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<document id="FB312D91FEA6F7BDE0F71CE44AC6DC14" ID-DOI="10.37520/aemnp.2020.038" ID-ISSN="1804-6487" ID-Zenodo-Dep="5177688" ID-ZooBank="1E75F2CD-A21E-4980-AFD1-11D5525E6CB7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628618608376" checkinUser="carolina" docAuthor="Shen, Jia-Wei, Choi, Su-Ho &amp; Leschen, Richard A. B." docDate="2020" docId="9C7187FFFFB8802BAF95A8E9FE129E6F" docLanguage="en" docName="ActaEntMusNatPrag.60.2.553-564.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 60 (2)" docSource="http://dx.doi.org/10.37520/aemnp.2020.038" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Phormiobius ramsayi Shen &amp; Choi &amp; Leschen 2020, sp. nov." docType="treatment" docUuid="424DB161-C438-4225-AEC0-39D226FF0E4A" docUuidSource="ZooBank" docVersion="3" lastPageNumber="563" masterDocId="6048FF87FFB08021AC5DAD34FFE59975" masterDocTitle="Revision of the New Zealand endemic genus Phormiobius (Coleoptera: Staphylinidae: Pselaphinae)" masterLastPageNumber="564" masterPageNumber="553" pageNumber="561" updateTime="1699047842021" updateUser="plazi" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="FB8B06410D248A9D04E4647736C77C2D">Revision of the New Zealand endemic genus Phormiobius (Coleoptera: Staphylinidae: Pselaphinae)</mods:title>
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<mods:namePart id="CB65B6F352663AE1392F10E1D7B65372">Shen, Jia-Wei</mods:namePart>
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<mods:namePart id="3BA41C11E6D8547EE60762081DA6607A">Choi, Su-Ho</mods:namePart>
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<mods:namePart id="04C66FEDEA4F01BCC65AE4C19B7842D2">Leschen, Richard A. B.</mods:namePart>
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<paragraph id="146736E9FFB88029AF95A8E9FAF59C80" blockId="8.[968,1296,1501,1525]" box="[968,1296,1501,1525]" pageId="8" pageNumber="561">
<heading id="4F2F8185FFB88029AF95A8E9FAF59C80" box="[968,1296,1501,1525]" centered="true" fontSize="10" level="2" pageId="8" pageNumber="561" reason="3">
<emphasis id="26ACEAFBFFB88029AF95A8E9FAF59C80" bold="true" box="[968,1296,1501,1525]" pageId="8" pageNumber="561">
<taxonomicName id="D3D84D6AFFB88029AF95A8E9FB519C80" ID-CoL="8Y8F5" authority="Shen &amp; Choi &amp; Leschen, 2020" authorityName="Shen &amp; Choi &amp; Leschen" authorityYear="2020" box="[968,1204,1501,1525]" class="Insecta" family="Staphylinidae" genus="Phormiobius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="561" phylum="Arthropoda" rank="species" species="ramsayi" status="sp. nov.">
<emphasis id="26ACEAFBFFB88029AF95A8E9FB519C80" bold="true" box="[968,1204,1501,1525]" italics="true" pageId="8" pageNumber="561">Phormiobius ramsayi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3D9F5780FFB88029A8E6A8EAFAF59C80" box="[1211,1296,1502,1525]" pageId="8" pageNumber="561" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="5CC26562FFB88029A867A8C8FB789F7A" box="[1082,1181,1532,1551]" pageId="8" pageNumber="561" type="description">
<paragraph id="146736E9FFB88029A867A8C8FB789F7A" blockId="8.[828,1437,1532,1646]" box="[1082,1181,1532,1551]" pageId="8" pageNumber="561">
(
<figureCitation id="8CE32A6CFFB88029A81DA8C8FB9D9F7A" box="[1088,1144,1532,1551]" captionStart="Fig" captionStartId="9.[1199,1230,1886,1905]" captionTargetBox="[161,1167,1249,2063]" captionTargetId="figure-79@9.[148,1182,1237,2079]" captionTargetPageId="9" captionText="Fig.8. Diagnostic characters of Phormiobius ramsayi sp. nov. A habitus; B abdomen, in ventral view; C metatrochanter; D metatibia. Scale bars: AB = 0.5 mm, CD = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5177704" httpUri="https://zenodo.org/record/5177704/files/figure.png" pageId="8" pageNumber="561">Figs 8</figureCitation>
