treatments-xml/data/03/F4/87/03F487CEE47BFFC6FF235331FCDC441E.xml
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<document id="3A539D3A00DAA32C26691C9B2F3BC9D0" ID-DOI="10.11646/zootaxa.5351.5.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="8392291" ID-ZooBank="C6C7769C-94F6-496A-9293-2F0E1E201330" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1695993006982" checkinUser="plazi" docAuthor="Davranoglou, Leonidas-Romanos, Baňař, Petr, Kakiopoulos, George, Balázs, Attila, Avtzis, Dimitrios N. &amp; Hlaváč, Peter" docDate="2023" docId="03F487CEE47BFFC6FF235331FCDC441E" docLanguage="en" docName="zootaxa.5351.5.4.pdf" docOrigin="Zootaxa 5351 (5)" docSource="http://dx.doi.org/10.11646/zootaxa.5351.5.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Afropselaphus tymficus Davranoglou, Hlavac &amp; Banar 2023, sp. n." docType="treatment" docVersion="2" lastPageNumber="565" masterDocId="FFCDFFB6E478FFC0FFB4541CFFC84257" masterDocTitle="Three new species of Afropselaphus Jeannel, 1950 (Staphylinidae: Pselaphinae) from Greece and a redescription of Pselaphogenius treskanus (Karaman, 1940)" masterLastPageNumber="570" masterPageNumber="559" pageNumber="562" updateTime="1695993641248" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="B6C9BD7AC3652C26CB8BF573CA95561E">Three new species of Afropselaphus Jeannel, 1950 (Staphylinidae: Pselaphinae) from Greece and a redescription of Pselaphogenius treskanus (Karaman, 1940)</mods:title>
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<mods:namePart id="27787BCEC4DD4DA97E60E17655C8CFEA">Davranoglou, Leonidas-Romanos</mods:namePart>
<mods:affiliation id="5BCE982688A4142374C0A06DE4203A57">Oxford University Museum of Natural History, University of Oxford, Parks Road, Oxford OX 1 3 PW, UK;</mods:affiliation>
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<mods:namePart id="899341B9967CEE2C07079C0A80D13448">Baňař, Petr</mods:namePart>
<mods:affiliation id="E33A343F58366824E3C0E786D102A43E">Department of Entomology, Moravian Museum, Zelný trh 6, 659 37 Brno, Czech Republic;</mods:affiliation>
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<mods:namePart id="2D48FFA0C931508599C2A6C3348D80F4">Kakiopoulos, George</mods:namePart>
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<mods:affiliation id="12011D94ACF5014221143F66C76D5639">Ilidos 60 - 62 street, 115 27, Athens, Greece; https: // orcid. org / 0000 - 0003 - 3486 - 5099</mods:affiliation>
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<mods:namePart id="F9DC9AE1CB1DBB7438CF1AD07B18C68A">Balázs, Attila</mods:namePart>
<mods:affiliation id="DC0FAECD1FF9745A6882A28EABC41F07">Department of Zoology, Fisheries, Hydrobiology and Apiculture, Mendel University in Brno, Zemědělská 1, 613 00 Brno, Czech Republic;</mods:affiliation>
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<mods:namePart id="CD68AB8091A61F557D14E5493E0B4FC9">Avtzis, Dimitrios N.</mods:namePart>
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<mods:affiliation id="36C418544D16E7183C901E41414EB7B8">Forest Research Institute, Hellenic Agricultural Organization Demeter, 570 06 Vassilika-Thessaloniki, Greece; dimitrios. avtzis @ gmail. com; https: // orcid. org / 0000 - 0002 - 7772 - 6892</mods:affiliation>
<mods:nameIdentifier id="AC30EEA33B6C16637D7B0C46040877D0" type="email">dimitrios.avtzis@gmail.com</mods:nameIdentifier>
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<mods:namePart id="C719ED8BA20E2B00278463444581F87E">Hlaváč, Peter</mods:namePart>
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<mods:affiliation id="979562354FC9A13CCADD25D26F434683">Department of Entomology, National Museum, Cirkusová 1740, CZ- 193 00 Praha 9 - Horní Počernice; peterclaviger @ gmail. com; https: // orcid. org / 0000 - 0001 - 5060 - 0811</mods:affiliation>
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<taxonomicName id="4C5D4D5BE47BFFC3FF235331FCF54510" authority="Davranoglou, Hlavac &amp; Banar" authorityName="Davranoglou, Hlavac &amp; Banar" authorityYear="2023" box="[151,829,1837,1864]" class="Insecta" family="Staphylinidae" genus="Afropselaphus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="562" phylum="Arthropoda" rank="species" species="tymficus" status="sp. n.">
<emphasis id="B929EACAE47BFFC3FF235331FE7E4510" bold="true" box="[151,438,1837,1864]" italics="true" pageId="3" pageNumber="562">Afropselaphus tymficus</emphasis>
Davranoglou, Hlaváč &amp; Baňař
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,
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<paragraph id="8BE236D8E47BFFC3FF23534DFE49453B" blockId="3.[151,908,1837,1900]" box="[151,385,1873,1900]" pageId="3" pageNumber="562">
(
<figureCitation id="13662A5DE47BFFC3FF2B534DFF31453C" box="[159,249,1873,1899]" captionStart="FIGURE 2" captionStartId="3.[151,250,1695,1719]" captionTargetBox="[389,1197,381,1672]" captionTargetId="figure-114@3.[389,1197,381,1672]" captionTargetPageId="3" captionText="FIGURE 2. Habitus photos of examined species of Pselaphini.A) A. taygetensis sp. n.; B) A. tymficus sp. n.; C) A. euboicus sp. n.; D) Pselaphogenius treskanus (Karaman, 1940). Scale bars correspond to 250 um." figureDoi="http://doi.org/10.5281/zenodo.8392297" httpUri="https://zenodo.org/record/8392297/files/figure.png" pageId="3" pageNumber="562">Figs 2B</figureCitation>
<figureCitation id="13662A5DE47BFFC3FEBD534DFEE2453C" box="[265,298,1873,1899]" captionStart="FIGURE 6" captionStartId="7.[151,250,1240,1264]" captionTargetBox="[325,1263,237,1217]" captionTargetId="figure-53@7.[325,1263,237,1217]" captionTargetPageId="7" captionText="FIGURE 6. Lateral view of metaventrite in the examined Pselaphini, using scanning electron microscopy.A) A. taygetensis sp. n.; B) A. tymficus sp. n.; C) A. euboicus sp. n.; D) Pselaphogenius treskanus (Karaman, 1940). Scale bars correspond to 100 um." figureDoi="http://doi.org/10.5281/zenodo.8392309" httpUri="https://zenodo.org/record/8392309/files/figure.png" pageId="3" pageNumber="562">6B</figureCitation>
,
<figureCitation id="13662A5DE47BFFC3FE81534EFEB1453B" box="[309,377,1874,1900]" captionStart="FIGURE 7" captionStartId="8.[151,250,1268,1292]" captionTargetBox="[208,1366,170,1243]" captionTargetId="figure-26@8.[208,1379,166,1244]" captionTargetPageId="8" captionText="FIGURE 7. Photomicrographs showing the dorsal aedeagal surface of the examined Pselaphini. A) A. taygetensis sp. n.; B) same, schematic; C) A. tymficus sp. n.; D) same, schematic; E) A. euboicus sp. n.; F) same, schematic; G) Pselaphogenius treskanus (Karaman, 1940); F) same, schematic. Blue dashed lines indicate membranous structures. All scale bars correspond to 200 um." figureDoi="http://doi.org/10.5281/zenodo.8392311" httpUri="https://zenodo.org/record/8392311/files/figure.png" pageId="3" pageNumber="562">7C, D</figureCitation>
)
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.
