261 lines
31 KiB
XML
261 lines
31 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.50.504" ID-GBIF-Dataset="2501a246-28f1-48c4-9a23-433c3c71cb22" ID-PMC="PMC3088018" ID-Pensoft-Pub="1313-2970-50-29" ID-PubMed="21594115" ModsDocAuthor="" ModsDocDate="2010" ModsDocID="1313-2970-50-29" ModsDocOrigin="ZooKeys 50" ModsDocTitle="The centipede genus Eupolybothrus Verhoeff, 1907 (Chilopoda: Lithobiomorpha: Lithobiidae) in North Africa, a cybertaxonomic revision, with a key to all species in the genus and the first use of DNA barcoding for the group" checkinTime="1451251067348" checkinUser="pensoft" docAuthor="Stoev, Pavel, Akkari, Nesrine, Zapparoli, Marzio, Porco, David, Enghoff, Henrik, Edgecombe, Gregory D., Georgiev, Teodor & Penev, Lyubomir" docDate="2010" docId="9C7EECF71E65861D9F6A13A28AB2069C" docLanguage="en" docName="ZooKeys 50: 29-77" docOrigin="ZooKeys 50" docSource="http://dx.doi.org/10.3897/zookeys.50.504" docTitle="Eupolybothrus Verhoeff 1907" docType="treatment" docVersion="4" lastPageNumber="34" masterDocId="8945F465A765FFFEFFE1FF8CFFF44F7A" masterDocTitle="The centipede genus Eupolybothrus Verhoeff, 1907 (Chilopoda: Lithobiomorpha: Lithobiidae) in North Africa, a cybertaxonomic revision, with a key to all species in the genus and the first use of DNA barcoding for the group" masterLastPageNumber="77" masterPageNumber="29" pageNumber="34" updateTime="1668160318181" updateUser="ExternalLinkService">
|
|
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
|
<mods:titleInfo>
|
|
<mods:title>The centipede genus Eupolybothrus Verhoeff, 1907 (Chilopoda: Lithobiomorpha: Lithobiidae) in North Africa, a cybertaxonomic revision, with a key to all species in the genus and the first use of DNA barcoding for the group</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Stoev, Pavel</mods:namePart>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Akkari, Nesrine</mods:namePart>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Zapparoli, Marzio</mods:namePart>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Porco, David</mods:namePart>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Enghoff, Henrik</mods:namePart>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Edgecombe, Gregory D.</mods:namePart>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Georgiev, Teodor</mods:namePart>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Penev, Lyubomir</mods:namePart>
|
|
</mods:name>
|
|
<mods:typeOfResource>text</mods:typeOfResource>
|
|
<mods:relatedItem type="host">
|
|
<mods:titleInfo>
|
|
<mods:title>ZooKeys</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:part>
|
|
<mods:date>2010</mods:date>
|
|
<mods:detail type="volume">
|
|
<mods:number>50</mods:number>
|
|
</mods:detail>
|
|
<mods:extent unit="page">
|
|
<mods:start>29</mods:start>
|
|
<mods:end>77</mods:end>
|
|
</mods:extent>
|
|
</mods:part>
|
|
</mods:relatedItem>
|
|
<mods:location>
|
|
<mods:url>http://dx.doi.org/10.3897/zookeys.50.504</mods:url>
|
|
</mods:location>
|
|
<mods:classification>journal article</mods:classification>
|
|
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.50.504</mods:identifier>
|
|
<mods:identifier type="Pensoft-Pub">1313-2970-50-29</mods:identifier>
|
|
</mods:mods>
|
|
<treatment ID-GBIF-Taxon="159359447" LSID="urn:lsid:plazi:treatment:9C7EECF71E65861D9F6A13A28AB2069C" httpUri="http://treatment.plazi.org/id/9C7EECF71E65861D9F6A13A28AB2069C" lastPageId="6" lastPageNumber="34" pageId="5" pageNumber="34">
|
|
<subSubSection pageId="5" pageNumber="34" type="nomenclature">
|
|
<paragraph pageId="5" pageNumber="34">
|
|
<taxonomicName LSID="urn:lsid:zoobank.org:act:9A79187E-FC75-4CB7-B634-ABBD4B471D2A" authority="Verhoeff, 1907" authorityName="Verhoeff" authorityYear="1907" class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolybothrus" order="Lithobiomorpha" pageId="5" pageNumber="34" phylum="Arthropoda" rank="genus">Eupolybothrus Verhoeff, 1907</taxonomicName>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="5" pageNumber="34" type="type species">
|
|
<paragraph pageId="5" pageNumber="34">Type species.</paragraph>
|
|
<paragraph pageId="5" pageNumber="34">
|
|