,
<figureCitation id="8CE32A6CFFB88029A8DCA8C8FB729F7A" box="[1153,1175,1532,1551]" captionStart="Fig" captionStartId="11.[150,182,1070,1089]" captionTargetBox="[162,753,198,1052]" captionTargetId="figure-849@11.[150,758,192,1053]" captionTargetPageId="11" captionText="Fig. 10. Geographic distribution of Phormiobius brouni sp. nov., P. graceae sp. nov., P. halli Broun, 1917, P. matau sp. nov., P. pseudhalli sp. nov. and P. ramsayi sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5177706" httpUri="https://zenodo.org/record/5177706/files/figure.png" pageId="8" pageNumber="561">10</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="5CC26562FFB88029AF61AB12FB139F1B" pageId="8" pageNumber="561" type="materials_examined">
<paragraph id="146736E9FFB88029AF61AB12FB139F1B" blockId="8.[828,1437,1532,1646]" pageId="8" pageNumber="561">
<emphasis id="26ACEAFBFFB88029AF61AB12FC269F4C" bold="true" box="[828,963,1574,1593]" pageId="8" pageNumber="561">Type material.</emphasis>
Hඈඅඈඍඒඉൾ: J, point-mounted (
<collectionCode id="72C9AE2CFFB88029A881AB12FAF99F4C" box="[1244,1308,1574,1593]" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="8" pageNumber="561" type="Other institutional type">NZAC</collectionCode>
):
<collectionCode id="72C9AE2CFFB88029A972AB12FA869F4C" box="[1327,1379,1574,1593]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="8" pageNumber="561">NEW</collectionCode>
<collectionCode id="72C9AE2CFFB88029A931AB12FC929F26" pageId="8" pageNumber="561">ZEA- LAND</collectionCode>
WO Mahoenui Gribbens Road
<date id="60661029FFB88029A820AB74FB049F26" box="[1149,1249,1600,1619]" pageId="8" pageNumber="561" value="1977-06-26">26 Jun 1977</date>
G.W.Ramsay // Litter 77/81. Pൺ*©ൺඍඒඉൾ:
<specimenCount id="02DEFD60FFB88029AFB7AB6FFC1F9F1B" box="[1002,1018,1621,1646]" pageId="8" pageNumber="561" type="female">1 ♀</specimenCount>
, same label data as
<typeStatus id="CB63884BFFB88029A8F9AB6FFB149F1B" box="[1188,1265,1627,1646]" pageId="8" pageNumber="561" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="5CC26562FFB8802BAF61ABB3FE089C4F" lastPageId="10" lastPageNumber="563" pageId="8" pageNumber="561" type="description">
<paragraph id="146736E9FFB88029AF61ABB3FAD09E2B" blockId="8.[828,1437,1671,2078]" pageId="8" pageNumber="561">
<emphasis id="26ACEAFBFFB88029AF61ABB3FC239FEB" bold="true" box="[828,966,1671,1694]" pageId="8" pageNumber="561">Description.</emphasis>
Body (
<figureCitation id="8CE32A6CFFB88029A84CABB3FB879FEB" box="[1041,1122,1671,1694]" captionStart="Fig" captionStartId="9.[1199,1230,1886,1905]" captionTargetBox="[161,1167,1249,2063]" captionTargetId="figure-79@9.[148,1182,1237,2079]" captionTargetPageId="9" captionText="Fig.8. Diagnostic characters of Phormiobius ramsayi sp. nov. A habitus; B abdomen, in ventral view; C metatrochanter; D metatibia. Scale bars: AB = 0.5 mm, CD = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5177704" httpUri="https://zenodo.org/record/5177704/files/figure.png" pageId="8" pageNumber="561">Fig. 8A</figureCitation>
) length
<quantity id="D3209B0CFFB88029A8EBABB3FAF39FEB" box="[1206,1302,1671,1694]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="8" pageNumber="561" unit="mm" value="1.9">1.90 mm</quantity>
, about 2.19× as long as wide (BL/AW = 2.19). Head about 1.29× longer than wide (HL/HW = 1.29). Eyes with 15 facets. Pronotum about as long as wide (PL/PW = 0.97). Elytra wider than long, about 0.58× as long as wide (EL/EW = 0.58) and about 0.76× as long as pronotum (EL/PL = 0.76). Abdomen about 1.21× longer than wide (AL/AW = 0.94).
