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<document id="DF7C659EFCAA1281D7B14429222EBE24" ID-CLB-Dataset="26476" ID-DOI="10.11646/zootaxa.4615.3.9" ID-GBIF-Dataset="1a339476-3c23-4b65-b776-47deaa6fa4dc" ID-ISSN="1175-5326" ID-Zenodo-Dep="3246228" ID-ZooBank="FEDBF302-2E13-488E-B03D-7D195812FD22" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1560509736834" checkinUser="plazi" docAuthor="Joharchi, Omid, Khaustov, Alexander A. &amp; Ermilov, Sergey G." docDate="2019" docId="9C63D231FF9FFFBE0693FAFEFBFE4289" docLanguage="en" docName="zootaxa.4615.3.9.pdf" docOrigin="Zootaxa 4615 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gaeolaelaps lankaensis Joharchi &amp; Khaustov &amp; Ermilov 2019, sp. nov." docType="treatment" docVersion="8" lastPageNumber="569" masterDocId="605AAA49FF9EFFB80604FF87FFA34142" masterDocTitle="Two new species of Gaeolaelaps Evans &amp; Till (Acari: Laelapidae) from Sri Lanka" masterLastPageNumber="576" masterPageNumber="563" pageNumber="564" updateTime="1698734918697" updateUser="ExternalLinkService">
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<mods:title id="5291D235EE405DB263AEE297B9EE2DD6">Two new species of Gaeolaelaps Evans &amp; Till (Acari: Laelapidae) from Sri Lanka</mods:title>
</mods:titleInfo>
<mods:name id="D0AB48F99A748E7BE014E19A7D40E366" type="personal">
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<mods:namePart id="EAD945D62F4881ED2FD3310D11D8E5A6">Joharchi, Omid</mods:namePart>
</mods:name>
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<mods:namePart id="FECFA3EEDCA012E005946A8D346022B2">Khaustov, Alexander A.</mods:namePart>
</mods:name>
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<mods:namePart id="ADD383E4265105097D65F9CB9D10D3AE">Ermilov, Sergey G.</mods:namePart>
</mods:name>
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<mods:titleInfo id="D42DA6C02BB456EB566A0979AFFF230A">
<mods:title id="2E848EF5C2898AC54EBDAEAC46D1B20B">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="1E5F34898F1C61B13493F9FA4FCC7D71">
<mods:date id="DC781C70FF218DB0BD0AB8CB68F0EB30">2019</mods:date>
<mods:detail id="8F14081028F7E265C4D0AD5771A44B18" type="pubDate">
<mods:number id="0CAE18D7769A02BA42A8FF018A8A6586">2019-06-14</mods:number>
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<mods:number id="883738C9D46707CB0D8464B13B0433C1">4615</mods:number>
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<mods:classification id="58DB2F1960181F16986E4A9524093EB0">journal article</mods:classification>
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<mods:identifier id="123EB6C9CE24B8E0BC2FC9717337A2D4" type="DOI">10.11646/zootaxa.4615.3.9</mods:identifier>
<mods:identifier id="4FA4E09F3B4D72A0D33185CDEC28418D" type="GBIF-Dataset">1a339476-3c23-4b65-b776-47deaa6fa4dc</mods:identifier>
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<treatment id="9C63D231FF9FFFBE0693FAFEFBFE4289" ID-DOI="http://doi.org/10.5281/zenodo.5921145" ID-GBIF-Taxon="158223222" ID-Zenodo-Dep="5921145" LSID="urn:lsid:plazi:treatment:9C63D231FF9FFFBE0693FAFEFBFE4289" httpUri="http://treatment.plazi.org/id/9C63D231FF9FFFBE0693FAFEFBFE4289" lastPageId="6" lastPageNumber="569" pageId="1" pageNumber="564">
<subSubSection id="5CD030ACFF9FFFB90693FAFEFDBB44D1" box="[151,536,1401,1428]" pageId="1" pageNumber="564" type="nomenclature">
<paragraph id="14756327FF9FFFB90693FAFEFDBB44D1" blockId="1.[151,536,1401,1463]" box="[151,536,1401,1428]" pageId="1" pageNumber="564">
<heading id="4F3DD44BFF9FFFB90693FAFEFDBB44D1" bold="true" box="[151,536,1401,1428]" fontSize="11" level="1" pageId="1" pageNumber="564" reason="1">
<emphasis id="26BEBF35FF9FFFB90693FAFEFDBB44D1" bold="true" box="[151,536,1401,1428]" pageId="1" pageNumber="564">
<taxonomicName id="D3CA18A4FF9FFFB90693FAFEFE1744D6" authority="Joharchi &amp; Khaustov &amp; Ermilov, 2019" authorityName="Joharchi &amp; Khaustov &amp; Ermilov" authorityYear="2019" box="[151,436,1401,1428]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="1" pageNumber="564" phylum="Arthropoda" rank="species" species="lankaensis" status="sp. nov.">
<emphasis id="26BEBF35FF9FFFB90693FAFEFE1744D6" bold="true" box="[151,436,1401,1428]" italics="true" pageId="1" pageNumber="564">Gaeolaelaps lankaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3D8D024EFF9FFFB907BFFAFEFDBB44D1" box="[443,536,1401,1427]" pageId="1" pageNumber="564" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="5CD030ACFF9FFFB90693FA1AFE9E44F5" box="[151,317,1437,1463]" pageId="1" pageNumber="564" type="description">
<paragraph id="14756327FF9FFFB90693FA1AFE9E44F5" blockId="1.[151,536,1401,1463]" box="[151,317,1437,1463]" pageId="1" pageNumber="564">
(
<figureCitation id="8CF17FA2FF9FFFB9069BFA1AFE9744F5" box="[159,308,1437,1463]" captionStart-0="FIGURES 15" captionStart-1="FIGURES 613" captionStart-2="FIGURE 1417" captionStartId-0="3.[151,264,1959,1984]" captionStartId-1="4.[151,264,1929,1954]" captionStartId-2="5.[151,250,1949,1974]" captionTargetBox-0="[191,1383,188,1893]" captionTargetBox-1="[192,1390,181,1895]" captionTargetBox-2="[165,1427,241,1901]" captionTargetId-0="figure@3.[151,1434,166,1939]" captionTargetId-1="figure@4.[177,1410,166,1909]" captionTargetId-2="figure@5.[151,1434,197,1927]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURES 15. Gaeolaelaps lankaensis sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, subcapitulum; 4, epistome; 5, chelicera." captionText-1="FIGURES 613. DIC micrographs of Gaeolaelaps lankaensis sp. nov., female. 6, idiosoma in dorsal view; 7, idiosoma in ventral view; 8, sternal shield; 9, genital shield and opisthogastric area; 10, opisthonotal region; 11, epistome; 12, palp; 13, chelicera." captionText-2="FIGURE 1417. Gaeolaelaps lankaensis sp. nov., female. 14, leg I (trochanter-tibia); 15, leg II; 16, leg III; 17, leg IV." httpUri-0="https://zenodo.org/record/3246230/files/figure.png" httpUri-1="https://zenodo.org/record/3246232/files/figure.png" httpUri-2="https://zenodo.org/record/3246236/files/figure.png" pageId="1" pageNumber="564">Figures 117</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="5CD030ACFF9FFFB90693FA62FB524705" pageId="1" pageNumber="564" type="materials_examined">
<paragraph id="14756327FF9FFFB90693FA62FB524705" blockId="1.[151,1437,1508,2039]" pageId="1" pageNumber="564">
<emphasis id="26BEBF35FF9FFFB90693FA62FE1A44BD" bold="true" box="[151,441,1508,1535]" pageId="1" pageNumber="564">Type material examined</emphasis>
:
<materialsCitation id="A4A2697AFF9FFFB907CCFA62FDF94705" ID-GBIF-Occurrence="2265605711" collectingDate="2019-01-24" collectionCode="TSUMZ" collectorName="O. Joharchi &amp; S. G. Ermilov &amp; A. A. Khaustov &amp; Thambadola Ella" country="Sri Lanka" elevation="42" latitude="6.45" location="Polgampola" longLatPrecision="1304" longitude="80.2" pageId="1" pageNumber="564" specimenCount="1" stateProvince="Sabaragamuwa" typeStatus="holotype">
<typeStatus id="CB71DD85FF9FFFB907CCFA62FD9044BD" box="[456,563,1509,1535]" pageId="1" pageNumber="564" type="holotype">Holotype</typeStatus>
, female,
<collectingCountry id="6CDD23B7FF9FFFB9049EFA62FCA844BD" box="[666,779,1509,1535]" name="Sri Lanka" pageId="1" pageNumber="564">Sri Lanka</collectingCountry>
,
<collectingRegion id="D60EADC5FF9FFFB90512FA62FB8C44BD" box="[790,1071,1509,1535]" country="Sri Lanka" name="Sabaragamuwa" pageId="1" pageNumber="564">Sabaragamuwa Province</collectingRegion>
,
<location id="111535FCFF9FFFB9023FFA62FB6744BD" LSID="urn:lsid:plazi:treatment:9C63D231FF9FFFBE0693FAFEFBFE4289:111535FCFF9FFFB9023FFA62FB6744BD" box="[1083,1220,1509,1535]" country="Sri Lanka" latitude="6.45" longLatPrecision="1304" longitude="80.2" name="Polgampola" pageId="1" pageNumber="564" stateProvince="Sabaragamuwa">Polgampola</location>
,
<geoCoordinate id="71FE05E0FF9FFFB902CBFA62FA9B44BD" box="[1231,1336,1509,1535]" degrees="06" direction="north" minutes="27" orientation="latitude" pageId="1" pageNumber="564" precision="925" value="6.45">06°27 N</geoCoordinate>
,
<geoCoordinate id="71FE05E0FF9FFFB90340FA62FF094761" degrees="080" direction="east" minutes="12" orientation="longitude" pageId="1" pageNumber="564" precision="925" value="80.2">080°12 E</geoCoordinate>
, alt.
