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<document id="38D75FB2105CFCEA5AC72E8A9B6E0D2C" ID-DOI="10.5281/zenodo.197288" ID-GBIF-Dataset="256e66c3-8f88-4aa5-89a8-a29aab35101f" ID-ISSN="1175-5326" ID-Zenodo-Dep="197288" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460264471404" checkinUser="plazi" docAuthor="Mwabvu, Tarombera, Hamer, Michelle, Slotow, Rob &amp; Barraclough, David" docDate="2010" docId="080E878B1C71770EFF144EE32BEA7E0F" docLanguage="en" docName="zt02567p049.pdf" docOrigin="Zootaxa 2567" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Archispirostreptus lugubris Brolemann 1901" docType="treatment" docVersion="8" lastPageNumber="18" masterDocId="F437FFF31C60771FFF834F7E28557937" masterDocTitle="A revision of the taxonomy and distribution of Archispirostreptus Silvestri 1895 (Diplopoda, Spirostreptida, Spirostreptidae), and description of a new spirostreptid genus with three new species" masterLastPageNumber="49" masterPageNumber="1" pageNumber="18" updateTime="1698245499894" updateUser="plazi">
<mods:mods id="2A1F1A8EEA41AF9E266504990CE7D7DD" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="0CCE483907C86C80557159D19018DDDF">A revision of the taxonomy and distribution of Archispirostreptus Silvestri 1895 (Diplopoda, Spirostreptida, Spirostreptidae), and description of a new spirostreptid genus with three new species</mods:title>
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<mods:namePart id="399F85A43F1307B6384B12A04E6C3BE6">Mwabvu, Tarombera</mods:namePart>
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<mods:namePart id="170B8482CCB89A2589BAF7244F7FA589">Hamer, Michelle</mods:namePart>
</mods:name>
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<mods:namePart id="15DDF458D3FA69D09E6270F3232631AF">Slotow, Rob</mods:namePart>
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<mods:namePart id="F94C99DB0469ECCBFF0AF0868D5F0178">Barraclough, David</mods:namePart>
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<mods:title id="8CD598C7B45544646216C2249D5D0B82">Zootaxa</mods:title>
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<mods:date id="A5F4438BFBCBB73E959D8322798E0F9E">2010</mods:date>
<mods:detail id="08374948E093B1CA308635C22EE2CBF2" type="volume">
<mods:number id="4F180EAE782DF141E9FC43D8118AB02A">2567</mods:number>
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<mods:identifier id="47B2763948D25CDAC383B3698595B548" type="DOI">10.5281/zenodo.197288</mods:identifier>
<mods:identifier id="5291B53ACD1B487250C002F0F3019A8F" type="GBIF-Dataset">256e66c3-8f88-4aa5-89a8-a29aab35101f</mods:identifier>
<mods:identifier id="38BFA8C04CD67887B5BE69922B7B5150" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="8E008ED2BBA0F1E36885132FFF8E540C" type="Zenodo-Dep">197288</mods:identifier>
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<treatment id="080E878B1C71770EFF144EE32BEA7E0F" ID-DOI="http://doi.org/10.5281/zenodo.6207855" ID-GBIF-Taxon="119394069" ID-Zenodo-Dep="6207855" LSID="urn:lsid:plazi:treatment:080E878B1C71770EFF144EE32BEA7E0F" httpUri="http://treatment.plazi.org/id/080E878B1C71770EFF144EE32BEA7E0F" lastPageNumber="18" pageId="17" pageNumber="18">
<subSubSection id="C8BD65161C71770EFF144EE3288E78ED" pageId="17" pageNumber="18" type="nomenclature">
<paragraph id="8018369D1C71770EFF144EE32A9D7880" blockId="17.[151,712,413,474]" box="[151,712,413,439]" pageId="17" pageNumber="18">
<heading id="DB5081F11C71770EFF144EE32A9D7880" bold="true" box="[151,712,413,439]" fontSize="11" level="1" pageId="17" pageNumber="18" reason="1">
<taxonomicName id="47A74D1E1C71770EFF144EE32A9D7880" ID-CoL="G9QB" authority="Brolemann 1901" authorityName="Brolemann" authorityYear="1901" box="[151,712,413,439]" class="Diplopoda" family="Spirostreptidae" genus="Archispirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="lugubris">
<emphasis id="B2D3EA8F1C71770EFF144EE32A9D7880" bold="true" box="[151,712,413,439]" pageId="17" pageNumber="18">
<emphasis id="B2D3EA8F1C71770EFF144EE329B77880" bold="true" box="[151,482,413,439]" italics="true" pageId="17" pageNumber="18">Archispirostreptus lugubris</emphasis>
