treatments-xml/data/35/7D/77/357D771BFF83FF89EDE2FEB1FF23FEAA.xml
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<document ID-DOI="10.1206/352.1" ID-ISSN="0003-0090" ID-Zenodo-Dep="5405335" approvalRequired="503" approvalRequired_for_taxonomicNames="62" approvalRequired_for_textStreams="429" approvalRequired_for_treatments="12" checkinTime="1630345222680" checkinUser="tatiana" docAuthor="Nesbitt, Sterling J." docDate="2011" docId="357D771BFF83FF89EDE2FEB1FF23FEAA" docLanguage="en" docName="B352.pdf" docOrigin="Bulletin of the American Museum of Natural History 2011 (352)" docSource="http://www.bioone.org/doi/abs/10.1206/352.1" docStyle="DocumentStyle:C5E2DA72A22EF33813C92A197453A310.3:BulAmeMusNatHis.2011-.journal_article.0cover" docStyleId="C5E2DA72A22EF33813C92A197453A310" docStyleName="BulAmeMusNatHis.2011-.journal_article.0cover" docStyleVersion="3" docTitle="Terrestrisuchus gracilis Crush 1984" docType="treatment" docVersion="6" lastPageNumber="42" masterDocId="C9440F63FFABFFA0EF17FFF9FFBFFF8E" masterDocTitle="The Early Evolution Of Archosaurs: Relationships And The Origin Of Major Clades" masterLastPageNumber="292" masterPageNumber="1" pageNumber="41" updateTime="1631052925311" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title>The Early Evolution Of Archosaurs: Relationships And The Origin Of Major Clades</mods:title>
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<mods:namePart>Nesbitt, Sterling J.</mods:namePart>
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<mods:date>2011</mods:date>
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<mods:number>2011-04-29</mods:number>
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<mods:number>2011</mods:number>
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<paragraph blockId="40.[757,1173,328,351]" box="[757,1173,328,351]" pageId="40" pageNumber="41">
<heading box="[757,1173,328,351]" centered="true" fontSize="9" level="2" pageId="40" pageNumber="41" reason="2">
<taxonomicName authority="Crush, 1984" authorityName="Crush" authorityYear="1984" box="[757,1173,328,351]" class="Reptilia" family="Saltoposuchidae" genus="Terrestrisuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prolacertiformes" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="gracilis">
<emphasis box="[757,1021,328,351]" italics="true" pageId="40" pageNumber="41">Terrestrisuchus gracilis</emphasis>
<bibRefCitation author="Crush, P. J." box="[1031,1173,329,351]" pageId="40" pageNumber="41" pagination="131 - 157" refId="ref188115" refString="Crush, P. J. 1984. A Late upper Triassic sphenosuchid crocodilian from Wales. Palaeontology 27: 131 - 157." type="journal article" year="1984">Crush, 1984</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
<paragraph blockId="40.[701,1228,373,1759]" pageId="40" pageNumber="41">
AGE:?Rhaetian, Late Triassic (
<bibRefCitation author="Robinson, P. L." pageId="40" pageNumber="41" pagination="283 - 293" refId="ref198843" refString="Robinson, P. L. 1957 a. An unusual sauropsid dentition. Zoological Journal of the Linnean Society 43: 283 - 293." type="journal article" year="1957">Robinson 1957a</bibRefCitation>
,
<bibRefCitation author="Robinson, P. L." box="[842,915,404,426]" pageId="40" pageNumber="41" pagination="260 - 282" refId="ref198869" refString="Robinson, P. L. 1957 b. The Mesozoic fissures of the Bristol Channel area and their vertebrate fauna. Zoological Journal of the Linnean Society 43: 260 - 282." type="journal article" year="1957">1957b</bibRefCitation>
,
<bibRefCitation author="Whiteside, D. I. &amp; J. E. A. Marshall" pageId="40" pageNumber="41" pagination="105 - 147" refId="ref203020" refString="Whiteside, D. I., and J. E. A. Marshall. 2008. The age, fauna and palaeoenvironment of the Late Triassic fissure deposits of Tytherington, south Gloucestershire, UK. Geological Magazine 145: 105 - 147." type="journal article" year="2008">Whiteside and Marshall, 2008</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="40" pageNumber="41" type="description">
<paragraph blockId="40.[701,1228,373,1759]" pageId="40" pageNumber="41">
OCCURRENCE: Fissure fills in the Carboniferous limestone of the Pant-y-ffynon Quarry, Cowbridge, Glamorgan,
<collectingCountry box="[1036,1110,523,545]" name="United Kingdom" pageId="40" pageNumber="41">Wales</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection pageId="40" pageNumber="41" type="materials_examined">
<paragraph blockId="40.[701,1228,373,1759]" pageId="40" pageNumber="41">
<typeStatus box="[730,858,552,576]" pageId="40" pageNumber="41">HOLOTYPE</typeStatus>
: BMNH R7557 (formerly P 47/ 21 and counter part P 47/22).