<elevation id="9FE78414FF9FFFB906E1F98EFE804761" box="[229,291,1545,1571]" metricMagnitude="1" metricUnit="m" metricValue="4.2" pageId="1" pageNumber="564" unit="m" value="42.0">
<quantity id="D332CEC2FF9FFFB906E1F98EFE804761" box="[229,291,1545,1571]" metricMagnitude="1" metricUnit="m" metricValue="4.2" pageId="1" pageNumber="564" unit="m" value="42.0">42 m</quantity>
</elevation>
,
<collectingDate id="7030BC0FFF9FFFB90735F98EFE514761" box="[305,498,1545,1571]" pageId="1" pageNumber="564" value="2019-01-24">
<date id="607445E7FF9FFFB90735F98EFE514761" box="[305,498,1545,1571]" pageId="1" pageNumber="564" value="2019-01-24">24 January 2019</date>
</collectingDate>
,
<collectorName id="B93F06F1FF9FFFB907FBF98EFD254761" box="[511,646,1545,1571]" pageId="1" pageNumber="564">O. Joharchi</collectorName>
,
<collectorName id="B93F06F1FF9FFFB90497F98EFC894761" box="[659,810,1545,1571]" pageId="1" pageNumber="564">S.G. Ermilov</collectorName>
&amp;
<collectorName id="B93F06F1FF9FFFB90555F98EFC5F4761" box="[849,1020,1545,1571]" pageId="1" pageNumber="564">A.A. Khaustov</collectorName>
coll., from forest soil-litter near the
<collectorName id="B93F06F1FF9FFFB90693F9AAFEF44705" box="[151,343,1581,1607]" pageId="1" pageNumber="564">Thambadola Ella</collectorName>
waterfall (in
<collectionCode id="72DBFBE2FF9FFFB907EAF9AAFDF14705" box="[494,594,1581,1607]" pageId="1" pageNumber="564">TSUMZ</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="A4A2697AFF9FFFB90461F9AAFB4E4705" ID-GBIF-Occurrence="2265605709" box="[613,1261,1581,1607]" collectingDate="2019-01-24" collectionCode="TSUMZ" collectorName="O. Joharchi &amp; S. G. Ermilov &amp; A. A. Khaustov &amp; Thambadola Ella" country="Sri Lanka" elevation="42" latitude="6.45" location="Polgampola" longLatPrecision="1304" longitude="80.2" pageId="1" pageNumber="564" specimenCount="1" stateProvince="Sabaragamuwa" typeStatus="paratype">
<typeStatus id="CB71DD85FF9FFFB90461F9AAFD694705" box="[613,714,1581,1607]" pageId="1" pageNumber="564" type="paratype">Paratype</typeStatus>
, one female, same data as holotype (in
<collectionCode id="72DBFBE2FF9FFFB90285F9AAFB464705" box="[1153,1253,1581,1607]" pageId="1" pageNumber="564">TSUMZ</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="5CD030ACFF9FFFB906C3F9D7FF744691" pageId="1" pageNumber="564" type="diagnosis">
<paragraph id="14756327FF9FFFB906C3F9D7FF744691" blockId="1.[151,1437,1508,2039]" pageId="1" pageNumber="564">
<emphasis id="26BEBF35FF9FFFB906C3F9D7FE994729" bold="true" box="[199,314,1616,1643]" pageId="1" pageNumber="564">Diagnosis</emphasis>
(female). Dorsal shield without distinct reticulate ornamentation over whole surface, narrowing from level of setae
<emphasis id="26BEBF35FF9FFFB90731F9F2FEED47CD" box="[309,334,1653,1679]" italics="true" pageId="1" pageNumber="564">r3</emphasis>
, progressively tapering until
<emphasis id="26BEBF35FF9FFFB9049EF9F2FD1747CD" box="[666,692,1653,1679]" italics="true" pageId="1" pageNumber="564">s6</emphasis>
, then subparallel from
<emphasis id="26BEBF35FF9FFFB905BFF9F2FC7B47CD" box="[955,984,1653,1679]" italics="true" pageId="1" pageNumber="564">S1</emphasis>
to level of
<emphasis id="26BEBF35FF9FFFB9025FF9F2FBD447CD" box="[1115,1143,1653,1679]" italics="true" pageId="1" pageNumber="564">S4</emphasis>
until rounded posteriorly, bearing 35 pairs of moderately long and simple setae (22 pairs of podonotal setae on shield and 13 pairs of opisthonotal setae on shield,
<emphasis id="26BEBF35FF9FFFB90783F93AFE064795" box="[391,421,1725,1751]" italics="true" pageId="1" pageNumber="564">Z5</emphasis>
&amp;
<emphasis id="26BEBF35FF9FFFB907CCF93AFE474795" box="[456,484,1725,1751]" italics="true" pageId="1" pageNumber="564">S5</emphasis>
outside the shield). Tritosternum with pair of very long pilose laciniae, Sternal shield smooth almost throughout; bearing two pairs of smooth pointed setae,
<emphasis id="26BEBF35FF9FFFB905BAF966FC7C47B9" box="[958,991,1761,1787]" italics="true" pageId="1" pageNumber="564">st1</emphasis>
outside the shield, metasternal poroids (
<date id="607445E7FF9FFFB906A5F882FF60465D" box="[161,195,1797,1823]" pageId="1" pageNumber="564">
<emphasis id="26BEBF35FF9FFFB906A5F882FF60465D" box="[161,195,1797,1823]" italics="true" pageId="1" pageNumber="564">iv3</emphasis>
</date>
) located on minute platelets, ratio of shield length/width (at broadest level) 1.1. Genital shield ratio of length/ width (at broadest level) 2.8, with irregular longitudinal lines in anterior half≃enclosing two diagonal lines forming an inverted-V in posterior ≃ part of surface. Anal shield ratio of length/width (at broadest level) 1.1, cribrum small. Peritreme thin and short, barely reaching anterior margin of coxa II, post-stigmatal section well≃reduced. Palp tarsal claw three-tined, fixed digit of chelicera with four teeth distally and a larger proximal tooth, femur II with 10 setae.
</paragraph>
</subSubSection>
<subSubSection id="5CD030ACFF9FFFBE06C3F85BFD9C4195" lastPageId="6" lastPageNumber="569" pageId="1" pageNumber="564" type="description">
<paragraph id="14756327FF9FFFB906C3F85BFDA546B5" blockId="1.[151,1437,1508,2039]" box="[199,518,2012,2039]" pageId="1" pageNumber="564">
<emphasis id="26BEBF35FF9FFFB906C3F85BFEFA46B5" bold="true" box="[199,345,2012,2039]" pageId="1" pageNumber="564">Description.</emphasis>
Female (n = 2)
</paragraph>
<paragraph id="14756327FF9CFFBA06C3FF1EFE4F4359" blockId="2.[151,1437,152,1979]" pageId="2" pageNumber="565">
<emphasis id="26BEBF35FF9CFFBA06C3FF1EFE2041F1" box="[199,387,153,179]" italics="true" pageId="2" pageNumber="565">Dorsal idiosoma</emphasis>
(
<figureCitation id="8CF17FA2FF9CFFBA0796FF1EFE7941F1" box="[402,474,153,179]" captionStart="FIGURES 15" captionStartId="3.[151,264,1959,1984]" captionTargetBox="[191,1383,188,1893]" captionTargetId="figure@3.[151,1434,166,1939]" captionTargetPageId="3" captionText="FIGURES 15. Gaeolaelaps lankaensis sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, subcapitulum; 4, epistome; 5, chelicera." httpUri="https://zenodo.org/record/3246230/files/figure.png" pageId="2" pageNumber="565">Figs 1</figureCitation>
,
<figureCitation id="8CF17FA2FF9CFFBA07ECFF1EFD8741F1" box="[488,548,153,179]" captionStart="FIGURES 613" captionStartId="4.[151,264,1929,1954]" captionTargetBox="[192,1390,181,1895]" captionTargetId="figure@4.[177,1410,166,1909]" captionTargetPageId="4" captionText="FIGURES 613. DIC micrographs of Gaeolaelaps lankaensis sp. nov., female. 6, idiosoma in dorsal view; 7, idiosoma in ventral view; 8, sternal shield; 9, genital shield and opisthogastric area; 10, opisthonotal region; 11, epistome; 12, palp; 13, chelicera." httpUri="https://zenodo.org/record/3246232/files/figure.png" pageId="2" pageNumber="565">6, 10</figureCitation>
). Idiosoma length 266270, width 151158. Dorsal shield 253257 long, 120 127 wide at
<emphasis id="26BEBF35FF9CFFBA0729FF3AFEE54195" box="[301,326,189,215]" italics="true" pageId="2" pageNumber="565">r2</emphasis>
level, covering most of dorsal idiosoma, 8085 wide at level of
<emphasis id="26BEBF35FF9CFFBA0243FF3AFBC04195" box="[1095,1123,189,215]" italics="true" pageId="2" pageNumber="565">S1</emphasis>
; without distinct reticulate ornamentation over whole surface, narrowing from level of setae
<emphasis id="26BEBF35FF9CFFBA056DFF66FC2041B9" box="[873,899,225,251]" italics="true" pageId="2" pageNumber="565">r3</emphasis>
, progressively tapering until
<emphasis id="26BEBF35FF9CFFBA02C2FF66FB7C41B9" box="[1222,1247,225,251]" italics="true" pageId="2" pageNumber="565">s6</emphasis>
, then subparallel from
<emphasis id="26BEBF35FF9CFFBA06D0FE82FF52405D" box="[212,241,261,287]" italics="true" pageId="2" pageNumber="565">S1</emphasis>
to level of
<emphasis id="26BEBF35FF9CFFBA076BFE82FE28405D" box="[367,395,261,287]" italics="true" pageId="2" pageNumber="565">S4</emphasis>
, until rounded posteriorly. Shield with 35 pairs of moderately long and simple setae, 22 pairs of podonotal setae on shield (
<emphasis id="26BEBF35FF9CFFBA07E4FEAEFDB14001" box="[480,530,297,323]" italics="true" pageId="2" pageNumber="565">j16</emphasis>
;
<emphasis id="26BEBF35FF9CFFBA0424FEAEFDF64001" box="[544,597,297,323]" italics="true" pageId="2" pageNumber="565">z16</emphasis>
;
<emphasis id="26BEBF35FF9CFFBA0467FEAEFD3B4001" box="[611,664,297,323]" italics="true" pageId="2" pageNumber="565">s16</emphasis>
