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<document id="E428C275A514A992C3710841ED7AA0CA" ID-DOI="10.11646/zootaxa.4032.4.1" ID-GBIF-Dataset="1459e41f-b82d-4c88-be4d-ee86e329f9c7" ID-ISSN="1175-5326" ID-Zenodo-Dep="243485" ID-ZooBank="AFEF8567-B7C6-4F54-AC51-4C990389BF1D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461294283733" checkinUser="plazi" docAuthor="Kustov, Semen &amp; Zherebilo, Danil" docDate="2015" docId="03F487D4FF841A7CFF04FD8CB954FA0F" docLanguage="en" docName="zt04032p369.pdf" docOrigin="Zootaxa 4032 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Wiedemannia (Chamaedipsia) beckeri Mik" docType="treatment" docVersion="9" lastPageNumber="354" masterDocId="FFCDFFACFF871A7FFF93FFA0BD01FFCD" masterDocTitle="New data on the genus Wiedemannia Zetterstedt (Diptera: Empididae) from the Caucasus with description of four new species" masterLastPageNumber="369" masterPageNumber="351" pageNumber="354" updateTime="1698616197258" updateUser="plazi">
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<mods:title id="6A57702F66FB1A7408ABD1DED529EB51">New data on the genus Wiedemannia Zetterstedt (Diptera: Empididae) from the Caucasus with description of four new species</mods:title>
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<mods:namePart id="A31FBC9CEEE66EBB32440752AF9F3FC3">Kustov, Semen</mods:namePart>
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<mods:namePart id="8D208AF390A2E771AFD4FD2A4289C26F">Zherebilo, Danil</mods:namePart>
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<mods:date id="F91A46CD44FA13FF1BD8DBDBE0907DA0">2015</mods:date>
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<treatment id="03F487D4FF841A7CFF04FD8CB954FA0F" ID-DOI="http://doi.org/10.5281/zenodo.5691015" ID-GBIF-Taxon="119642280" ID-Zenodo-Dep="5691015" LSID="urn:lsid:plazi:treatment:03F487D4FF841A7CFF04FD8CB954FA0F" httpUri="http://treatment.plazi.org/id/03F487D4FF841A7CFF04FD8CB954FA0F" lastPageNumber="354" pageId="3" pageNumber="354">
<subSubSection id="C3476549FF841A7CFF04FD8CBDEAFDAA" pageId="3" pageNumber="354" type="nomenclature">
<paragraph id="8BE236C2FF841A7CFF04FD8CBFB2FD8B" blockId="3.[151,691,556,615]" box="[151,691,556,582]" pageId="3" pageNumber="354">
<heading id="D0AA81AEFF841A7CFF04FD8CBFB2FD8B" bold="true" box="[151,691,556,582]" fontSize="11" level="1" pageId="3" pageNumber="354" reason="1">
<taxonomicName id="4C5D4D41FF841A7CFF04FD8CBFB2FD8B" ID-CoL="8FM5W" authority="Mik" authorityName="Mik" box="[151,691,556,582]" class="Insecta" family="Empididae" genus="Wiedemannia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="354" phylum="Arthropoda" rank="species" species="beckeri" subGenus="Chamaedipsia">
<emphasis id="B929EAD0FF841A7CFF04FD8CBFB2FD8B" bold="true" box="[151,691,556,582]" pageId="3" pageNumber="354">
<emphasis id="B929EAD0FF841A7CFF04FD8CBC3FFD8B" bold="true" box="[151,318,556,582]" italics="true" pageId="3" pageNumber="354">Wiedemannia</emphasis>
(