</paragraph>
<paragraph id="146736E9FFB8802BAF07AA52FD049C6F" blockId="8.[828,1437,1671,2078]" lastBlockId="10.[151,761,1187,1818]" lastPageId="10" lastPageNumber="563" pageId="8" pageNumber="561">
<emphasis id="26ACEAFBFFB88029AF07AA52FC7D9E0B" bold="true" box="[858,920,1894,1918]" italics="true" pageId="8" pageNumber="561">Male.</emphasis>
Eyes with 15 facets.Abdominal concavities (
<figureCitation id="8CE32A6CFFB88029A92CAA53FCBE9EEB" captionStart="Fig" captionStartId="9.[1199,1230,1886,1905]" captionTargetBox="[161,1167,1249,2063]" captionTargetId="figure-79@9.[148,1182,1237,2079]" captionTargetPageId="9" captionText="Fig.8. Diagnostic characters of Phormiobius ramsayi sp. nov. A habitus; B abdomen, in ventral view; C metatrochanter; D metatibia. Scale bars: AB = 0.5 mm, CD = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5177704" httpUri="https://zenodo.org/record/5177704/files/figure.png" pageId="8" pageNumber="561">Fig. 8B</figureCitation>
) present on V1, V2, V3 and V4, and all inner surfaces polished; C1 and C2 well-impressed with distinct edges, C2 deep and lacking spines, greatest width longer than length of metatrochanter and slightly widened posteriorly; C3 broader than C2 and lacking spines, C4 very weak. Metatrochanter (
<figureCitation id="8CE32A6CFFBA802BAD10A997FE479DCF" box="[333,418,1187,1210]" captionStart="Fig" captionStartId="9.[1199,1230,1886,1905]" captionTargetBox="[161,1167,1249,2063]" captionTargetId="figure-79@9.[148,1182,1237,2079]" captionTargetPageId="9" captionText="Fig.8. Diagnostic characters of Phormiobius ramsayi sp. nov. A habitus; B abdomen, in ventral view; C metatrochanter; D metatibia. Scale bars: AB = 0.5 mm, CD = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5177704" httpUri="https://zenodo.org/record/5177704/files/figure.png" pageId="10" pageNumber="563">Fig. 8C</figureCitation>
) with inner face concave with well-developed longitudinal carina; metafemora weakly curved at apex; metatibia (
<figureCitation id="8CE32A6CFFBA802BADF5A9D7FE1D9D8F" box="[424,504,1251,1274]" captionStart="Fig" captionStartId="9.[1199,1230,1886,1905]" captionTargetBox="[161,1167,1249,2063]" captionTargetId="figure-79@9.[148,1182,1237,2079]" captionTargetPageId="9" captionText="Fig.8. Diagnostic characters of Phormiobius ramsayi sp. nov. A habitus; B abdomen, in ventral view; C metatrochanter; D metatibia. Scale bars: AB = 0.5 mm, CD = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5177704" httpUri="https://zenodo.org/record/5177704/files/figure.png" pageId="10" pageNumber="563">Fig. 8D</figureCitation>
) with small apical spine. Tergite 5 about 1.5× wider than long, broadly rounded.