;
<emphasis id="26BEBF35FF9CFFBA04A3FEAEFD7F4001" box="[679,732,297,323]" italics="true" pageId="2" pageNumber="565">r25</emphasis>
),
<emphasis id="26BEBF35FF9CFFBA04F6FEAEFCA84001" box="[754,779,297,323]" italics="true" pageId="2" pageNumber="565">
r
<quantity id="D332CEC2FF9CFFBA04F9FEAEFCA84001" box="[765,779,297,323]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="2" pageNumber="565" unit="in" value="6.0">6</quantity>
</emphasis>
in lateral soft cuticle, and 13 pairs of opisthonotal setae on shield (
<emphasis id="26BEBF35FF9CFFBA06EDFECAFE834025" box="[233,288,333,359]" italics="true" pageId="2" pageNumber="565">J15</emphasis>
;
<emphasis id="26BEBF35FF9CFFBA072AFECAFEC44025" box="[302,359,333,359]" italics="true" pageId="2" pageNumber="565">Z14</emphasis>
;
<emphasis id="26BEBF35FF9CFFBA0771FECAFE0E4025" box="[373,429,333,359]" italics="true" pageId="2" pageNumber="565">S14</emphasis>
),
<emphasis id="26BEBF35FF9CFFBA07C7FECAFE424025" box="[451,481,333,359]" italics="true" pageId="2" pageNumber="565">Z5</emphasis>
&amp;
<emphasis id="26BEBF35FF9CFFBA0407FECAFDBC4025" box="[515,543,333,359]" italics="true" pageId="2" pageNumber="565">
S
<quantity id="D332CEC2FF9CFFBA0415FECAFDBC4025" box="[529,543,333,359]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="2" pageNumber="565" unit="in" value="5.0">5</quantity>
</emphasis>
in soft cuticle posterior to shield (
<figureCitation id="8CF17FA2FF9CFFBA059CFECAFC4A4025" box="[920,1001,333,359]" captionStart="FIGURES 613" captionStartId="4.[151,264,1929,1954]" captionTargetBox="[192,1390,181,1895]" captionTargetId="figure@4.[177,1410,166,1909]" captionTargetPageId="4" captionText="FIGURES 613. DIC micrographs of Gaeolaelaps lankaensis sp. nov., female. 6, idiosoma in dorsal view; 7, idiosoma in ventral view; 8, sternal shield; 9, genital shield and opisthogastric area; 10, opisthonotal region; 11, epistome; 12, palp; 13, chelicera." httpUri="https://zenodo.org/record/3246232/files/figure.png" pageId="2" pageNumber="565">Fig. 10</figureCitation>
),
<emphasis id="26BEBF35FF9CFFBA05FAFEC9FBB94025" box="[1022,1050,334,359]" italics="true" pageId="2" pageNumber="565">Zx</emphasis>
setae and unpaired supernumerary seta (
<emphasis id="26BEBF35FF9CFFBA06D6FEF5FF4840C9" box="[210,235,370,395]" italics="true" pageId="2" pageNumber="565">Jx</emphasis>
) absent (
<figureCitation id="8CF17FA2FF9CFFBA0755FEF6FE0040C9" box="[337,419,369,395]" captionStart="FIGURES 613" captionStartId="4.[151,264,1929,1954]" captionTargetBox="[192,1390,181,1895]" captionTargetId="figure@4.[177,1410,166,1909]" captionTargetPageId="4" captionText="FIGURES 613. DIC micrographs of Gaeolaelaps lankaensis sp. nov., female. 6, idiosoma in dorsal view; 7, idiosoma in ventral view; 8, sternal shield; 9, genital shield and opisthogastric area; 10, opisthonotal region; 11, epistome; 12, palp; 13, chelicera." httpUri="https://zenodo.org/record/3246232/files/figure.png" pageId="2" pageNumber="565">Fig. 10</figureCitation>
). All setae smooth and uniform in length (podonotal setae 1822; opisthonotal setae 1518) and thickness, except
<emphasis id="26BEBF35FF9CFFBA0796FE12FE0C40ED" box="[402,431,405,431]" italics="true" pageId="2" pageNumber="565">Z5</emphasis>
(2729),
<emphasis id="26BEBF35FF9CFFBA0424FE12FD9E40ED" box="[544,573,405,431]" italics="true" pageId="2" pageNumber="565">S5</emphasis>
(2426),
<emphasis id="26BEBF35FF9CFFBA04A9FE12FD6540ED" box="[685,710,405,431]" italics="true" pageId="2" pageNumber="565">z1</emphasis>
(1517) and
<emphasis id="26BEBF35FF9CFFBA0565FE12FCDF40ED" box="[865,892,405,431]" italics="true" pageId="2" pageNumber="565">J5</emphasis>
(2224, with one or two barbs,
<figureCitation id="8CF17FA2FF9CFFBA02EAFE12FAE240ED" box="[1262,1345,405,431]" captionStart="FIGURES 613" captionStartId="4.[151,264,1929,1954]" captionTargetBox="[192,1390,181,1895]" captionTargetId="figure@4.[177,1410,166,1909]" captionTargetPageId="4" captionText="FIGURES 613. DIC micrographs of Gaeolaelaps lankaensis sp. nov., female. 6, idiosoma in dorsal view; 7, idiosoma in ventral view; 8, sternal shield; 9, genital shield and opisthogastric area; 10, opisthonotal region; 11, epistome; 12, palp; 13, chelicera." httpUri="https://zenodo.org/record/3246232/files/figure.png" pageId="2" pageNumber="565">Fig. 10</figureCitation>
), dorsal setae decreasing slightly in length from anterior to posterior. Shield with about eight pairs of discernible porelike structures, five on podonotum (
<emphasis id="26BEBF35FF9CFFBA0431FE5AFDFA40B5" box="[565,601,477,503]" italics="true" pageId="2" pageNumber="565">id1</emphasis>
,
<emphasis id="26BEBF35FF9CFFBA046DFE5AFD2E40B5" box="[617,653,477,503]" italics="true" pageId="2" pageNumber="565">id4</emphasis>
,
<emphasis id="26BEBF35FF9CFFBA0499FE5AFD6240B5" box="[669,705,477,503]" italics="true" pageId="2" pageNumber="565">id5</emphasis>
,
<emphasis id="26BEBF35FF9CFFBA04D5FE5AFD5840B5" box="[721,763,477,503]" italics="true" pageId="2" pageNumber="565">gd1</emphasis>
,
<emphasis id="26BEBF35FF9CFFBA0508FE5AFC9540B5" box="[780,822,477,503]" italics="true" pageId="2" pageNumber="565">gd6</emphasis>
) and three on opisthonotum (
<emphasis id="26BEBF35FF9CFFBA028AFE5AFB6540B5" box="[1166,1222,477,503]" italics="true" pageId="2" pageNumber="565">idm6</emphasis>
,
<emphasis id="26BEBF35FF9CFFBA02D2FE5AFAA140B5" box="[1238,1282,477,503]" italics="true" pageId="2" pageNumber="565">idl3</emphasis>
,
<emphasis id="26BEBF35FF9CFFBA0316FE5AFA9D40B5" box="[1298,1342,477,503]" italics="true" pageId="2" pageNumber="565">idl4</emphasis>
), others indistinct, see
<figureCitation id="8CF17FA2FF9CFFBA073CFD86FE024359" box="[312,417,513,539]" captionStart="FIGURES 15" captionStartId="3.[151,264,1959,1984]" captionTargetBox="[191,1383,188,1893]" captionTargetId="figure@3.[151,1434,166,1939]" captionTargetPageId="3" captionText="FIGURES 15. Gaeolaelaps lankaensis sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, subcapitulum; 4, epistome; 5, chelicera." httpUri="https://zenodo.org/record/3246230/files/figure.png" pageId="2" pageNumber="565">Figures 1</figureCitation>
and
<figureCitation id="8CF17FA2FF9CFFBA07DCFD86FE4A4359" box="[472,489,513,539]" captionStart="FIGURES 613" captionStartId="4.[151,264,1929,1954]" captionTargetBox="[192,1390,181,1895]" captionTargetId="figure@4.[177,1410,166,1909]" captionTargetPageId="4" captionText="FIGURES 613. DIC micrographs of Gaeolaelaps lankaensis sp. nov., female. 6, idiosoma in dorsal view; 7, idiosoma in ventral view; 8, sternal shield; 9, genital shield and opisthogastric area; 10, opisthonotal region; 11, epistome; 12, palp; 13, chelicera." httpUri="https://zenodo.org/record/3246232/files/figure.png" pageId="2" pageNumber="565">6</figureCitation>
.
</paragraph>
<paragraph id="14756327FF9CFFBA06C3FDA2FE8D444D" blockId="2.[151,1437,152,1979]" pageId="2" pageNumber="565">
<emphasis id="26BEBF35FF9CFFBA06C3FDA2FE29437D" box="[199,394,549,575]" italics="true" pageId="2" pageNumber="565">Ventral idiosoma</emphasis>
(
<figureCitation id="8CF17FA2FF9CFFBA0799FDA2FE49437D" box="[413,490,549,575]" captionStart="FIGURES 15" captionStartId="3.[151,264,1959,1984]" captionTargetBox="[191,1383,188,1893]" captionTargetId="figure@3.[151,1434,166,1939]" captionTargetPageId="3" captionText="FIGURES 15. Gaeolaelaps lankaensis sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, subcapitulum; 4, epistome; 5, chelicera." httpUri="https://zenodo.org/record/3246230/files/figure.png" pageId="2" pageNumber="565">Figs 2</figureCitation>
,
<figureCitation id="8CF17FA2FF9CFFBA07FEFDA2FD8B437D" box="[506,552,549,575]" captionStart="FIGURES 613" captionStartId="4.[151,264,1929,1954]" captionTargetBox="[192,1390,181,1895]" captionTargetId="figure@4.[177,1410,166,1909]" captionTargetPageId="4" captionText="FIGURES 613. DIC micrographs of Gaeolaelaps lankaensis sp. nov., female. 6, idiosoma in dorsal view; 7, idiosoma in ventral view; 8, sternal shield; 9, genital shield and opisthogastric area; 10, opisthonotal region; 11, epistome; 12, palp; 13, chelicera." httpUri="https://zenodo.org/record/3246232/files/figure.png" pageId="2" pageNumber="565">79</figureCitation>