<emphasis id="B929EAD0FF841A7CFEDDFD8CBCFBFD8B" bold="true" box="[334,506,556,582]" italics="true" pageId="3" pageNumber="354">Chamaedipsia</emphasis>
)
<emphasis id="B929EAD0FF841A7CFD9FFD8CBF65FD8B" bold="true" box="[524,612,556,582]" italics="true" pageId="3" pageNumber="354">beckeri</emphasis>
(Mik)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BE236C2FF841A7CFF04FDEEBDEAFDAA" blockId="3.[151,691,556,615]" box="[151,235,590,615]" pageId="3" pageNumber="354">
(
<figureCitation id="13662A47FF841A7CFF0CFDEEBDE1FDAA" box="[159,224,590,615]" captionStart="FIGURES 1 6" captionStartId="4.[151,263,1947,1969]" captionTargetBox="[227,1339,208,1887]" captionTargetId="figure@4.[205,1369,193,1904]" captionTargetPageId="4" captionText="FIGURES 1 6. Male terminalia of Wiedemannia, lateral view. 1, W. beckeri (Mik); 2, W. braueri (Mik); 3, W. caucasica Joost; 4, W. chvali Joost; 5, W. fallaciosa (Loew); 6, W. ivkovicae sp. nov. Scale bar = 0.1 mm. Abbreviations: cerc pl—cercal plate; cl cerc—clasping cercus, distph—distiphallus, epand—epandrium, hypd—hypandrium, ph—phallus." httpUri="https://zenodo.org/record/243486/files/figure.png" pageId="3" pageNumber="354">Fig. 1</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3476549FF841A7CFF04FD36BF16FD61" box="[151,535,662,684]" pageId="3" pageNumber="354" type="reference_group">
<paragraph id="8BE236C2FF841A7CFF04FD36BF16FD61" blockId="3.[151,535,662,684]" box="[151,535,662,684]" pageId="3" pageNumber="354">
<treatmentCitationGroup id="AB4D11ECFF841A7CFF04FD36BF16FD61" box="[151,535,662,684]" pageId="3" pageNumber="354">
<taxonomicName id="4C5D4D41FF841A7CFF04FD36BF12FD61" ID-CoL="8BFNR" authority="Mik, 1889: 71" authorityName="Mik" authorityPageNumber="71" authorityYear="1889" box="[151,531,662,684]" class="Insecta" family="Empididae" genus="Chamaedipsia" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="354" phylum="Arthropoda" rank="species" species="beckeri">
<emphasis id="B929EAD0FF841A7CFF04FD36BC7BFD61" box="[151,378,662,684]" italics="true" pageId="3" pageNumber="354">Chamaedipsia beckeri</emphasis>
<treatmentCitation id="0AFC10D3FF841A7CFE13FD36BF12FD61" author="Mik" box="[384,531,662,684]" page="71" pageId="3" pageNumber="354" year="1889">
<bibRefCitation id="EFCC4B33FF841A7CFE13FD36BCECFD61" author="Mik" box="[384,493,662,684]" pageId="3" pageNumber="354" refString="Mik, J. (1889) Eine neue, aus den Beskiden stammende Art der alten Gattung Clinocera Meig. Wiener Entomologische Zeitung, 8, 71 - 72, 150 - 152." type="journal article" year="1889">Mik, 1889</bibRefCitation>
: 71
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3476549FF841A7CFF04FD79BE4AFBDF" pageId="3" pageNumber="354" type="materials_examined">
<paragraph id="8BE236C2FF841A7CFF04FD79BE4AFBDF" blockId="3.[151,1437,729,1474]" pageId="3" pageNumber="354">
<emphasis id="B929EAD0FF841A7CFF04FD79BC78FD3F" bold="true" box="[151,377,729,754]" pageId="3" pageNumber="354">Material examined</emphasis>
.
