibRefCitation author="Bonaparte, J. F." box="[93,294,391,413]" pageId="49" pageNumber="50" pagination="808 - 820" refId="ref184976" refString="Bonaparte, J. F. 1976. Pisanosaurus mertii Casimiquela and the origin of the Ornithischia. Journal of Paleontology 50: 808 - 820." type="journal article" year="1976">Bonaparte, 1976</bibRefCitation>
;
<bibRefCitation author="Weishampel, D. B. &amp; L. M. Witmer" pageId="49" pageNumber="50" pagination="486 - 497" refId="ref202678" refString="Weishampel, D. B., and L. M. Witmer. 1990. Heterodontosauridae. In D. B. Weishampel, P. Dodson, and H. Osmolska (editors), The Dinosauria: 486 - 497. Berkeley: University of California Press." type="book chapter" year="1990">Weishampel and Witmer, 1990</bibRefCitation>
;
<bibRefCitation author="Sereno, P. C." box="[173,350,420,442]" pageId="49" pageNumber="50" pagination="168 - 197" refId="ref200069" refString="Sereno, P. C. 1991 b. Lesothosaurus, '' Fabrosaurids, '' and the early evolution of Ornithischia. Journal of Vertebrate Paleontology 11: 168 - 197." type="journal article" year="1991">Sereno, 1991b</bibRefCitation>
;
<bibRefCitation author="Irmis, R. B. &amp; W. G. Parker &amp; S. J. Nesbitt &amp; J. Liu" box="[372,614,420,442]" pageId="49" pageNumber="50" pagination="3 - 22" refId="ref192454" refString="Irmis, R. B., W. G. Parker, S. J. Nesbitt, and J. Liu. 2007 a. Ornithischian dinosaurs: the Triassic record. Historical Biology 18: 3 - 22." type="journal article" year="2007">Irmis et al., 2007a</bibRefCitation>
;
<bibRefCitation author="Butler, R. J. &amp; L. B. Porro &amp; D. B. Norman" box="[93,322,450,472]" pageId="49" pageNumber="50" pagination="702 - 711" refId="ref186334" refString="Butler, R. J., L. B. Porro, and D. B. Norman. 2008 b. A juvenile skull of the primitive ornithischian dinosaur Heterodontosaurus tucki from the ' Stormberg' of southern Africa. Journal of Vertebrate Paleontology 28: 702 - 711." type="journal article" year="2008">Butler et al., 2008b</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
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