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, ♁:
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, with one label: “GREECE [GR 014]
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W PAPINGO /
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/ sifting in
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forest; along /
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,
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app. extr. /
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,
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&amp;
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lgt.” [white, printed] (
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).
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:
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1 ♁♁,
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, 1 ex, same data as for holotype (
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,
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;
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1 ♁, 11 exx, with one label: “
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[GR 013]
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/ OROs TYMFI,
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W PAPINGO /
<geoCoordinate id="EE69501FE47CFFC4FE6754C2FDBC42AE" box="[467,628,222,249]" degrees="39" direction="east" minutes="57" orientation="longitude" pageId="4" pageNumber="563" precision="1" seconds="55.0" value="39.96528">39°57´55.0´´E</geoCoordinate>
20°42´39.2´´ / sifting in narrow canyon along / stream,
<quantity id="4CA59B3DE47CFFC4FB5F54C3FAFD42AE" box="[1259,1333,223,249]" metricMagnitude="2" metricUnit="m" metricValue="9.07" pageId="4" pageNumber="563" unit="m" value="907.0">
<elevation id="0070D1EBE47CFFC4FB5F54C3FAFD42AE" box="[1259,1333,223,249]" metricMagnitude="2" metricUnit="m" metricValue="9.07" pageId="4" pageNumber="563" unit="m" value="907.0">907 m</elevation>
</quantity>
, Winkler app. extr. /
<collectorName id="26A8530EE47CFFC4FEA0551FFEA6434A" box="[276,366,259,285]" pageId="4" pageNumber="563">P.Baňař</collectorName>
,
<collectorName id="26A8530EE47CFFC4FECD551FFE13434A" box="[377,475,259,285]" pageId="4" pageNumber="563">P.Hlaváč</collectorName>
&amp;
<collectorName id="26A8530EE47CFFC4FE4A551FFDA3434A" box="[510,619,259,285]" pageId="4" pageNumber="563">A. Balázs</collectorName>
lgt.” [white, printed] (
<collectionCode id="ED4CAE1DE47CFFC4FCD2551FFC09434A" box="[870,961,259,285]" country="Czech Republic" httpUri="http://grbio.org/cool/w50r-ge1v" name="Moravske Muzeum [Moravian Museum]" pageId="4" pageNumber="563">MMBC</collectionCode>
,
<collectionCode id="ED4CAE1DE47CFFC4FC79551FFBD0434A" box="[973,1048,259,285]" pageId="4" pageNumber="563">PCPH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF226650E47CFFC4FF2352F8FABA4577" ID-DOI="http://doi.org/10.5281/zenodo.8392301" ID-Zenodo-Dep="8392301" httpUri="https://zenodo.org/record/8392301/files/figure.png" pageId="4" pageNumber="563" startId="4.[151,250,1764,1788]" targetBox="[325,1263,345,1741]" targetPageId="4" targetType="figure">
<paragraph id="8BE236D8E47CFFC4FF2352F8FABA4577" blockId="4.[151,1436,1764,1824]" pageId="4" pageNumber="563">
<emphasis id="B929EACAE47CFFC4FF2352F8FEDA44AB" bold="true" box="[151,274,1764,1788]" pageId="4" pageNumber="563">FIGURE 3.</emphasis>
Scanning electron microscopy image of dorsal view of head in the examined
<taxonomicName id="4C5D4D5BE47CFFC4FBAE52F8FB4F44AB" authorityName="Jeannel" authorityYear="1950" box="[1050,1159,1764,1788]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="563" phylum="Arthropoda" rank="tribe" tribe="Pselaphini">Pselaphini</taxonomicName>
.A)
<taxonomicName id="4C5D4D5BE47CFFC4FB1952F9FAF244AB" authority="Davranoglou &amp; Baňař &amp; Kakiopoulos &amp; Balázs &amp; Avtzis &amp; Hlaváč, 2023" authorityName="Davranoglou &amp; Baňař &amp; Kakiopoulos &amp; Balázs &amp; Avtzis &amp; Hlaváč" authorityYear="2023" box="[1197,1338,1765,1788]" class="Insecta" family="Staphylinidae" genus="Afropselaphus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="563" phylum="Arthropoda" rank="species" species="taygetensis" status="sp. n.">
<emphasis id="B929EACAE47CFFC4FB1952F9FAF244AB" box="[1197,1338,1765,1788]" italics="true" pageId="4" pageNumber="563">A. taygetensis</emphasis>
</taxonomicName>
<emphasis id="B929EACAE47CFFC4FA8A52F8FABE44AB" bold="true" box="[1342,1398,1764,1788]" pageId="4" pageNumber="563">
<taxonomicNameLabel id="A21A57B1E47CFFC4FA8A52F8FABE44AB" box="[1342,1398,1764,1788]" pageId="4" pageNumber="563" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
; B)
<taxonomicName id="4C5D4D5BE47CFFC4FF235315FECF4577" authority="Davranoglou &amp; Baňař &amp; Kakiopoulos &amp; Balázs &amp; Avtzis &amp; Hlaváč, 2023" authorityName="Davranoglou &amp; Baňař &amp; Kakiopoulos &amp; Balázs &amp; Avtzis &amp; Hlaváč" authorityYear="2023" box="[151,263,1800,1824]" class="Insecta" family="Staphylinidae" genus="Afropselaphus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="563" phylum="Arthropoda" rank="species" species="tymficus" status="sp. n.">