</paragraph>
<caption id="40A76661FFB98028ACCBA9BCFAA39DC3" ID-DOI="http://doi.org/10.5281/zenodo.5177702" ID-Zenodo-Dep="5177702" httpUri="https://zenodo.org/record/5177702/files/figure.png" pageId="9" pageNumber="562" startId="9.[150,181,1160,1179]" targetBox="[234,1359,214,1138]" targetPageId="9">
<paragraph id="146736E9FFB98028ACCBA9BCFAA39DC3" blockId="9.[150,1436,1160,1206]" pageId="9" pageNumber="562">
Fig. 7. Diagnostic characters of
<taxonomicName id="D3D84D6AFFB98028ADFAA9BCFD969DEE" authority="Shen &amp; Choi &amp; Leschen, 2020" authorityName="Shen &amp; Choi &amp; Leschen" authorityYear="2020" box="[423,627,1160,1179]" class="Insecta" family="Staphylinidae" genus="Phormiobius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="562" phylum="Arthropoda" rank="species" species="pseudhalli" status="sp. nov.">
<emphasis id="26ACEAFBFFB98028ADFAA9BCFD969DEE" box="[423,627,1160,1179]" italics="true" pageId="9" pageNumber="562">Phormiobius pseudhalli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3D9F5780FFB98028AE25A9BCFD5E9DEE" box="[632,699,1160,1179]" pageId="9" pageNumber="562" rank="species">sp. nov.</taxonomicNameLabel>
A habitus; B abdomen, in ventral view; C metatibia; D metatrochanter; E T8; F V8 and T9; G aedeagus, in dorsal view; H same, in lateral view; I same, in ventral view. Scale bars: AB = 0.5 mm, CI = 0.1 mm.
</paragraph>
</caption>
<caption id="40A76661FFB98028A8F2AA6AFA769164" ID-DOI="http://doi.org/10.5281/zenodo.5177704" ID-Zenodo-Dep="5177704" httpUri="https://zenodo.org/record/5177704/files/figure.png" pageId="9" pageNumber="562" startId="9.[1199,1230,1886,1905]" targetBox="[161,1167,1249,2063]" targetPageId="9">
<paragraph id="146736E9FFB98028A8F2AA6AFA769164" blockId="9.[1199,1437,1886,2065]" pageId="9" pageNumber="562">
Fig.8. Diagnostic characters of
<taxonomicName id="D3D84D6AFFB98028A890AA4DFA789EF9" authority="Shen &amp; Choi &amp; Leschen, 2020" authorityName="Shen &amp; Choi &amp; Leschen" authorityYear="2020" box="[1229,1437,1913,1932]" class="Insecta" family="Staphylinidae" genus="Phormiobius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="562" phylum="Arthropoda" rank="species" species="ramsayi" status="sp. nov.">
<emphasis id="26ACEAFBFFB98028A890AA4DFA789EF9" box="[1229,1437,1913,1932]" italics="true" pageId="9" pageNumber="562">Phormiobius ramsayi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3D9F5780FFB98028A8F2AAA0FB1D9ED2" box="[1199,1272,1940,1959]" pageId="9" pageNumber="562" rank="species">sp. nov.</taxonomicNameLabel>
A habitus; B abdomen, in ventral view; C metatrochanter; D metatibia. Scale bars: AB = 0.5 mm, CD = 0.1 mm.