). Tritosternum with pair of very long pilose laciniae (6871), fused basally (46), columnar base 2022 × 911 wide; presternal area lightly sclerotised, with a few transverse curved lines. Sternal shield (length 9599) narrowest between coxae II (6063), widest between coxa IIIII (8486), with straight anterior and posterior margin, with two pairs of smooth pointed setae (
<emphasis id="26BEBF35FF9CFFBA05B8FD16FC7F43E9" box="[956,988,657,683]" italics="true" pageId="2" pageNumber="565">
st
<date id="607445E7FF9CFFBA05CBFD16FC7F43E9" box="[975,988,657,683]" pageId="2" pageNumber="565" value="1924-02-26">2</date>
</emphasis>
2426,
<emphasis id="26BEBF35FF9CFFBA023EFD16FBF843E9" box="[1082,1115,657,683]" italics="true" pageId="2" pageNumber="565">
st
<date id="607445E7FF9CFFBA0249FD16FBF843E9" box="[1101,1115,657,683]" pageId="2" pageNumber="565" value="1925-03-27">3</date>
</emphasis>
2527),
<emphasis id="26BEBF35FF9CFFBA02C6FD16FB4143E9" box="[1218,1250,657,683]" italics="true" pageId="2" pageNumber="565">st1</emphasis>
(2022) outside the shield (
<figureCitation id="8CF17FA2FF9CFFBA0712FD32FEF8438D" box="[278,347,693,719]" captionStart="FIGURES 613" captionStartId="4.[151,264,1929,1954]" captionTargetBox="[192,1390,181,1895]" captionTargetId="figure@4.[177,1410,166,1909]" captionTargetPageId="4" captionText="FIGURES 613. DIC micrographs of Gaeolaelaps lankaensis sp. nov., female. 6, idiosoma in dorsal view; 7, idiosoma in ventral view; 8, sternal shield; 9, genital shield and opisthogastric area; 10, opisthonotal region; 11, epistome; 12, palp; 13, chelicera." httpUri="https://zenodo.org/record/3246232/files/figure.png" pageId="2" pageNumber="565">Fig. 8</figureCitation>
), setae never reaching base of next posterior setae, one pair of lyrifissures (
<date id="607445E7FF9CFFBA02C4FD32FB41438D" box="[1216,1250,693,719]" pageId="2" pageNumber="565">
<emphasis id="26BEBF35FF9CFFBA02C4FD32FB41438D" box="[1216,1250,693,719]" italics="true" pageId="2" pageNumber="565">iv1</emphasis>
</date>
) adjacent to anterior margin, and a pair of sub-oval poroids (
<date id="607445E7FF9CFFBA049BFD5EFD6143B1" box="[671,706,729,755]" pageId="2" pageNumber="565">
<emphasis id="26BEBF35FF9CFFBA049BFD5EFD6143B1" box="[671,706,729,755]" italics="true" pageId="2" pageNumber="565">iv2</emphasis>
</date>
) between
<emphasis id="26BEBF35FF9CFFBA053CFD5EFCFB43B1" box="[824,856,729,755]" italics="true" pageId="2" pageNumber="565">st2</emphasis>
and
<emphasis id="26BEBF35FF9CFFBA0595FD5EFC1143B1" box="[913,946,729,755]" italics="true" pageId="2" pageNumber="565">st3</emphasis>
, surface of sternal shield without reticulate ornamentation, almost smooth, except some irregular longitudinal lines laterally (
<figureCitation id="8CF17FA2FF9CFFBA022DFD7AFBCE4255" box="[1065,1133,765,791]" captionStart="FIGURES 613" captionStartId="4.[151,264,1929,1954]" captionTargetBox="[192,1390,181,1895]" captionTargetId="figure@4.[177,1410,166,1909]" captionTargetPageId="4" captionText="FIGURES 613. DIC micrographs of Gaeolaelaps lankaensis sp. nov., female. 6, idiosoma in dorsal view; 7, idiosoma in ventral view; 8, sternal shield; 9, genital shield and opisthogastric area; 10, opisthonotal region; 11, epistome; 12, palp; 13, chelicera." httpUri="https://zenodo.org/record/3246232/files/figure.png" pageId="2" pageNumber="565">Fig. 8</figureCitation>
), lateral margins alongside coxa IIIII fused with endopodal plates. Metasternal setae
<emphasis id="26BEBF35FF9CFFBA0537FCA6FCF74279" box="[819,852,801,827]" italics="true" pageId="2" pageNumber="565">st4</emphasis>
(1820) inserted on soft cuticle posterior to sternal shield and metasternal poroids (
<date id="607445E7FF9CFFBA0404FCC2FD81421D" box="[512,546,837,863]" pageId="2" pageNumber="565">
<emphasis id="26BEBF35FF9CFFBA0404FCC2FD81421D" box="[512,546,837,863]" italics="true" pageId="2" pageNumber="565">iv3</emphasis>
</date>
) located on minute platelets (Fig, 9). Endopodal plates III/IV elongate, narrow and curved. Genital shield tongue-shaped, length 8993, maximum width 3133, posterior margin slightly tapered, surface reticulated with irregular longitudinal lines in anterior which enclosing two diagonal lines forming an inverted-V in posterior part and a pair of simple setae
<emphasis id="26BEBF35FF9CFFBA04EEFC36FCA84289" box="[746,779,945,971]" italics="true" pageId="2" pageNumber="565">st5</emphasis>
(1315) (
<figureCitation id="8CF17FA2FF9CFFBA057CFC36FC184289" box="[888,955,945,971]" captionStart="FIGURES 613" captionStartId="4.[151,264,1929,1954]" captionTargetBox="[192,1390,181,1895]" captionTargetId="figure@4.[177,1410,166,1909]" captionTargetPageId="4" captionText="FIGURES 613. DIC micrographs of Gaeolaelaps lankaensis sp. nov., female. 6, idiosoma in dorsal view; 7, idiosoma in ventral view; 8, sternal shield; 9, genital shield and opisthogastric area; 10, opisthonotal region; 11, epistome; 12, palp; 13, chelicera." httpUri="https://zenodo.org/record/3246232/files/figure.png" pageId="2" pageNumber="565">Fig. 9</figureCitation>
). Shield flanked by a pair of minute narrow platelets; paragenital poroids located on soft cuticle lateral to shield near seta
<emphasis id="26BEBF35FF9CFFBA05F7FC52FBB742AD" box="[1011,1044,981,1007]" italics="true" pageId="2" pageNumber="565">st5</emphasis>
. Anal shield subtriangular, anterior margin rounded and triangular posteriorly, length 3436, width 3032, anterior half with lineate ornamentation, with a pair of lateral pores (
<emphasis id="26BEBF35FF9CFFBA07CCFB9AFE534575" box="[456,496,1053,1079]" italics="true" pageId="2" pageNumber="565">gv3</emphasis>
), para-anal setae (1416) slightly shorter than post-anal seta (1517), cribrum small. Opisthogastric cuticle with six pairs of smooth and uniform in length setae (1518) (
<emphasis id="26BEBF35FF9CFFBA0267FBC6FB634519" box="[1123,1216,1089,1115]" italics="true" pageId="2" pageNumber="565">Jv1Jv4</emphasis>
,
<emphasis id="26BEBF35FF9CFFBA02CBFBC6FA8F4519" box="[1231,1324,1089,1115]" italics="true" pageId="2" pageNumber="565">Zv1 Zv</emphasis>
2), except
<emphasis id="26BEBF35FF9CFFBA0693FBE2FF1D453D" box="[151,190,1125,1151]" italics="true" pageId="2" pageNumber="565">Jv3</emphasis>
(2426) and
<emphasis id="26BEBF35FF9CFFBA0752FBE2FEDE453D" box="[342,381,1125,1151]" italics="true" pageId="2" pageNumber="565">Jv4</emphasis>
(3133). Exopodal and parapodal platelets strip-like behind coxae IV. Peritreme short, barely reaching anterior margin of coxa II, peritrematal shield with small protrusion on outer margin, peritrematal shield narrow, free from exopodal shields, each shield bearing five discernible pore-like structures, a lyrifissure
<emphasis id="26BEBF35FF9CFFBA0345FB29FAF44585" box="[1345,1367,1198,1223]" italics="true" pageId="2" pageNumber="565">ip</emphasis>
and a gland pore
<emphasis id="26BEBF35FF9CFFBA0711FB55FE9245A9" box="[277,305,1234,1259]" italics="true" pageId="2" pageNumber="565">gp</emphasis>
at level of between coxae II and III, two lyrifissures
<emphasis id="26BEBF35FF9CFFBA0587FB55FC3A45A9" box="[899,921,1234,1259]" italics="true" pageId="2" pageNumber="565">ip</emphasis>
and a gland pore
<emphasis id="26BEBF35FF9CFFBA0265FB55FBDE45A9" box="[1121,1149,1234,1259]" italics="true" pageId="2" pageNumber="565">gp</emphasis>
on post-stigmatic section (
<figureCitation id="8CF17FA2FF9CFFBA069BFB72FF4B444D" box="[159,232,1269,1295]" captionStart="FIGURES 15" captionStartId="3.[151,264,1959,1984]" captionTargetBox="[191,1383,188,1893]" captionTargetId="figure@3.[151,1434,166,1939]" captionTargetPageId="3" captionText="FIGURES 15. Gaeolaelaps lankaensis sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, subcapitulum; 4, epistome; 5, chelicera." httpUri="https://zenodo.org/record/3246230/files/figure.png" pageId="2" pageNumber="565">Figs 2</figureCitation>
,
<figureCitation id="8CF17FA2FF9CFFBA06F0FB72FE81444D" box="[244,290,1269,1295]" captionStart="FIGURES 613" captionStartId="4.[151,264,1929,1954]" captionTargetBox="[192,1390,181,1895]" captionTargetId="figure@4.[177,1410,166,1909]" captionTargetPageId="4" captionText="FIGURES 613. DIC micrographs of Gaeolaelaps lankaensis sp. nov., female. 6, idiosoma in dorsal view; 7, idiosoma in ventral view; 8, sternal shield; 9, genital shield and opisthogastric area; 10, opisthonotal region; 11, epistome; 12, palp; 13, chelicera." httpUri="https://zenodo.org/record/3246232/files/figure.png" pageId="2" pageNumber="565">79</figureCitation>
).