<emphasis id="B929EACAE47CFFC4FF235315FECF4577" box="[151,263,1800,1824]" italics="true" pageId="4" pageNumber="563">A. tymficus</emphasis>
</taxonomicName>
<emphasis id="B929EACAE47CFFC4FEBA5314FE8F4577" bold="true" box="[270,327,1800,1824]" pageId="4" pageNumber="563">
<taxonomicNameLabel id="A21A57B1E47CFFC4FEBA5314FE8F4577" box="[270,327,1800,1824]" pageId="4" pageNumber="563" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
; C)
<taxonomicName id="4C5D4D5BE47CFFC4FEC05315FE224577" authority="Davranoglou &amp; Baňař &amp; Kakiopoulos &amp; Balázs &amp; Avtzis &amp; Hlaváč, 2023" authorityName="Davranoglou &amp; Baňař &amp; Kakiopoulos &amp; Balázs &amp; Avtzis &amp; Hlaváč" authorityYear="2023" box="[372,490,1800,1824]" class="Insecta" family="Staphylinidae" genus="Afropselaphus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="563" phylum="Arthropoda" rank="species" species="euboicus" status="sp. n.">
<emphasis id="B929EACAE47CFFC4FEC05315FE224577" box="[372,490,1800,1824]" italics="true" pageId="4" pageNumber="563">A. euboicus</emphasis>
</taxonomicName>
<emphasis id="B929EACAE47CFFC4FE455314FDE24577" bold="true" box="[497,554,1800,1824]" pageId="4" pageNumber="563">
<taxonomicNameLabel id="A21A57B1E47CFFC4FE455314FDE24577" box="[497,554,1800,1824]" pageId="4" pageNumber="563" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
; D)
<taxonomicName id="4C5D4D5BE47CFFC4FDEC5314FC974577" baseAuthorityName="Karaman" baseAuthorityYear="1940" box="[600,863,1800,1824]" class="Insecta" family="Staphylinidae" genus="Pselaphogenius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="563" phylum="Arthropoda" rank="species" species="treskanus">
<emphasis id="B929EACAE47CFFC4FDEC5314FC974577" box="[600,863,1800,1824]" italics="true" pageId="4" pageNumber="563">Pselaphogenius treskanus</emphasis>
</taxonomicName>
(Karaman, 1940). Scale bars correspond to 100 um.
</paragraph>
</caption>
<subSubSection id="C3476553E47CFFC4FF735352FC3345AE" pageId="4" pageNumber="563" type="diagnosis">
<paragraph id="8BE236D8E47CFFC4FF735352FC3345AE" blockId="4.[151,1437,1870,2041]" pageId="4" pageNumber="563">
<emphasis id="B929EACAE47CFFC4FF735352FE8A453F" bold="true" box="[199,322,1870,1896]" pageId="4" pageNumber="563">Diagnosis.</emphasis>
Head 1.4 times longer than wide, scape 2.3 times longer than wide, median gular ridge with a distinct median bulge (
<figureCitation id="13662A5DE47CFFC4FE2C536EFE3845DA" box="[408,496,1906,1933]" captionStart="FIGURE 4" captionStartId="5.[151,250,1185,1209]" captionTargetBox="[355,1231,181,1161]" captionTargetId="figure-17@5.[355,1231,181,1161]" captionTargetPageId="5" captionText="FIGURE 4. Scanning electron microscopy image of lateral view of head in the examined Pselaphini.A) A. taygetensis sp. n.; B) A. tymficus sp. n.; C) A. euboicus sp. n.; D) Pselaphogenius treskanus (Karaman, 1940). Scale bars correspond to 100 um." figureDoi="http://doi.org/10.5281/zenodo.8392303" httpUri="https://zenodo.org/record/8392303/files/figure.png" pageId="4" pageNumber="563">Fig. 4B</figureCitation>
), tergite 1 (IV) 1.61.7 times broader than long, sternite 2 (IV) 2.3 times wider than long; metaventrite of male produced into distinct subtriangular spine (
<figureCitation id="13662A5DE47CFFC4FC71538AFBE845E7" box="[965,1056,1942,1968]" captionStart="FIGURE 6" captionStartId="7.[151,250,1240,1264]" captionTargetBox="[325,1263,237,1217]" captionTargetId="figure-53@7.[325,1263,237,1217]" captionTargetPageId="7" captionText="FIGURE 6. Lateral view of metaventrite in the examined Pselaphini, using scanning electron microscopy.A) A. taygetensis sp. n.; B) A. tymficus sp. n.; C) A. euboicus sp. n.; D) Pselaphogenius treskanus (Karaman, 1940). Scale bars correspond to 100 um." figureDoi="http://doi.org/10.5281/zenodo.8392309" httpUri="https://zenodo.org/record/8392309/files/figure.png" pageId="4" pageNumber="563">Fig. 6B</figureCitation>
); central spine of internal sac of aedeagus with three simple branches adjacent to a denticle-bearing sclerite (
<figureCitation id="13662A5DE47CFFC4FC5A53A6FBA14582" box="[1006,1129,1978,2005]" captionStart="FIGURE 7" captionStartId="8.[151,250,1268,1292]" captionTargetBox="[208,1366,170,1243]" captionTargetId="figure-26@8.[208,1379,166,1244]" captionTargetPageId="8" captionText="FIGURE 7. Photomicrographs showing the dorsal aedeagal surface of the examined Pselaphini. A) A. taygetensis sp. n.; B) same, schematic; C) A. tymficus sp. n.; D) same, schematic; E) A. euboicus sp. n.; F) same, schematic; G) Pselaphogenius treskanus (Karaman, 1940); F) same, schematic. Blue dashed lines indicate membranous structures. All scale bars correspond to 200 um." figureDoi="http://doi.org/10.5281/zenodo.8392311" httpUri="https://zenodo.org/record/8392311/files/figure.png" pageId="4" pageNumber="563">Fig. 7C, D</figureCitation>
); second spine of aedeagus simple, overlaying membrane bearing large number of denticles (