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Lithobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lithobius grossipes" order="Lithobiomorpha" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="grossipes">Lithobius grossipes</taxonomicName>
|
|
C.L. Koch, 1847, by subsequent designation of
|
|
<bibRefCitation author="Chamberlin, RV" journalOrPublisher="Bulletin of the Museum of Comparative Zoology" pageId="35" pageNumber="64" pagination="385 - 437" title="The Ethopolidae of America north of Mexico." url="http://www.biodiversitylibrary.org/page/2810046549" volume="57" year="1925">Chamberlin (1925)</bibRefCitation>
|
|
. Type locality: Triest, Italy.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="5" pageNumber="34" type="diagnosis">
|
|
<paragraph pageId="5" pageNumber="34">Diagnosis.</paragraph>
|
|
<paragraph pageId="5" pageNumber="34">
|
|
Medium- to large-sized
|
|
<taxonomicName family="Lithobiidae" lsidName="" pageId="5" pageNumber="34" rank="family">Lithobiidae</taxonomicName>
|
|
(body length 16-48 mm) with numerous irregularly arranged pores on the last four pairs of coxae; antennal articles always more than 20, from 38 to around 80; ocelli numerous, usually from 1+16 to 1+24, absent or reduced in some troglobitic species; porodont situated laterad to the forcipular coxosternal teeth; tergites with or without posterior triangular projections, tarsi of all legs bipartite; forcipular coxosternite with 5+5-14+14 teeth (usually from 7+7 to 10+10); female gonopod with 2 spurs and simple claw, male gonopods single or bipartite.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="5" pageNumber="34" type="assigned valid species (20)">
|
|
<paragraph pageId="5" pageNumber="34">Assigned valid species (20).</paragraph>
|
|
<paragraph pageId="5" pageNumber="34">
|
|
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolybothrus (Eupolybothrus) andreevi" order="Lithobiomorpha" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="andreevi" subGenus="Eupolybothrus">Eupolybothrus (Eupolybothrus) andreevi</taxonomicName>
|
|
Matic, 1964,
|
|
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolybothrus (Schizopolybothrus) caesar" order="Lithobiomorpha" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="caesar" subGenus="Schizopolybothrus">Eupolybothrus (Schizopolybothrus) caesar</taxonomicName>
|
|
(Verhoeff, 1899),
|
|
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolybothrus (Propolybothrus) dolops" order="Lithobiomorpha" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="dolops" subGenus="Propolybothrus">Eupolybothrus (Propolybothrus) dolops</taxonomicName>
|
|
Zapparoli, 1998,
|
|
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolybothrus (Schizopolybothrus) excellens" order="Lithobiomorpha" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="excellens" subGenus="Schizopolybothrus">Eupolybothrus (Schizopolybothrus) excellens</taxonomicName>
|
|
(Silvestri, 1894),
|
|
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolybothrus (Eupolybothrus) fasciatus" order="Lithobiomorpha" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="fasciatus" subGenus="Eupolybothrus">Eupolybothrus (Eupolybothrus) fasciatus</taxonomicName>
|
|
(Newport, 1845),
|
|
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolybothrus (Eupolybothrus) gloriastygis" order="Lithobiomorpha" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="gloriastygis" subGenus="Eupolybothrus">Eupolybothrus (Eupolybothrus) gloriastygis</taxonomicName>
|
|
(Absolon, 1916),
|
|
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolybothrus (Eupolybothrus) grossipes" order="Lithobiomorpha" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="grossipes" subGenus="Eupolybothrus">Eupolybothrus (Eupolybothrus) grossipes</taxonomicName>
|
|
(C.L. Koch, 1847),
|
|
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolybothrus kahfi" order="Lithobiomorpha" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="kahfi">Eupolybothrus kahfi</taxonomicName>
|
|
Stoev & Akkari sp. n.,
|
|
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolybothrus (Parapolybothrus) herzegowinensis" order="Lithobiomorpha" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="herzegowinensis" subGenus="Parapolybothrus">Eupolybothrus (Parapolybothrus) herzegowinensis</taxonomicName>
|
|
(Verhoeff, 1900),
|
|