</paragraph>
<paragraph id="14756327FF9CFFBA06C3FA9EFD6A47D9" blockId="2.[151,1437,152,1979]" pageId="2" pageNumber="565">
<taxonomicName id="D3CA18A4FF9CFFBA06C3FA9EFEF54471" authorityName="Krefft" authorityYear="1871" box="[199,342,1305,1331]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="2" pageNumber="565" phylum="Nematoda" rank="genus">
<emphasis id="26BEBF35FF9CFFBA06C3FA9EFEF54471" box="[199,342,1305,1331]" italics="true" pageId="2" pageNumber="565">Gnathosoma</emphasis>
</taxonomicName>
(
<figureCitation id="8CF17FA2FF9CFFBA0761FA9EFE694471" box="[357,458,1304,1331]" captionStart="FIGURES 15" captionStartId="3.[151,264,1959,1984]" captionTargetBox="[191,1383,188,1893]" captionTargetId="figure@3.[151,1434,166,1939]" captionTargetPageId="3" captionText="FIGURES 15. Gaeolaelaps lankaensis sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, subcapitulum; 4, epistome; 5, chelicera." httpUri="https://zenodo.org/record/3246230/files/figure.png" pageId="2" pageNumber="565">Figs 35</figureCitation>
,
<figureCitation id="8CF17FA2FF9CFFBA07D1FA9EFDBE4471" box="[469,541,1305,1331]" captionStart="FIGURES 613" captionStartId="4.[151,264,1929,1954]" captionTargetBox="[192,1390,181,1895]" captionTargetId="figure@4.[177,1410,166,1909]" captionTargetPageId="4" captionText="FIGURES 613. DIC micrographs of Gaeolaelaps lankaensis sp. nov., female. 6, idiosoma in dorsal view; 7, idiosoma in ventral view; 8, sternal shield; 9, genital shield and opisthogastric area; 10, opisthonotal region; 11, epistome; 12, palp; 13, chelicera." httpUri="https://zenodo.org/record/3246232/files/figure.png" pageId="2" pageNumber="565">1113</figureCitation>
). Hypostomal groove with six transverse rows of denticles, each row with about 819 small teeth, and smooth anterior and posterior transverse lines (
<figureCitation id="8CF17FA2FF9CFFBA05ADFABAFC4D4415" box="[937,1006,1341,1367]" captionStart="FIGURES 15" captionStartId="3.[151,264,1959,1984]" captionTargetBox="[191,1383,188,1893]" captionTargetId="figure@3.[151,1434,166,1939]" captionTargetPageId="3" captionText="FIGURES 15. Gaeolaelaps lankaensis sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, subcapitulum; 4, epistome; 5, chelicera." httpUri="https://zenodo.org/record/3246230/files/figure.png" pageId="2" pageNumber="565">Fig. 3</figureCitation>
). Hypostome with four pairs of setae, internal posterior hypostomal setae
<emphasis id="26BEBF35FF9CFFBA042CFAE6FDE74439" box="[552,580,1377,1403]" italics="true" pageId="2" pageNumber="565">h1</emphasis>
longest (2931),
<emphasis id="26BEBF35FF9CFFBA050EFAE6FC844439" box="[778,807,1377,1403]" italics="true" pageId="2" pageNumber="565">h2</emphasis>
(1315),
<emphasis id="26BEBF35FF9CFFBA0591FAE6FC124439" box="[917,945,1377,1403]" italics="true" pageId="2" pageNumber="565">h3</emphasis>
(2225), palpcoxal setae (
<emphasis id="26BEBF35FF9CFFBA02D8FAE5FB554439" box="[1244,1270,1378,1403]" italics="true" pageId="2" pageNumber="565">pc</emphasis>
) (1719) (
<figureCitation id="8CF17FA2FF9CFFBA036BFAE6FF0444DD" captionStart="FIGURES 15" captionStartId="3.[151,264,1959,1984]" captionTargetBox="[191,1383,188,1893]" captionTargetId="figure@3.[151,1434,166,1939]" captionTargetPageId="3" captionText="FIGURES 15. Gaeolaelaps lankaensis sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, subcapitulum; 4, epistome; 5, chelicera." httpUri="https://zenodo.org/record/3246230/files/figure.png" pageId="2" pageNumber="565">Fig. 3</figureCitation>
). Anterior margin of epistome undulating, convex central part with few denticles, almost smooth, lateral processes densely denticulate (
<figureCitation id="8CF17FA2FF9CFFBA0784FA2EFE684481" box="[384,459,1449,1475]" captionStart="FIGURES 15" captionStartId="3.[151,264,1959,1984]" captionTargetBox="[191,1383,188,1893]" captionTargetId="figure@3.[151,1434,166,1939]" captionTargetPageId="3" captionText="FIGURES 15. Gaeolaelaps lankaensis sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, subcapitulum; 4, epistome; 5, chelicera." httpUri="https://zenodo.org/record/3246230/files/figure.png" pageId="2" pageNumber="565">Figs 4</figureCitation>
,
<figureCitation id="8CF17FA2FF9CFFBA07DCFA2EFE564481" box="[472,501,1449,1475]" captionStart="FIGURES 613" captionStartId="4.[151,264,1929,1954]" captionTargetBox="[192,1390,181,1895]" captionTargetId="figure@4.[177,1410,166,1909]" captionTargetPageId="4" captionText="FIGURES 613. DIC micrographs of Gaeolaelaps lankaensis sp. nov., female. 6, idiosoma in dorsal view; 7, idiosoma in ventral view; 8, sternal shield; 9, genital shield and opisthogastric area; 10, opisthonotal region; 11, epistome; 12, palp; 13, chelicera." httpUri="https://zenodo.org/record/3246232/files/figure.png" pageId="2" pageNumber="565">11</figureCitation>
). Corniculi robust and horn-like, reaching mid-level of palp femur. Internal malae with median and lateral projections, fringed, inner lobes touching with outer lobes; labrum with pilose surface. Chaetotaxy of palps: trochanter 2, femur 5, genu 6, tibia 14, tarsus 15, all setae smooth and needle-like except
<emphasis id="26BEBF35FF9CFFBA0360FA76FAD94749" box="[1380,1402,1521,1547]" italics="true" pageId="2" pageNumber="565">al</emphasis>
on palp femur and
<emphasis id="26BEBF35FF9CFFBA0740F992FEF9476D" box="[324,346,1557,1583]" italics="true" pageId="2" pageNumber="565">al</emphasis>
,
<emphasis id="26BEBF35FF9CFFBA0762F992FE20476D" box="[358,387,1557,1583]" italics="true" pageId="2" pageNumber="565">d1</emphasis>
on palp genu thickened and apically spatulate, palp tarsal claw three-tined (
<figureCitation id="8CF17FA2FF9CFFBA02C1F992FAB3476D" box="[1221,1296,1557,1583]" captionStart="FIGURES 613" captionStartId="4.[151,264,1929,1954]" captionTargetBox="[192,1390,181,1895]" captionTargetId="figure@4.[177,1410,166,1909]" captionTargetPageId="4" captionText="FIGURES 613. DIC micrographs of Gaeolaelaps lankaensis sp. nov., female. 6, idiosoma in dorsal view; 7, idiosoma in ventral view; 8, sternal shield; 9, genital shield and opisthogastric area; 10, opisthonotal region; 11, epistome; 12, palp; 13, chelicera." httpUri="https://zenodo.org/record/3246232/files/figure.png" pageId="2" pageNumber="565">Fig 12</figureCitation>
). Fixed digit of chelicera with four teeth of varying sizes distally and a larger proximal tooth, a setaceous pilus dentilis, dorsal cheliceral seta prostrate, arthrodial membrane with a rounded flap and normal filaments, cheliceral lyrifissures distinct, movable digit with two teeth (
<figureCitation id="8CF17FA2FF9CFFBA044CF906FD3247D9" box="[584,657,1664,1691]" captionStart="FIGURES 15" captionStartId="3.[151,264,1959,1984]" captionTargetBox="[191,1383,188,1893]" captionTargetId="figure@3.[151,1434,166,1939]" captionTargetPageId="3" captionText="FIGURES 15. Gaeolaelaps lankaensis sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, subcapitulum; 4, epistome; 5, chelicera." httpUri="https://zenodo.org/record/3246230/files/figure.png" pageId="2" pageNumber="565">Figs 5</figureCitation>
,
<figureCitation id="8CF17FA2FF9CFFBA0499F906FD1F47D9" box="[669,700,1665,1691]" captionStart="FIGURES 613" captionStartId="4.[151,264,1929,1954]" captionTargetBox="[192,1390,181,1895]" captionTargetId="figure@4.[177,1410,166,1909]" captionTargetPageId="4" captionText="FIGURES 613. DIC micrographs of Gaeolaelaps lankaensis sp. nov., female. 6, idiosoma in dorsal view; 7, idiosoma in ventral view; 8, sternal shield; 9, genital shield and opisthogastric area; 10, opisthonotal region; 11, epistome; 12, palp; 13, chelicera." httpUri="https://zenodo.org/record/3246232/files/figure.png" pageId="2" pageNumber="565">13</figureCitation>
).
</paragraph>
<paragraph id="14756327FF9CFFBA06C3F921FB3C46F9" blockId="2.[151,1437,152,1979]" pageId="2" pageNumber="565">
<emphasis id="26BEBF35FF9CFFBA06C3F921FF5F47FD" box="[199,252,1702,1727]" italics="true" pageId="2" pageNumber="565">Legs</emphasis>
(
<figureCitation id="8CF17FA2FF9CFFBA070FF922FE2E47FD" box="[267,397,1701,1727]" captionStart="FIGURE 1417" captionStartId="5.[151,250,1949,1974]" captionTargetBox="[165,1427,241,1901]" captionTargetId="figure@5.[151,1434,197,1927]" captionTargetPageId="5" captionText="FIGURE 1417. Gaeolaelaps lankaensis sp. nov., female. 14, leg I (trochanter-tibia); 15, leg II; 16, leg III; 17, leg IV." httpUri="https://zenodo.org/record/3246236/files/figure.png" pageId="2" pageNumber="565">Figs 1417</figureCitation>
). Legs II and III short (223229, 161167), I and IV longer (322329, 266273). Chaetotaxy normal for free living
<taxonomicName id="D3CA18A4FF9CFFBA078AF94EFDAE47A1" box="[398,525,1737,1763]" class="Arachnida" family="Laelapidae" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="565" phylum="Arthropoda" rank="family">Laelapidae</taxonomicName>
: Leg I (
<figureCitation id="8CF17FA2FF9CFFBA0465F94EFD1747A1" box="[609,692,1737,1763]" captionStart="FIGURE 1417" captionStartId="5.[151,250,1949,1974]" captionTargetBox="[165,1427,241,1901]" captionTargetId="figure@5.[151,1434,197,1927]" captionTargetPageId="5" captionText="FIGURE 1417. Gaeolaelaps lankaensis sp. nov., female. 14, leg I (trochanter-tibia); 15, leg II; 16, leg III; 17, leg IV." httpUri="https://zenodo.org/record/3246236/files/figure.png" pageId="2" pageNumber="565">Fig. 14</figureCitation>
): coxa 0 0/1, 0/1 0, trochanter 1 0/1, 1/2 1, femur 2 2/1, 3/3 2, genu 2 3/2, 3/1 2, tibia 2 3/2, 3/1 2. Leg II (
<figureCitation id="8CF17FA2FF9CFFBA0444F96AFD374645" box="[576,660,1772,1799]" captionStart="FIGURE 1417" captionStartId="5.[151,250,1949,1974]" captionTargetBox="[165,1427,241,1901]" captionTargetId="figure@5.[151,1434,197,1927]" captionTargetPageId="5" captionText="FIGURE 1417. Gaeolaelaps lankaensis sp. nov., female. 14, leg I (trochanter-tibia); 15, leg II; 16, leg III; 17, leg IV." httpUri="https://zenodo.org/record/3246236/files/figure.png" pageId="2" pageNumber="565">Fig. 15</figureCitation>
): coxa 0 0/1, 0/1 0, trochanter 1 1/1, 0/2 0, femur 1 3/1, 2/2 1, genu 2 3/1, 2/1 2, tibia 2 2/1, 2/1 2 (all ventral setae thickened). Leg III (
<figureCitation id="8CF17FA2FF9CFFBA057AF896FC70466E" box="[894,979,1809,1836]" captionStart="FIGURE 1417" captionStartId="5.[151,250,1949,1974]" captionTargetBox="[165,1427,241,1901]" captionTargetId="figure@5.[151,1434,197,1927]" captionTargetPageId="5" captionText="FIGURE 1417. Gaeolaelaps lankaensis sp. nov., female. 14, leg I (trochanter-tibia); 15, leg II; 16, leg III; 17, leg IV." httpUri="https://zenodo.org/record/3246236/files/figure.png" pageId="2" pageNumber="565">Fig. 16</figureCitation>
): coxa 0 0/1, 0/1 0, trochanter 1 0/1, 0/2 1, femur 1 2/1, 1/0 1, genu 2 2/1, 2/1 1 (all ventral setae thickened), tibia: 2 1/1, 2/1 1. Leg IV (
<figureCitation id="8CF17FA2FF9CFFBA02E1F8B2FA99460D" box="[1253,1338,1845,1871]" captionStart="FIGURE 1417" captionStartId="5.[151,250,1949,1974]" captionTargetBox="[165,1427,241,1901]" captionTargetId="figure@5.[151,1434,197,1927]" captionTargetPageId="5" captionText="FIGURE 1417. Gaeolaelaps lankaensis sp. nov., female. 14, leg I (trochanter-tibia); 15, leg II; 16, leg III; 17, leg IV." httpUri="https://zenodo.org/record/3246236/files/figure.png" pageId="2" pageNumber="565">Fig. 17</figureCitation>
): coxa 0 0/1, 0/0 0, trochanter 1 0/2, 0/1 1, femur 1 2/1, 1/0 1, genu 2 2/1, 3/0 1 (
<emphasis id="26BEBF35FF9CFFBA05B8F8DDFC754631" box="[956,982,1882,1907]" italics="true" pageId="2" pageNumber="565">av</emphasis>
thickened), tibia 2 1/1, 3/1 2 (all ventral setae thickened). Tarsi II IV with 18 setae (3 3/2, 3/2 3 +
<emphasis id="26BEBF35FF9CFFBA050BF8F9FC9346D5" box="[783,816,1918,1943]" italics="true" pageId="2" pageNumber="565">mv</emphasis>
,
<emphasis id="26BEBF35FF9CFFBA0539F8FAFCFC46D5" box="[829,863,1917,1943]" italics="true" pageId="2" pageNumber="565">md</emphasis>
). All pretarsi with well developed paired claws and rounded pulvilli and a long thin stalk, ventral and lateral setae on tarsus II and III thickened.