<figureCitation id="13662A5DE47CFFC4FCC753C2FC2645AE" box="[883,1006,2014,2041]" captionStart="FIGURE 7" captionStartId="8.[151,250,1268,1292]" captionTargetBox="[208,1366,170,1243]" captionTargetId="figure-26@8.[208,1379,166,1244]" captionTargetPageId="8" captionText="FIGURE 7. Photomicrographs showing the dorsal aedeagal surface of the examined Pselaphini. A) A. taygetensis sp. n.; B) same, schematic; C) A. tymficus sp. n.; D) same, schematic; E) A. euboicus sp. n.; F) same, schematic; G) Pselaphogenius treskanus (Karaman, 1940); F) same, schematic. Blue dashed lines indicate membranous structures. All scale bars correspond to 200 um." figureDoi="http://doi.org/10.5281/zenodo.8392311" httpUri="https://zenodo.org/record/8392311/files/figure.png" pageId="4" pageNumber="563">Fig. 7C, D</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF226650E47DFFC5FF2350BDFABA468B" ID-DOI="http://doi.org/10.5281/zenodo.8392303" ID-Zenodo-Dep="8392303" httpUri="https://zenodo.org/record/8392303/files/figure.png" pageId="5" pageNumber="564" startId="5.[151,250,1185,1209]" targetBox="[355,1231,181,1161]" targetPageId="5" targetType="figure">
<paragraph id="8BE236D8E47DFFC5FF2350BDFABA468B" blockId="5.[151,1436,1185,1245]" pageId="5" pageNumber="564">
<emphasis id="B929EACAE47DFFC5FF2350BDFEDA46EE" bold="true" box="[151,274,1185,1209]" pageId="5" pageNumber="564">FIGURE 4.</emphasis>
Scanning electron microscopy image of lateral view of head in the examined
<taxonomicName id="4C5D4D5BE47DFFC5FBAE50BDFB4F46EE" authorityName="Jeannel" authorityYear="1950" box="[1050,1159,1185,1209]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="564" phylum="Arthropoda" rank="tribe" tribe="Pselaphini">Pselaphini</taxonomicName>
. A)
<taxonomicName id="4C5D4D5BE47DFFC5FB1A50BDFAF246EE" authority="Davranoglou &amp; Baňař &amp; Kakiopoulos &amp; Balázs &amp; Avtzis &amp; Hlaváč, 2023" authorityName="Davranoglou &amp; Baňař &amp; Kakiopoulos &amp; Balázs &amp; Avtzis &amp; Hlaváč" authorityYear="2023" box="[1198,1338,1185,1209]" class="Insecta" family="Staphylinidae" genus="Afropselaphus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="564" phylum="Arthropoda" rank="species" species="taygetensis" status="sp. n.">
<emphasis id="B929EACAE47DFFC5FB1A50BDFAF246EE" box="[1198,1338,1185,1209]" italics="true" pageId="5" pageNumber="564">A. taygetensis</emphasis>
</taxonomicName>
<emphasis id="B929EACAE47DFFC5FA8B50BDFABF46EE" bold="true" box="[1343,1399,1185,1209]" pageId="5" pageNumber="564">
<taxonomicNameLabel id="A21A57B1E47DFFC5FA8B50BDFABF46EE" box="[1343,1399,1185,1209]" pageId="5" pageNumber="564" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
; B)
<taxonomicName id="4C5D4D5BE47DFFC5FF2350D9FECF468A" authority="Davranoglou &amp; Baňař &amp; Kakiopoulos &amp; Balázs &amp; Avtzis &amp; Hlaváč, 2023" authorityName="Davranoglou &amp; Baňař &amp; Kakiopoulos &amp; Balázs &amp; Avtzis &amp; Hlaváč" authorityYear="2023" box="[151,263,1221,1245]" class="Insecta" family="Staphylinidae" genus="Afropselaphus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="564" phylum="Arthropoda" rank="species" species="tymficus" status="sp. n.">
<emphasis id="B929EACAE47DFFC5FF2350D9FECF468A" box="[151,263,1221,1245]" italics="true" pageId="5" pageNumber="564">A. tymficus</emphasis>
</taxonomicName>
<emphasis id="B929EACAE47DFFC5FEBA50D9FE8F468A" bold="true" box="[270,327,1221,1245]" pageId="5" pageNumber="564">
<taxonomicNameLabel id="A21A57B1E47DFFC5FEBA50D9FE8F468A" box="[270,327,1221,1245]" pageId="5" pageNumber="564" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
; C)
<taxonomicName id="4C5D4D5BE47DFFC5FEC050D9FE22468A" authority="Davranoglou &amp; Baňař &amp; Kakiopoulos &amp; Balázs &amp; Avtzis &amp; Hlaváč, 2023" authorityName="Davranoglou &amp; Baňař &amp; Kakiopoulos &amp; Balázs &amp; Avtzis &amp; Hlaváč" authorityYear="2023" box="[372,490,1221,1245]" class="Insecta" family="Staphylinidae" genus="Afropselaphus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="564" phylum="Arthropoda" rank="species" species="euboicus" status="sp. n.">
<emphasis id="B929EACAE47DFFC5FEC050D9FE22468A" box="[372,490,1221,1245]" italics="true" pageId="5" pageNumber="564">A. euboicus</emphasis>
</taxonomicName>
<emphasis id="B929EACAE47DFFC5FE4550D9FDE2468A" bold="true" box="[497,554,1221,1245]" pageId="5" pageNumber="564">
<taxonomicNameLabel id="A21A57B1E47DFFC5FE4550D9FDE2468A" box="[497,554,1221,1245]" pageId="5" pageNumber="564" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
; D)
<taxonomicName id="4C5D4D5BE47DFFC5FDEC50D9FC97468A" baseAuthorityName="Karaman" baseAuthorityYear="1940" box="[600,863,1221,1245]" class="Insecta" family="Staphylinidae" genus="Pselaphogenius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="564" phylum="Arthropoda" rank="species" species="treskanus">
<emphasis id="B929EACAE47DFFC5FDEC50D9FC97468A" box="[600,863,1221,1245]" italics="true" pageId="5" pageNumber="564">Pselaphogenius treskanus</emphasis>
</taxonomicName>
(Karaman, 1940). Scale bars correspond to 100 um.