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolybothrus (Parapolybothrus) imperialis" order="Lithobiomorpha" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="imperialis" subGenus="Parapolybothrus">Eupolybothrus (Parapolybothrus) imperialis</taxonomicName>
|
|
(Meinert, 1872),
|
|
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolybothrus (Schizopolybothrus) leostygis" order="Lithobiomorpha" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="leostygis" subGenus="Schizopolybothrus">Eupolybothrus (Schizopolybothrus) leostygis</taxonomicName>
|
|
(Verhoeff, 1899),
|
|
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolybothrus (Eupolybothrus) litoralis" order="Lithobiomorpha" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="litoralis" subGenus="Eupolybothrus">Eupolybothrus (Eupolybothrus) litoralis</taxonomicName>
|
|
(L. Koch, 1867),
|
|
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolybothrus (Eupolybothrus) longicornis" order="Lithobiomorpha" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="longicornis" subGenus="Eupolybothrus">Eupolybothrus (Eupolybothrus) longicornis</taxonomicName>
|
|
(Risso, 1826),
|
|
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolybothrus (Allopolybothrus) nudicornis" order="Lithobiomorpha" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="nudicornis" subGenus="Allopolybothrus">Eupolybothrus (Allopolybothrus) nudicornis</taxonomicName>
|
|
(Gervais, 1837),
|
|
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolybothrus (Parapolybothrus) obrovensis" order="Lithobiomorpha" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="obrovensis" subGenus="Parapolybothrus">Eupolybothrus (Parapolybothrus) obrovensis</taxonomicName>
|
|
(Verhoeff, 1930),
|
|
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolybothrus (Mesobothrus) transsylvanicus" order="Lithobiomorpha" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="transsylvanicus" subGenus="Mesobothrus">Eupolybothrus (Mesobothrus) transsylvanicus</taxonomicName>
|
|
(Latzel, 1882),
|
|
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolybothrus (Schizopolybothrus) tabularum" order="Lithobiomorpha" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="tabularum" subGenus="Schizopolybothrus">Eupolybothrus (Schizopolybothrus) tabularum</taxonomicName>
|
|
Verhoeff, 1937,
|
|
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolybothrus (Leptopolybothrus) tridentinus" order="Lithobiomorpha" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="tridentinus" subGenus="Leptopolybothrus">Eupolybothrus (Leptopolybothrus) tridentinus</taxonomicName>
|
|
(Fanzago, 1874),
|
|
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolybothrus (Propolybothrus) werneri" order="Lithobiomorpha" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="werneri" subGenus="Propolybothrus">Eupolybothrus (Propolybothrus) werneri</taxonomicName>
|
|
(Attems, 1902),
|
|
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolybothrus (Mesobothrus) zeus" order="Lithobiomorpha" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="zeus" subGenus="Mesobothrus">Eupolybothrus (Mesobothrus) zeus</taxonomicName>
|
|
(Verhoeff, 1901).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="5" pageNumber="34" type="taxa of uncertain taxonomic status (10)">
|
|
<paragraph pageId="5" pageNumber="34">Taxa of uncertain taxonomic status (10).</paragraph>
|
|
<paragraph pageId="5" pageNumber="34">
|
|
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolybothrus (Schizopolybothrus) acherontis" order="Lithobiomorpha" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="acherontis" subGenus="Schizopolybothrus">Eupolybothrus (Schizopolybothrus) acherontis</taxonomicName>
|
|
(Verhoeff, 1900),
|
|
<taxonomicName genus="Acherontis" lsidName="acherontis wardaranus" pageId="5" pageNumber="34" rank="species" species="wardaranus">Eupolybothrus acherontis wardaranus</taxonomicName>
|
|
(Verhoeff, 1937),
|
|
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolybothrus (Mesobothrus) macedonicus" order="Lithobiomorpha" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="macedonicus" subGenus="Mesobothrus">Eupolybothrus (Mesobothrus) macedonicus</taxonomicName>
|
|
(Verhoeff, 1943),
|
|
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolybothrus osellai" order="Lithobiomorpha" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="osellai">Eupolybothrus osellai</taxonomicName>
