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<mods:title id="5BD1381F22CBFD4161BBC005152A8B02">Systematics of the family Carditidae (Bivalvia: Archiheterodonta) in the Cenozoic of Argentina</mods:title>
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<subSubSection id="C3476516B81EB475FF687567FE32F81C" pageId="25" pageNumber="76" type="nomenclature">
<paragraph id="8BE2369DB81EB475FF687567FDF3F8FE" blockId="25.[151,643,1862,1922]" box="[151,643,1862,1888]" pageId="25" pageNumber="76">
<heading id="D0AA81F1B81EB475FF687567FDF3F8FE" bold="true" box="[151,643,1862,1888]" fontSize="11" level="1" pageId="25" pageNumber="76" reason="1">
<taxonomicName id="4C5D4D1EB81EB475FF687567FDF3F8FE" authority="(Feruglio, 1954)" baseAuthorityName="Feruglio" baseAuthorityYear="1954" box="[151,643,1862,1888]" class="Bivalvia" family="Carditidae" genus="Scalaricardita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carditoida" pageId="25" pageNumber="76" phylum="Mollusca" rank="species" species="laciarina">
<emphasis id="B929EA8FB81EB475FF687567FDF3F8FE" bold="true" box="[151,643,1862,1888]" pageId="25" pageNumber="76">
<emphasis id="B929EA8FB81EB475FF687567FEC6F8FE" bold="true" box="[151,438,1862,1888]" italics="true" pageId="25" pageNumber="76">Scalaricardita laciarina</emphasis>
(
<bibRefCitation id="EFCC4B6CB81EB475FE3A7567FD0BF8FE" author="Feruglio" box="[453,635,1862,1888]" pageId="25" pageNumber="76" refString="Feruglio, E. (1954) Alcune nuove forme di brachiopodi e molluschi del Terziario e Cretaceo superiore della Patagonia. Pubblicazioni dell' Istituto Geologico della Universita di Torino, 3, 1 - 45." type="journal article" year="1954">Feruglio, 1954</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BE2369DB81EB475FF687548FE32F81C" blockId="25.[151,643,1862,1922]" box="[151,322,1897,1922]" pageId="25" pageNumber="76">Figure 6.1416</paragraph>
</subSubSection>
<subSubSection id="C3476516B81EB475FF687590FB2AF878" pageId="25" pageNumber="76" type="description">
<paragraph id="8BE2369DB81EB475FF687590FDE9F859" blockId="25.[151,1114,1969,2022]" box="[151,665,1969,1991]" pageId="25" pageNumber="76">
v1933
<taxonomicName id="4C5D4D1EB81EB475FF237590FE2EF859" authority="Pérez &amp; Del Río, 2017" authorityName="Pérez &amp; Del Río" authorityYear="2017" box="[220,350,1969,1991]" class="Bivalvia" family="Carditidae" genus="Venericardia" higherTaxonomySource="CoL" kingdom="Animalia" order="Carditida" pageId="25" pageNumber="76" phylum="Mollusca" rank="genus" status="n.f.">
<emphasis id="B929EA8FB81EB475FF237590FE2EF859" box="[220,350,1969,1991]" italics="true" pageId="25" pageNumber="76">Venericardia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A57F4B81EB475FE9B7590FEF7F859" box="[356,391,1969,1991]" pageId="25" pageNumber="76" rank="form">n.f.</taxonomicNameLabel>
Feruglio, p. 96, figs. 811.
</paragraph>
<paragraph id="8BE2369DB81EB475FF6875F1FB2AF878" blockId="25.[151,1114,1969,2022]" box="[151,1114,2000,2022]" pageId="25" pageNumber="76">
*1954
<taxonomicName id="4C5D4D1EB81EB475FF2375F1FD50F878" authority="Feruglio" authorityName="Feruglio" box="[220,544,2000,2022]" class="Bivalvia" family="Carditidae" genus="Venericardia" higherTaxonomySource="CoL" kingdom="Animalia" order="Carditida" pageId="25" pageNumber="76" phylum="Mollusca" rank="species" species="laciarina">
<emphasis id="B929EA8FB81EB475FF2375F1FEB1F878" box="[220,449,2000,2022]" italics="true" pageId="25" pageNumber="76">Venericardia laciarina</emphasis>
Feruglio
</taxonomicName>
, p. 1011, pl. 1, figs 5ab, 6ab, pl. 2, figs. 14a,b, 56.
</paragraph>
</subSubSection>
<subSubSection id="C3476516B81DB476FF6872B6FCA7FF2E" box="[151,983,151,176]" pageId="26" pageNumber="77" type="materials_examined">
<paragraph id="8BE2369DB81DB476FF6872B6FCA7FF2E" blockId="26.[151,1437,151,1400]" box="[151,983,151,176]" pageId="26" pageNumber="77">
<emphasis id="B929EA8FB81DB476FF6872B6FE14FF2E" bold="true" box="[151,356,151,176]" pageId="26" pageNumber="77">Material studied.</emphasis>
MACN-Pi 2213 (nine left valves and five right valves).
</paragraph>
</subSubSection>
<subSubSection id="C3476516B81DB476FF38729DFBB0FF66" pageId="26" pageNumber="77" type="diagnosis">
<paragraph id="8BE2369DB81DB476FF38729DFBB0FF66" blockId="26.[151,1437,151,1400]" pageId="26" pageNumber="77">
<emphasis id="B929EA8FB81DB476FF38729DFE31FF4B" bold="true" box="[199,321,188,213]" pageId="26" pageNumber="77">Diagnosis.</emphasis>
Shell with subtriangular outline. Umbo subcentrally. Lunule faintly defined. Left anterior tooth large. External sculpture of 25 to 27 low radial ribs more conspicuous towards anterior margin.
</paragraph>
</subSubSection>
<subSubSection id="C3476516B81DB476FF387325FF9EFD6E" pageId="26" pageNumber="77" type="description">
<paragraph id="8BE2369DB81DB476FF387325FD9CFE16" blockId="26.[151,1437,151,1400]" pageId="26" pageNumber="77">
<emphasis id="B929EA8FB81DB476FF387325FE29FE83" bold="true" box="[199,345,260,285]" pageId="26" pageNumber="77">Description.</emphasis>
Shell small-sized (maximum size of
<quantity id="4CA59B78B81DB476FD067325FC39FE82" box="[761,841,260,284]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="26" pageNumber="77" unit="mm" value="17.0">17 mm</quantity>
) with subtriangular outline, posterior margin slightly truncated forming a gentle angle as it meets the ventral one, dorsal margin straight and elongate sloping pronouncedly from beaks to posterior end, ventral and anterior margins rounded. Umbo placed near centre of the valves. Lunule small, narrow, flat, shallow and weak.
</paragraph>
<paragraph id="8BE2369DB81DB476FF3873B5FBD9FDA3" blockId="26.[151,1437,151,1400]" pageId="26" pageNumber="77">Right hinge with a concave ventral edge; anterior tooth very small, elongate, short, inclined forward, in contact with lunular margin; middle tooth triangular, small, inclined posteriorly base broad and with straight anterior and slightly posterior sides, higher towards base on lateral view; posterior tooth very thin and straight, extending to the apex of middle tooth. Left hinge with slightly curved ventral edge; anterior tooth triangular, elongate, high, inclined forward with straight anterior and posterior sides; posterior tooth elongate, wide and straight.</paragraph>
<paragraph id="8BE2369DB81DB476FF387066FF9EFD6E" blockId="26.[151,1437,151,1400]" pageId="26" pageNumber="77">External sculpture of 25 to 27 entire radial ribs, low and wide, slightly wider towards ventral marginwith subelliptic transverse section covered with subrectangular and small closely-spaced and barely noticeable nodes; shallow and narrow intercostal spaces with subtriangular transverse section. Pallial line placed about a quarter of total valve height. Inner ventral margin strongly crenulated, crenulations subrectangular, truncated, covering entire margin.</paragraph>
</subSubSection>
<subSubSection id="C3476516B81DB476FF3870DDFB54FC59" pageId="26" pageNumber="77" type="discussion">
<paragraph id="8BE2369DB81DB476FF3870DDFB47FCC2" blockId="26.[151,1437,151,1400]" pageId="26" pageNumber="77">
<emphasis id="B929EA8FB81DB476FF3870DDFE4BFC8B" bold="true" box="[199,315,764,789]" pageId="26" pageNumber="77">Remarks.</emphasis>
<bibRefCitation id="EFCC4B6CB81DB476FEBD70DDFE9BFC8A" author="Feruglio" box="[322,491,764,789]" pageId="26" pageNumber="77" refString="Feruglio, E. (1933) I terrazzi marini della Patagonia. Giornali di Geologia. Annali del Museo Geologico di Bologna, VIII bis, 1 - 288." type="book chapter" year="1933">Feruglio (1933</bibRefCitation>
,
<bibRefCitation id="EFCC4B6CB81DB476FE0870DDFD40FC8B" author="Feruglio" box="[503,560,764,789]" pageId="26" pageNumber="77" refString="Feruglio, E. (1954) Alcune nuove forme di brachiopodi e molluschi del Terziario e Cretaceo superiore della Patagonia. Pubblicazioni dell' Istituto Geologico della Universita di Torino, 3, 1 - 45." type="journal article" year="1954">1954</bibRefCitation>
) described and illustrated specimens of this carditid recorderd in Cerro Laciar (lower Pliocene,
<collectingRegion id="4999F87FB81DB476FE9F7101FD22FCA6" box="[352,594,799,824]" country="Argentina" name="Santa Cruz" pageId="26" pageNumber="77">Santa Cruz Province</collectingRegion>
) but that
<typeStatus id="54E6883FB81DB476FD327101FD8DFCA6" box="[717,765,800,824]" pageId="26" pageNumber="77">type</typeStatus>
material was not available to the authors. However, the Feruglio collection housed at the MACN hosts 12 valves belonging in
<taxonomicName id="4C5D4D1EB81DB476FC527164FB40FCC3" box="[941,1072,836,861]" class="Bivalvia" family="Carditidae" genus="Venericardia" higherTaxonomySource="CoL" kingdom="Animalia" order="Carditida" pageId="26" pageNumber="77" phylum="Mollusca" rank="species" species="laciarina">
<emphasis id="B929EA8FB81DB476FC527164FB40FCC3" box="[941,1072,836,861]" italics="true" pageId="26" pageNumber="77">S. laciarina</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BE2369DB81DB476FF387146FB54FC59" blockId="26.[151,1437,151,1400]" pageId="26" pageNumber="77">
<taxonomicName id="4C5D4D1EB81DB476FF387146FEA6FC1E" baseAuthorityName="Feruglio" baseAuthorityYear="1954" box="[199,470,871,896]" class="Bivalvia" family="Carditidae" genus="Scalaricardita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carditoida" pageId="26" pageNumber="77" phylum="Mollusca" rank="species" species="laciarina">
<emphasis id="B929EA8FB81DB476FF387146FEA6FC1E" box="[199,470,871,896]" italics="true" pageId="26" pageNumber="77">Scalaricardita laciarina</emphasis>
</taxonomicName>
is assigned to
<taxonomicName id="4C5D4D1EB81DB476FD827146FC6EFC1E" box="[637,798,871,896]" class="Bivalvia" family="Carditidae" genus="Scalaricardita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carditoida" pageId="26" pageNumber="77" phylum="Mollusca" rank="genus">
<emphasis id="B929EA8FB81DB476FD827146FC6EFC1E" box="[637,798,871,896]" italics="true" pageId="26" pageNumber="77">Scalaricardita</emphasis>
</taxonomicName>
because of its subtriangular shell with slightly truncated posterior margin, a triangular and wide right middle tooth, 23 to 27 low and wide radial ribs covered by subrectangular, elongate and closely-spaced nodes and narrow intercostal spaces.
</paragraph>
</subSubSection>
<subSubSection id="C3476516B81DB476FF3871F5FE77FAE6" pageId="26" pageNumber="77" type="description">
<paragraph id="8BE2369DB81DB476FF3871F5FDDFFBAA" blockId="26.[151,1437,151,1400]" pageId="26" pageNumber="77">
<taxonomicName id="4C5D4D1EB81DB476FF3871F5FEABFC73" baseAuthorityName="Feruglio" baseAuthorityYear="1954" box="[199,475,980,1005]" class="Bivalvia" family="Carditidae" genus="Scalaricardita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carditoida" pageId="26" pageNumber="77" phylum="Mollusca" rank="species" species="laciarina">
<emphasis id="B929EA8FB81DB476FF3871F5FEABFC73" box="[199,475,980,1005]" italics="true" pageId="26" pageNumber="77">Scalaricardita laciarina</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="4C5D4D1EB81DB476FCDB71F4FCEDFC73" authorityName="J. de C. Sowerby" authorityYear="1825" box="[804,925,980,1005]" class="Bivalvia" family="Carditidae" genus="Venericardia" higherTaxonomySource="CoL" kingdom="Animalia" order="Carditida" pageId="26" pageNumber="77" phylum="Mollusca" rank="species" species="scalaris">
<emphasis id="B929EA8FB81DB476FCDB71F4FCEDFC73" box="[804,925,980,1005]" italics="true" pageId="26" pageNumber="77">S. scalaris</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFCC4B6CB81DB476FC4E71F5FB98FC72" author="Janssen" box="[945,1256,980,1005]" pageId="26" pageNumber="77" refString="Janssen, A. &amp; Moerdijk, P. (2004) Revision of Pliocene representatives of Cyclocardia (Bivalvia, Carditidae) from the North Sea Basin. Basteria, 68, 77 - 85." type="journal article" year="2004">Janssen &amp; Moerdijk 2004</bibRefCitation>
: figs. 4ab) by having a larger shell, subcentrally placed umbo, less-defined lunule, larger left anterior tooth, more pronounced anterior radial ribs and wider intercostal spaces.
</paragraph>
<paragraph id="8BE2369DB81DB476FF38761EFD91FBE3" blockId="26.[151,1437,151,1400]" pageId="26" pageNumber="77">
<taxonomicName id="4C5D4D1EB81DB476FF38761EFEA5FBC6" baseAuthorityName="Feruglio" baseAuthorityYear="1954" box="[199,469,1087,1112]" class="Bivalvia" family="Carditidae" genus="Scalaricardita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carditoida" pageId="26" pageNumber="77" phylum="Mollusca" rank="species" species="laciarina">
<emphasis id="B929EA8FB81DB476FF38761EFEA5FBC6" box="[199,469,1087,1112]" italics="true" pageId="26" pageNumber="77">Scalaricardita laciarina</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C5D4D1EB81DB476FD977661FD81FBC6" baseAuthorityName="Tate" baseAuthorityYear="1886" box="[616,753,1088,1112]" class="Bivalvia" family="Carditidae" genus="Scalaricardita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carditoida" pageId="26" pageNumber="77" phylum="Mollusca" rank="species" species="compacta">
<emphasis id="B929EA8FB81DB476FD977661FD81FBC6" box="[616,753,1088,1112]" italics="true" pageId="26" pageNumber="77">S. compacta</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFCC4B6CB81DB476FD007661FB77FBC6" author="Tate" box="[767,1031,1087,1112]" pageId="26" pageNumber="77" refString="Tate, R. (1886) The lamellibranchs of the older Tertiary of Australia. Part I. Transactions and Proceedings of the Royal Society of South Australia, 8, 96 - 158." type="journal article" year="1886">Tate 1886: pl. 2, fig. 13</bibRefCitation>
) by having more subtriangular shell with more prominent and subcentrally placed umbo.
</paragraph>
<paragraph id="8BE2369DB81DB476FF3876A6FB10FB5B" blockId="26.[151,1437,151,1400]" pageId="26" pageNumber="77">
<taxonomicName id="4C5D4D1EB81DB476FF3876A6FE96FB3E" box="[199,486,1159,1184]" class="Bivalvia" family="Carditidae" genus="Scalaricardita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carditoida" pageId="26" pageNumber="77" phylum="Mollusca" rank="species" species="laciariana">
<emphasis id="B929EA8FB81DB476FF3876A6FE96FB3E" box="[199,486,1159,1184]" italics="true" pageId="26" pageNumber="77">Scalaricardita laciariana</emphasis>
</taxonomicName>
can be separated from
<taxonomicName id="4C5D4D1EB81DB476FD0776A9FCDAFB3E" baseAuthorityName="Chapman &amp; Crespin" baseAuthorityYear="1928" box="[760,938,1159,1184]" class="Bivalvia" family="Carditidae" genus="Scalaricardita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carditoida" pageId="26" pageNumber="77" phylum="Mollusca" rank="species" species="subcompacta">
<emphasis id="B929EA8FB81DB476FD0776A9FCDAFB3E" box="[760,938,1159,1184]" italics="true" pageId="26" pageNumber="77">S. subcompacta</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFCC4B6CB81DB476FC4376A6FA27FB3E" author="Ludbrook" box="[956,1367,1159,1184]" pageId="26" pageNumber="77" refString="Ludbrook, N. H. (1954) The molluscan fauna of the Pliocene strata underlying the Adelaide Plains. Part II - Pelecypoda. Transactions of the Royal Society of South Australia, 78, 18 - 80." type="journal article" year="1954">Ludbrook 1954: pl. 2, figs. 5 and 8</bibRefCitation>
) by a more subtriangular shell with subcentrally placed umbo and larger right anterior tooth.
</paragraph>
<paragraph id="8BE2369DB81DB476FF3876EEFE0FFA93" blockId="26.[151,1437,151,1400]" pageId="26" pageNumber="77">
This species differs from
<taxonomicName id="4C5D4D1EB81DB476FE1276F1FD08FB76" authorityName="Bartrum &amp; Powell" authorityYear="1928" box="[493,632,1231,1256]" class="Bivalvia" family="Carditidae" genus="Pleuromeris" kingdom="Animalia" order="Carditoida" pageId="26" pageNumber="77" phylum="Mollusca" rank="species" species="miniscula">
<emphasis id="B929EA8FB81DB476FE1276F1FD08FB76" box="[493,632,1231,1256]" italics="true" pageId="26" pageNumber="77">S. miniscula</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFCC4B6CB81DB476FD7776F1FC8BFB76" author="Bartrum" box="[648,1019,1231,1256]" pageId="26" pageNumber="77" refString="Bartrum, J. A. &amp; Powell, A. W. (1928) Mollusca from Kaawa Creek Beds, West Coast, South of Waikato River. Transactions of the New Zealand Institute, 59, 139 - 162." type="journal article" year="1928">Bartrum &amp; Powell 1928: fig. 12</bibRefCitation>
) by having more subtriangular shell and more radial ribs.
</paragraph>
<paragraph id="8BE2369DB81DB476FF387736FEA5FACB" blockId="26.[151,1437,151,1400]" pageId="26" pageNumber="77">
<taxonomicName id="4C5D4D1EB81DB476FF387736FEA6FAAE" baseAuthorityName="Feruglio" baseAuthorityYear="1954" box="[199,470,1303,1328]" class="Bivalvia" family="Carditidae" genus="Scalaricardita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carditoida" pageId="26" pageNumber="77" phylum="Mollusca" rank="species" species="laciarina">
<emphasis id="B929EA8FB81DB476FF387736FEA6FAAE" box="[199,470,1303,1328]" italics="true" pageId="26" pageNumber="77">Scalaricardita laciarina</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C5D4D1EB81DB476FD947739FC68FAAE" baseAuthorityName="Ihering" baseAuthorityYear="1907" box="[619,792,1304,1328]" class="Bivalvia" family="Carditidae" genus="Scalaricardita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carditoida" pageId="26" pageNumber="77" phylum="Mollusca" rank="species" species="camaronesia">
<emphasis id="B929EA8FB81DB476FD947739FC68FAAE" box="[619,792,1304,1328]" italics="true" pageId="26" pageNumber="77">S. camaronesia</emphasis>
</taxonomicName>
by having a larger shell, less anteriorly placed umbo and narrower right middle tooth.
</paragraph>
<paragraph id="8BE2369DB81DB476FF68777EFE77FAE6" blockId="26.[151,1437,151,1400]" box="[151,263,1375,1400]" pageId="26" pageNumber="77">
[
<figureCitation id="13662A18B81DB476FF62777EFF8FFAE6" box="[157,255,1375,1400]" captionStart="FIGURE 3" captionStartId="11.[151,250,1830,1852]" captionTargetBox="[165,1403,448,1796]" captionTargetId="figure@11.[151,1436,435,1809]" captionTargetPageId="11" captionText="FIGURE 3. (1 3) Cyclocardia cannada (Ihering, 1907). 1, LV, Holotype MACN-Pi 344, from Cañadón de los Artilleros. 2, LV, MACN-Pi 344, from Cañadón de los Artilleros. 3, RV, MACN-Pi 348, from Yegua Quemada. (4 6) Cyclocardia compressa (Reeve, 1843). 4, LV, MACN-Pi 363, from Puerto Deseado. 5, LV, MACN-Pi 363, from Puerto Deseado. 6, RV, MACN-Pi 363, from Puerto Deseado. (7 9, 11) Cyclocardia dalek n. sp. from Playa La Mina. 7, LV, Holotype MACN-Pi 5770. 8, LV, Holotype MACN-Pi 5770. 9, RV, Paratype MACN-Pi 5771. 11, RV, Paratype MACN-Pi 5771. (10, 12 13) Cyclocardia nortensis (del Río, 1986), from Punta Norte. 10, LV, Holotype CPBA 13242. 12, RV, Paratype CPBA 13247. 13, RV, Paratype CPBA 13247. Scale bar represents 10 mm." httpUri="https://zenodo.org/record/1035564/files/figure.png" pageId="26" pageNumber="77">Figure 3</figureCitation>
]
</paragraph>
</subSubSection>
</treatment>
</document>