</paragraph>
<paragraph blockId="40.[701,1228,373,1759]" box="[730,1202,611,635]" pageId="40" pageNumber="41">
REFERRED MATERIAL: See
<bibRefCitation author="Crush, P. J." box="[1052,1197,613,635]" pageId="40" pageNumber="41" pagination="131 - 157" refId="ref188115" refString="Crush, P. J. 1984. A Late upper Triassic sphenosuchid crocodilian from Wales. Palaeontology 27: 131 - 157." type="journal article" year="1984">Crush, 1984</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="40" pageNumber="41" type="discussion">
<paragraph blockId="40.[701,1228,373,1759]" pageId="40" pageNumber="41">
REMARKS:
<taxonomicName baseAuthorityName="Crush" baseAuthorityYear="1984" box="[867,1041,643,666]" class="Reptilia" family="Saltoposuchidae" genus="Terrestrisuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prolacertiformes" pageId="40" pageNumber="41" phylum="Chordata" rank="genus">
<emphasis box="[867,1041,643,666]" italics="true" pageId="40" pageNumber="41">Terrestrisuchus</emphasis>
</taxonomicName>
was named for and based on material collected from fissure fills in a Carboniferous limestone in
<collectingCountry box="[1150,1224,703,725]" name="United Kingdom" pageId="40" pageNumber="41">Wales</collectingCountry>
. The abundant taxon is known from dozens of specimens, from articulated and disarticulated crania, and postcrania. A few of the three-dimensionally preserved bones were prepared out of the matrix and formed the basis of the description by
<bibRefCitation author="Crush, P. J." box="[1060,1224,880,902]" pageId="40" pageNumber="41" pagination="131 - 157" refId="ref188115" refString="Crush, P. J. 1984. A Late upper Triassic sphenosuchid crocodilian from Wales. Palaeontology 27: 131 - 157." type="journal article" year="1984">Crush (1984)</bibRefCitation>
. Although much of the skull was described by
<bibRefCitation author="Crush, P. J." box="[701,855,939,961]" pageId="40" pageNumber="41" pagination="131 - 157" refId="ref188115" refString="Crush, P. J. 1984. A Late upper Triassic sphenosuchid crocodilian from Wales. Palaeontology 27: 131 - 157." type="journal article" year="1984">Crush (1984)</bibRefCitation>
, portions of the skull, including much of the braincase, nasals, and premaxilla, remain unknown. Originally, the
<typeStatus pageId="40" pageNumber="41">holotype</typeStatus>
and referred material were housed at University College, London, but they were transferred to and reside at the Natural History Museum (BMNH).