</paragraph>
</caption>
<subSubSection id="C3476553E47DFFC6FF735117FC834727" lastPageId="6" lastPageNumber="565" pageId="5" pageNumber="564" type="description">
<paragraph id="8BE236D8E47DFFC5FF735117FE9C47E2" blockId="5.[151,1437,1291,1965]" pageId="5" pageNumber="564">
<emphasis id="B929EACAE47DFFC5FF735117FE914772" bold="true" box="[199,345,1291,1317]" pageId="5" pageNumber="564">Description.</emphasis>
Body length
<quantity id="4CA59B3DE47DFFC5FE5A5117FD8E4771" box="[494,582,1291,1318]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="5" pageNumber="564" unit="mm" value="1.9">1.9 mm</quantity>
. Coloration largely reddish-brown, maxillary palpi and legs lighter, posterior margin of elytra much darker, almost black (
<figureCitation id="13662A5DE47DFFC5FD3F5133FD2B471E" box="[651,739,1327,1353]" captionStart="FIGURE 2" captionStartId="3.[151,250,1695,1719]" captionTargetBox="[389,1197,381,1672]" captionTargetId="figure-114@3.[389,1197,381,1672]" captionTargetPageId="3" captionText="FIGURE 2. Habitus photos of examined species of Pselaphini.A) A. taygetensis sp. n.; B) A. tymficus sp. n.; C) A. euboicus sp. n.; D) Pselaphogenius treskanus (Karaman, 1940). Scale bars correspond to 250 um." figureDoi="http://doi.org/10.5281/zenodo.8392297" httpUri="https://zenodo.org/record/8392297/files/figure.png" pageId="5" pageNumber="564">Fig. 2B</figureCitation>
). Pilosity of body sparse; head, legs, and antennae covered by sparse, adjacent golden setae; elytra with few rows of setae, which are denser on elytral marginal line; with row of setae on distal margin of elytra; proximal margin of tergite II with very dense pilosity (
<figureCitation id="13662A5DE47DFFC5FBE8516BFB7B47C6" box="[1116,1203,1399,1425]" captionStart="FIGURE 2" captionStartId="3.[151,250,1695,1719]" captionTargetBox="[389,1197,381,1672]" captionTargetId="figure-114@3.[389,1197,381,1672]" captionTargetPageId="3" captionText="FIGURE 2. Habitus photos of examined species of Pselaphini.A) A. taygetensis sp. n.; B) A. tymficus sp. n.; C) A. euboicus sp. n.; D) Pselaphogenius treskanus (Karaman, 1940). Scale bars correspond to 250 um." figureDoi="http://doi.org/10.5281/zenodo.8392297" httpUri="https://zenodo.org/record/8392297/files/figure.png" pageId="5" pageNumber="564">Fig. 2B</figureCitation>
), remaining segment largely glabrous.
</paragraph>
<paragraph id="8BE236D8E47DFFC5FF7351A3FA8744DA" blockId="5.[151,1437,1291,1965]" pageId="5" pageNumber="564">
Head 1.4 times longer than wide, each eye composed of 7 ommatidia (
<figureCitation id="13662A5DE47DFFC5FC6C51A3FBF2478E" box="[984,1082,1471,1497]" captionStart="FIGURE 3" captionStartId="4.[151,250,1764,1788]" captionTargetBox="[325,1263,345,1741]" captionTargetId="figure-99@4.[325,1263,345,1741]" captionTargetPageId="4" captionText="FIGURE 3. Scanning electron microscopy image of dorsal view of head in the examined Pselaphini.A) A. taygetensis sp. n.; B) A. tymficus sp. n.; C) A. euboicus sp. n.; D) Pselaphogenius treskanus (Karaman, 1940). Scale bars correspond to 100 um." figureDoi="http://doi.org/10.5281/zenodo.8392301" httpUri="https://zenodo.org/record/8392301/files/figure.png" pageId="5" pageNumber="564">Figs. 3B</figureCitation>
,
<figureCitation id="13662A5DE47DFFC5FBF151A3FBA1478E" box="[1093,1129,1471,1497]" captionStart="FIGURE 4" captionStartId="5.[151,250,1185,1209]" captionTargetBox="[355,1231,181,1161]" captionTargetId="figure-17@5.[355,1231,181,1161]" captionTargetPageId="5" captionText="FIGURE 4. Scanning electron microscopy image of lateral view of head in the examined Pselaphini.A) A. taygetensis sp. n.; B) A. tymficus sp. n.; C) A. euboicus sp. n.; D) Pselaphogenius treskanus (Karaman, 1940). Scale bars correspond to 100 um." figureDoi="http://doi.org/10.5281/zenodo.8392303" httpUri="https://zenodo.org/record/8392303/files/figure.png" pageId="5" pageNumber="564">4B</figureCitation>
); median gular ridge with a distinct median protrusion (
<figureCitation id="13662A5DE47DFFC5FE7A51FFFDEE47AA" box="[462,550,1507,1533]" captionStart="FIGURE 4" captionStartId="5.[151,250,1185,1209]" captionTargetBox="[355,1231,181,1161]" captionTargetId="figure-17@5.[355,1231,181,1161]" captionTargetPageId="5" captionText="FIGURE 4. Scanning electron microscopy image of lateral view of head in the examined Pselaphini.A) A. taygetensis sp. n.; B) A. tymficus sp. n.; C) A. euboicus sp. n.; D) Pselaphogenius treskanus (Karaman, 1940). Scale bars correspond to 100 um." figureDoi="http://doi.org/10.5281/zenodo.8392303" httpUri="https://zenodo.org/record/8392303/files/figure.png" pageId="5" pageNumber="564">Fig. 4B</figureCitation>
); antennae
<quantity id="4CA59B3DE47DFFC5FD1251FFFD3447A9" box="[678,764,1507,1534]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="5" pageNumber="564" unit="mm" value="0.9">0.9 mm</quantity>