</paragraph>
</caption>
<caption id="40A76661FFBA802BACCAA902FD2E9D3C" box="[151,715,1078,1097]" pageId="10" pageNumber="563" startId="10.[151,182,1078,1097]">
<paragraph id="146736E9FFBA802BACCAA902FD2E9D3C" blockId="10.[151,715,1078,1097]" box="[151,715,1078,1097]" pageId="10" pageNumber="563">
Fig. 9. Lectotype and its labels of
<taxonomicName id="D3D84D6AFFBA802BADE7A902FD229D3C" authority="Broun, 1917" authorityName="Broun" authorityYear="1917" box="[442,711,1078,1097]" class="Insecta" family="Staphylinidae" genus="Phormiobius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="563" phylum="Arthropoda" rank="species" species="halli">
<emphasis id="26ACEAFBFFBA802BADE7A902FDB09D3C" box="[442,597,1078,1097]" italics="true" pageId="10" pageNumber="563">Phormiobius halli</emphasis>
<bibRefCitation id="70494B18FFBA802BAE07A902FD229D3C" author="BROUN T." box="[602,711,1078,1097]" pageId="10" pageNumber="563" pagination="347 - 474" refId="ref10283" refString="BROUN T. 1917: Descriptions of new genera and species of Coleoptera. Bulletin of the New Zealand Institute Wellington 1: 347 - 474." type="journal article" year="1917">Broun, 1917</bibRefCitation>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="146736E9FFBA802BACE8A816FE089C4F" blockId="10.[151,761,1187,1818]" box="[181,493,1314,1338]" pageId="10" pageNumber="563">
<emphasis id="26ACEAFBFFBA802BACE8A816FEEA9C4F" bold="true" box="[181,271,1314,1338]" italics="true" pageId="10" pageNumber="563">Female.</emphasis>
Eyes with 14 facets.
</paragraph>
</subSubSection>
<subSubSection id="5CC26562FFBA802BACCAA877FDC59FEF" pageId="10" pageNumber="563" type="diagnosis">
<paragraph id="146736E9FFBA802BACCAA877FDC59FEF" blockId="10.[151,761,1187,1818]" pageId="10" pageNumber="563">
<emphasis id="26ACEAFBFFBA802BACCAA877FE6F9C2F" bold="true" box="[151,394,1347,1370]" pageId="10" pageNumber="563">Differential diagnosis.</emphasis>
This species can be easily distinguished from its congeners by males having a well-developed concavity on V1 and metatrochanter with inner face concave with a well-developed longitudinal carina. It is most like
<taxonomicName id="D3D84D6AFFBA802BAC95A8F7FEDF9CAF" authority="Shen &amp; Choi &amp; Leschen, 2020" authorityName="Shen &amp; Choi &amp; Leschen" authorityYear="2020" box="[200,314,1475,1498]" class="Insecta" family="Staphylinidae" genus="Phormiobius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="563" phylum="Arthropoda" rank="species" species="graceae" status="sp. nov.">
<emphasis id="26ACEAFBFFBA802BAC95A8F7FEDF9CAF" box="[200,314,1475,1498]" italics="true" pageId="10" pageNumber="563">P. graceae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3D9F5780FFBA802BAD1EA8F7FE729CAF" box="[323,407,1475,1498]" pageId="10" pageNumber="563" rank="species">sp. nov.</taxonomicNameLabel>
by C3 lacking apical spines but can be separated from it by the metatrochanteral process having the concave inner face and V1 lacking a concavity. Additional male characters that characterize this species are as follows: C2 deep and polished with edge conspicuously demarcated with side rims and without apical spines and metatibia without small apical spine.
</paragraph>
</subSubSection>
<subSubSection id="5CC26562FFBA802BACCAAB94FE8E9F82" pageId="10" pageNumber="563" type="etymology">
<paragraph id="146736E9FFBA802BACCAAB94FE8E9F82" blockId="10.[151,761,1187,1818]" pageId="10" pageNumber="563">
<emphasis id="26ACEAFBFFBA802BACCAAB94FEFD9FC2" bold="true" box="[151,280,1696,1719]" pageId="10" pageNumber="563">Etymology.</emphasis>
This species is dedicated to Graham Ramsay, entomologist, and collector of the new species; it is a noun in genitive singular.
</paragraph>
</subSubSection>
<subSubSection id="5CC26562FFBA802BACCAAA37FE129E6F" box="[151,503,1795,1818]" pageId="10" pageNumber="563" type="distribution">
<paragraph id="146736E9FFBA802BACCAAA37FE129E6F" blockId="10.[151,761,1187,1818]" box="[151,503,1795,1818]" pageId="10" pageNumber="563">
<emphasis id="26ACEAFBFFBA802BACCAAA37FECF9E6F" bold="true" box="[151,298,1795,1818]" pageId="10" pageNumber="563">Distribution.</emphasis>
North Island: WO.
</paragraph>
</subSubSection>
</treatment>
</document>