</paragraph>
<caption id="40B533AFFF9DFFBB0693F820FEB046A6" httpUri="https://zenodo.org/record/3246230/files/figure.png" pageId="3" pageNumber="566" startId="3.[151,264,1959,1984]" targetBox="[191,1383,188,1893]" targetPageId="3">
<paragraph id="14756327FF9DFFBB0693F820FEB046A6" blockId="3.[151,1436,1959,2020]" pageId="3" pageNumber="566">
<emphasis id="26BEBF35FF9DFFBB0693F820FE994682" bold="true" box="[151,314,1959,1984]" pageId="3" pageNumber="566">FIGURES 15.</emphasis>
<taxonomicName id="D3CA18A4FF9DFFBB073BF82FFD8C4682" authority="Joharchi &amp; Khaustov &amp; Ermilov, 2019" authorityName="Joharchi &amp; Khaustov &amp; Ermilov" authorityYear="2019" box="[319,559,1960,1984]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="566" phylum="Arthropoda" rank="species" species="lankaensis" status="sp. nov.">
<emphasis id="26BEBF35FF9DFFBB073BF82FFD8C4682" box="[319,559,1960,1984]" italics="true" pageId="3" pageNumber="566">Gaeolaelaps lankaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3D8D024EFF9DFFBB0431F82FFD274682" box="[565,644,1960,1984]" pageId="3" pageNumber="566" rank="species">
<emphasis id="26BEBF35FF9DFFBB0431F82FFD274682" bold="true" box="[565,644,1960,1984]" pageId="3" pageNumber="566">sp. nov.</emphasis>
</taxonomicNameLabel>
, female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, subcapitulum; 4, epistome; 5, chelicera.
</paragraph>
</caption>
<caption id="40B533AFFF9AFFBC0693F80EFF5946A8" httpUri="https://zenodo.org/record/3246232/files/figure.png" pageId="4" pageNumber="567" startId="4.[151,264,1929,1954]" targetBox="[192,1390,181,1895]" targetPageId="4">
<paragraph id="14756327FF9AFFBC0693F80EFF5946A8" blockId="4.[151,1437,1929,2026]" pageId="4" pageNumber="567">
<emphasis id="26BEBF35FF9AFFBC0693F80EFEE946E0" bold="true" box="[151,330,1929,1954]" pageId="4" pageNumber="567">FIGURES 613.</emphasis>
DIC micrographs of
<taxonomicName id="D3CA18A4FF9AFFBC0429F80DFC8246E0" authority="Joharchi &amp; Khaustov &amp; Ermilov, 2019" authorityName="Joharchi &amp; Khaustov &amp; Ermilov" authorityYear="2019" box="[557,801,1930,1954]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="567" phylum="Arthropoda" rank="species" species="lankaensis" status="sp. nov.">
<emphasis id="26BEBF35FF9AFFBC0429F80DFC8246E0" box="[557,801,1930,1954]" italics="true" pageId="4" pageNumber="567">Gaeolaelaps lankaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3D8D024EFF9AFFBC052EF80DFCDE46E0" box="[810,893,1930,1954]" pageId="4" pageNumber="567" rank="species">
<emphasis id="26BEBF35FF9AFFBC052EF80DFCDE46E0" bold="true" box="[810,893,1930,1954]" pageId="4" pageNumber="567">sp. nov.</emphasis>
</taxonomicNameLabel>
, female. 6, idiosoma in dorsal view; 7, idiosoma in ventral view; 8, sternal shield; 9, genital shield and opisthogastric area; 10, opisthonotal region; 11, epistome; 12, palp; 13, chelicera.
</paragraph>
</caption>
<caption id="40B533AFFF9BFFBD0693F81AFAEF46F7" box="[151,1356,1949,1974]" httpUri="https://zenodo.org/record/3246236/files/figure.png" pageId="5" pageNumber="568" startId="5.[151,250,1949,1974]" targetBox="[165,1427,241,1901]" targetPageId="5">
<paragraph id="14756327FF9BFFBD0693F81AFAEF46F7" blockId="5.[151,1356,1949,1974]" box="[151,1356,1949,1974]" pageId="5" pageNumber="568">
<emphasis id="26BEBF35FF9BFFBD0693F81AFEE546F4" bold="true" box="[151,326,1949,1974]" pageId="5" pageNumber="568">FIGURE 1417.</emphasis>
<taxonomicName id="D3CA18A4FF9BFFBD0748F81AFD9E46F7" authority="Joharchi &amp; Khaustov &amp; Ermilov, 2019" authorityName="Joharchi &amp; Khaustov &amp; Ermilov" authorityYear="2019" box="[332,573,1949,1973]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="568" phylum="Arthropoda" rank="species" species="lankaensis" status="sp. nov.">
<emphasis id="26BEBF35FF9BFFBD0748F81AFD9E46F7" box="[332,573,1949,1973]" italics="true" pageId="5" pageNumber="568">Gaeolaelaps lankaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3D8D024EFF9BFFBD0440F819FD3746F4" box="[580,660,1950,1974]" pageId="5" pageNumber="568" rank="species">
<emphasis id="26BEBF35FF9BFFBD0440F819FD3746F4" bold="true" box="[580,660,1950,1974]" pageId="5" pageNumber="568">sp. nov.</emphasis>
</taxonomicNameLabel>
, female. 14, leg I (trochanter-tibia); 15, leg II; 16, leg III; 17, leg IV.
</paragraph>
</caption>
<paragraph id="14756327FF98FFBE06C3FF1DFCCB41F1" blockId="6.[151,1437,153,971]" box="[199,872,153,179]" pageId="6" pageNumber="569">
<emphasis id="26BEBF35FF98FFBE06C3FF1DFE6C41F1" box="[199,463,154,179]" italics="true" pageId="6" pageNumber="569">Insemination structures</emphasis>
: Not seen, apparently unsclerotised.
</paragraph>
<paragraph id="14756327FF98FFBE06C3FF3AFD9C4195" blockId="6.[151,1437,153,971]" box="[199,575,188,215]" pageId="6" pageNumber="569">
<emphasis id="26BEBF35FF98FFBE06C3FF3AFEA74195" bold="true" box="[199,260,189,215]" pageId="6" pageNumber="569">Male</emphasis>
and
<emphasis id="26BEBF35FF98FFBE073EFF3BFE614195" bold="true" box="[314,450,188,215]" pageId="6" pageNumber="569">immatures.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="5CD030ACFF98FFBE06C3FF66FC3641B9" box="[199,917,225,251]" pageId="6" pageNumber="569" type="etymology">
<paragraph id="14756327FF98FFBE06C3FF66FC3641B9" blockId="6.[151,1437,153,971]" box="[199,917,225,251]" pageId="6" pageNumber="569">
<emphasis id="26BEBF35FF98FFBE06C3FF66FEED41B9" bold="true" box="[199,334,225,251]" pageId="6" pageNumber="569">Etymology.</emphasis>
The specific name is derived from the
<typeStatus id="CB71DD85FF98FFBE0500FF66FC9741B9" box="[772,820,225,251]" pageId="6" pageNumber="569">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="5CD030ACFF98FFBE06C3FE83FBFE4289" pageId="6" pageNumber="569" type="diagnosis">
<paragraph id="14756327FF98FFBE06C3FE83FF5A421D" blockId="6.[151,1437,153,971]" pageId="6" pageNumber="569">
<emphasis id="26BEBF35FF98FFBE06C3FE83FE77405D" bold="true" box="[199,468,260,287]" pageId="6" pageNumber="569">Differential diagnosis.</emphasis>
The new species is unique within
<emphasis id="26BEBF35FF98FFBE0584FE82FBAD405D" box="[896,1038,261,287]" italics="true" pageId="6" pageNumber="569">Gaeolaelaps</emphasis>
by the following combination of characters: palp tarsal claw three-tined (
<figureCitation id="8CF17FA2FF98FFBE046FFEAEFD614001" box="[619,706,297,323]" captionStart="FIGURES 613" captionStartId="4.[151,264,1929,1954]" captionTargetBox="[192,1390,181,1895]" captionTargetId="figure@4.[177,1410,166,1909]" captionTargetPageId="4" captionText="FIGURES 613. DIC micrographs of Gaeolaelaps lankaensis sp. nov., female. 6, idiosoma in dorsal view; 7, idiosoma in ventral view; 8, sternal shield; 9, genital shield and opisthogastric area; 10, opisthonotal region; 11, epistome; 12, palp; 13, chelicera." httpUri="https://zenodo.org/record/3246232/files/figure.png" pageId="6" pageNumber="569">Fig. 12</figureCitation>
),
<emphasis id="26BEBF35FF98FFBE04DFFEAEFD5F4001" box="[731,764,297,323]" italics="true" pageId="6" pageNumber="569">st1</emphasis>
located outside sternal shield (
<figureCitation id="8CF17FA2FF98FFBE026FFEAEFB114001" box="[1131,1202,297,323]" captionStart="FIGURES 613" captionStartId="4.[151,264,1929,1954]" captionTargetBox="[192,1390,181,1895]" captionTargetId="figure@4.[177,1410,166,1909]" captionTargetPageId="4" captionText="FIGURES 613. DIC micrographs of Gaeolaelaps lankaensis sp. nov., female. 6, idiosoma in dorsal view; 7, idiosoma in ventral view; 8, sternal shield; 9, genital shield and opisthogastric area; 10, opisthonotal region; 11, epistome; 12, palp; 13, chelicera." httpUri="https://zenodo.org/record/3246232/files/figure.png" pageId="6" pageNumber="569">Fig. 8</figureCitation>
), dorsal shield with 35 pairs of moderately long and simple setae,