<collectingCountry id="F34A7652FF841A7CFE1BFD79BCEFFD3F" box="[392,494,729,754]" name="Russia" pageId="3" pageNumber="354">
<emphasis id="B929EAD0FF841A7CFE1BFD79BCEFFD3F" bold="true" box="[392,494,729,754]" pageId="3" pageNumber="354">RUSSIA</emphasis>
</collectingCountry>
: Krasnodarskiy Territory, Nature Reserve “Kamyshanova Polyana” (5),
<date id="FFE31002FF841A7CFA8EFD79B899FD3F" box="[1309,1432,729,754]" pageId="3" pageNumber="354" value="2013-06-24">24.vi.2013</date>
, S.Yu. Kustov (1 ♀); same data,
<date id="FFE31002FF841A7CFE69FD5DBF75FCDB" box="[506,628,765,790]" pageId="3" pageNumber="354" value="2013-06-24">24.vi.2013</date>
, S. Kustov (
<specimenCount id="9D5BFD4BFF841A7CFC93FD5EBE2BFCDB" box="[768,810,766,790]" pageId="3" pageNumber="354" type="generic">1 ♂</specimenCount>
); same data,
<date id="FFE31002FF841A7CFC2DFD5DB921FCDB" box="[958,1056,765,790]" pageId="3" pageNumber="354" value="2015-05-03">3.v.2015</date>
, S.Yu. Kustov, D. Zherebilo (
<specimenCount id="9D5BFD4BFF841A7CFAFEFD5EB896FCD8" box="[1389,1431,766,790]" pageId="3" pageNumber="354" type="generic">1 ♂</specimenCount>
, 1 ♀).
<collectingCountry id="F34A7652FF841A7CFF71FC81BC68FCF7" box="[226,361,801,826]" name="Georgia" pageId="3" pageNumber="354">
<emphasis id="B929EAD0FF841A7CFF71FC81BC68FCF7" bold="true" box="[226,361,801,826]" pageId="3" pageNumber="354">GEORGIA</emphasis>
</collectingCountry>
: Khevsureti (48),
<date id="FFE31002FF841A7CFDD6FC81BFC6FCF7" box="[581,711,801,826]" pageId="3" pageNumber="354" value="2014-07-21">21.vii.2014</date>
, D. Zherebilo (
<specimenCount id="9D5BFD4BFF841A7CFC12FC82BEB1FCF7" box="[897,944,802,826]" pageId="3" pageNumber="354" type="generic">1 ♂</specimenCount>
); Samtskhe-Javakheti (29),
<date id="FFE31002FF841A7CFB6BFC81B870FCF7" box="[1272,1393,801,826]" pageId="3" pageNumber="354" value="2014-04-20">20.iv.2014</date>
, D. Zherebilo (
<specimenCount id="9D5BFD4BFF841A7CFE84FCE6BC44FC90" box="[279,325,838,862]" pageId="3" pageNumber="354" type="generic">2 ♂</specimenCount>
2 ♀); Lechkhumi Ridge (41),
<date id="FFE31002FF841A7CFD3CFCE5BE26FC93" box="[687,807,837,862]" pageId="3" pageNumber="354" value="2014-04-29">29.iv.2014</date>
, D. Zherebilo (
<specimenCount id="9D5BFD4BFF841A7CFC4DFCE6B90AFC93" box="[990,1035,838,862]" pageId="3" pageNumber="354" type="generic">1 ♂</specimenCount>
); env. Lahepa (23),
<date id="FFE31002FF841A7CFB6AFCE5B873FC93" box="[1273,1394,837,862]" pageId="3" pageNumber="354" value="2014-04-29">29.iv.2014</date>
, D. Zherebilo (
<specimenCount id="9D5BFD4BFF841A7CFE85FCCABC3FFC4C" box="[278,318,874,898]" pageId="3" pageNumber="354" type="generic">1 ♂</specimenCount>
, 2 ♀); Mingrelia (28),
<date id="FFE31002FF841A7CFDACFCCABFA3FC4F" box="[575,674,874,898]" pageId="3" pageNumber="354" value="2014-05-02">2.v.2014</date>
, D. Zherebilo (
<specimenCount id="9D5BFD4BFF841A7CFCDFFCC9BE77FC4C" box="[844,886,873,898]" pageId="3" pageNumber="354" type="generic">5 ♂</specimenCount>
, 3 ♀); ridge Ofitsari (32),
<date id="FFE31002FF841A7CFB0CFCC9B800FC4F" box="[1183,1281,873,898]" pageId="3" pageNumber="354" value="2014-05-05">5.v.2014</date>
, D. Zherebilo (