</paragraph>
<paragraph blockId="40.[701,1228,373,1759]" pageId="40" pageNumber="41">
Soon after
<taxonomicName baseAuthorityName="Crush" baseAuthorityYear="1984" box="[888,1062,1146,1169]" class="Reptilia" family="Saltoposuchidae" genus="Terrestrisuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prolacertiformes" pageId="40" pageNumber="41" phylum="Chordata" rank="genus">
<emphasis box="[888,1062,1146,1169]" italics="true" pageId="40" pageNumber="41">Terrestrisuchus</emphasis>
</taxonomicName>
was named,
<bibRefCitation author="Benton, M. J. &amp; J. M. Clark" box="[701,1031,1176,1198]" pageId="40" pageNumber="41" pagination="295 - 338" refId="ref184635" refString="Benton, M. J., and J. M. Clark. 1988. Archosaur phylogeny and the relationships of the Crocodylia. In M. J. Benton (editor), The phylogeny and classification of the tetrapods. Vol. 1. Amphibians and reptiles, 295 - 338. Oxford: Clarendon Press." type="book chapter" year="1988">Benton and Clark (1988)</bibRefCitation>
proposed that the taxon may be synonymous with
<taxonomicName authorityName="von Huene" authorityYear="1921" class="Reptilia" family="Saltoposuchidae" genus="Saltoposuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prolacertiformes" pageId="40" pageNumber="41" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="40" pageNumber="41">Saltoposuchus</emphasis>
</taxonomicName>
from the Stubensandstein (Norian) of
<collectingRegion box="[852,1094,1265,1287]" country="Germany" name="Baden-Wuerttemberg" pageId="40" pageNumber="41">Baden-Wurttemberg</collectingRegion>
,
<collectingCountry box="[1110,1224,1265,1286]" name="Germany" pageId="40" pageNumber="41">Germany</collectingCountry>
.
<bibRefCitation author="Benton, M. J. &amp; J. M. Clark" box="[701,1026,1294,1316]" pageId="40" pageNumber="41" pagination="295 - 338" refId="ref184635" refString="Benton, M. J., and J. M. Clark. 1988. Archosaur phylogeny and the relationships of the Crocodylia. In M. J. Benton (editor), The phylogeny and classification of the tetrapods. Vol. 1. Amphibians and reptiles, 295 - 338. Oxford: Clarendon Press." type="book chapter" year="1988">Benton and Clark (1988)</bibRefCitation>
rightfully criticized the single maxillary character cited by
<bibRefCitation author="Crush, P. J." box="[756,925,1353,1375]" pageId="40" pageNumber="41" pagination="131 - 157" refId="ref188115" refString="Crush, P. J. 1984. A Late upper Triassic sphenosuchid crocodilian from Wales. Palaeontology 27: 131 - 157." type="journal article" year="1984">Crush (1984)</bibRefCitation>
to separate the two taxa.
<bibRefCitation author="Sereno, P. C. &amp; R. Wild" box="[781,1101,1383,1405]" pageId="40" pageNumber="41" pagination="435 - 458" refId="ref200310" refString="Sereno, P. C., and R. Wild. 1992. Procompsognathus: theropod, '' thecodont' ' or both? Journal of Vertebrate Paleontology 12: 435 - 458." type="journal article" year="1992">Sereno and Wild (1992)</bibRefCitation>
defended the position that the two taxa should be separated, but as demonstrated by
<bibRefCitation author="Clark, J. M. &amp; H. D. Sues &amp; D. S. Berman" pageId="40" pageNumber="41" pagination="683 - 704" refId="ref187498" refString="Clark, J. M., H. D. Sues, and D. S. Berman. 2000. A new specimen of Hesperosuchus agilis from the Upper Triassic of New Mexico and the interrelationships of basal crocodylomorph archosaurs. Journal of Vertebrate Paleontology 20: 683 - 704." type="journal article" year="2000">Clark et al. (2000)</bibRefCitation>
, many of the differences named are noncomparable between the two taxa.