<emphasis id="26BEBF35FF98FFBE04B5FECAFD6C4025" box="[689,719,333,359]" italics="true" pageId="6" pageNumber="569">Z5</emphasis>
&amp;
<emphasis id="26BEBF35FF98FFBE04FCFECAFCB74025" box="[760,788,333,359]" italics="true" pageId="6" pageNumber="569">S5</emphasis>
on soft cuticle posterior to shield, without
<emphasis id="26BEBF35FF98FFBE0308FEC9FA8B4025" box="[1292,1320,334,359]" italics="true" pageId="6" pageNumber="569">Zx</emphasis>
setae and unpaired supernumerary seta (
<emphasis id="26BEBF35FF98FFBE07F2FEF5FDAC40C9" box="[502,527,370,395]" italics="true" pageId="6" pageNumber="569">Jx</emphasis>
) (
<figureCitation id="8CF17FA2FF98FFBE042EFEF6FDD740C9" box="[554,628,369,395]" captionStart="FIGURES 15" captionStartId="3.[151,264,1959,1984]" captionTargetBox="[191,1383,188,1893]" captionTargetId="figure@3.[151,1434,166,1939]" captionTargetPageId="3" captionText="FIGURES 15. Gaeolaelaps lankaensis sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, subcapitulum; 4, epistome; 5, chelicera." httpUri="https://zenodo.org/record/3246230/files/figure.png" pageId="6" pageNumber="569">Figs 1</figureCitation>
,
<figureCitation id="8CF17FA2FF98FFBE0482FEF6FD3540CE" box="[646,662,369,396]" captionStart="FIGURES 613" captionStartId="4.[151,264,1929,1954]" captionTargetBox="[192,1390,181,1895]" captionTargetId="figure@4.[177,1410,166,1909]" captionTargetPageId="4" captionText="FIGURES 613. DIC micrographs of Gaeolaelaps lankaensis sp. nov., female. 6, idiosoma in dorsal view; 7, idiosoma in ventral view; 8, sternal shield; 9, genital shield and opisthogastric area; 10, opisthonotal region; 11, epistome; 12, palp; 13, chelicera." httpUri="https://zenodo.org/record/3246232/files/figure.png" pageId="6" pageNumber="569">6</figureCitation>
), tritosternum with pair of very long pilose laciniae (
<figureCitation id="8CF17FA2FF98FFBE0303FEF6FAEC40C9" box="[1287,1359,369,395]" captionStart="FIGURES 15" captionStartId="3.[151,264,1959,1984]" captionTargetBox="[191,1383,188,1893]" captionTargetId="figure@3.[151,1434,166,1939]" captionTargetPageId="3" captionText="FIGURES 15. Gaeolaelaps lankaensis sp. nov., female. 1, dorsal idiosoma; 2, ventral idiosoma; 3, subcapitulum; 4, epistome; 5, chelicera." httpUri="https://zenodo.org/record/3246230/files/figure.png" pageId="6" pageNumber="569">Fig. 2</figureCitation>
), peritreme short (barely reaching anterior margin of coxa II) and femur of leg II with ten (1 3/1, 2/2 1,
<emphasis id="26BEBF35FF98FFBE0319FE12FAE240ED" box="[1309,1345,405,431]" italics="true" pageId="6" pageNumber="569">al2</emphasis>
absent) setae instead of the usual 11. In the key to
<emphasis id="26BEBF35FF98FFBE0489FE3EFCB84091" box="[653,795,441,467]" italics="true" pageId="6" pageNumber="569">Gaeolaelaps</emphasis>
species with short peritremes provided in
<bibRefCitation id="705B1ED6FF98FFBE030CFE3EFEA840BA" author="Vatankhah, F. &amp; Nemati, A. &amp; Esfandiari, M. &amp; Shishehbor, P." pageId="6" pageNumber="569" pagination="566 - 574" refId="ref9024" refString="Vatankhah, F., Nemati, A., Esfandiari, M. &amp; Shishehbor, P. (2016) Description of a new species of Gaeolaelaps (Acari: Laelapidae) from Iran, with a key to world species of the genus with short peritremes. Zootaxa, 4121 (5), 566 - 574. https: // doi. org / 10.11646 / zootaxa. 4121.5.6" type="journal article" year="2016">
Vatankhah
<emphasis id="26BEBF35FF98FFBE038CFE3DFF0E40B5" italics="true" pageId="6" pageNumber="569">et al</emphasis>
. (2016)
</bibRefCitation>
, this species came to couplet 11, but the uncommon character state of 35 pairs of dorsal shield setae, makes couplet choice problematic. In the classification of
<bibRefCitation id="705B1ED6FF98FFBE0527FD86FC0D4359" author="Karg, W." box="[803,942,513,539]" pageId="6" pageNumber="569" pagination="233 - 256" refId="ref8287" refString="Karg, W. (1982) Zur Kenntnis der Raubmilbeng Hypoaspis Canestrini, 1884 (Acarina, Parasitiformes). Mitteilungen aus dem Zoologischen Museum in Berlin, 58, 233 - 256. https: // doi. org / 10.1002 / mmnz. 4830580204" type="journal article" year="1982">Karg (1982)</bibRefCitation>
,
<taxonomicName id="D3CA18A4FF98FFBE05BEFD86FBF14359" authorityName="Joharchi &amp; Khaustov &amp; Ermilov" authorityYear="2019" box="[954,1106,513,539]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="569" phylum="Arthropoda" rank="species" species="lankaensis">
<emphasis id="26BEBF35FF98FFBE05BEFD86FC6D4359" box="[954,974,513,539]" italics="true" pageId="6" pageNumber="569">G</emphasis>
.
<emphasis id="26BEBF35FF98FFBE05D8FD86FBF14359" box="[988,1106,513,539]" italics="true" pageId="6" pageNumber="569">lankaensis</emphasis>
</taxonomicName>
falls into the
<emphasis id="26BEBF35FF98FFBE02E8FD85FAE64359" box="[1260,1349,514,539]" italics="true" pageId="6" pageNumber="569">angusta</emphasis>
species group of
<taxonomicName id="D3CA18A4FF98FFBE0706FDA1FD80437D" authority="" authorityName="Evans &amp; Till" authorityYear="1966" box="[258,547,549,575]" class="Arachnida" family="Laelapidae" genus="Hypoaspis" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="569" phylum="Arthropoda" rank="subGenus" subGenus="Gaeolaelaps">
<emphasis id="26BEBF35FF98FFBE0706FDA1FEDB437D" box="[258,376,550,575]" italics="true" pageId="6" pageNumber="569">Hypoaspis</emphasis>
(
<emphasis id="26BEBF35FF98FFBE078EFDA2FDBB437D" box="[394,536,549,575]" italics="true" pageId="6" pageNumber="569">Gaeolaelaps</emphasis>
)
</taxonomicName>
, because its dorsal shield narrows from level of setae
<emphasis id="26BEBF35FF98FFBE0296FDA2FB08437D" box="[1170,1195,549,575]" italics="true" pageId="6" pageNumber="569">r3</emphasis>
, and the shield lacks
<emphasis id="26BEBF35FF98FFBE0693FDCDFF104321" box="[151,179,586,611]" italics="true" pageId="6" pageNumber="569">Zx</emphasis>
setae on the opisthonotal region.
<taxonomicName id="D3CA18A4FF98FFBE0434FDCEFC984321" authorityName="Joharchi &amp; Khaustov &amp; Ermilov" authorityYear="2019" box="[560,827,585,611]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="569" phylum="Arthropoda" rank="species" species="lankaensis">
<emphasis id="26BEBF35FF98FFBE0434FDCEFC984321" box="[560,827,585,611]" italics="true" pageId="6" pageNumber="569">Gaeolaelaps lankaensis</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="D3CA18A4FF98FFBE05D4FDCEFAEA4326" authority="(Bhattacharyya, 1968)" authorityName="Bhattacharyya" authorityYear="1968" baseAuthorityName="Bhattacharyya" baseAuthorityYear="1968" box="[976,1353,585,612]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="569" phylum="Arthropoda" rank="species" species="tarsalis">
<emphasis id="26BEBF35FF98FFBE05D4FDCEFC474321" box="[976,996,585,611]" italics="true" pageId="6" pageNumber="569">G</emphasis>
.
<emphasis id="26BEBF35FF98FFBE05F7FDCEFBE44321" box="[1011,1095,585,611]" italics="true" pageId="6" pageNumber="569">tarsalis</emphasis>
(
<bibRefCitation id="705B1ED6FF98FFBE0252FDCEFAE24326" author="Bhattacharyya, S. K." box="[1110,1345,585,612]" pageId="6" pageNumber="569" pagination="527 - 549" refId="ref7317" refString="Bhattacharyya, S. K. (1968) Studies on Indian mites (Acarina: Mesostigmata). 6. Six records and descriptions of nine new species. Acarologia, 10, 527 - 549." type="journal article" year="1968">Bhattacharyya, 1968</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="D3CA18A4FF98FFBE0385FDCEFDBC43C5" authority="(Womersley, 1956)" authorityName="Womersley" authorityYear="1956" baseAuthorityName="Womersley" baseAuthorityYear="1956" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="569" phylum="Arthropoda" rank="species" species="queenslandicus">
<emphasis id="26BEBF35FF98FFBE0385FDCEFA364321" box="[1409,1429,585,611]" italics="true" pageId="6" pageNumber="569">G</emphasis>
.