long; scape cylindrical, 2.3 times longer than wide, and 2.3 times longer than pedicel; pedicel about as long as wide; pedicel and antennomeres 38 rounded, subequal in length; antennomeres 910 about 1.4 times longer than length of each antennomere 38; terminal antennomere (11) about 1.3 times as long as 910 combined, and 1.9 times longer than wide (
<figureCitation id="13662A5DE47DFFC5FC265253FC21443D" box="[914,1001,1615,1642]" captionStart="FIGURE 5" captionStartId="6.[151,250,1008,1032]" captionTargetBox="[361,1225,181,984]" captionTargetId="figure-26@6.[361,1225,181,984]" captionTargetPageId="6" captionText="FIGURE 5. Scanning electron microscopy image of antenna in the examined Pselaphini.A) A. taygetensis sp. n.; B) A. tymficus sp. n.; C) A. euboicus sp. n.; D) Pselaphogenius treskanus (Karaman, 1940). Scale bars correspond to 200 um." figureDoi="http://doi.org/10.5281/zenodo.8392305" httpUri="https://zenodo.org/record/8392305/files/figure.png" pageId="5" pageNumber="564">Fig. 5B</figureCitation>
). Maxillary palpomere 1 shortest, 2 2.1 times longer than 1; 3 extremely short and rounded; palpomere 4 1.9 times longer than 2, apex club-shaped.
</paragraph>
<paragraph id="8BE236D8E47DFFC5FF735284FBA644AD" blockId="5.[151,1437,1291,1965]" pageId="5" pageNumber="564">
Pronotum
<quantity id="4CA59B3DE47DFFC5FE88528BFE5A44E5" box="[316,402,1687,1714]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="564" unit="mm" value="0.4">0.4 mm</quantity>
wide, smooth, about 1.5 times longer than wide; slightly constricted proximally and distally, broadest medially; about 1.1 times longer than head; metaventrite produced into a distinct subtriangular protuberance (
<figureCitation id="13662A5DE47DFFC5FF2B52C3FF3044AE" box="[159,248,1759,1785]" captionStart="FIGURE 6" captionStartId="7.[151,250,1240,1264]" captionTargetBox="[325,1263,237,1217]" captionTargetId="figure-53@7.[325,1263,237,1217]" captionTargetPageId="7" captionText="FIGURE 6. Lateral view of metaventrite in the examined Pselaphini, using scanning electron microscopy.A) A. taygetensis sp. n.; B) A. tymficus sp. n.; C) A. euboicus sp. n.; D) Pselaphogenius treskanus (Karaman, 1940). Scale bars correspond to 100 um." figureDoi="http://doi.org/10.5281/zenodo.8392309" httpUri="https://zenodo.org/record/8392309/files/figure.png" pageId="5" pageNumber="564">Fig. 6B</figureCitation>
). Elytra about 1.4 times broader than long, and 1.1 times longer than pronotum
</paragraph>
<paragraph id="8BE236D8E47DFFC5FF73531FFE3B4516" blockId="5.[151,1437,1291,1965]" pageId="5" pageNumber="564">Abdomen about as broad as elytra; tergite 1 (IV) (excluding paratergites) 1.2 times wider than long; sternite 2 (IV) 1.9 times wider than long.</paragraph>
<paragraph id="8BE236D8E47DFFC5FF735357FEB545FA" blockId="5.[151,1437,1291,1965]" pageId="5" pageNumber="564">
Aedeagus (
<figureCitation id="13662A5DE47DFFC5FEF35357FE0B4532" box="[327,451,1867,1893]" captionStart="FIGURE 7" captionStartId="8.[151,250,1268,1292]" captionTargetBox="[208,1366,170,1243]" captionTargetId="figure-26@8.[208,1379,166,1244]" captionTargetPageId="8" captionText="FIGURE 7. Photomicrographs showing the dorsal aedeagal surface of the examined Pselaphini. A) A. taygetensis sp. n.; B) same, schematic; C) A. tymficus sp. n.; D) same, schematic; E) A. euboicus sp. n.; F) same, schematic; G) Pselaphogenius treskanus (Karaman, 1940); F) same, schematic. Blue dashed lines indicate membranous structures. All scale bars correspond to 200 um." figureDoi="http://doi.org/10.5281/zenodo.8392311" httpUri="https://zenodo.org/record/8392311/files/figure.png" pageId="5" pageNumber="564">Fig. 7C, D</figureCitation>
) with central spine of internal sac of aedeagus with three simple branches adjacent to a denticle-bearing sclerite (
<figureCitation id="13662A5DE47DFFC5FE785373FD8045DE" box="[460,584,1903,1929]" captionStart="FIGURE 7" captionStartId="8.[151,250,1268,1292]" captionTargetBox="[208,1366,170,1243]" captionTargetId="figure-26@8.[208,1379,166,1244]" captionTargetPageId="8" captionText="FIGURE 7. Photomicrographs showing the dorsal aedeagal surface of the examined Pselaphini. A) A. taygetensis sp. n.; B) same, schematic; C) A. tymficus sp. n.; D) same, schematic; E) A. euboicus sp. n.; F) same, schematic; G) Pselaphogenius treskanus (Karaman, 1940); F) same, schematic. Blue dashed lines indicate membranous structures. All scale bars correspond to 200 um." figureDoi="http://doi.org/10.5281/zenodo.8392311" httpUri="https://zenodo.org/record/8392311/files/figure.png" pageId="5" pageNumber="564">Fig. 7C, D</figureCitation>
); second spine of aedeagus simple, overlaying a membrane bearing a large number of denticles.
</paragraph>
<caption id="DF226650E47EFFC6FF2357ECFB33467B" ID-DOI="http://doi.org/10.5281/zenodo.8392305" ID-Zenodo-Dep="8392305" httpUri="https://zenodo.org/record/8392305/files/figure.png" pageId="6" pageNumber="565" startId="6.[151,250,1008,1032]" targetBox="[361,1225,181,984]" targetPageId="6" targetType="figure">
<paragraph id="8BE236D8E47EFFC6FF2357ECFB33467B" blockId="6.[151,1436,1008,1068]" pageId="6" pageNumber="565">
<emphasis id="B929EACAE47EFFC6FF2357ECFEDA465F" bold="true" box="[151,274,1008,1032]" pageId="6" pageNumber="565">FIGURE 5.</emphasis>
Scanning electron microscopy image of antenna in the examined
<taxonomicName id="4C5D4D5BE47EFFC6FC1157ECFBDA465F" authorityName="Jeannel" authorityYear="1950" box="[933,1042,1008,1032]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="565" phylum="Arthropoda" rank="tribe" tribe="Pselaphini">Pselaphini</taxonomicName>
. A)
<taxonomicName id="4C5D4D5BE47EFFC6FB8C57EDFB0D465F" authority="Davranoglou &amp; Baňař &amp; Kakiopoulos &amp; Balázs &amp; Avtzis &amp; Hlaváč, 2023" authorityName="Davranoglou &amp; Baňař &amp; Kakiopoulos &amp; Balázs &amp; Avtzis &amp; Hlaváč" authorityYear="2023" box="[1080,1221,1009,1032]" class="Insecta" family="Staphylinidae" genus="Afropselaphus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="565" phylum="Arthropoda" rank="species" species="taygetensis" status="sp. n.">
<emphasis id="B929EACAE47EFFC6FB8C57EDFB0D465F" box="[1080,1221,1009,1032]" italics="true" pageId="6" pageNumber="565">A. taygetensis</emphasis>
</taxonomicName>
<emphasis id="B929EACAE47EFFC6FB7E57ECFACA465F" bold="true" box="[1226,1282,1008,1032]" pageId="6" pageNumber="565">
<taxonomicNameLabel id="A21A57B1E47EFFC6FB7E57ECFACA465F" box="[1226,1282,1008,1032]" pageId="6" pageNumber="565" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
; B)
<taxonomicName id="4C5D4D5BE47EFFC6FA9957EDFA54465F" authority="Davranoglou &amp; Baňař &amp; Kakiopoulos &amp; Balázs &amp; Avtzis &amp; Hlaváč, 2023" authorityName="Davranoglou &amp; Baňař &amp; Kakiopoulos &amp; Balázs &amp; Avtzis &amp; Hlaváč" authorityYear="2023" box="[1325,1436,1008,1032]" class="Insecta" family="Staphylinidae" genus="Afropselaphus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="565" phylum="Arthropoda" rank="species" species="tymficus" status="sp. n.">
<emphasis id="B929EACAE47EFFC6FA9957EDFA54465F" box="[1325,1436,1008,1032]" italics="true" pageId="6" pageNumber="565">A. tymficus</emphasis>
</taxonomicName>
<emphasis id="B929EACAE47EFFC6FF235008FF18467B" bold="true" box="[151,208,1044,1068]" pageId="6" pageNumber="565">
<taxonomicNameLabel id="A21A57B1E47EFFC6FF235008FF18467B" box="[151,208,1044,1068]" pageId="6" pageNumber="565" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
; C)
<taxonomicName id="4C5D4D5BE47EFFC6FF495009FEBB467B" authority="Davranoglou &amp; Baňař &amp; Kakiopoulos &amp; Balázs &amp; Avtzis &amp; Hlaváč, 2023" authorityName="Davranoglou &amp; Baňař &amp; Kakiopoulos &amp; Balázs &amp; Avtzis &amp; Hlaváč" authorityYear="2023" box="[253,371,1044,1068]" class="Insecta" family="Staphylinidae" genus="Afropselaphus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="565" phylum="Arthropoda" rank="species" species="euboicus" status="sp. n.">
<emphasis id="B929EACAE47EFFC6FF495009FEBB467B" box="[253,371,1044,1068]" italics="true" pageId="6" pageNumber="565">A. euboicus</emphasis>
</taxonomicName>
<emphasis id="B929EACAE47EFFC6FECE5008FE7B467B" bold="true" box="[378,435,1044,1068]" pageId="6" pageNumber="565">
<taxonomicNameLabel id="A21A57B1E47EFFC6FECE5008FE7B467B" box="[378,435,1044,1068]" pageId="6" pageNumber="565" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
; D)
<taxonomicName id="4C5D4D5BE47EFFC6FE555008FD20467B" baseAuthorityName="Karaman" baseAuthorityYear="1940" box="[481,744,1044,1068]" class="Insecta" family="Staphylinidae" genus="Pselaphogenius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="565" phylum="Arthropoda" rank="species" species="treskanus">
<emphasis id="B929EACAE47EFFC6FE555008FD20467B" box="[481,744,1044,1068]" italics="true" pageId="6" pageNumber="565">Pselaphogenius treskanus</emphasis>
</taxonomicName>
(Karaman, 1940). Scale bars correspond to 200 um.
</paragraph>
</caption>
<paragraph id="8BE236D8E47EFFC6FF735047FCB5471A" blockId="6.[151,1437,1114,1609]" pageId="6" pageNumber="565">
<emphasis id="B929EACAE47EFFC6FF735047FC794622" bold="true" box="[199,945,1114,1141]" pageId="6" pageNumber="565">
Measurements of
<typeStatus id="54E6887AE47EFFC6FE175047FDC24622" box="[419,522,1115,1141]" pageId="6" pageNumber="565">holotype</typeStatus>
(in mm)—W = width, L = length.