<bibRefCitation author="Clark, J. M. &amp; H. D. Sues &amp; D. S. Berman" box="[701,935,1530,1553]" pageId="40" pageNumber="41" pagination="683 - 704" refId="ref187498" refString="Clark, J. M., H. D. Sues, and D. S. Berman. 2000. A new specimen of Hesperosuchus agilis from the Upper Triassic of New Mexico and the interrelationships of basal crocodylomorph archosaurs. Journal of Vertebrate Paleontology 20: 683 - 704." type="journal article" year="2000">Clark et al. (2000)</bibRefCitation>
cited a few differences but were unsure in the end whether the taxa were different. Most recently,
<bibRefCitation author="Allen, D." pageId="40" pageNumber="41" pagination="29" refId="ref183508" refString="Allen, D. 2003. When Terrestrisuchus gracilis reaches puberty it becomes Saltoposuchus connectens! Journal of Vertebrate Paleontology suppl. to 23: 29 A pp." type="journal article" year="2003">Allen (2003)</bibRefCitation>
suggested that
<taxonomicName baseAuthorityName="Crush" baseAuthorityYear="1984" box="[970,1144,1618,1641]" class="Reptilia" family="Saltoposuchidae" genus="Terrestrisuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prolacertiformes" pageId="40" pageNumber="41" phylum="Chordata" rank="genus">
<emphasis box="[970,1144,1618,1641]" italics="true" pageId="40" pageNumber="41">Terrestrisuchus</emphasis>
</taxonomicName>
was a juvenile of
<taxonomicName authorityName="von Huene" authorityYear="1921" box="[841,1002,1648,1671]" class="Reptilia" family="Saltoposuchidae" genus="Saltoposuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prolacertiformes" pageId="40" pageNumber="41" phylum="Chordata" rank="genus">
<emphasis box="[841,1002,1648,1671]" italics="true" pageId="40" pageNumber="41">Saltoposuchus</emphasis>
</taxonomicName>
. Given the uncertainties of the taxonomy of the two taxa, I score only material of
<taxonomicName baseAuthorityName="Crush" baseAuthorityYear="1984" box="[1000,1174,1707,1730]" class="Reptilia" family="Saltoposuchidae" genus="Terrestrisuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prolacertiformes" pageId="40" pageNumber="41" phylum="Chordata" rank="genus">
<emphasis box="[1000,1174,1707,1730]" italics="true" pageId="40" pageNumber="41">Terrestrisuchus</emphasis>
</taxonomicName>
described by
<bibRefCitation author="Crush, P. J." box="[837,996,1737,1759]" pageId="40" pageNumber="41" pagination="131 - 157" refId="ref188115" refString="Crush, P. J. 1984. A Late upper Triassic sphenosuchid crocodilian from Wales. Palaeontology 27: 131 - 157." type="journal article" year="1984">Crush (1984)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="41" pageNumber="42" type="key">
<paragraph blockId="41.[93,620,211,292]" pageId="41" pageNumber="42">
KEY REFERENCES:
<bibRefCitation author="Crush, P. J." box="[365,518,213,235]" pageId="41" pageNumber="42" pagination="131 - 157" refId="ref188115" refString="Crush, P. J. 1984. A Late upper Triassic sphenosuchid crocodilian from Wales. Palaeontology 27: 131 - 157." type="journal article" year="1984">Crush, 1984</bibRefCitation>
;
<bibRefCitation author="Sereno, P. C. &amp; R. Wild" pageId="41" pageNumber="42" pagination="435 - 458" refId="ref200310" refString="Sereno, P. C., and R. Wild. 1992. Procompsognathus: theropod, '' thecodont' ' or both? Journal of Vertebrate Paleontology 12: 435 - 458." type="journal article" year="1992">Sereno and Wild, 1992</bibRefCitation>
;
<bibRefCitation author="Clark, J. M. &amp; H. D. Sues &amp; D. S. Berman" box="[306,531,242,264]" pageId="41" pageNumber="42" pagination="683 - 704" refId="ref187498" refString="Clark, J. M., H. D. Sues, and D. S. Berman. 2000. A new specimen of Hesperosuchus agilis from the Upper Triassic of New Mexico and the interrelationships of basal crocodylomorph archosaurs. Journal of Vertebrate Paleontology 20: 683 - 704." type="journal article" year="2000">Clark et al., 2000</bibRefCitation>
;
<bibRefCitation author="Allen, D." pageId="41" pageNumber="42" pagination="29" refId="ref183508" refString="Allen, D. 2003. When Terrestrisuchus gracilis reaches puberty it becomes Saltoposuchus connectens! Journal of Vertebrate Paleontology suppl. to 23: 29 A pp." type="journal article" year="2003">Allen, 2003</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>