<emphasis id="26BEBF35FF98FFBE0693FDEAFEE743C5" box="[151,324,621,647]" italics="true" pageId="6" pageNumber="569">queenslandicus</emphasis>
(
<bibRefCitation id="705B1ED6FF98FFBE0755FDEAFDB443C5" author="Womersley, H." box="[337,535,620,647]" pageId="6" pageNumber="569" pagination="505 - 599" refId="ref9223" refString="Womersley, H. (1956) On some new Acarina-Mesostigmata from Australia, New Zealand and New Guinea. Journal of the Linnean Society of London, Zoology, 42, 505 - 599. https: // doi. org / 10.1111 / j. 1096 - 3642.1956. tb 02218. x" type="journal article" year="1956">Womersley, 1956</bibRefCitation>
)
</taxonomicName>
(we followed here
<bibRefCitation id="705B1ED6FF98FFBE04F1FDEAFC6943C5" author="Nemati, A. &amp; Gwiazdowicz, D. &amp; Khalili-Moghadam, A." box="[757,970,621,647]" pageId="6" pageNumber="569" pagination="710 - 734" refId="ref8765" refString="Nemati, A., Gwiazdowicz, D. &amp; Khalili-Moghadam, A. (2018) New data on the knowledge of Gaeolaelaps mites (Acari: Mesostigmata: Laelapidae). Acarologia, 58, 710 - 734. https: // doi. org / 10.24349 / acarologia / 20184266" type="journal article" year="2018">
Nemati
<emphasis id="26BEBF35FF98FFBE054AFDE9FCDD43C5" box="[846,894,621,647]" italics="true" pageId="6" pageNumber="569">et al</emphasis>
., 2018
</bibRefCitation>
and consider
<taxonomicName id="D3CA18A4FF98FFBE0260FDEAFADC43C5" authority="(Karg, 1965)" authorityName="Karg" authorityYear="1965" baseAuthorityName="Karg" baseAuthorityYear="1965" box="[1124,1407,620,647]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="569" phylum="Arthropoda" rank="species" species="angustus">
<emphasis id="26BEBF35FF98FFBE0260FDEAFBDB43C5" box="[1124,1144,621,647]" italics="true" pageId="6" pageNumber="569">G</emphasis>
.
<emphasis id="26BEBF35FF98FFBE0282FDE9FB4943C5" box="[1158,1258,622,647]" italics="true" pageId="6" pageNumber="569">angustus</emphasis>
(
<bibRefCitation id="705B1ED6FF98FFBE02F3FDEAFAD543C5" author="Karg, W." box="[1271,1398,620,647]" pageId="6" pageNumber="569" pagination="193 - 340" refId="ref8200" refString="Karg, W. (1965) Larvalsystematische und phylogenetische Untersuchung sowie Revision des Systems der Gamasina Leach, 1915 (Acarina, Parasitiformes). Mitteilungen aus dem Zoologischen Museum in Berlin, 41, 193 - 340. https: // doi. org / 10.1002 / mmnz. 19650410207" type="journal article" year="1965">Karg, 1965</bibRefCitation>
)
</taxonomicName>
as a junior synonymy of
<taxonomicName id="D3CA18A4FF98FFBE0795FD16FDC343E9" baseAuthorityName="Womersley" baseAuthorityYear="1956" box="[401,608,657,683]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="569" phylum="Arthropoda" rank="species" species="queenslandicus">
<emphasis id="26BEBF35FF98FFBE0795FD16FE0643E9" box="[401,421,657,683]" italics="true" pageId="6" pageNumber="569">G</emphasis>
.
<emphasis id="26BEBF35FF98FFBE07B7FD16FDC343E9" box="[435,608,657,683]" italics="true" pageId="6" pageNumber="569">queenslandicus</emphasis>
</taxonomicName>
) in general appearance and also having a three-tined palp tarsal claw, but
<taxonomicName id="D3CA18A4FF98FFBE0693FD32FE02438D" authorityName="Joharchi &amp; Khaustov &amp; Ermilov" authorityYear="2019" box="[151,417,693,719]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="569" phylum="Arthropoda" rank="species" species="lankaensis">
<emphasis id="26BEBF35FF98FFBE0693FD32FE02438D" box="[151,417,693,719]" italics="true" pageId="6" pageNumber="569">Gaeolaelaps lankaensis</emphasis>
</taxonomicName>
can be easily distinguished from these two species by having 35 pairs of setae on the dorsal shield, short peritreme (barely reaching anterior margin of coxa II), without spine-like seta on femur of leg II, and with ten seate (
<emphasis id="26BEBF35FF98FFBE0742FD7AFEC94255" box="[326,362,765,791]" italics="true" pageId="6" pageNumber="569">al2</emphasis>
absent) (vs. 37 or 38 pairs in
<taxonomicName id="D3CA18A4FF98FFBE04C1FD7AFC364255" baseAuthorityName="Womersley" baseAuthorityYear="1956" box="[709,917,765,791]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="569" phylum="Arthropoda" rank="species" species="queenslandicus">
<emphasis id="26BEBF35FF98FFBE04C1FD7AFD7A4255" box="[709,729,765,791]" italics="true" pageId="6" pageNumber="569">G</emphasis>
.
<emphasis id="26BEBF35FF98FFBE04ECFD7AFC364255" box="[744,917,765,791]" italics="true" pageId="6" pageNumber="569">queenslandicus</emphasis>
</taxonomicName>
and 39 pairs in
<taxonomicName id="D3CA18A4FF98FFBE0255FD7AFB6A4255" baseAuthorityName="Bhattacharyya" baseAuthorityYear="1968" box="[1105,1225,765,791]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="569" phylum="Arthropoda" rank="species" species="tarsalis">
<emphasis id="26BEBF35FF98FFBE0255FD7AFBC64255" box="[1105,1125,765,791]" italics="true" pageId="6" pageNumber="569">G</emphasis>
.
<emphasis id="26BEBF35FF98FFBE0271FD7AFB6A4255" box="[1141,1225,765,791]" italics="true" pageId="6" pageNumber="569">tarsalis</emphasis>
</taxonomicName>
, peritreme long in both species (reach well to coxa I) and also femur II armed with spine-like setae in both species, with 11 setae, (
<emphasis id="26BEBF35FF98FFBE037CFCA6FA3F4279" box="[1400,1436,801,827]" italics="true" pageId="6" pageNumber="569">al2</emphasis>
present).
</paragraph>
<paragraph id="14756327FF98FFBE06C3FCEEFBFE4289" blockId="6.[151,1437,153,971]" pageId="6" pageNumber="569">
The new species also resembles
<taxonomicName id="D3CA18A4FF98FFBE0434FCEEFCD642C1" authority="(Berlese, 1917)" authorityName="Berlese" authorityYear="1917" baseAuthorityName="Berlese" baseAuthorityYear="1917" box="[560,885,873,899]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="569" phylum="Arthropoda" rank="species" species="atomarius">
<emphasis id="26BEBF35FF98FFBE0434FCEEFDE742C1" box="[560,580,873,899]" italics="true" pageId="6" pageNumber="569">G</emphasis>
.
<emphasis id="26BEBF35FF98FFBE0455FCEDFD6042C1" box="[593,707,874,899]" italics="true" pageId="6" pageNumber="569">atomarius</emphasis>
(
<bibRefCitation id="705B1ED6FF98FFBE04D5FCEEFCCE42C1" author="Berlese, A." box="[721,877,873,899]" pageId="6" pageNumber="569" pagination="125 - 177" refId="ref7231" refString="Berlese, A. (1917) Centuria seconda di Acari nuovi. Redia, 12, 125 - 177." type="journal article" year="1917">Berlese, 1917</bibRefCitation>
)
</taxonomicName>
(
<emphasis id="26BEBF35FF98FFBE0581FCEDFC6042C1" box="[901,963,874,899]" italics="true" pageId="6" pageNumber="569">sensu</emphasis>
<bibRefCitation id="705B1ED6FF98FFBE05CDFCEEFAA842C1" author="Van Aswegen, P. I. M. &amp; Loots, G. C." box="[969,1291,873,899]" pageId="6" pageNumber="569" pagination="167 - 213" refId="ref8968" refString="Van Aswegen, P. I. M. &amp; Loots, G. C. (1970) A taxonomic study of the genus Hypoaspis Canestrini sens. lat. (Acari: Laelapinae) in the Ethiopian region. Publicacoes Culturais da Companhia de Diamantes de Angola, 82, 167 - 213." type="journal article" year="1970">Van Aswegen &amp; Loots, 1970</bibRefCitation>
) and
<taxonomicName id="D3CA18A4FF98FFBE034DFCEEFE0E42E5" authority="Kazemi et al., 2014" authorityName="Kazemi" authorityYear="2014" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="569" phylum="Arthropoda" rank="species" species="khajooii">
<emphasis id="26BEBF35FF98FFBE034DFCEEFAFE42C1" box="[1353,1373,873,899]" italics="true" pageId="6" pageNumber="569">G</emphasis>
.
<emphasis id="26BEBF35FF98FFBE036EFCEEFF6942E5" italics="true" pageId="6" pageNumber="569">khajooii</emphasis>
<bibRefCitation id="705B1ED6FF98FFBE06D5FC0AFE0E42E5" author="Kazemi, Sh. &amp; Rajaei, A. &amp; Beaulieu, F." box="[209,429,909,935]" pageId="6" pageNumber="569" pagination="501 - 530" refId="ref8567" refString="Kazemi, Sh., Rajaei, A. &amp; Beaulieu, F. (2014) Two new species of Gaeolaelaps (Acari: Mesostigmata: Laelapidae) from Iran, with a revised generic concept and notes on significant morphological characters in the genus. Zootaxa, 3861 (6), 501 - 530. https: // doi. org / 10.11646 / zootaxa. 3861.6.1" type="journal article" year="2014">
Kazemi
<emphasis id="26BEBF35FF98FFBE072BFC09FEC342E5" box="[303,352,909,935]" italics="true" pageId="6" pageNumber="569">et al</emphasis>
., 2014
</bibRefCitation>
</taxonomicName>
which have a similar dorsal shield, but it can be easily distinguished from both species by having 35 setae on dorsal shield (
<quantity id="D332CEC2FF98FFBE0409FC36FDE44289" box="[525,583,945,971]" metricMagnitude="-1" metricUnit="m" metricValue="9.652" pageId="6" pageNumber="569" unit="in" value="38.0">38 in</quantity>
<taxonomicName id="D3CA18A4FF98FFBE044AFC36FD414289" baseAuthorityName="Berlese" baseAuthorityYear="1917" box="[590,738,945,971]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="569" phylum="Arthropoda" rank="species" species="atomarius">
<emphasis id="26BEBF35FF98FFBE044AFC36FDC14289" box="[590,610,945,971]" italics="true" pageId="6" pageNumber="569">G</emphasis>
.
<emphasis id="26BEBF35FF98FFBE0474FC35FD414289" box="[624,738,946,971]" italics="true" pageId="6" pageNumber="569">atomarius</emphasis>
</taxonomicName>
, z3 absent and
<quantity id="D332CEC2FF98FFBE058BFC36FC6B4289" box="[911,968,945,971]" metricMagnitude="-1" metricUnit="m" metricValue="9.906" pageId="6" pageNumber="569" unit="in" value="39.0">39 in</quantity>
<taxonomicName id="D3CA18A4FF98FFBE05CBFC36FBEE4289" box="[975,1101,945,971]" class="Arachnida" family="Laelapidae" genus="Gaeolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="569" phylum="Arthropoda" rank="species" species="khajooii">
<emphasis id="26BEBF35FF98FFBE05CBFC36FC404289" box="[975,995,945,971]" italics="true" pageId="6" pageNumber="569">G</emphasis>
.
<emphasis id="26BEBF35FF98FFBE05F5FC36FBEE4289" box="[1009,1101,945,971]" italics="true" pageId="6" pageNumber="569">khajooii</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>