</emphasis>
Body, L—1.87; head, L—0.37, W—0.26; antenna, L—0.9; scapus, L—0.16, W—0.07; pedicel, L—0.07, W—0.06; antennomere III, L—0.06; antennomere IV, L—0.05; antennomere V, L—0.06; antennomere VI, L—0.06; antennomere VII, L—0.05; antennomere VIII, L—0.05; antennomere IX, L—0.07; antennomere X, L—0.07; antennomere XI, L—0.19, W—0.1; maxillary palpomere I, L—0.09; maxillary palpomere II, L—0.19; maxillary palpomere III, L—0.05; maxillary palpomere IV, L—0.36; pronotum, L—0.4, W—0.27; elytra, L—0.45, W—0.31; tergite IV, L—0.45, W—0.69; tergite IV (excluding paratergites) W—0.56; ventrite IV, L—0.39, W—0.75.
</paragraph>
<paragraph id="8BE236D8E47EFFC6FF73514BFC834727" blockId="6.[151,1437,1114,1609]" box="[199,843,1366,1393]" pageId="6" pageNumber="565">
<emphasis id="B929EACAE47EFFC6FF73514BFEED4726" bold="true" box="[199,293,1367,1393]" pageId="6" pageNumber="565">Female.</emphasis>
Similar to male, but with a smooth metaventrite.
</paragraph>
</subSubSection>
<subSubSection id="C3476553E47EFFC6FF735167FE9F478A" pageId="6" pageNumber="565" type="etymology">
<paragraph id="8BE236D8E47EFFC6FF735167FB9847C3" blockId="6.[151,1437,1114,1609]" box="[199,1104,1402,1429]" pageId="6" pageNumber="565">
<emphasis id="B929EACAE47EFFC6FF735167FE8647C2" bold="true" box="[199,334,1403,1429]" pageId="6" pageNumber="565">Etymology.</emphasis>
The new species is named after Mt. Tymfi, where it was discovered.
</paragraph>
<paragraph id="8BE236D8E47EFFC6FF735182FE9F478A" blockId="6.[151,1437,1114,1609]" pageId="6" pageNumber="565">
<emphasis id="B929EACAE47EFFC6FF735182FEE247EF" bold="true" box="[199,298,1438,1464]" pageId="6" pageNumber="565">Habitat.</emphasis>
Specimens were collected in dense leaf-litter accumulations from a mixed deciduous forest on Mt. Tymfi. (
<figureCitation id="13662A5DE47EFFC6FF4651DEFE81478A" box="[242,329,1474,1501]" captionStart="FIGURE 8" captionStartId="9.[151,250,1185,1209]" captionTargetBox="[344,1243,181,1161]" captionTargetId="figure-17@9.[344,1243,181,1161]" captionTargetPageId="9" captionText="FIGURE 8. Habitat photos of examined species. A) Type locality of A. taygetensis sp. n. in Kastoreion; B) type locality of A. tymficus sp. n.; in Mt Tymfi; C) leaf litter where A. euboicus sp. n. was collected from in Mt Dirfis, the type locality; D) habitat in Mt Vitsi, where Pselaphogenius treskanus (Karaman, 1940) was collected from." figureDoi="http://doi.org/10.5281/zenodo.8392315" httpUri="https://zenodo.org/record/8392315/files/figure.png" pageId="6" pageNumber="565">Fig. 8B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3476553E47EFFC6FF7351FAFCDB4457" box="[199,787,1510,1537]" pageId="6" pageNumber="565" type="distribution">
<paragraph id="8BE236D8E47EFFC6FF7351FAFCDB4457" blockId="6.[151,1437,1114,1609]" box="[199,787,1510,1537]" pageId="6" pageNumber="565">
<emphasis id="B929EACAE47EFFC6FF7351FAFEA94457" bold="true" box="[199,353,1510,1536]" pageId="6" pageNumber="565">Distribution.</emphasis>
<collectingCountry id="F34A7648E47EFFC6FED351FBFE7E4456" box="[359,438,1511,1537]" name="Greece" pageId="6" pageNumber="565">Greece</collectingCountry>
(so far endemic to Mt. Tymfi).
</paragraph>
</subSubSection>
<subSubSection id="C3476553E47EFFC6FF735217FCDC441E" pageId="6" pageNumber="565" type="discussion">
<paragraph id="8BE236D8E47EFFC6FF735217FCDC441E" blockId="6.[151,1437,1114,1609]" pageId="6" pageNumber="565">
<emphasis id="B929EACAE47EFFC6FF735217FEF34472" bold="true" box="[199,315,1547,1573]" pageId="6" pageNumber="565">Remarks.</emphasis>
The complex sclerites of the internal sac of aedeagus of
<taxonomicName id="4C5D4D5BE47EFFC6FC135217FBEA4472" authority="Davranoglou &amp; Baňař &amp; Kakiopoulos &amp; Balázs &amp; Avtzis &amp; Hlaváč, 2023" authorityName="Davranoglou &amp; Baňař &amp; Kakiopoulos &amp; Balázs &amp; Avtzis &amp; Hlaváč" authorityYear="2023" box="[935,1058,1547,1573]" class="Insecta" family="Staphylinidae" genus="Afropselaphus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="565" phylum="Arthropoda" rank="species" species="tymficus" status="sp. n.">
<emphasis id="B929EACAE47EFFC6FC135217FBEA4472" box="[935,1058,1547,1573]" italics="true" pageId="6" pageNumber="565">A. tymficus</emphasis>
</taxonomicName>
<emphasis id="B929EACAE47EFFC6FB9C5217FBAE4472" bold="true" box="[1064,1126,1547,1573]" pageId="6" pageNumber="565">
<taxonomicNameLabel id="A21A57B1E47EFFC6FB9C5217FBAE4472" box="[1064,1126,1547,1573]" pageId="6" pageNumber="565" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
are strongly autapomorphic and no close parallels can be found in Balkan congeners.
</paragraph>
</subSubSection>
</treatment>
</document>