|
|
Matic, Floca & Hurezeanu, 1992,
|
|
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolybothrus ruffoi" order="Lithobiomorpha" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="ruffoi">Eupolybothrus ruffoi</taxonomicName>
|
|
Matic, Floca & Hurezeanu, 1992,
|
|
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolybothrus sketi" order="Lithobiomorpha" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="sketi">Eupolybothrus sketi</taxonomicName>
|
|
Matic, 1979,
|
|
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolybothrus (Schizopolybothrus) spiniger" order="Lithobiomorpha" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="spiniger" subGenus="Schizopolybothrus">Eupolybothrus (Schizopolybothrus) spiniger</taxonomicName>
|
|
(Latzel, 1888),
|
|
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolybothrus (Schizopolybothrus) stygis" order="Lithobiomorpha" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="stygis" subGenus="Schizopolybothrus">Eupolybothrus (Schizopolybothrus) stygis</taxonomicName>
|
|
(Folkmanova, 1940),
|
|
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolybothrus valkanovi" order="Lithobiomorpha" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="valkanovi">Eupolybothrus valkanovi</taxonomicName>
|
|
(Kaczmarek, 1973),
|
|
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolybothrus (Propolybothrus) verrucosus" order="Lithobiomorpha" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="verrucosus" subGenus="Propolybothrus">Eupolybothrus (Propolybothrus) verrucosus</taxonomicName>
|
|
(Sseliwanoff, 1876).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection lastPageId="6" lastPageNumber="35" pageId="5" pageNumber="34" type="remarks.">
|
|
<paragraph pageId="5" pageNumber="34">Remarks.</paragraph>
|
|
<paragraph lastPageId="6" lastPageNumber="35" pageId="5" pageNumber="34">
|
|
Several taxa assigned to
|
|
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolybothrus" order="Lithobiomorpha" pageId="5" pageNumber="34" phylum="Arthropoda" rank="genus">Eupolybothrus</taxonomicName>
|
|
remain species inquirendae. Here we briefly review the current status of these taxa.
|
|
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolybothrus stygis" order="Lithobiomorpha" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="stygis">Eupolybothrus stygis</taxonomicName>
|
|
was described from Iljina
|
|
<normalizedToken originalValue="pećina">pecina</normalizedToken>
|
|
(cave) near Trebinje in Bosnia and Herzegovina (
|
|
<bibRefCitation author="Folkmanova, B" journalOrPublisher="Vestnik ceskoslovenske zoologicke spolecnosti v Praze" pageId="36" pageNumber="65" pagination="47 - 58" title="O novych balkanskych jeskennich Chilopodech ve sberech Dr. K. Absolona." volume="8" year="1940">Folkmanova 1940</bibRefCitation>
|
|
), and
|
|
<bibRefCitation author="Stoev, P" journalOrPublisher="Historia naturalis bulgarica" pageId="39" pageNumber="68" pagination="93 - 107" title="On the centipedes (Chilopoda) of the Republic of Macedonia." volume="13" year="2001 b">Stoev (2001b)</bibRefCitation>
|
|
suggested that it could be identical with
|
|
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolybothrus leostygis" order="Lithobiomorpha" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="leostygis">Eupolybothrus leostygis</taxonomicName>
|
|
, a troglobitic species
|
|
<pageBreakToken pageId="6" pageNumber="35" start="start">known</pageBreakToken>
|
|
from the same area (
|
|
<bibRefCitation author="Eason, EH" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="36" pageNumber="65" pagination="111 - 144" title="The identity of the European and Mediterranean species of Lithobiidae (Chilopoda) described by K. W. Verhoeff and now represented by material preserved in the British Museum (Natural History)." volume="77" year="1983">Eason 1983</bibRefCitation>
|
|
). In the key below it keys out together with
|
|
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolybothrus acherontis" order="Lithobiomorpha" pageId="6" pageNumber="35" phylum="Arthropoda" rank="species" species="acherontis">Eupolybothrus acherontis</taxonomicName>
|
|
, another poorly known species from Bosnia and Herzegovina.