(
<bibRefCitation id="E4364B6C1C71770EFE724EE32A957880" author="Brolemann" box="[497,704,413,439]" pageId="17" pageNumber="18" refString="Brolemann, H. W. (1901) Materiali per la conoscenza della fauna Eritrea, raccolti dal Dr. P. Magretti. Bollettino della Societa Entomologica Italiana, 33, 26 - 35." type="journal article" year="1901">Brölemann 1901</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8018369D1C71770EFF144EBE288E78ED" blockId="17.[151,712,413,474]" box="[151,219,448,474]" pageId="17" pageNumber="18">
<figureCitation id="189C2A181C71770EFF144EBE288E78ED" box="[151,219,448,474]" captionStart="FIGURE 6 a, b" captionStartId="18.[151,255,1979,2003]" captionTargetBox="[319,1251,208,1929]" captionTargetId="figure@18.[310,1261,194,1939]" captionTargetPageId="18" captionText="FIGURE 6 a, b. Archispirostreptus lugubris (MNHN, 1 B 106). a, oral view of gonopods; b, aboral view of gonopods." httpUri="https://zenodo.org/record/197299/files/figure.png" pageId="17" pageNumber="18">Fig. 6</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C8BD65161C71770EFF144D742BF17B56" pageId="17" pageNumber="18" type="reference_group">
<paragraph id="8018369D1C71770EFF144D742A1D7B76" blockId="17.[151,932,522,609]" pageId="17" pageNumber="18">
<taxonomicName id="47A74D1E1C71770EFF144D742A697B16" ID-CoL="6ZCHM" authority="Brolemann 1901" authorityName="Brolemann" authorityYear="1901" box="[151,572,522,545]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="lugubris">
<emphasis id="B2D3EA8F1C71770EFF144D7429D67B16" box="[151,387,522,545]" italics="true" pageId="17" pageNumber="18">Spirostreptus lugubris</emphasis>
<bibRefCitation id="E4364B6C1C71770EFE0A4D742A697B16" author="Brolemann" box="[393,572,522,545]" pageId="17" pageNumber="18" refString="Brolemann, H. W. (1901) Materiali per la conoscenza della fauna Eritrea, raccolti dal Dr. P. Magretti. Bollettino della Societa Entomologica Italiana, 33, 26 - 35." type="journal article" year="1901">Brölemann 1901</bibRefCitation>
</taxonomicName>
, p. 31–34, figs 8–11.
<taxonomicName id="47A74D1E1C71770EFF144D542A1D7B76" ID-CoL="6L3K5" authority="Attems 1914" authorityName="Attems" authorityYear="1914" box="[151,584,554,577]" class="Diplopoda" family="Spirostreptidae" genus="Graphidostreptus" kingdom="Animalia" order="Spirostreptida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="lugubris">
<emphasis id="B2D3EA8F1C71770EFF144D5429E77B76" box="[151,434,554,577]" italics="true" pageId="17" pageNumber="18">Graphidostreptus lugubris</emphasis>
:
<bibRefCitation id="E4364B6C1C71770EFE3C4D542A1D7B76" author="Attems" box="[447,584,554,577]" pageId="17" pageNumber="18" refString="Attems, C. (1914) Afrikanische Spirostreptiden nebst Ueberblick uber die Spirostreptiden orbis terrarum. Zoologica, 25, 1 - 233." type="journal article" year="1914">Attems 1914</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8018369D1C71770EFF144D342BF17B56" blockId="17.[151,932,522,609]" box="[151,932,586,609]" pageId="17" pageNumber="18">
<taxonomicName id="47A74D1E1C71770EFF144D342A307B56" ID-CoL="G9QB" authority="Schubart 1951" authorityName="Schubart" authorityYear="1951" box="[151,613,586,609]" class="Diplopoda" family="Spirostreptidae" genus="Archispirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="lugubris">
<emphasis id="B2D3EA8F1C71770EFF144D3429EF7B56" box="[151,442,586,609]" italics="true" pageId="17" pageNumber="18">Archispirostreptus lugubris</emphasis>
:
<bibRefCitation id="E4364B6C1C71770EFE4B4D342A307B56" author="Schubart" box="[456,613,586,609]" pageId="17" pageNumber="18" refString="Schubart, O. (1951) Uber einige Diplopoden vom Bergmassiv Air in der Sud-Sahara, gesammelt von L. Chopard und A. Villiers im Jahre 1947. Bulletin de I'Institut Francaise d'Afrique Noire, 13, 116 - 125." type="journal article" year="1951">Schubart 1951</bibRefCitation>
</taxonomicName>
;
<bibRefCitation id="E4364B6C1C71770EFDF34D342B5A7B56" author="Hoffman" box="[624,783,586,609]" pageId="17" pageNumber="18" refString="Hoffman, R. L. (1965) A new species in the diplopod genus Archispirostreptus from south western Arabia. Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg, 3 (51), 17 - 23." type="journal article" year="1965">Hoffman 1965</bibRefCitation>
;
<bibRefCitation id="E4364B6C1C71770EFC994D342BF17B56" author="Krabbe" box="[794,932,586,609]" pageId="17" pageNumber="18" refString="Krabbe, E. (1982) Systematik der Spirostreptidae (Diplopoda, Spirostreptomorpha). Abhandlungen des Naturwissenschaftlichen Vereins in Hamburg. Verlag Paul Parey. Hamburg, 476 pp." type="book" year="1982">Krabbe 1982</bibRefCitation>
</paragraph>
</subSubSection>
<subSubSection id="C8BD65161C71770EFF144DF12CCC7A28" pageId="17" pageNumber="18" type="materials_examined">
<paragraph id="8018369D1C71770EFF144DF12D3E7B9D" blockId="17.[151,1437,655,1848]" box="[151,1387,655,682]" pageId="17" pageNumber="18">
<emphasis id="B2D3EA8F1C71770EFF144DF129187B9E" bold="true" box="[151,333,655,681]" pageId="17" pageNumber="18">
<typeStatus id="5F1C883F1C71770EFF144DF128877B9E" box="[151,210,655,681]" pageId="17" pageNumber="18">Type</typeStatus>
material:
</emphasis>
<typeStatus id="5F1C883F1C71770EFED54DEE299D7B9D" box="[342,456,656,682]" pageId="17" pageNumber="18" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F8B0760D1C71770EFE504DEE2A0A7B9D" box="[467,607,656,682]" name="Ethiopia" pageId="17" pageNumber="18">ETHIOPIA</collectingCountry>
: 1 3, Sabarguma Allata,
<date id="F419105D1C71770EFC064DEE2C537B9D" box="[901,1030,656,682]" pageId="17" pageNumber="18" value="1900-03-15">15.iii.1900</date>
, P. Magretti (
<collectionCode id="E6B6AE581C71770EFB224DEE2CA87B9D" LSID="urn:lsid:biocol.org:col:34988" box="[1185,1277,656,682]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="17" pageNumber="18">MNHN</collectionCode>
1B 106).
</paragraph>
<paragraph id="8018369D1C71770EFF464DCB2CCC7A28" blockId="17.[151,1437,655,1848]" pageId="17" pageNumber="18">
<emphasis id="B2D3EA8F1C71770EFF464DCB2A1D7BF8" bold="true" box="[197,584,693,719]" pageId="17" pageNumber="18">Additional material examined:</emphasis>
<collectingCountry id="F8B0760D1C71770EFDD34DCB2AFB7BF8" box="[592,686,693,719]" name="Niger" pageId="17" pageNumber="18">NIGER</collectingCountry>
: 1 3, Monts Baguezane, Irrabellaben,
<date id="F419105D1C71770EFBFB4DCB2D5D7BF8" box="[1144,1288,693,719]" pageId="17" pageNumber="18" value="1947-08-26">26.viii.1947</date>
, L. Chopard &amp; A. Villiers (
<collectionCode id="E6B6AE581C71770EFEC34DA329C97BC0" LSID="urn:lsid:biocol.org:col:34988" box="[320,412,733,759]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="17" pageNumber="18">MNHN</collectionCode>
1B 108) (gonopods missing).
<collectingCountry id="F8B0760D1C71770EFC874DA32BDA7BC0" box="[772,911,733,759]" name="Senegal" pageId="17" pageNumber="18">SENEGAL</collectingCountry>
: 2 3, Bambey [14
<geoCoordinate id="E593505A1C71770EFBE44DA42CFB7BC0" box="[1127,1198,730,759]" direction="north" orientation="latitude" pageId="17" pageNumber="18" precision="925" value="0.71666664">0 43'N</geoCoordinate>
, 16
<geoCoordinate id="E593505A1C71770EFB5B4DA42D727BC0" box="[1240,1319,730,759]" direction="west" orientation="longitude" pageId="17" pageNumber="18" precision="925" value="-0.23333333">0 14'W</geoCoordinate>
], (
<collectionCode id="E6B6AE581C71770EFAC04DA32DC97BC0" box="[1347,1436,733,759]" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="17" pageNumber="18">MRAC</collectionCode>
20504); 1 3, 1 Ƥ, Gouloubou [13
<geoCoordinate id="E593505A1C71770EFDA24C7D2A327A28" box="[545,615,771,799]" direction="north" orientation="latitude" pageId="17" pageNumber="18" precision="925" value="0.45">0 27'N</geoCoordinate>
, 13
<geoCoordinate id="E593505A1C71770EFD134C7D2A8A7A28" box="[656,735,771,799]" direction="west" orientation="longitude" pageId="17" pageNumber="18" precision="925" value="-0.71666664">0 43'W</geoCoordinate>
],
<date id="F419105D1C71770EFD704C7B2B007A28" box="[755,853,773,799]" pageId="17" pageNumber="18" value="1979-01-02">2.i.1979</date>
, D. Thys (
<collectionCode id="E6B6AE581C71770EFC4C4C7B2C7E7A28" LSID="urn:lsid:biocol.org:col:34988" box="[975,1067,773,799]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="17" pageNumber="18">MNHN</collectionCode>
1B 224).
</paragraph>
</subSubSection>
<subSubSection id="C8BD65161C71770EFF464C542B847A5D" pageId="17" pageNumber="18" type="diagnosis">
<paragraph id="8018369D1C71770EFF464C542B847A5D" blockId="17.[151,1437,655,1848]" pageId="17" pageNumber="18">
<emphasis id="B2D3EA8F1C71770EFF464C5429127A73" bold="true" box="[197,327,810,836]" pageId="17" pageNumber="18">Diagnosis:</emphasis>
Sternite apex flat; oral fold of metaplicae abruptly narrow at level of telopodite knee (
<figureCitation id="189C2A181C71770EFABB4C552DDB7A72" box="[1336,1422,811,837]" captionStart="FIGURE 6 a, b" captionStartId="18.[151,255,1979,2003]" captionTargetBox="[319,1251,208,1929]" captionTargetId="figure@18.[310,1261,194,1939]" captionTargetPageId="18" captionText="FIGURE 6 a, b. Archispirostreptus lugubris (MNHN, 1 B 106). a, oral view of gonopods; b, aboral view of gonopods." httpUri="https://zenodo.org/record/197299/files/figure.png" pageId="17" pageNumber="18">Fig. 6a</figureCitation>
). Lateral metaplical process slanting gently towards paracoxite (
<figureCitation id="189C2A181C71770EFCF74C2E2B9D7A5D" box="[884,968,848,874]" captionStart="FIGURE 6 a, b" captionStartId="18.[151,255,1979,2003]" captionTargetBox="[319,1251,208,1929]" captionTargetId="figure@18.[310,1261,194,1939]" captionTargetPageId="18" captionText="FIGURE 6 a, b. Archispirostreptus lugubris (MNHN, 1 B 106). a, oral view of gonopods; b, aboral view of gonopods." httpUri="https://zenodo.org/record/197299/files/figure.png" pageId="17" pageNumber="18">Fig. 6a</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C8BD65161C71770EFF464C0B2B547C64" pageId="17" pageNumber="18" type="description">
<paragraph id="8018369D1C71770EFF464C0B2C3C7AB8" blockId="17.[151,1437,655,1848]" box="[197,1129,885,911]" pageId="17" pageNumber="18">
<emphasis id="B2D3EA8F1C71770EFF464C0B290A7AB8" bold="true" box="[197,351,885,911]" pageId="17" pageNumber="18">Description:</emphasis>
<emphasis id="B2D3EA8F1C71770EFEE44C0829C37AB8" box="[359,406,886,911]" italics="true" pageId="17" pageNumber="18">Size</emphasis>
: Body length
<quantity id="475F9B781C71770EFDBF4C0B2A8E7AB8" box="[572,731,885,911]" metricMagnitude="-1" metricUnit="m" metricValue="1.285" metricValueMax="1.55" metricValueMin="1.02" pageId="17" pageNumber="18" unit="mm" value="128.5" valueMax="155.0" valueMin="102.0">102–155 mm</quantity>
; maximum body width
<quantity id="475F9B781C71770EFC724C0B2C317AB8" box="[1009,1124,885,911]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.3" metricValueMin="0.8" pageId="17" pageNumber="18" unit="mm" value="10.5" valueMax="13.0" valueMin="8.0">8–13 mm</quantity>
.
</paragraph>
<paragraph id="8018369D1C71770EFF464CE52A737A82" blockId="17.[151,1437,655,1848]" box="[197,550,923,949]" pageId="17" pageNumber="18">
<emphasis id="B2D3EA8F1C71770EFF464CE529907A83" box="[197,453,923,948]" italics="true" pageId="17" pageNumber="18">Number of body rings</emphasis>
: 55–64.
</paragraph>
<paragraph id="8018369D1C71770EFF464CBF2BC17AED" blockId="17.[151,1437,655,1848]" box="[197,916,960,986]" pageId="17" pageNumber="18">
<emphasis id="B2D3EA8F1C71770EFF464CBF294D7AED" box="[197,280,961,986]" italics="true" pageId="17" pageNumber="18">Colour</emphasis>
: Body black or dark brown; legs and antennae brown.
</paragraph>
<paragraph id="8018369D1C71770EFF464C982C887AC8" blockId="17.[151,1437,655,1848]" box="[197,1245,997,1023]" pageId="17" pageNumber="18">
<emphasis id="B2D3EA8F1C71770EFF464C98294E7AC8" box="[197,283,998,1023]" italics="true" pageId="17" pageNumber="18">Collum</emphasis>
: Anterior produced into lobe, with 2–3 complete striae and 1–2 incomplete striae.
</paragraph>
<paragraph id="8018369D1C71770EFF464B702D767D1F" blockId="17.[151,1437,655,1848]" box="[197,1315,1035,1064]" pageId="17" pageNumber="18">
<emphasis id="B2D3EA8F1C71770EFF464B702AEB7D10" box="[197,702,1035,1063]" italics="true" pageId="17" pageNumber="18">Pre-femoral process of 1st pair of male legs</emphasis>
: Basally round with short tapering apical extension.
</paragraph>
<paragraph id="8018369D1C71770EFF464B4D2CDB7D7A" blockId="17.[151,1437,655,1848]" box="[197,1166,1075,1101]" pageId="17" pageNumber="18">
<emphasis id="B2D3EA8F1C71770EFF464B4D29677D7B" box="[197,306,1075,1100]" italics="true" pageId="17" pageNumber="18">Gonopod</emphasis>
: (
<figureCitation id="189C2A181C71770EFECA4B4D299B7D7A" box="[329,462,1075,1101]" captionStart="FIGURE 6 a, b" captionStartId="18.[151,255,1979,2003]" captionTargetBox="[319,1251,208,1929]" captionTargetId="figure@18.[310,1261,194,1939]" captionTargetPageId="18" captionText="FIGURE 6 a, b. Archispirostreptus lugubris (MNHN, 1 B 106). a, oral view of gonopods; b, aboral view of gonopods." httpUri="https://zenodo.org/record/197299/files/figure.png" pageId="17" pageNumber="18">Figs 6a, 6b</figureCitation>
) Sternum broad proximally, apex flat; paracoxite triangular.
</paragraph>
<paragraph id="8018369D1C71770EFF464B272A477D44" blockId="17.[151,1437,655,1848]" box="[197,530,1113,1139]" pageId="17" pageNumber="18">Width of proplicae constant.</paragraph>
<paragraph id="8018369D1C71770EFF464B002AFA7D8A" blockId="17.[151,1437,655,1848]" pageId="17" pageNumber="18">Metaplicae wide proximally; opposite metaplicae wider apart distal to sternite apex; oral fold of metaplicae narrow at level of telopodite knee.</paragraph>
<paragraph id="8018369D1C71770EFF464BB72BE77C3F" blockId="17.[151,1437,655,1848]" pageId="17" pageNumber="18">
Lateral metaplical process slanting towards paracoxite and broadly joined to metaplica. Lateral metaplical process at obtuse angle with apical metaplical process (
<figureCitation id="189C2A181C71770EFC9D4B902BF07C3F" box="[798,933,1262,1288]" captionStart="FIGURE 6 a, b" captionStartId="18.[151,255,1979,2003]" captionTargetBox="[319,1251,208,1929]" captionTargetId="figure@18.[310,1261,194,1939]" captionTargetPageId="18" captionText="FIGURE 6 a, b. Archispirostreptus lugubris (MNHN, 1 B 106). a, oral view of gonopods; b, aboral view of gonopods." httpUri="https://zenodo.org/record/197299/files/figure.png" pageId="17" pageNumber="18">Figs 6a, 6b</figureCitation>
).
</paragraph>
<paragraph id="8018369D1C71770EFF464A6D2C497C1A" blockId="17.[151,1437,655,1848]" box="[197,1052,1299,1325]" pageId="17" pageNumber="18">Distally apical metaplical process convex medially and rounded apically.</paragraph>
<paragraph id="8018369D1C71770EFF464A472B547C64" blockId="17.[151,1437,655,1848]" box="[197,769,1337,1363]" pageId="17" pageNumber="18">Origin of antetorsal processes at telopodite knee.</paragraph>
</subSubSection>
<subSubSection id="C8BD65161C71770EFF464A23293F7CAA" pageId="17" pageNumber="18" type="distribution">
<paragraph id="8018369D1C71770EFF464A23293F7CAA" blockId="17.[151,1437,655,1848]" pageId="17" pageNumber="18">
<emphasis id="B2D3EA8F1C71770EFF464A2329397C40" bold="true" box="[197,364,1373,1399]" pageId="17" pageNumber="18">Distribution:</emphasis>
Known from
<collectingCountry id="F8B0760D1C71770EFD974A202BDC7C4F" box="[532,905,1374,1400]" name="Democratic Republic of the Congo" pageId="17" pageNumber="18">Democratic Republic of Congo</collectingCountry>
,
<collectingCountry id="F8B0760D1C71770EFC154A202BA27C4F" box="[918,1015,1374,1400]" name="Somalia" pageId="17" pageNumber="18">Somalia</collectingCountry>
(
<bibRefCitation id="E4364B6C1C71770EFB854A202CF37C4F" author="Krabbe" box="[1030,1190,1374,1400]" pageId="17" pageNumber="18" refString="Krabbe, E. (1982) Systematik der Spirostreptidae (Diplopoda, Spirostreptomorpha). Abhandlungen des Naturwissenschaftlichen Vereins in Hamburg. Verlag Paul Parey. Hamburg, 476 pp." type="book" year="1982">Krabbe 1982</bibRefCitation>
),
<collectingCountry id="F8B0760D1C71770EFB384A202D557C4F" box="[1211,1280,1374,1400]" name="Niger" pageId="17" pageNumber="18">Niger</collectingCountry>
,
<collectingCountry id="F8B0760D1C71770EFA8E4A202D3F7C4F" box="[1293,1386,1374,1400]" name="Senegal" pageId="17" pageNumber="18">Senegal</collectingCountry>
and
<collectingCountry id="F8B0760D1C71770EFF144AFD28AD7CAA" box="[151,248,1411,1437]" name="Ethiopia" pageId="17" pageNumber="18">Ethiopia</collectingCountry>
(
<figureCitation id="189C2A181C71770EFE8B4AFD29087CAA" box="[264,349,1411,1437]" captionStart="FIGURE 15" captionStartId="44.[151,255,1699,1723]" captionTargetBox="[293,1333,514,1636]" captionTargetId="figure@44.[184,1403,453,1675]" captionTargetPageId="44" captionText="FIGURE 15. Distribution of Archispirostreptus and Cacuminostreptus species." httpUri="https://zenodo.org/record/197317/files/figure.png" pageId="17" pageNumber="18">Fig. 15</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C8BD65161C71770EFF464AD62BEA7E0F" pageId="17" pageNumber="18" type="discussion">
<paragraph id="8018369D1C71770EFF464AD62DC17F6F" blockId="17.[151,1437,655,1848]" pageId="17" pageNumber="18">
<emphasis id="B2D3EA8F1C71770EFF464AD629157CF5" bold="true" box="[197,320,1448,1474]" pageId="17" pageNumber="18">Remarks:</emphasis>
Abrupt narrowing of the oral fold of the metaplicae has only been observed in the gonopods of
<taxonomicName id="47A74D1E1C71770EFF144AB028AB7CD0" box="[151,254,1486,1511]" class="Diplopoda" family="Spirostreptidae" genus="Archispirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="lugubris">
<emphasis id="B2D3EA8F1C71770EFF144AB028AB7CD0" box="[151,254,1486,1511]" italics="true" pageId="17" pageNumber="18">lugubris</emphasis>
</taxonomicName>
. The telopodite is similar to those of
<taxonomicName id="47A74D1E1C71770EFD564AB02B0C7CD0" box="[725,857,1486,1511]" class="Diplopoda" family="Spirostreptidae" genus="Archispirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="dodsoni">
<emphasis id="B2D3EA8F1C71770EFD564AB02B0C7CD0" box="[725,857,1486,1511]" italics="true" pageId="17" pageNumber="18">A. dodsoni</emphasis>
</taxonomicName>
,
<taxonomicName id="47A74D1E1C71770EFCE94AB02C767CD0" box="[874,1059,1486,1511]" class="Diplopoda" family="Spirostreptidae" genus="Archispirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="tumuliporus">
<emphasis id="B2D3EA8F1C71770EFCE94AB02C767CD0" box="[874,1059,1486,1511]" italics="true" pageId="17" pageNumber="18">A. tumuliporus</emphasis>
</taxonomicName>
,
<taxonomicName id="47A74D1E1C71770EFBB04AB02CCC7CD0" box="[1075,1177,1486,1511]" class="Diplopoda" family="Spirostreptidae" genus="Archispirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="gigas">
<emphasis id="B2D3EA8F1C71770EFBB04AB02CCC7CD0" box="[1075,1177,1486,1511]" italics="true" pageId="17" pageNumber="18">A. gigas</emphasis>
</taxonomicName>
,
<taxonomicName id="47A74D1E1C71770EFB294AB02D117CD0" box="[1194,1348,1486,1511]" class="Diplopoda" family="Spirostreptidae" genus="Archispirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="divergens">
<emphasis id="B2D3EA8F1C71770EFB294AB02D117CD0" box="[1194,1348,1486,1511]" italics="true" pageId="17" pageNumber="18">A. divergens</emphasis>
</taxonomicName>
and
<taxonomicName id="47A74D1E1C71770EFA014AB028BC7F3B" class="Diplopoda" family="Spirostreptidae" genus="Archispirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="smithii">
<emphasis id="B2D3EA8F1C71770EFA014AB028BC7F3B" italics="true" pageId="17" pageNumber="18">A. smithii</emphasis>
</taxonomicName>
; and the lateral metaplical process resembles that of
<taxonomicName id="47A74D1E1C71770EFCE94A8D2BBB7F3B" box="[874,1006,1523,1548]" class="Diplopoda" family="Spirostreptidae" genus="Archispirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="beccarii">
<emphasis id="B2D3EA8F1C71770EFCE94A8D2BBB7F3B" box="[874,1006,1523,1548]" italics="true" pageId="17" pageNumber="18">A. beccarii</emphasis>
</taxonomicName>
,
<taxonomicName id="47A74D1E1C71770EFC7D4A8D2CE57F3B" box="[1022,1200,1523,1548]" class="Diplopoda" family="Spirostreptidae" genus="Archispirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="tumuliporus">
<emphasis id="B2D3EA8F1C71770EFC7D4A8D2CE57F3B" box="[1022,1200,1523,1548]" italics="true" pageId="17" pageNumber="18">A. tumuliporus</emphasis>
</taxonomicName>
,
<taxonomicName id="47A74D1E1C71770EFB424A8D2D127F3B" box="[1217,1351,1523,1548]" class="Diplopoda" family="Spirostreptidae" genus="Archispirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="syriacus">
<emphasis id="B2D3EA8F1C71770EFB424A8D2D127F3B" box="[1217,1351,1523,1548]" italics="true" pageId="17" pageNumber="18">A. syriacus</emphasis>
</taxonomicName>
and
<taxonomicName id="47A74D1E1C71770EFA014A8D28A07F05" class="Diplopoda" family="Spirostreptidae" genus="Archispirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="dodsoni">
<emphasis id="B2D3EA8F1C71770EFA014A8D28A07F05" italics="true" pageId="17" pageNumber="18">A. dodsoni</emphasis>
</taxonomicName>
more than
<taxonomicName id="47A74D1E1C71770EFEFE4967298A7F05" box="[381,479,1561,1586]" class="Diplopoda" family="Spirostreptidae" genus="Archispirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="gigas">
<emphasis id="B2D3EA8F1C71770EFEFE4967298A7F05" box="[381,479,1561,1586]" italics="true" pageId="17" pageNumber="18">A. gigas</emphasis>
</taxonomicName>
,
<taxonomicName id="47A74D1E1C71770EFE6C49672AD67F05" box="[495,643,1561,1586]" class="Diplopoda" family="Spirostreptidae" genus="Archispirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="divergens">
<emphasis id="B2D3EA8F1C71770EFE6C49672AD67F05" box="[495,643,1561,1586]" italics="true" pageId="17" pageNumber="18">A. divergens</emphasis>
</taxonomicName>
or
<taxonomicName id="47A74D1E1C71770EFD2E49672B6C7F05" box="[685,825,1561,1586]" class="Diplopoda" family="Spirostreptidae" genus="Archispirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="phillipsii">
<emphasis id="B2D3EA8F1C71770EFD2E49672B6C7F05" box="[685,825,1561,1586]" italics="true" pageId="17" pageNumber="18">A. phillipsii</emphasis>
</taxonomicName>
. We have also established that some material of
<taxonomicName id="47A74D1E1C71770EFA00496728AC7F60" class="Diplopoda" family="Spirostreptidae" genus="Archispirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="lugubris">
<emphasis id="B2D3EA8F1C71770EFA00496728AC7F60" italics="true" pageId="17" pageNumber="18">A. lugubris</emphasis>
</taxonomicName>
(MNHN 1B 224) from
<collectingCountry id="F8B0760D1C71770EFD9349402A257F6F" box="[528,624,1598,1624]" name="Senegal" pageId="17" pageNumber="18">Senegal</collectingCountry>
, housed in MNHN had been incorrectly identified as
<taxonomicName id="47A74D1E1C71770EFB6349402DD87F60" box="[1248,1421,1598,1624]" class="Diplopoda" family="Spirostreptidae" genus="Archispirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="tumuliporus">
<emphasis id="B2D3EA8F1C71770EFB6349402CA77F60" box="[1248,1266,1598,1623]" italics="true" pageId="17" pageNumber="18">A</emphasis>
.
<emphasis id="B2D3EA8F1C71770EFA8349402DD87F60" box="[1280,1421,1598,1623]" italics="true" pageId="17" pageNumber="18">tumuliporus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8018369D1C71770EFF46491D2BEA7E0F" blockId="17.[151,1437,655,1848]" pageId="17" pageNumber="18">
Two subspecies—
<taxonomicName id="47A74D1E1C71770EFE3A491D2C677F4A" authority="Attems 1914" authorityName="Attems" authorityYear="1914" box="[441,1074,1635,1661]" class="Diplopoda" family="Spirostreptidae" genus="Graphidostreptus" kingdom="Animalia" order="Spirostreptida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="subSpecies" species="lugubris" subSpecies="maior">
<emphasis id="B2D3EA8F1C71770EFE3A491D2B2E7F4B" box="[441,891,1635,1660]" italics="true" pageId="17" pageNumber="18">Graphidostreptus lugubris maior</emphasis>
<bibRefCitation id="E4364B6C1C71770EFC04491D2C677F4A" author="Attems" box="[903,1074,1635,1661]" pageId="17" pageNumber="18" refString="Attems, C. (1914) Afrikanische Spirostreptiden nebst Ueberblick uber die Spirostreptiden orbis terrarum. Zoologica, 25, 1 - 233." type="journal article" year="1914">Attems 1914</bibRefCitation>
</taxonomicName>
, p. 157, figs 155–156 and
<taxonomicName id="47A74D1E1C71770EFF1449F72ABA7F94" authority="Schubart 1951" authorityName="Schubart" authorityYear="1951" box="[151,751,1673,1699]" class="Diplopoda" family="Spirostreptidae" genus="Graphidostreptus" kingdom="Animalia" order="Spirostreptida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="subSpecies" species="lugubris" subSpecies="villiersi">
<emphasis id="B2D3EA8F1C71770EFF1449F72A627F95" box="[151,567,1673,1698]" italics="true" pageId="17" pageNumber="18">Graphidostreptus lugubris villiersi</emphasis>
<bibRefCitation id="E4364B6C1C71770EFDBC49F72ABA7F94" author="Schubart" box="[575,751,1673,1699]" pageId="17" pageNumber="18" refString="Schubart, O. (1951) Uber einige Diplopoden vom Bergmassiv Air in der Sud-Sahara, gesammelt von L. Chopard und A. Villiers im Jahre 1947. Bulletin de I'Institut Francaise d'Afrique Noire, 13, 116 - 125." type="journal article" year="1951">Schubart 1951</bibRefCitation>
</taxonomicName>
, p. 119–124, figs 1–5—from southern Sahara desert have been described.
<taxonomicName id="47A74D1E1C71770EFEE149D02BA17FFF" authority="Schubart 1951" authorityName="Schubart" authorityYear="1951" box="[354,1012,1710,1736]" class="Diplopoda" family="Spirostreptidae" genus="Archispirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="subSpecies" species="lugubris" subSpecies="villiersi">
<emphasis id="B2D3EA8F1C71770EFEE149D02B757FF0" box="[354,800,1710,1735]" italics="true" pageId="17" pageNumber="18">Archispirostreptus lugubris villiersi</emphasis>
(
<bibRefCitation id="E4364B6C1C71770EFCB149D02BBF7FFF" author="Schubart" box="[818,1002,1710,1736]" pageId="17" pageNumber="18" refString="Schubart, O. (1951) Uber einige Diplopoden vom Bergmassiv Air in der Sud-Sahara, gesammelt von L. Chopard und A. Villiers im Jahre 1947. Bulletin de I'Institut Francaise d'Afrique Noire, 13, 116 - 125." type="journal article" year="1951">Schubart 1951</bibRefCitation>
)
</taxonomicName>
is a smaller variation of the
<typeStatus id="5F1C883F1C71770EFADC49D02DCC7FFF" box="[1375,1433,1710,1736]" pageId="17" pageNumber="18">type</typeStatus>
. Furthermore, the apical metaplical process of the gonopod is shorter and the lateral metaplical process is longer, with a pointed apex, than in
<taxonomicName id="47A74D1E1C71770EFDB849872B5F7E25" box="[571,778,1785,1810]" class="Diplopoda" family="Spirostreptidae" genus="Archispirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="subSpecies" species="lugubris" subSpecies="maior">
<emphasis id="B2D3EA8F1C71770EFDB849872B5F7E25" box="[571,778,1785,1810]" italics="true" pageId="17" pageNumber="18">A. lugubris maior</emphasis>
</taxonomicName>
and the other subspecies
<taxonomicName id="47A74D1E1C71770EFBB549872D777E25" box="[1078,1314,1785,1810]" class="Diplopoda" family="Spirostreptidae" genus="Archispirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="subSpecies" species="lugubris" subSpecies="lugubris">
<emphasis id="B2D3EA8F1C71770EFBB549872D777E25" box="[1078,1314,1785,1810]" italics="true" pageId="17" pageNumber="18">A. lugubris lugubris</emphasis>
</taxonomicName>
. Based on gonopod morphology these subspecies represent recognizable forms.
</paragraph>
</subSubSection>
</treatment>
</document>
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