<specimenCount id="9D5BFD4BFF841A7CFF31FC2EBDCAFC6B" box="[162,203,910,934]" pageId="3" pageNumber="354" type="generic">1 ♂</specimenCount>
); Shavsheti ridge (24),
<date id="FFE31002FF841A7CFE42FC2DBF3AFC6B" box="[465,571,909,934]" pageId="3" pageNumber="354" value="2015-04-04">4.iv.2015</date>
, A.V. Zherebilo (
<specimenCount id="9D5BFD4BFF841A7CFD6DFC2EBE27FC68" box="[766,806,910,934]" pageId="3" pageNumber="354" type="generic">1 ♂</specimenCount>
, 1 ♀); env. Chinkadziebi (25),
<date id="FFE31002FF841A7CFB13FC2DB9EAFC6B" box="[1152,1259,909,934]" pageId="3" pageNumber="354" value="2015-04-06">6.iv.2015</date>
, A.V. Zherebilo (
<specimenCount id="9D5BFD4BFF841A7CFF31FC12BDCCFC07" box="[162,205,946,970]" pageId="3" pageNumber="354" type="generic">1 ♂</specimenCount>
); env. Zedakinchha (34),
<date id="FFE31002FF841A7CFE7CFC11BF5BFC07" box="[495,602,945,970]" pageId="3" pageNumber="354" value="2015-04-09">9.iv.2015</date>
, A.V. Zherebilo (
<specimenCount id="9D5BFD4BFF841A7CFCB3FC12BE48FC04" box="[800,841,946,970]" pageId="3" pageNumber="354" type="generic">1 ♂</specimenCount>
, 1 ♀); env. Chkumi (36),
<date id="FFE31002FF841A7CFBE2FC11B9E9FC07" box="[1137,1256,945,970]" pageId="3" pageNumber="354" value="2015-04-11">11.iv.2015</date>
, A.V. Zherebilo (1 ♀).
<emphasis id="B929EAD0FF841A7CFF73FC75BC78FC23" bold="true" box="[224,377,981,1006]" pageId="3" pageNumber="354">ABKHAZIA</emphasis>
: env. Sukhum (18),
<date id="FFE31002FF841A7CFDC8FC75BFD7FC23" box="[603,726,981,1006]" pageId="3" pageNumber="354" value="2013-06-27">27.vi.2013</date>
, D. Zherebilo (
<specimenCount id="9D5BFD4BFF841A7CFC12FC76BEABFC23" box="[897,938,982,1006]" pageId="3" pageNumber="354" type="generic">1 ♂</specimenCount>
); River Bzyb (11),
<date id="FFE31002FF841A7CFB12FC75B803FC23" box="[1153,1282,981,1006]" pageId="3" pageNumber="354" value="2013-07-11">11.vii.2013</date>
, D. Zherebilo (
<specimenCount id="9D5BFD4BFF841A7CFF33FC5ABDCBFBDC" box="[160,202,1018,1042]" pageId="3" pageNumber="354" type="generic">3 ♂</specimenCount>
, 1 ♀); River Pshitsa (9),
<date id="FFE31002FF841A7CFE77FC59BF64FBDF" box="[484,613,1017,1042]" pageId="3" pageNumber="354" value="2013-07-11">11.vii.2013</date>
, D. Zherebilo (
<specimenCount id="9D5BFD4BFF841A7CFC81FC5ABE3CFBDF" box="[786,829,1018,1042]" pageId="3" pageNumber="354" type="generic">1 ♂</specimenCount>
).
</paragraph>
</subSubSection>
<subSubSection id="C3476549FF841A7CFF54FBBDB88EFB6C" pageId="3" pageNumber="354" type="diagnosis">
<paragraph id="8BE236C2FF841A7CFF54FBBDB88EFB6C" blockId="3.[151,1437,729,1474]" pageId="3" pageNumber="354">
<emphasis id="B929EAD0FF841A7CFF54FBBDBC3BFBFB" bold="true" box="[199,314,1053,1078]" pageId="3" pageNumber="354">Diagnosis</emphasis>
. This species differs by acrostichals divergent around the prescutellar depression, 5 pairs of dorsocentrals with interspersed shorter setae, olive-brown scutum (postpronotal lobe and lower margin of notopleuron slightly lighter) with triangular patch of blue pruinescence on prescutellar depression, fore femur apically without strong anterior setae; clasping cercus unilobed, elongated, broader basally and narrowed to apex.
</paragraph>
</subSubSection>
<subSubSection id="C3476549FF841A7CFF54FB0DBCEDFAC3" pageId="3" pageNumber="354" type="distribution">
<paragraph id="8BE236C2FF841A7CFF54FB0DBCEDFAC3" blockId="3.[151,1437,729,1474]" pageId="3" pageNumber="354">
<emphasis id="B929EAD0FF841A7CFF54FB0DBC61FB0B" bold="true" box="[199,352,1197,1222]" pageId="3" pageNumber="354">Distribution.</emphasis>
PALAEARCTIC: Europe:
<collectingCountry id="F34A7652FF841A7CFD3DFB0DBE05FB0B" box="[686,772,1197,1222]" name="Austria" pageId="3" pageNumber="354">Austria</collectingCountry>
,
<collectingCountry id="F34A7652FF841A7CFC84FB0DB93FFB0B" box="[791,1086,1197,1222]" name="Bosnia and Herzegovina" pageId="3" pageNumber="354">Bosnia and Herzegovina</collectingCountry>
, French mainland,
<collectingCountry id="F34A7652FF841A7CFABCFB0EB899FB0B" box="[1327,1432,1198,1222]" name="Germany" pageId="3" pageNumber="354">Germany</collectingCountry>
,
<collectingCountry id="F34A7652FF841A7CFF04FB72BDFBFB27" box="[151,250,1234,1258]" name="Hungary" pageId="3" pageNumber="354">Hungary</collectingCountry>
, Italian mainland,
<collectingCountry id="F34A7652FF841A7CFE42FB71BF23FB27" box="[465,546,1233,1258]" name="Poland" pageId="3" pageNumber="354">Poland</collectingCountry>
,
<collectingCountry id="F34A7652FF841A7CFDBCFB71BF96FB27" box="[559,663,1233,1258]" name="Romania" pageId="3" pageNumber="354">Romania</collectingCountry>
,
<collectingCountry id="F34A7652FF841A7CFD36FB71BE0BFB27" box="[677,778,1233,1258]" name="Slovakia" pageId="3" pageNumber="354">Slovakia</collectingCountry>
,
<collectingCountry id="F34A7652FF841A7CFC84FB71BE7DFB27" box="[791,892,1233,1258]" name="Slovenia" pageId="3" pageNumber="354">Slovenia</collectingCountry>
,
<collectingCountry id="F34A7652FF841A7CFC1AFB71B913FB27" box="[905,1042,1233,1258]" name="Switzerland" pageId="3" pageNumber="354">Switzerland</collectingCountry>
; Caucasus:
<collectingCountry id="F34A7652FF841A7CFB08FB71B9E7FB27" box="[1179,1254,1233,1258]" name="Russia" pageId="3" pageNumber="354">Russia</collectingCountry>
(Krasnodarskiy Territory), Abkhazia,
<collectingCountry id="F34A7652FF841A7CFE18FB55BCE9FAC3" box="[395,488,1269,1294]" name="Georgia" pageId="3" pageNumber="354">Georgia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3476549FF841A7CFF54FAB9B954FA0F" pageId="3" pageNumber="354" type="discussion">
<paragraph id="8BE236C2FF841A7CFF54FAB9B954FA0F" blockId="3.[151,1437,729,1474]" pageId="3" pageNumber="354">
<emphasis id="B929EAD0FF841A7CFF54FAB9BC35FAFF" bold="true" box="[199,308,1305,1330]" pageId="3" pageNumber="354">Remarks</emphasis>
.
<bibRefCitation id="EFCC4B33FF841A7CFED7FABABCD3FAFF" author="Joost" box="[324,466,1306,1330]" pageId="3" pageNumber="354" refString="Joost, W. (1981) Beitrag zur Kenntnis der Hemerodromiiniae des Kaukasus (I) (Diptera, Empididae). Reichenbachia, 19 (31), 183 - 191." type="journal article" year="1981">Joost (1981)</bibRefCitation>
first reported this species from the Caucasus, collected the stream Yusengi (
<collectingCountry id="F34A7652FF841A7CFADBFAB9B899FAFF" box="[1352,1432,1305,1330]" name="Russia" pageId="3" pageNumber="354">Russia</collectingCountry>
: Kabardino-Balkaria Republic). In the Catalogue of Palaearctic
<taxonomicName id="4C5D4D41FF841A7CFCCCFA9DB9DAFA9B" authority="Chvala &amp; Wagner 1989" authorityName="Chvala &amp; Wagner" authorityYear="1989" box="[863,1243,1341,1366]" kingdom="Plantae" order="Diptera" pageId="3" pageNumber="354" rank="order">
Diptera (
<bibRefCitation id="EFCC4B33FF841A7CFC51FA9DB9D2FA9B" author="Chvala" box="[962,1235,1341,1366]" pageId="3" pageNumber="354" refString="Chvala, M. &amp; Wagner, R. (1989) Hybotidae. In: Catalogue of Palaearctic Diptera. Volume 6. Therevidae-Empididae. Elsevier Science Publishing, pp. 228 - 336." type="book" year="1989">Chvála &amp; Wagner 1989</bibRefCitation>
)
</taxonomicName>
, this species was mistakenly indicated only from the territory of
<collectingCountry id="F34A7652FF841A7CFD24FAC1BE15FAB7" box="[695,788,1377,1402]" name="Georgia" pageId="3" pageNumber="354">Georgia</collectingCountry>
. Therefore
<taxonomicName id="4C5D4D41FF841A7CFC0BFAC2B9CFFAB4" box="[920,1230,1377,1402]" class="Insecta" family="Empididae" genus="Chamaedipsia" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="354" phylum="Arthropoda" rank="species" species="beckeri" subGenus="Chamaedipsia">
<emphasis id="B929EAD0FF841A7CFC0BFAC2BEB3FAB4" box="[920,946,1378,1401]" italics="true" pageId="3" pageNumber="354">W.</emphasis>
(
<emphasis id="B929EAD0FF841A7CFC55FAC1B969FAB4" box="[966,1128,1377,1401]" italics="true" pageId="3" pageNumber="354">Chamaedipsia</emphasis>
)
<emphasis id="B929EAD0FF841A7CFBE8FAC1B9CFFAB4" box="[1147,1230,1377,1401]" italics="true" pageId="3" pageNumber="354">beckeri</emphasis>
</taxonomicName>
is noted from the territories of
<collectingCountry id="F34A7652FF841A7CFEBBFA25BC83FA53" box="[296,386,1413,1438]" name="Georgia" pageId="3" pageNumber="354">Georgia</collectingCountry>
and Abkhazia for the first time.
<taxonomicName id="4C5D4D41FF841A7CFD7EFA25BEDEFA50" box="[749,991,1413,1437]" class="Insecta" family="Empididae" genus="Wiedemannia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="354" phylum="Arthropoda" rank="species" species="beckeri">
<emphasis id="B929EAD0FF841A7CFD7EFA25BEDEFA50" box="[749,991,1413,1437]" italics="true" pageId="3" pageNumber="354">Wiedemannia beckeri</emphasis>
</taxonomicName>
is common from mountain rivers of the Caucasus from elevations between
<quantity id="4CA59B27FF841A7CFDB1FA09BFAEFA0C" box="[546,687,1449,1474]" metricMagnitude="3" metricUnit="m" metricValue="1.595" metricValueMax="2.83" metricValueMin="0.36" pageId="3" pageNumber="354" unit="m" value="1595.0" valueMax="2830.0" valueMin="360.0">3602830 m</quantity>
a.s.l. and is active from April to July.
</paragraph>
</subSubSection>
</treatment>
</document>