|
|
<bibRefCitation author="Stoev, P" journalOrPublisher="Arthropoda Selecta" pageId="39" pageNumber="68" pagination="199 - 206" title="On centipedes (Chilopoda) of Albania, 2." volume="9" year="2001 a">Stoev (2001a</bibRefCitation>
|
|
,
|
|
<bibRefCitation author="Stoev, P" journalOrPublisher="Historia naturalis bulgarica" pageId="39" pageNumber="68" pagination="93 - 107" title="On the centipedes (Chilopoda) of the Republic of Macedonia." volume="13" year="2001 b">b</bibRefCitation>
|
|
) noted that
|
|
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolybothrus spiniger" order="Lithobiomorpha" pageId="6" pageNumber="35" phylum="Arthropoda" rank="species" species="spiniger">Eupolybothrus spiniger</taxonomicName>
|
|
,
|
|
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolybothrus acherontis" order="Lithobiomorpha" pageId="6" pageNumber="35" phylum="Arthropoda" rank="species" species="acherontis">Eupolybothrus acherontis</taxonomicName>
|
|
and
|
|
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolybothrus acherontis" order="Lithobiomorpha" pageId="6" pageNumber="35" phylum="Arthropoda" rank="species" species="acherontis">Eupolybothrus acherontis</taxonomicName>
|
|
wardaranus could be identical with
|
|
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolybothrus caesar" order="Lithobiomorpha" pageId="6" pageNumber="35" phylum="Arthropoda" rank="species" species="caesar">Eupolybothrus caesar</taxonomicName>
|
|
. Being the oldest available name, in case of synonymy
|
|
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolybothrus spiniger" order="Lithobiomorpha" pageId="6" pageNumber="35" phylum="Arthropoda" rank="species" species="spiniger">Eupolybothrus spiniger</taxonomicName>
|
|
would have priority over
|
|
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolybothrus caesar" order="Lithobiomorpha" pageId="6" pageNumber="35" phylum="Arthropoda" rank="species" species="caesar">Eupolybothrus caesar</taxonomicName>
|
|
. Thanks to Verena Stagl, curator of myriapods at NHMW, we were able to obtain a photograph of the prefemur of the ultimate leg pair of male
|
|
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolybothrus spiniger" order="Lithobiomorpha" pageId="6" pageNumber="35" phylum="Arthropoda" rank="species" species="spiniger">Eupolybothrus spiniger</taxonomicName>
|
|
which shows no differences with that of
|
|
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolybothrus caesar" order="Lithobiomorpha" pageId="6" pageNumber="35" phylum="Arthropoda" rank="species" species="caesar">Eupolybothrus caesar</taxonomicName>
|
|
. However, until we personally examine the types we prefer to treat
|
|
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolybothrus spiniger" order="Lithobiomorpha" pageId="6" pageNumber="35" phylum="Arthropoda" rank="species" species="spiniger">Eupolybothrus spiniger</taxonomicName>
|
|
and
|
|
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolybothrus caesar" order="Lithobiomorpha" pageId="6" pageNumber="35" phylum="Arthropoda" rank="species" species="caesar">Eupolybothrus caesar</taxonomicName>
|
|
as separate species.
|
|
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolybothrus valkanovi" order="Lithobiomorpha" pageId="6" pageNumber="35" phylum="Arthropoda" rank="species" species="valkanovi">Eupolybothrus valkanovi</taxonomicName>
|
|
was based on a single female with unusually short gonopodial spurs found near Asenovgrad, Rhodope Mts in Bulgaria (
|
|
<bibRefCitation author="Kaczmarek, J" journalOrPublisher="Bulletin de la Societe des Amis des Sciences et des Lettres de Poznan, Serie D" pageId="37" pageNumber="66" pagination="181 - 191" title="Beitraege zur Kenntnis bulgarischer Chilopoden. Teil V. Das Genus Polybothrus." volume="14" year="1973">Kaczmarek 1973</bibRefCitation>
|
|
). According to
|
|
<bibRefCitation author="Stoev, P" journalOrPublisher="Series Faunistica" pageId="39" pageNumber="68" volume="25" volumeTitle="A Catalogue and Key to the centipedes (Chilopoda) of Bulgaria." year="2002">Stoev (2002)</bibRefCitation>
|
|
it is most likely conspecific with the morphologically similar
|
|
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolybothrus transsylvanicus" order="Lithobiomorpha" pageId="6" pageNumber="35" phylum="Arthropoda" rank="species" species="transsylvanicus">Eupolybothrus transsylvanicus</taxonomicName>
|
|
which is also known to occur in the area.
|
|
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolybothrus sketi" order="Lithobiomorpha" pageId="6" pageNumber="35" phylum="Arthropoda" rank="species" species="sketi">Eupolybothrus sketi</taxonomicName>
|
|
was described from male and female specimens found in the Jakupica Mts, in the Former Yugoslav Republic of Macedonia (
|
|
<bibRefCitation author="Matic, Z" journalOrPublisher="Bioloski vestnik, Ljubljana" pageId="38" pageNumber="67" pagination="147 - 155" title="Nouveautes sur la faune des Chilopodes de Yougoslavie." volume="27" year="1979">Matic 1979</bibRefCitation>
|
|
). It is listed under the possible synonyms of
|
|
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolybothrus transsylvanicus" order="Lithobiomorpha" pageId="6" pageNumber="35" phylum="Arthropoda" rank="species" species="transsylvanicus">Eupolybothrus transsylvanicus</taxonomicName>
|
|
by
|
|
<bibRefCitation author="Minelli, A" journalOrPublisher="Lavori della Societa Italiana di Biogeografia, Nuova Serie" pageId="38" pageNumber="67" title="CHILOBASE. A web resource for Chilopoda taxonomy." url="http://chilobase.bio.unipd.it" year="2006">Minelli (2006)</bibRefCitation>
|
|
but its status is yet to be clarified. Morphologically, it is most closely related to
|
|
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolybothrus zeus" order="Lithobiomorpha" pageId="6" pageNumber="35" phylum="Arthropoda" rank="species" species="zeus">Eupolybothrus zeus</taxonomicName>
|
|
from Greece.
|
|
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolybothrus tabularum" order="Lithobiomorpha" pageId="6" pageNumber="35" phylum="Arthropoda" rank="species" species="tabularum">Eupolybothrus tabularum</taxonomicName>
|
|
was synonymised under
|
|
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolybothrus excellens" order="Lithobiomorpha" pageId="6" pageNumber="35" phylum="Arthropoda" rank="species" species="excellens">Eupolybothrus excellens</taxonomicName>
|
|
by
|
|
<bibRefCitation author="Minelli, A" journalOrPublisher="Lavori della Societa Italiana di Biogeografia, Nuova Serie" pageId="38" pageNumber="67" pagination="373 - 411" title="I Chilopodi della regione ligure con particolare riguardo al popolamento delle Alpi Liguri." volume="9" year="1985">Minelli and Zapparoli (1985)</bibRefCitation>
|
|
but was recently found to be a good species. A paper on this subject is currently in preparation by M. Zapparoli and will be published elsewhere. Although both species share some traits in common (like 15VCm spine), the long median protuberance on the prefemur of leg 15 in males convincingly distinguishes
|
|
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolybothrus excellens" order="Lithobiomorpha" pageId="6" pageNumber="35" phylum="Arthropoda" rank="species" species="excellens">Eupolybothrus excellens</taxonomicName>
|
|
from
|
|
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolybothrus tabularum" order="Lithobiomorpha" pageId="6" pageNumber="35" phylum="Arthropoda" rank="species" species="tabularum">Eupolybothrus tabularum</taxonomicName>
|
|
(see key below).
|
|
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolybothrus macedonicus" order="Lithobiomorpha" pageId="6" pageNumber="35" phylum="Arthropoda" rank="species" species="macedonicus">Eupolybothrus macedonicus</taxonomicName>
|
|
is hitherto known only from its type locality, Temna cave near Loutraki, North Greece (
|
|
<bibRefCitation author="Zapparoli, M" journalOrPublisher="Fragmenta entomologica" pageId="40" pageNumber="69" pagination="1 - 146" title="A catalogue of the centipedes from Greece (Chilopoda)." volume="34" year="2002">Zapparoli 2002</bibRefCitation>
|
|
). Likewise,
|
|
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolybothrus verrucosus" order="Lithobiomorpha" pageId="6" pageNumber="35" phylum="Arthropoda" rank="species" species="verrucosus">Eupolybothrus verrucosus</taxonomicName>
|
|
is presently known only from its original description based on a single female specimen from Moldova (
|
|
<bibRefCitation author="Minelli, A" journalOrPublisher="Lavori della Societa Italiana di Biogeografia, Nuova Serie" pageId="38" pageNumber="67" title="CHILOBASE. A web resource for Chilopoda taxonomy." url="http://chilobase.bio.unipd.it" year="2006">Minelli 2006</bibRefCitation>
|
|
). The taxonomic status of both species remains uncertain. The status of
|
|
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolybothrus cloudsley-thompsoni" order="Lithobiomorpha" pageId="6" pageNumber="35" phylum="Arthropoda" rank="species" species="cloudsley-thompsoni">Eupolybothrus cloudsley-thompsoni</taxonomicName>
|
|
,
|
|
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolybothrus osellai" order="Lithobiomorpha" pageId="6" pageNumber="35" phylum="Arthropoda" rank="species" species="osellai">Eupolybothrus osellai</taxonomicName>
|
|
and
|
|
<taxonomicName class="Chilopoda" family="Lithobiidae" genus="Eupolybothrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolybothrus ruffoi" order="Lithobiomorpha" pageId="6" pageNumber="35" phylum="Arthropoda" rank="species" species="ruffoi">Eupolybothrus ruffoi</taxonomicName>
|
|
is addressed under Discussion.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |