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99 lines
12 KiB
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<document id="8CAA32E91CC48955D78D64A45E48F3A0" ID-DOI="10.15407/zoo2023.04.311" ID-ISSN="2707-7268" ID-Zenodo-Dep="10831776" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1699900057102" checkinUser="felipe" docAuthor="Yanenko, V. & Kovalchuk, О." docDate="2023" docId="485B878D917A0F3AF3DEFC06FD19F9B8" docLanguage="en" docName="Zoodiversity.57.4.311-322.pdf" docOrigin="Zoodiversity 57 (4)" docSource="http://dx.doi.org/10.15407/zoo2023.04.311" docStyle="DocumentStyle:65E6F982F72656C606C1639D3996B172.2:Zoodiversity.2020-.journal_article" docStyleId="65E6F982F72656C606C1639D3996B172" docStyleName="Zoodiversity.2020-.journal_article" docStyleVersion="2" docTitle="Testudo chernovi Khosatzky 1948" docType="treatment" docVersion="3" lastPageNumber="315" masterDocId="B462FFF5917E0F3EF378FFD3FFBAFFBB" masterDocTitle="Late Miocene Turtles Of Grytsiv (Western Ukraine) With Rodent Gnaw Marks On The Carapace Surface" masterLastPageNumber="322" masterPageNumber="311" pageNumber="315" updateTime="1710775831372" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-4.0">
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<mods:title id="123E4EE3B62C173F246BC922C1A32C4B">Late Miocene Turtles Of Grytsiv (Western Ukraine) With Rodent Gnaw Marks On The Carapace Surface</mods:title>
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<mods:name id="424907A8D379A37BCC4E666C8268ED51" type="personal">
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<mods:namePart id="9C15D6FD99B7907FF59F5ECA4BE29BC2">Yanenko, V.</mods:namePart>
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<mods:affiliation id="0DF646CC69BEEEB19185389F98C0C904">Department of Palaeontology, National Museum of Natural History, National Academy of Sciences of Ukraine, B. Khmelnytskyi st., 15, Kyiv 01054 Ukraine</mods:affiliation>
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<mods:namePart id="4577A3D7F99436E80156AE6A5C84D918">Kovalchuk, О.</mods:namePart>
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<mods:affiliation id="659DA54531ADDB916BBD0AB5181FC668">Department of Palaeontology, National Museum of Natural History, National Academy of Sciences of Ukraine, B. Khmelnytskyi st., 15, Kyiv 01054 Ukraine & Department of Palaeozoology, Faculty of Biological Sciences, University of Wrocław, Sienkiewicza St., 21, Wrocław 50 - 335, Poland</mods:affiliation>
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<mods:typeOfResource id="ABE08DCF814B271DB97A795E76A009C4">text</mods:typeOfResource>
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<mods:title id="9A258362869AEC56A6F90B2F93F6137A">Zoodiversity</mods:title>
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<mods:date id="4C6B4040AEEC416FD65FDB26505CC810">2023</mods:date>
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<mods:url id="8273308AA76AA0F02E202150D6AE24FD">http://dx.doi.org/10.15407/zoo2023.04.311</mods:url>
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<mods:identifier id="1A93C9312C4BC0E603F0CB323F6FDA79" type="DOI">10.15407/zoo2023.04.311</mods:identifier>
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<mods:identifier id="FAF23339FAFA135297A2CF93BC176486" type="ISSN">2707-7268</mods:identifier>
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<treatment id="485B878D917A0F3AF3DEFC06FD19F9B8" ID-DOI="http://doi.org/10.5281/zenodo.10166254" ID-Zenodo-Dep="10166254" LSID="urn:lsid:plazi:treatment:485B878D917A0F3AF3DEFC06FD19F9B8" httpUri="http://treatment.plazi.org/id/485B878D917A0F3AF3DEFC06FD19F9B8" lastPageNumber="315" pageId="4" pageNumber="315">
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<subSubSection id="88E86510917A0F3AF3DEFC06FD86FC4B" box="[166,572,981,1008]" pageId="4" pageNumber="315" type="nomenclature">
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<paragraph id="C04D369B917A0F3AF3DEFC06FD86FC4B" blockId="4.[166,572,981,1008]" box="[166,572,981,1008]" pageId="4" pageNumber="315">
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<heading id="9B0581F7917A0F3AF3DEFC06FD86FC4B" box="[166,572,981,1008]" fontSize="36" level="2" pageId="4" pageNumber="315" reason="3">
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<taxonomicName id="07F24D18917A0F3AF3DEFC06FD86FC4B" ID-CoL="8FPL8" authority="Khosatzky, 1948" authorityName="Khosatzky" authorityYear="1948" box="[166,572,981,1008]" class="Testudines" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Cryptodira" pageId="4" pageNumber="315" phylum="Chordata" rank="species" species="chernovi">
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<emphasis id="F286EA89917A0F3AF3DEFC06FECFFC4B" bold="true" box="[166,373,981,1008]" italics="true" pageId="4" pageNumber="315">Testudo chernovi</emphasis>
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Khosatzky, 1948
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="88E86510917A0F3AF3A3FBD5FD19F9B8" pageId="4" pageNumber="315" type="materials_examined">
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<paragraph id="C04D369B917A0F3AF3A3FBD5FC3AFBA7" blockId="4.[219,896,1030,1052]" box="[219,896,1030,1052]" pageId="4" pageNumber="315">
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Material. Nuchals,
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<collectionCode id="A6E3AE5E917A0F3AF2DFFBD5FDA0FBA7" box="[423,538,1030,1052]" pageId="4" pageNumber="315">NMNHU-P</collectionCode>
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AR 406 (
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<figureCitation id="58C92A1E917A0F3AF10AFBD5FD1FFBA7" box="[626,677,1030,1052]" captionStart="Fig" captionStartId="3.[174,207,1146,1168]" captionTargetBox="[238,1169,209,1101]" captionTargetId="figure-162@3.[210,1196,174,1124]" captionTargetPageId="3" captionText="Fig. 2. Turtle remains from the late Miocene of Grytsiv: A–B — Emys sp., xiphiplastron NMNHU-P AR402/4 (A), marginal scute NMNHU-P AR 402/1 (B); C–E — Melanochelys cf. M. moldavica Chkhikvadze, 1983, costal plate NMNHU-P AR 403 (C), right xiphiplastron NMNHU-P AR 405 (D–E); F, I — Trionychidae indet., shell fragments NMNHU-P AR 401/1 (F), AR 401/2 (I); G–H — Testudo chernovi Khosatzky, 1948, nuchal NMNHU-P AR 406 (G), type specimen after Khosatzky (1948 a: FIg.1), modiFIed (not to scale). Scale bars equal 5 mm in A–C, F–G and I, 10 mm in D and E." figureDoi="http://doi.org/10.5281/zenodo.10831780" httpUri="https://zenodo.org/record/10831780/files/figure.png" pageId="4" pageNumber="315">FIg. 2</figureCitation>
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, G), AR 407 (
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<figureCitation id="58C92A1E917A0F3AF05DFBD5FCE2FBA7" box="[805,856,1030,1052]" captionStart="Fig" captionStartId="5.[173,206,1136,1158]" captionTargetBox="[293,1112,184,1114]" captionTargetId="figure-281@5.[272,1134,174,1136]" captionTargetPageId="5" captionText="Fig. 3. The nuchal plate of Testudo chernovi NMNHU-P AR 407 from Grytsiv with gnaw marks (A); B–C — Machichnus inrosus isp. n., enlarged affected area (B) and a set of individual striae (C); D — combined SEM image of the mandible of Anomalomys grytsivensis Nesin & Kovalchuk, 2021 (NMNHU-P 22/2703) showing the labial surface of the lower incisor with longitudinal ridges." figureDoi="http://doi.org/10.5281/zenodo.10831782" httpUri="https://zenodo.org/record/10831782/files/figure.png" pageId="4" pageNumber="315">FIg. 3</figureCitation>
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, A).
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</paragraph>
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<paragraph id="C04D369B917A0F3AF3A3FB91FB3BFB52" blockId="4.[166,1225,1090,1539]" pageId="4" pageNumber="315">
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Description. The anterior edge of the bones is slightly notched. The cervical scute is narrow, arrow-shaped and wedged on the inner side; its length reaches 11.0 mm. Lateral wings of the nuchal plate are not covered with pleural scutes. The midline length of the specimens AR 406 and AR 407 is equal to 31.6 and
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<quantity id="070A9B7E917A0F3AF181FB7FFCD8FB7C" box="[761,866,1196,1223]" metricMagnitude="-2" metricUnit="m" metricValue="3.61" pageId="4" pageNumber="315" unit="mm" value="36.1">36.1 mm</quantity>
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, the greatest width — 38.0 and
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<quantity id="070A9B7E917A0F3AF3DEFB1CFEAAFB51" box="[166,272,1231,1258]" metricMagnitude="-2" metricUnit="m" metricValue="4.62" pageId="4" pageNumber="315" unit="mm" value="46.2">46.2 mm</quantity>
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, respectively. The length to width ratio is 0.83 (AR 406) and 0.78 (AR 407).
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</paragraph>
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<paragraph id="C04D369B917A0F3AF3A3FB21FEF7FA21" blockId="4.[166,1225,1090,1539]" pageId="4" pageNumber="315">
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Remarks. The specimens are identical in morphology and shape to those of
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<taxonomicName id="07F24D18917A0F3AF716FB20FF63FA94" authorityName="Khosatzky" authorityYear="1948" class="Testudines" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Cryptodira" pageId="4" pageNumber="315" phylum="Chordata" rank="species" species="chernovi">
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<emphasis id="F286EA89917A0F3AF716FB20FF63FA94" italics="true" pageId="4" pageNumber="315">T. chernovi</emphasis>
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</taxonomicName>
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from the Early Pliocene of
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<collectingCountry id="B8E5760B917A0F3AF16DFAC5FDCFFA8B" box="[533,629,1302,1328]" name="Ukraine" pageId="4" pageNumber="315">Ukraine</collectingCountry>
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(
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<bibRefCitation id="A4634B6A917A0F3AF1FBFAC5FCE0FA8B" author="Khosatzky, L. I." box="[643,858,1302,1328]" pageId="4" pageNumber="315" pagination="92 - 96" refId="ref6016" refString="Khosatzky, L. I. 1948 a. New species of the genus Testudo Linne (Testudines, Reptilia) from the Pliocene of Ukraine. Byulleten Komissii po izucheniyu chetvertichnogo perioda, 11, 92 - 96 [In Russian]" type="journal article" year="1948">Khosatzky, 1948 a</bibRefCitation>
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), although they are smaller and possess a shorter cervical scute. The nuchal of the closely related species
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<taxonomicName id="07F24D18917A0F3AF770FAEAFE06FACD" authority="Khosatzky, 1948" authorityName="Khosatzky" authorityYear="1948" class="Testudines" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Cryptodira" pageId="4" pageNumber="315" phylum="Chordata" rank="species" species="kuchurganica">
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<emphasis id="F286EA89917A0F3AF770FAEAFF48FACD" italics="true" pageId="4" pageNumber="315">Testudo kuchurganica</emphasis>
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Khosatzky, 1948
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</taxonomicName>
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has a straight anterior edge and its lateral wings are covered with pleural scutes.
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</paragraph>
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<paragraph id="C04D369B917A0F3AF3A3FA70FD19F9B8" blockId="4.[166,1225,1090,1539]" pageId="4" pageNumber="315">
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A series of characteristic scratches has been observed on one of the specimens considered (fig. 3, B–C). It was interpreted as traces of gnawing or rasping (
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<bibRefCitation id="A4634B6A917A0F3AF0AFFA15FB7BFA5B" author="Mikulas, R. & Kadlecova, E. & Fejfar, O. & Dvorak, Z." box="[983,1217,1478,1504]" pageId="4" pageNumber="315" pagination="113 - 127" refId="ref6992" refString="Mikulas, R., Kadlecova, E., Fejfar, O. & Dvorak, Z. 2006. Three New Ichnogenera of Biting and Gnawing Traces on Reptilian and Mammalian Bones: A Case Study from the Miocene of the Czech Republic. Ichnos: An International Journal for Plant and Animal Traces, 13 (3), 113 - 127, http: // dx. doi. org / 10.1080 / 10420940600850729" type="journal article" year="2006">Mikuláš et al., 2006</bibRefCitation>
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) and described as a new ichnospecies below.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |