treatments-xml/data/03/BF/87/03BF87BA0C5F756BB7C9A9CC328BFE63.xml

<document id="36058FE0ACFBABDF99DA3A78F12E9F4A" ID="10.11646/zootaxa.4263.3.2SLASH11006" ID-CLB-Dataset="33074" ID-DOI="10.11646/zootaxa.4263.3.2" ID-GBIF-Dataset="e98df155-1049-42c3-b779-f7272da2cc2f" ID-ISSN="1175-5326" ID-Zenodo-Dep="573630" ID-ZooBank="093CCBCF-E709-4C9D-8697-BF18BFC635F1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1494401628858" checkinUser="plazi" docAuthor="Fontoura, Paulo, Rubal, Marcos &amp; Veiga, Puri" docDate="2017" docId="03BF87BA0C5F756BB7C9A9CC328BFE63" docLanguage="en" docName="zootaxa.4263.3.2SLASH11006.pdf" docOrigin="Zootaxa 4263 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ramazzottius littoreus Fontoura, Rubal &amp; Veiga, 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="458" masterDocId="FF86FFC20C5D7563B75EA913300AFFEB" masterDocTitle="Two new species of Tardigrada (Eutardigrada: Ramazzottiidae, Macrobiotidae) from the supralittoral zone of the Atlantic Iberian Peninsula rocky shores" masterLastPageNumber="466" masterPageNumber="450" pageNumber="452" updateTime="1698439436455" updateUser="ExternalLinkService">
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<mods:title id="2BD9B556FD0B65D4003413CAF0A959C8">Two new species of Tardigrada (Eutardigrada: Ramazzottiidae, Macrobiotidae) from the supralittoral zone of the Atlantic Iberian Peninsula rocky shores</mods:title>
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<mods:namePart id="BA6CE7DD5278FC37EE578AA6CD6CB0EA">Fontoura, Paulo</mods:namePart>
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<mods:namePart id="B41DE9EDA6C2BFC1ED8012C911DBCBA0">Rubal, Marcos</mods:namePart>
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<mods:namePart id="D5AA75CDBE8016F14425816AC966741B">Veiga, Puri</mods:namePart>
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<mods:date id="5B2872E5FEF98FB933340B2F74C7CC14">2017</mods:date>
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<treatment id="03BF87BA0C5F756BB7C9A9CC328BFE63" ID-DOI="http://doi.org/10.5281/zenodo.6015868" ID-GBIF-Taxon="130509336" ID-Zenodo-Dep="6015868" LSID="urn:lsid:plazi:treatment:03BF87BA0C5F756BB7C9A9CC328BFE63" httpUri="http://treatment.plazi.org/id/03BF87BA0C5F756BB7C9A9CC328BFE63" lastPageId="8" lastPageNumber="458" pageId="2" pageNumber="452">
<subSubSection id="C30C65270C5F7561B7C9A9CC322EFEF0" pageId="2" pageNumber="452" type="nomenclature">
<paragraph id="8BA936AC0C5F7561B7C9A9CC3202FF11" blockId="2.[151,548,223,283]" box="[151,520,223,250]" pageId="2" pageNumber="452">
<heading id="D0E181C00C5F7561B7C9A9CC3202FF11" bold="true" box="[151,520,223,250]" fontSize="11" level="1" pageId="2" pageNumber="452" reason="1">
<emphasis id="B962EABE0C5F7561B7C9A9CC3202FF11" bold="true" box="[151,520,223,250]" pageId="2" pageNumber="452">
<taxonomicName id="4C164D2F0C5F7561B7C9A9CC31A8FF12" box="[151,418,223,249]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="2" pageNumber="452" phylum="Tardigrada" rank="species" species="littoreus" status="sp. nov.">
<emphasis id="B962EABE0C5F7561B7C9A9CC31A8FF12" bold="true" box="[151,418,223,249]" italics="true" pageId="2" pageNumber="452">Ramazzottius littoreus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25157C50C5F7561B6F4A9F33202FF11" box="[426,520,224,250]" pageId="2" pageNumber="452" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA936AC0C5F7561B7C9A811322EFEF0" blockId="2.[151,548,223,283]" box="[151,548,258,283]" pageId="2" pageNumber="452">
(
<figureCitation id="132D2A290C5F7561B7C0A8113103FEF0" box="[158,265,258,283]" captionStart="FIGURE 1" captionStartId="2.[151,250,1782,1804]" captionTargetBox="[151,1436,554,1758]" captionTargetId="figure@2.[151,1436,554,1758]" captionTargetPageId="2" captionText="FIGURE 1. Ramazzottius littoreus sp. nov. holotype: habitus (A) Scale bar = 50 µm. Sculpture on dorsal cuticle of a paratype; arrows show dorsal muscle attachment points and asterisk shows elliptical cephalic organs (B). Legs and claws IV of the holotype; arrows show lateral gibbosities on hind legs, and arrowhead shows cuticle ornamentation (C). Scale bar (B, C) = 10 µm." httpUri="https://zenodo.org/record/573631/files/figure.png" pageId="2" pageNumber="452">Figures 1</figureCitation>
, 
<figureCitation id="132D2A290C5F7561B649A810312FFEF0" box="[279,293,259,283]" captionStart="FIGURE 2" captionStartId="4.[151,250,1689,1711]" captionTargetBox="[441,1155,608,1658]" captionTargetId="figure@4.[418,1168,589,1668]" captionTargetPageId="4" captionText="FIGURE 2. Schematic representation of muscle attachment points in Ramazzottius littoreus sp. nov. (A: ventral view; B: dorsal view) and Macrobiotus halophilus sp. nov. (C: ventral view; D: dorsal view). Numerals (I – XV) identify transversal rows." httpUri="https://zenodo.org/record/573632/files/figure.png" pageId="2" pageNumber="452">2</figureCitation>
A, B, 3, 4; 
<tableCitation id="C69403170C5F7561B6FFA81131F4FEF0" box="[417,510,258,283]" captionStart="TABLE 1" captionStartId="3.[151,239,440,463]" captionText="TABLE 1. Measurements (in µm) and pt values of selected morphological structures for specimens of Ramazzottius littoreus sp. nov. (N—number of specimens / structures measured; SD—Standard deviation; Range refers to the smallest and largest measured structure). Parameters of Thorpe normalisation are also indicated: a * = Y-intercept of the regression line of the Thorpe normalised structure versus buccal tube length; b = allometric exponent (values in bold indicate allometry)." pageId="2" pageNumber="452">Tables 1</tableCitation>
, 
<tableCitation id="C69403170C5F7561B553A8103216FEF0" box="[525,540,259,283]" captionStart="TABLE 2" captionStartId="7.[151,239,152,175]" captionText="TABLE 2. Measurements (in µm) of selected morphological structures for eggs of Ramazzottius littoreus sp. nov. (N—number of eggs / structures measured; SD—Standard deviation; Range refers to the smallest and largest measured structure)." pageId="2" pageNumber="452">2</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30C65270C5F7561B7C9A8593435FE18" pageId="2" pageNumber="452" type="materials_examined">
<paragraph id="8BA936AC0C5F7561B7C9A8593494FE6C" blockId="2.[151,1436,330,499]" pageId="2" pageNumber="452">
<emphasis id="B962EABE0C5F7561B7C9A859318BFE88" bold="true" box="[151,385,330,355]" pageId="2" pageNumber="452">Material examined.</emphasis>
Holotype (slide C. 
<date id="FFA8106C0C5F7561B50AA85932ADFE88" box="[596,679,330,355]" pageId="2" pageNumber="452" value="1960-07">VII-60</date>
); 62 paratypes (49 animals, slides C. 
<date id="FFA8106C0C5F7561B31FA859349AFE88" box="[1089,1168,330,355]" pageId="2" pageNumber="452" value="1960-07">VII-60</date>
to C. 
<date id="FFA8106C0C5F7561B395A8583511FE88" box="[1227,1307,331,355]" pageId="2" pageNumber="452" value="1974-07">VII-74</date>
and C. 
<date id="FFA8106C0C5F7561B237A85830B9FE6C" pageId="2" pageNumber="452" value="1976-07">VII- 76</date>
to C. 
<date id="FFA8106C0C5F7561B7B0A87C314AFE6C" box="[238,320,367,391]" pageId="2" pageNumber="452" value="1978-07">VII-78</date>
, and 13 eggs, of which three were embryonate, slides C. 
<date id="FFA8106C0C5F7561B4EAA87C3409FE6C" box="[948,1027,367,391]" pageId="2" pageNumber="452" value="1971-07">VII-71</date>
to C. 
<date id="FFA8106C0C5F7561B360A87C349BFE6C" box="[1086,1169,367,391]" pageId="2" pageNumber="452" value="1978-07">VII-78</date>
).
</paragraph>
<paragraph id="8BA936AC0C5F7561B799A8813368FE24" blockId="2.[151,1436,330,499]" pageId="2" pageNumber="452">
<materialsCitation id="3B7E3CF10C5F7561B799A8813368FE24" ID-GBIF-Occurrence="1502353012" collectorName="P. Fontoura" country="Portugal" county="University" location="Department" municipality="Sciences" pageId="2" pageNumber="452" specimenCount="3" stateProvince="Porto" typeStatus="holotype">
<emphasis id="B962EABE0C5F7561B799A8813184FE40" bold="true" box="[199,398,402,427]" pageId="2" pageNumber="452">
<typeStatus id="54AD880E0C5F7561B799A8813108FE40" box="[199,258,402,427]" pageId="2" pageNumber="452">Type</typeStatus>
repository.
</emphasis>
<typeStatus id="54AD880E0C5F7561B6C5A8813209FE40" box="[411,515,402,427]" pageId="2" pageNumber="452" type="holotype">Holotype</typeStatus>
and 
<typeStatus id="54AD880E0C5F7561B51BA88032BAFE40" box="[581,688,403,427]" pageId="2" pageNumber="452" type="paratype">paratypes</typeStatus>
are deposited in the 
<location id="8EC960770C5F7561B4EBA8803433FE41" LSID="urn:lsid:plazi:treatment:03BF87BA0C5F756BB7C9A9CC328BFE63:8EC960770C5F7561B4EBA8803433FE41" box="[949,1081,403,426]" country="Portugal" county="University" municipality="Sciences" name="Department" pageId="2" pageNumber="452" stateProvince="Porto">Department</location>
of 
<location id="8EC960770C5F7561B334A88134CEFE40" LSID="urn:lsid:plazi:treatment:03BF87BA0C5F756BB7C9A9CC328BFE63:8EC960770C5F7561B334A88134CEFE40" box="[1130,1220,402,427]" country="Portugal" county="University" municipality="Sciences" name="Biology" pageId="2" pageNumber="452" stateProvince="Porto">Biology</location>
of the 
<location id="8EC960770C5F7561B27AA8813572FE40" LSID="urn:lsid:plazi:treatment:03BF87BA0C5F756BB7C9A9CC328BFE63:8EC960770C5F7561B27AA8813572FE40" box="[1316,1400,402,427]" country="Portugal" county="University" municipality="Sciences" name="Faculty" pageId="2" pageNumber="452" stateProvince="Porto">Faculty</location>
of 
<collectingMunicipality id="6BCDACD60C5F7561B7C9A8A530F6FE24" box="[151,252,438,463]" pageId="2" pageNumber="452">Sciences</collectingMunicipality>
, 
<collectingCounty id="62C84E200C5F7561B659A8A53177FE24" box="[263,381,438,463]" pageId="2" pageNumber="452">University</collectingCounty>
of 
<collectingRegion id="49D2F84E0C5F7561B6FCA8A431E8FE25" box="[418,482,439,462]" country="Portugal" name="Porto" pageId="2" pageNumber="452">Porto</collectingRegion>
, 
<collectingCountry id="F301763C0C5F7561B6B3A8A53246FE24" box="[493,588,438,463]" name="Portugal" pageId="2" pageNumber="452">Portugal</collectingCountry>
(collection 
<collectorName id="26E3537A0C5F7561B58CA8A4335DFE24" box="[722,855,439,463]" pageId="2" pageNumber="452">P. Fontoura</collectorName>
).
</materialsCitation>
</paragraph>
<paragraph id="8BA936AC0C5F7561B799A8C93435FE18" blockId="2.[151,1436,330,499]" box="[199,1087,474,499]" pageId="2" pageNumber="452">
<emphasis id="B962EABE0C5F7561B799A8C9316FFE18" bold="true" box="[199,357,474,499]" pageId="2" pageNumber="452">Type locality.</emphasis>
43°41'37.77''N; 
<materialsCitation id="3B7E3CF10C5F7561B540A8C93436FE18" ID-GBIF-Occurrence="1502353011" box="[542,1084,474,499]" country="Spain" location="San Cibrao" longitude="-7.4370112" municipality="Lugo" pageId="2" pageNumber="452" specimenCount="1" stateProvince="Galicia">
<geoCoordinate id="EE22506B0C5F7561B540A8C932B3FE18" box="[542,697,474,499]" direction="west" orientation="longitude" pageId="2" pageNumber="452" precision="1" value="-7.4370112">7°26'13.24''W</geoCoordinate>
, 
<location id="8EC960770C5F7561B59BA8C8334FFE18" LSID="urn:lsid:plazi:treatment:03BF87BA0C5F756BB7C9A9CC328BFE63:8EC960770C5F7561B59BA8C8334FFE18" box="[709,837,474,499]" country="Spain" longitude="-7.4370112" municipality="Lugo" name="San Cibrao" pageId="2" pageNumber="452" stateProvince="Galicia">San Cibrao</location>
, 
<collectingMunicipality id="6BCDACD60C5F7561B40EA8C83384FE18" box="[848,910,475,499]" pageId="2" pageNumber="452">Lugo</collectingMunicipality>
, 
<collectingRegion id="49D2F84E0C5F7561B4C7A8C933E7FE18" box="[921,1005,474,499]" country="Spain" name="Galicia" pageId="2" pageNumber="452">Galicia</collectingRegion>
, 
<collectingCountry id="F301763C0C5F7561B4A6A8C93436FE18" box="[1016,1084,474,499]" name="Spain" pageId="2" pageNumber="452">Spain</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF6966240C5F7561B7C9AFE530CAF883" httpUri="https://zenodo.org/record/573631/files/figure.png" pageId="2" pageNumber="452" targetBox="[151,1436,554,1758]" targetPageId="2">
<paragraph id="8BA936AC0C5F7561B7C9AFE530CAF883" blockId="2.[151,1436,1782,1896]" pageId="2" pageNumber="452">
<emphasis id="B962EABE0C5F7561B7C9AFE53119F8E0" bold="true" box="[151,275,1782,1804]" pageId="2" pageNumber="452">FIGURE 1.</emphasis>
<taxonomicName id="4C164D2F0C5F7561B647AFE431F0F8E7" box="[281,506,1782,1804]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="2" pageNumber="452" phylum="Tardigrada" rank="species" species="littoreus" status="sp. nov.">
<emphasis id="B962EABE0C5F7561B647AFE431F0F8E7" box="[281,506,1782,1804]" italics="true" pageId="2" pageNumber="452">Ramazzottius littoreus</emphasis>
</taxonomicName>
<emphasis id="B962EABE0C5F7561B55EAFE5325AF8E0" bold="true" box="[512,592,1782,1803]" pageId="2" pageNumber="452">
<taxonomicNameLabel id="A25157C50C5F7561B55EAFE5325AF8E0" box="[512,592,1782,1803]" pageId="2" pageNumber="452" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
holotype: habitus (A) Scale bar = 50 µm. Sculpture on dorsal cuticle of a paratype; arrows show dorsal muscle attachment points and asterisk shows elliptical cephalic organs (B). Legs and claws IV of the holotype; arrows show lateral gibbosities on hind legs, and arrowhead shows cuticle ornamentation (C). Scale bar (B, C) = 10 µm.
</paragraph>
</caption>
<subSubSection id="C30C65270C5E7564B799A9843497FC17" lastPageId="7" lastPageNumber="457" pageId="3" pageNumber="453" type="diagnosis">
<paragraph id="8BA936AC0C5E7560B799A9843127FE63" blockId="3.[151,1436,151,392]" pageId="3" pageNumber="453">
<emphasis id="B962EABE0C5E7560B799A98431ABFF5B" bold="true" box="[199,417,151,176]" pageId="3" pageNumber="453">Specific diagnosis.</emphasis>
<taxonomicName id="4C164D2F0C5E7560B6F9A98B3236FF44" box="[423,572,152,175]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="3" pageNumber="453" phylum="Tardigrada" rank="genus">
<emphasis id="B962EABE0C5E7560B6F9A98B3236FF44" box="[423,572,152,175]" italics="true" pageId="3" pageNumber="453">Ramazzottius</emphasis>
</taxonomicName>
with granulated cuticle. Cuticular granules also present on legs. Eyes always absent. Elliptical cephalic organs present. Five longitudinal and nine transversal pinkish/brownish bands. Lateral gibbosities on hind legs. Claws of 
<taxonomicName id="4C164D2F0C5E7560B540A9F332B9FF1C" box="[542,691,224,247]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="3" pageNumber="453" phylum="Tardigrada" rank="genus">
<emphasis id="B962EABE0C5E7560B540A9F332B9FF1C" box="[542,691,224,247]" italics="true" pageId="3" pageNumber="453">Ramazzottius</emphasis>
</taxonomicName>
<typeStatus id="54AD880E0C5E7560B5E5A9F332E1FF13" box="[699,747,224,248]" pageId="3" pageNumber="453">type</typeStatus>
with accessory points on main branches and lunules on hind legs. Posterior part of mouth cavity with four dorsal and four ventral teeth. Anterior portion of the buccal tube bent ventrally. Pharyngeal bulb with two macroplacoids, first macroplacoid slightly longer than the second (1&gt;2). Free laid eggs, with cone-shaped processes of variable size with reticulated bulbous bases. Surface between processes finely dotted.
</paragraph>
<caption id="DF6966240C5E7560B7C9A8AB3162FDA4" pageId="3" pageNumber="453">
<paragraph id="8BA936AC0C5E7560B7C9A8AB3162FDA4" blockId="3.[151,1436,440,591]" pageId="3" pageNumber="453">
<emphasis id="B962EABE0C5E7560B7C9A8AB3107FE24" bold="true" box="[151,269,440,463]" pageId="3" pageNumber="453">TABLE 1.</emphasis>
Measurements (in µm) and 
<emphasis id="B962EABE0C5E7560B514A8AB3255FE24" box="[586,607,440,463]" italics="true" pageId="3" pageNumber="453">pt</emphasis>
values of selected morphological structures for specimens of 
<taxonomicName id="4C164D2F0C5E7560B250A8AB30E5FE04" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="3" pageNumber="453" phylum="Tardigrada" rank="species" species="littoreus" status="sp. nov.">
<emphasis id="B962EABE0C5E7560B250A8AB30E5FE04" italics="true" pageId="3" pageNumber="453">Ramazzottius littoreus</emphasis>
</taxonomicName>
<emphasis id="B962EABE0C5E7560B7A8A8CB3147FE04" bold="true" box="[246,333,472,495]" pageId="3" pageNumber="453">
<taxonomicNameLabel id="A25157C50C5E7560B7A8A8CB3147FE04" box="[246,333,472,495]" pageId="3" pageNumber="453" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(N—number of specimens/structures measured; SD—Standard deviation; Range refers to the smallest and largest measured structure). Parameters of Thorpe normalisation are also indicated: 
<emphasis id="B962EABE0C5E7560B3D0A8EB34A3FDE4" box="[1166,1193,504,527]" italics="true" pageId="3" pageNumber="453">a*</emphasis>
= Y-intercept of the regression line of the Thorpe normalised structure versus buccal tube length; 
<emphasis id="B962EABE0C5E7560B4B4AB0433FDFDC4" box="[1002,1015,535,559]" italics="true" pageId="3" pageNumber="453">b</emphasis>
= allometric exponent (values in bold indicate allometry).
</paragraph>
</caption>
<paragraph id="8BA936AC0C5E7567B7C1AB7932B6FE1F" blockId="3.[159,1217,618,640]" lastBlockId="4.[151,1437,151,501]" lastPageId="4" lastPageNumber="454" pageId="3" pageNumber="453">
CHARACTERS 
<typeStatus id="54AD880E0C5E7560B6BCAB79324AFD6B" box="[482,576,618,640]" pageId="3" pageNumber="453" type="holotype">Holotype</typeStatus>
Mean ± SD (Range); (N) Mean ± SD (Range); (N) 
<emphasis id="B962EABE0C597567B799A9843218FF5B" bold="true" box="[199,530,151,176]" pageId="4" pageNumber="454">
Description of the 
<typeStatus id="54AD880E0C597567B6FCA9843207FF5B" box="[418,525,151,176]" pageId="4" pageNumber="454" type="holotype">holotype</typeStatus>
:
</emphasis>
Body length 397 µm (table 1). Before being mounted on a slide five longitudinal and nine transversal pinkish/brownish bands separated by unpigmented areas are visible. Small and sparse clumps of dark pigment are present (
<figureCitation id="132D2A290C597567B6ADA9CC3230FF13" box="[499,570,223,248]" captionStart="FIGURE 1" captionStartId="2.[151,250,1782,1804]" captionTargetBox="[151,1436,554,1758]" captionTargetId="figure@2.[151,1436,554,1758]" captionTargetPageId="2" captionText="FIGURE 1. Ramazzottius littoreus sp. nov. holotype: habitus (A) Scale bar = 50 µm. Sculpture on dorsal cuticle of a paratype; arrows show dorsal muscle attachment points and asterisk shows elliptical cephalic organs (B). Legs and claws IV of the holotype; arrows show lateral gibbosities on hind legs, and arrowhead shows cuticle ornamentation (C). Scale bar (B, C) = 10 µm." httpUri="https://zenodo.org/record/573631/files/figure.png" pageId="4" pageNumber="454">Figs 1</figureCitation>
A, C). Eyes absent. Elliptical cephalic organs present (
<figureCitation id="132D2A290C597567B397A9CC3507FF13" box="[1225,1293,223,248]" captionStart="FIGURE 1" captionStartId="2.[151,250,1782,1804]" captionTargetBox="[151,1436,554,1758]" captionTargetId="figure@2.[151,1436,554,1758]" captionTargetPageId="2" captionText="FIGURE 1. Ramazzottius littoreus sp. nov. holotype: habitus (A) Scale bar = 50 µm. Sculpture on dorsal cuticle of a paratype; arrows show dorsal muscle attachment points and asterisk shows elliptical cephalic organs (B). Legs and claws IV of the holotype; arrows show lateral gibbosities on hind legs, and arrowhead shows cuticle ornamentation (C). Scale bar (B, C) = 10 µm." httpUri="https://zenodo.org/record/573631/files/figure.png" pageId="4" pageNumber="454">Fig. 1</figureCitation>
B, asterisk). Cuticular ornamentation consisting of granules (polygonal tubercles) extending dorso-laterally from the head to the caudal region; where it is more visible. In the posterior part of the body tubercles have a diameter of about 2.0–3.0 µm (
<figureCitation id="132D2A290C597567B792A85F311AFE8F" box="[204,272,332,357]" captionStart="FIGURE 1" captionStartId="2.[151,250,1782,1804]" captionTargetBox="[151,1436,554,1758]" captionTargetId="figure@2.[151,1436,554,1758]" captionTargetPageId="2" captionText="FIGURE 1. Ramazzottius littoreus sp. nov. holotype: habitus (A) Scale bar = 50 µm. Sculpture on dorsal cuticle of a paratype; arrows show dorsal muscle attachment points and asterisk shows elliptical cephalic organs (B). Legs and claws IV of the holotype; arrows show lateral gibbosities on hind legs, and arrowhead shows cuticle ornamentation (C). Scale bar (B, C) = 10 µm." httpUri="https://zenodo.org/record/573631/files/figure.png" pageId="4" pageNumber="454">Figs 1</figureCitation>
B, C, arrowhead). External side of legs I–III with small, difficult to see, granules (
<figureCitation id="132D2A290C597567B3EFA85F34FEFE8F" box="[1201,1268,332,357]" captionStart="FIGURE 3" captionStartId="5.[151,250,1337,1359]" captionTargetBox="[151,1436,195,1316]" captionTargetId="figure@5.[151,1436,195,1316]" captionTargetPageId="5" captionText="FIGURE 3. Ramazzottius littoreus sp. nov. (holotype). Bucco-pharyngeal apparatus in dorsal view (holotype); arrow shows posterior band with four dorsal teeth (A). Bucco-pharyngeal apparatus in ventral view (paratype); arrow shows posterior band with four ventral teeth (only three visible) (B). Claws III (holotype); arrow shows main branch of external claw with robust accessory points, and arrowhead shows cuticle granulation (C). Claws IV (holotype); arrows show lunules (D). All scale bars = 10 µm." httpUri="https://zenodo.org/record/573633/files/figure.png" pageId="4" pageNumber="454">Fig. 3</figureCitation>
C, arrowhead). Dorsal side of legs IV sclerotised and also covered with obvious granules. Lateral gibbosities present on hind legs (
<figureCitation id="132D2A290C597567B7C1A88730E8FE47" box="[159,226,404,429]" captionStart="FIGURE 1" captionStartId="2.[151,250,1782,1804]" captionTargetBox="[151,1436,554,1758]" captionTargetId="figure@2.[151,1436,554,1758]" captionTargetPageId="2" captionText="FIGURE 1. Ramazzottius littoreus sp. nov. holotype: habitus (A) Scale bar = 50 µm. Sculpture on dorsal cuticle of a paratype; arrows show dorsal muscle attachment points and asterisk shows elliptical cephalic organs (B). Legs and claws IV of the holotype; arrows show lateral gibbosities on hind legs, and arrowhead shows cuticle ornamentation (C). Scale bar (B, C) = 10 µm." httpUri="https://zenodo.org/record/573631/files/figure.png" pageId="4" pageNumber="454">Figs 1</figureCitation>
A, C, arrows). Ventral cuticle smooth. Granular nodes corresponding to muscle attachment points are only clearly seen under DIC, particularly in segmental folds deprived of tubercles (
<figureCitation id="132D2A290C597567B359A8A4344DFE3B" box="[1031,1095,439,464]" captionStart="FIGURE 1" captionStartId="2.[151,250,1782,1804]" captionTargetBox="[151,1436,554,1758]" captionTargetId="figure@2.[151,1436,554,1758]" captionTargetPageId="2" captionText="FIGURE 1. Ramazzottius littoreus sp. nov. holotype: habitus (A) Scale bar = 50 µm. Sculpture on dorsal cuticle of a paratype; arrows show dorsal muscle attachment points and asterisk shows elliptical cephalic organs (B). Legs and claws IV of the holotype; arrows show lateral gibbosities on hind legs, and arrowhead shows cuticle ornamentation (C). Scale bar (B, C) = 10 µm." httpUri="https://zenodo.org/record/573631/files/figure.png" pageId="4" pageNumber="454">Fig. 1</figureCitation>
B, arrows). They are arranged in ten ventral and eight dorsal rows (
<figureCitation id="132D2A290C597567B56FA8CF327DFE1F" box="[561,631,476,501]" captionStart="FIGURE 2" captionStartId="4.[151,250,1689,1711]" captionTargetBox="[441,1155,608,1658]" captionTargetId="figure@4.[418,1168,589,1668]" captionTargetPageId="4" captionText="FIGURE 2. Schematic representation of muscle attachment points in Ramazzottius littoreus sp. nov. (A: ventral view; B: dorsal view) and Macrobiotus halophilus sp. nov. (C: ventral view; D: dorsal view). Numerals (I – XV) identify transversal rows." httpUri="https://zenodo.org/record/573632/files/figure.png" pageId="4" pageNumber="454">Figs 2</figureCitation>
A, B).
</paragraph>
<paragraph id="8BA936AC0C5E7560B6BCAB86357DF8B8" pageId="3" pageNumber="453">
<table id="F916C40C0C5E8A9CB7C1AB87357DF899" box="[159,1399,660,1906]" gridcols="6" gridrows="18" pageId="3" pageNumber="453">
<tr id="352634EE0C5E8A9CB7C1AB87357DFD41" box="[159,1399,660,682]" gridrow="0" pageId="3" pageNumber="453" rowspan-0="1">
<th id="76F75D920C5E8A9CB6BCAB87326EFD41" box="[482,612,660,682]" gridcol="1" gridrow="0" pageId="3" pageNumber="453">
µm 
<emphasis id="B962EABE0C5E7560B555AB873225FD42" box="[523,559,660,681]" italics="true" pageId="3" pageNumber="453">(pt)</emphasis>
</th>
<th id="76F75D920C5E8A9CB5C8AB873386FD41" box="[662,908,660,682]" gridcol="2" gridrow="0" pageId="3" pageNumber="453">µm</th>
<th id="76F75D920C5E8A9CB49BAB8734EFFD41" box="[965,1253,660,682]" gridcol="3" gridrow="0" pageId="3" pageNumber="453">
<emphasis id="B962EABE0C5E7560B49BAB8633D3FD41" box="[965,985,661,682]" italics="true" pageId="3" pageNumber="453">pt</emphasis>
</th>
<th id="76F75D920C5E8A9CB25AAB87352EFD41" box="[1284,1316,660,682]" gridcol="4" gridrow="0" pageId="3" pageNumber="453">b</th>
<th id="76F75D920C5E8A9CB215AB87357DFD41" box="[1355,1399,660,682]" gridcol="5" gridrow="0" pageId="3" pageNumber="453">a*</th>
</tr>
<tr id="352634EE0C5E8A9CB7C1ABAE357DFD19" box="[159,1399,701,754]" gridrow="1" pageId="3" pageNumber="453">
<th id="76F75D920C5E8A9CB7C1ABAE31CAFD19" box="[159,448,701,754]" gridcol="0" gridrow="1" pageId="3" pageNumber="453">Body length</th>
<td id="76F75D920C5E8A9CB6BCABAE326EFD19" box="[482,612,701,754]" gridcol="1" gridrow="1" pageId="3" pageNumber="453">
397 (
<emphasis id="B962EABE0C5E7560B548ABAE326EFD38" box="[534,612,701,723]" italics="true" pageId="3" pageNumber="453">1044.7)</emphasis>
</td>
<td id="76F75D920C5E8A9CB5C8ABAE3386FD19" box="[662,908,701,754]" gridcol="2" gridrow="1" pageId="3" pageNumber="453">288.2 ± 55.2 (159–397); (N=23)</td>
<td id="76F75D920C5E8A9CB49BABAE34EFFD19" box="[965,1253,701,754]" gridcol="3" gridrow="1" pageId="3" pageNumber="453">
<emphasis id="B962EABE0C5E7560B49BABAE33F4FD38" box="[965,1022,701,723]" italics="true" pageId="3" pageNumber="453">932.3</emphasis>
± 
<emphasis id="B962EABE0C5E7560B34CABAE3434FD38" box="[1042,1086,701,723]" italics="true" pageId="3" pageNumber="453">95.6</emphasis>
(
<emphasis id="B962EABE0C5E7560B313ABAE34DFFD38" box="[1101,1237,701,723]" italics="true" pageId="3" pageNumber="453">779.4–1136.5</emphasis>
); (N=23)
</td>
<td id="76F75D920C5E8A9CB25AABAE352EFD19" box="[1284,1316,701,754]" gridcol="4" gridrow="1" pageId="3" pageNumber="453">-</td>
<td id="76F75D920C5E8A9CB215ABAE357DFD19" box="[1355,1399,701,754]" gridcol="5" gridrow="1" pageId="3" pageNumber="453">-</td>
</tr>
<tr id="352634EE0C5E8A9CB7C1AA16357DFCD1" box="[159,1399,773,826]" gridrow="2" pageId="3" pageNumber="453">
<th id="76F75D920C5E8A9CB7C1AA1631CAFCD1" box="[159,448,773,826]" gridcol="0" gridrow="2" pageId="3" pageNumber="453">Buccal tube length</th>
<td id="76F75D920C5E8A9CB6BCAA16326EFCD1" box="[482,612,773,826]" gridcol="1" gridrow="2" pageId="3" pageNumber="453">38.0 (-)</td>
<td id="76F75D920C5E8A9CB5C8AA163386FCD1" box="[662,908,773,826]" gridcol="2" gridrow="2" pageId="3" pageNumber="453">30.7± 3.4 (20.4–38.0); (N=24)</td>
<td id="76F75D920C5E8A9CB49BAA1634EFFCD1" box="[965,1253,773,826]" gridcol="3" gridrow="2" pageId="3" pageNumber="453">-</td>
<td id="76F75D920C5E8A9CB25AAA16352EFCD1" box="[1284,1316,773,826]" gridcol="4" gridrow="2" pageId="3" pageNumber="453">-</td>
<td id="76F75D920C5E8A9CB215AA16357DFCD1" box="[1355,1399,773,826]" gridcol="5" gridrow="2" pageId="3" pageNumber="453">-</td>
</tr>
<tr id="352634EE0C5E8A9CB7C1AA5E357DFC69" box="[159,1399,845,898]" gridrow="3" pageId="3" pageNumber="453">
<th id="76F75D920C5E8A9CB7C1AA5E31CAFC69" box="[159,448,845,898]" gridcol="0" gridrow="3" pageId="3" pageNumber="453">Buccal tube external width</th>
<td id="76F75D920C5E8A9CB6BCAA5E326EFC69" box="[482,612,845,898]" gridcol="1" gridrow="3" pageId="3" pageNumber="453">
2.5 (
<emphasis id="B962EABE0C5E7560B54EAA5E3225FC88" box="[528,559,845,867]" italics="true" pageId="3" pageNumber="453">6.6</emphasis>
)
</td>
<td id="76F75D920C5E8A9CB5C8AA5E3386FC69" box="[662,908,845,898]" gridcol="2" gridrow="3" pageId="3" pageNumber="453">2.3 ± 0.4 (1.2–3.0); (N=24)</td>
<td id="76F75D920C5E8A9CB49BAA5E34EFFC69" box="[965,1253,845,898]" gridcol="3" gridrow="3" pageId="3" pageNumber="453">
<emphasis id="B962EABE0C5E7560B49BAA5E33EFFC88" box="[965,997,845,867]" italics="true" pageId="3" pageNumber="453">7.4</emphasis>
± 
<emphasis id="B962EABE0C5E7560B4A7AA5E3413FC88" box="[1017,1049,845,867]" italics="true" pageId="3" pageNumber="453">0.8</emphasis>
(
<emphasis id="B962EABE0C5E7560B379AA5E3478FC88" box="[1063,1138,845,867]" italics="true" pageId="3" pageNumber="453">6.3–8.8</emphasis>
); (N=24)
</td>
<td id="76F75D920C5E8A9CB25AAA5E352EFC69" box="[1284,1316,845,898]" gridcol="4" gridrow="3" pageId="3" pageNumber="453">1.3</td>
<td id="76F75D920C5E8A9CB215AA5E357DFC69" box="[1355,1399,845,898]" gridcol="5" gridrow="3" pageId="3" pageNumber="453">2.3</td>
</tr>
<tr id="352634EE0C5E8A9CB7C1AA86357DFC21" box="[159,1399,917,970]" gridrow="4" pageId="3" pageNumber="453">
<th id="76F75D920C5E8A9CB7C1AA8631CAFC21" box="[159,448,917,970]" gridcol="0" gridrow="4" pageId="3" pageNumber="453">Buccal tube internal width</th>
<td id="76F75D920C5E8A9CB6BCAA86326EFC21" box="[482,612,917,970]" gridcol="1" gridrow="4" pageId="3" pageNumber="453">
1.4 (
<emphasis id="B962EABE0C5E7560B54EAA863225FC40" box="[528,559,917,939]" italics="true" pageId="3" pageNumber="453">3.7</emphasis>
)
</td>
<td id="76F75D920C5E8A9CB5C8AA863386FC21" box="[662,908,917,970]" gridcol="2" gridrow="4" pageId="3" pageNumber="453">1.1± 0.2 (0.6–1.5); (N=23)</td>
<td id="76F75D920C5E8A9CB49BAA8634EFFC21" box="[965,1253,917,970]" gridcol="3" gridrow="4" pageId="3" pageNumber="453">
<emphasis id="B962EABE0C5E7560B49BAA8633EFFC40" box="[965,997,917,939]" italics="true" pageId="3" pageNumber="453">3.6</emphasis>
± 
<emphasis id="B962EABE0C5E7560B4A7AA863413FC40" box="[1017,1049,917,939]" italics="true" pageId="3" pageNumber="453">0.4</emphasis>
(
<emphasis id="B962EABE0C5E7560B379AA863478FC40" box="[1063,1138,917,939]" italics="true" pageId="3" pageNumber="453">2.7–4.6</emphasis>
); (N=23)
</td>
<td id="76F75D920C5E8A9CB25AAA86352EFC21" box="[1284,1316,917,970]" gridcol="4" gridrow="4" pageId="3" pageNumber="453">
<emphasis id="B962EABE0C5E7560B25AAA863529FC41" bold="true" box="[1284,1315,917,938]" pageId="3" pageNumber="453">1.5</emphasis>
</td>
<td id="76F75D920C5E8A9CB215AA86357DFC21" box="[1355,1399,917,970]" gridcol="5" gridrow="4" pageId="3" pageNumber="453">1.1</td>
</tr>
<tr id="352634EE0C5E8A9CB7C1AACE357DFBF9" box="[159,1399,989,1042]" gridrow="5" pageId="3" pageNumber="453">
<th id="76F75D920C5E8A9CB7C1AACE31CAFBF9" box="[159,448,989,1042]" gridcol="0" gridrow="5" pageId="3" pageNumber="453">Stylet support insertion</th>
<td id="76F75D920C5E8A9CB6BCAACE326EFBF9" box="[482,612,989,1042]" gridcol="1" gridrow="5" pageId="3" pageNumber="453">
23.1 (
<emphasis id="B962EABE0C5E7560B543AACE3243FC18" box="[541,585,989,1011]" italics="true" pageId="3" pageNumber="453">60.8</emphasis>
)
</td>
<td id="76F75D920C5E8A9CB5C8AACE3386FBF9" box="[662,908,989,1042]" gridcol="2" gridrow="5" pageId="3" pageNumber="453">18.7± 2.0 (12.4–23.1); (N=24)</td>
<td id="76F75D920C5E8A9CB49BAACE34EFFBF9" box="[965,1253,989,1042]" gridcol="3" gridrow="5" pageId="3" pageNumber="453">
<emphasis id="B962EABE0C5E7560B49BAACE33FBFC18" box="[965,1009,989,1011]" italics="true" pageId="3" pageNumber="453">60.9</emphasis>
± 
<emphasis id="B962EABE0C5E7560B35BAACE342EFC18" box="[1029,1060,989,1011]" italics="true" pageId="3" pageNumber="453">1.3</emphasis>
(
<emphasis id="B962EABE0C5E7560B36AAACE3492FC18" box="[1076,1176,989,1011]" italics="true" pageId="3" pageNumber="453">58.0–62.9</emphasis>
); (N=24)
</td>
<td id="76F75D920C5E8A9CB25AAACE352EFBF9" box="[1284,1316,989,1042]" gridcol="4" gridrow="5" pageId="3" pageNumber="453">1.0</td>
<td id="76F75D920C5E8A9CB215AACE357DFBF9" box="[1355,1399,989,1042]" gridcol="5" gridrow="5" pageId="3" pageNumber="453">18.7</td>
</tr>
<tr id="352634EE0C5E8A9CB7C1AD36357DFBB1" box="[159,1399,1061,1114]" gridrow="6" pageId="3" pageNumber="453">
<th id="76F75D920C5E8A9CB7C1AD3631CAFBB1" box="[159,448,1061,1114]" gridcol="0" gridrow="6" pageId="3" pageNumber="453">First macroplacoid length</th>
<td id="76F75D920C5E8A9CB6BCAD36326EFBB1" box="[482,612,1061,1114]" gridcol="1" gridrow="6" pageId="3" pageNumber="453">
3.4 (
<emphasis id="B962EABE0C5E7560B54EAD363225FBD0" box="[528,559,1061,1083]" italics="true" pageId="3" pageNumber="453">9.0</emphasis>
)
</td>
<td id="76F75D920C5E8A9CB5C8AD363386FBB1" box="[662,908,1061,1114]" gridcol="2" gridrow="6" pageId="3" pageNumber="453">3.1± 0.4 (1.7–3.8); (N=24)</td>
<td id="76F75D920C5E8A9CB49BAD3634EFFBB1" box="[965,1253,1061,1114]" gridcol="3" gridrow="6" pageId="3" pageNumber="453">
<emphasis id="B962EABE0C5E7560B49BAD3633FBFBD0" box="[965,1009,1061,1083]" italics="true" pageId="3" pageNumber="453">10.2</emphasis>
± 
<emphasis id="B962EABE0C5E7560B35BAD36342EFBD0" box="[1029,1060,1061,1083]" italics="true" pageId="3" pageNumber="453">0.9</emphasis>
(
<emphasis id="B962EABE0C5E7560B36AAD363483FBD0" box="[1076,1161,1061,1083]" italics="true" pageId="3" pageNumber="453">8.3–11.9</emphasis>
); (N=24)
</td>
<td id="76F75D920C5E8A9CB25AAD36352EFBB1" box="[1284,1316,1061,1114]" gridcol="4" gridrow="6" pageId="3" pageNumber="453">1.2</td>
<td id="76F75D920C5E8A9CB215AD36357DFBB1" box="[1355,1399,1061,1114]" gridcol="5" gridrow="6" pageId="3" pageNumber="453">3.2</td>
</tr>
<tr id="352634EE0C5E8A9CB7C1AD7E357DFB49" box="[159,1399,1133,1186]" gridrow="7" pageId="3" pageNumber="453">
<th id="76F75D920C5E8A9CB7C1AD7E31CAFB49" box="[159,448,1133,1186]" gridcol="0" gridrow="7" pageId="3" pageNumber="453">Second macroplacoid length</th>
<td id="76F75D920C5E8A9CB6BCAD7E326EFB49" box="[482,612,1133,1186]" gridcol="1" gridrow="7" pageId="3" pageNumber="453">
3.3 (
<emphasis id="B962EABE0C5E7560B54EAD7E3225FB68" box="[528,559,1133,1155]" italics="true" pageId="3" pageNumber="453">8.7</emphasis>
)
</td>
<td id="76F75D920C5E8A9CB5C8AD7E3386FB49" box="[662,908,1133,1186]" gridcol="2" gridrow="7" pageId="3" pageNumber="453">2.7± 0.4 (1.7–3.3); (N=24)</td>
<td id="76F75D920C5E8A9CB49BAD7E34EFFB49" box="[965,1253,1133,1186]" gridcol="3" gridrow="7" pageId="3" pageNumber="453">
<emphasis id="B962EABE0C5E7560B49BAD7E33EFFB68" box="[965,997,1133,1155]" italics="true" pageId="3" pageNumber="453">8.9</emphasis>
± 
<emphasis id="B962EABE0C5E7560B4A7AD7E3413FB68" box="[1017,1049,1133,1155]" italics="true" pageId="3" pageNumber="453">0.7</emphasis>
(
<emphasis id="B962EABE0C5E7560B379AD7E3471FB68" box="[1063,1147,1133,1155]" italics="true" pageId="3" pageNumber="453">8.0–11.2</emphasis>
); (N=24)
</td>
<td id="76F75D920C5E8A9CB25AAD7E352EFB49" box="[1284,1316,1133,1186]" gridcol="4" gridrow="7" pageId="3" pageNumber="453">1.1</td>
<td id="76F75D920C5E8A9CB215AD7E357DFB49" box="[1355,1399,1133,1186]" gridcol="5" gridrow="7" pageId="3" pageNumber="453">2.8</td>
</tr>
<tr id="352634EE0C5E8A9CB7C1ADA6357DFB01" box="[159,1399,1205,1258]" gridrow="8" pageId="3" pageNumber="453">
<th id="76F75D920C5E8A9CB7C1ADA631CAFB01" box="[159,448,1205,1258]" gridcol="0" gridrow="8" pageId="3" pageNumber="453">Placoid row length</th>
<td id="76F75D920C5E8A9CB6BCADA6326EFB01" box="[482,612,1205,1258]" gridcol="1" gridrow="8" pageId="3" pageNumber="453">
8.1 (
<emphasis id="B962EABE0C5E7560B54EADA63236FB20" box="[528,572,1205,1227]" italics="true" pageId="3" pageNumber="453">21.3</emphasis>
)
</td>
<td id="76F75D920C5E8A9CB5C8ADA63386FB01" box="[662,908,1205,1258]" gridcol="2" gridrow="8" pageId="3" pageNumber="453">6.6± 0.9 (3.9–8.1); (N=24)</td>
<td id="76F75D920C5E8A9CB49BADA634EFFB01" box="[965,1253,1205,1258]" gridcol="3" gridrow="8" pageId="3" pageNumber="453">
<emphasis id="B962EABE0C5E7560B49BADA633FBFB20" box="[965,1009,1205,1227]" italics="true" pageId="3" pageNumber="453">21.6</emphasis>
± 
<emphasis id="B962EABE0C5E7560B35BADA6342EFB20" box="[1029,1060,1205,1227]" italics="true" pageId="3" pageNumber="453">1.1</emphasis>
(
<emphasis id="B962EABE0C5E7560B36AADA63492FB20" box="[1076,1176,1205,1227]" italics="true" pageId="3" pageNumber="453">19.1–23.5</emphasis>
); (N=24)
</td>
<td id="76F75D920C5E8A9CB25AADA6352EFB01" box="[1284,1316,1205,1258]" gridcol="4" gridrow="8" pageId="3" pageNumber="453">
<emphasis id="B962EABE0C5E7560B25AADA63529FB21" bold="true" box="[1284,1315,1205,1226]" pageId="3" pageNumber="453">1.2</emphasis>
</td>
<td id="76F75D920C5E8A9CB215ADA6357DFB01" box="[1355,1399,1205,1258]" gridcol="5" gridrow="8" pageId="3" pageNumber="453">6.7</td>
</tr>
<tr id="352634EE0C5E8A9CB7C1ADEE357DFAD9" box="[159,1399,1277,1330]" gridrow="9" pageId="3" pageNumber="453">
<th id="76F75D920C5E8A9CB7C1ADEE31CAFAD9" box="[159,448,1277,1330]" gridcol="0" gridrow="9" pageId="3" pageNumber="453">Externalclaw IV: primary branch</th>
<td id="76F75D920C5E8A9CB6BCADEE326EFAD9" box="[482,612,1277,1330]" gridcol="1" gridrow="9" pageId="3" pageNumber="453">
14.1 (
<emphasis id="B962EABE0C5E7560B543ADEE3243FAF8" box="[541,585,1277,1299]" italics="true" pageId="3" pageNumber="453">37.1</emphasis>
)
</td>
<td id="76F75D920C5E8A9CB5C8ADEE3386FAD9" box="[662,908,1277,1330]" gridcol="2" gridrow="9" pageId="3" pageNumber="453">11.3± 1.4 (7.7–14.1); (N=18)</td>
<td id="76F75D920C5E8A9CB49BADEE34EFFAD9" box="[965,1253,1277,1330]" gridcol="3" gridrow="9" pageId="3" pageNumber="453">
<emphasis id="B962EABE0C5E7560B49BADEE33FBFAF8" box="[965,1009,1277,1299]" italics="true" pageId="3" pageNumber="453">37.5</emphasis>
± 
<emphasis id="B962EABE0C5E7560B35BADEE342EFAF8" box="[1029,1060,1277,1299]" italics="true" pageId="3" pageNumber="453">1.8</emphasis>
(
<emphasis id="B962EABE0C5E7560B36AADEE3492FAF8" box="[1076,1176,1277,1299]" italics="true" pageId="3" pageNumber="453">33.6–41.6</emphasis>
); (N=18)
</td>
<td id="76F75D920C5E8A9CB25AADEE352EFAD9" box="[1284,1316,1277,1330]" gridcol="4" gridrow="9" pageId="3" pageNumber="453">
<emphasis id="B962EABE0C5E7560B25AADEE3529FAF9" bold="true" box="[1284,1315,1277,1298]" pageId="3" pageNumber="453">1.0</emphasis>
</td>
<td id="76F75D920C5E8A9CB215ADEE357DFAD9" box="[1355,1399,1277,1330]" gridcol="5" gridrow="9" pageId="3" pageNumber="453">11.5</td>
</tr>
<tr id="352634EE0C5E8A9CB7C1AC56357DFA91" box="[159,1399,1349,1402]" gridrow="10" pageId="3" pageNumber="453">
<th id="76F75D920C5E8A9CB7C1AC5631CAFA91" box="[159,448,1349,1402]" gridcol="0" gridrow="10" pageId="3" pageNumber="453">External claw I: base</th>
<td id="76F75D920C5E8A9CB6BCAC56326EFA91" box="[482,612,1349,1402]" gridcol="1" gridrow="10" pageId="3" pageNumber="453">
12.5 (
<emphasis id="B962EABE0C5E7560B543AC563243FAB0" box="[541,585,1349,1371]" italics="true" pageId="3" pageNumber="453">32.9</emphasis>
)
</td>
<td id="76F75D920C5E8A9CB5C8AC563386FA91" box="[662,908,1349,1402]" gridcol="2" gridrow="10" pageId="3" pageNumber="453">10.2± 1.4 (6.3–12.5); (N=15)</td>
<td id="76F75D920C5E8A9CB49BAC5634EFFA91" box="[965,1253,1349,1402]" gridcol="3" gridrow="10" pageId="3" pageNumber="453">
<emphasis id="B962EABE0C5E7560B49BAC5633FBFAB0" box="[965,1009,1349,1371]" italics="true" pageId="3" pageNumber="453">33.6</emphasis>
± 
<emphasis id="B962EABE0C5E7560B35BAC56342EFAB0" box="[1029,1060,1349,1371]" italics="true" pageId="3" pageNumber="453">2.2</emphasis>
(
<emphasis id="B962EABE0C5E7560B36AAC563492FAB0" box="[1076,1176,1349,1371]" italics="true" pageId="3" pageNumber="453">29.5–37.1</emphasis>
); (N=15)
</td>
<td id="76F75D920C5E8A9CB25AAC56352EFA91" box="[1284,1316,1349,1402]" gridcol="4" gridrow="10" pageId="3" pageNumber="453">1.0</td>
<td id="76F75D920C5E8A9CB215AC56357DFA91" box="[1355,1399,1349,1402]" gridcol="5" gridrow="10" pageId="3" pageNumber="453">10.6</td>
</tr>
<tr id="352634EE0C5E8A9CB7C1AC9E357DFA29" box="[159,1399,1421,1474]" gridrow="11" pageId="3" pageNumber="453">
<th id="76F75D920C5E8A9CB7C1AC9E31CAFA29" box="[159,448,1421,1474]" gridcol="0" gridrow="11" pageId="3" pageNumber="453">Internal claw I length</th>
<td id="76F75D920C5E8A9CB6BCAC9E326EFA29" box="[482,612,1421,1474]" gridcol="1" gridrow="11" pageId="3" pageNumber="453">
13.4 (
<emphasis id="B962EABE0C5E7560B543AC9E3243FA48" box="[541,585,1421,1443]" italics="true" pageId="3" pageNumber="453">35.3</emphasis>
)
</td>
<td id="76F75D920C5E8A9CB5C8AC9E3386FA29" box="[662,908,1421,1474]" gridcol="2" gridrow="11" pageId="3" pageNumber="453">10.9± 1.6 (7.3–13.4); (N=14)</td>
<td id="76F75D920C5E8A9CB49BAC9E34EFFA29" box="[965,1253,1421,1474]" gridcol="3" gridrow="11" pageId="3" pageNumber="453">
<emphasis id="B962EABE0C5E7560B49BAC9E33FBFA48" box="[965,1009,1421,1443]" italics="true" pageId="3" pageNumber="453">35.7</emphasis>
± 
<emphasis id="B962EABE0C5E7560B35BAC9E342EFA48" box="[1029,1060,1421,1443]" italics="true" pageId="3" pageNumber="453">3.0</emphasis>
(
<emphasis id="B962EABE0C5E7560B36AAC9E3492FA48" box="[1076,1176,1421,1443]" italics="true" pageId="3" pageNumber="453">32.1–41.1</emphasis>
); (N=14)
</td>
<td id="76F75D920C5E8A9CB25AAC9E352EFA29" box="[1284,1316,1421,1474]" gridcol="4" gridrow="11" pageId="3" pageNumber="453">1.0</td>
<td id="76F75D920C5E8A9CB215AC9E357DFA29" box="[1355,1399,1421,1474]" gridcol="5" gridrow="11" pageId="3" pageNumber="453">11.0</td>
</tr>
<tr id="352634EE0C5E8A9CB7C1ACC6357DF9E1" box="[159,1399,1493,1546]" gridrow="12" pageId="3" pageNumber="453">
<th id="76F75D920C5E8A9CB7C1ACC631CAF9E1" box="[159,448,1493,1546]" gridcol="0" gridrow="12" pageId="3" pageNumber="453">External claw II/III: primary branch</th>
<td id="76F75D920C5E8A9CB6BCACC6326EF9E1" box="[482,612,1493,1546]" gridcol="1" gridrow="12" pageId="3" pageNumber="453">
14.8 (
<emphasis id="B962EABE0C5E7560B543ACC63243FA00" box="[541,585,1493,1515]" italics="true" pageId="3" pageNumber="453">38.9</emphasis>
)
</td>
<td id="76F75D920C5E8A9CB5C8ACC63386F9E1" box="[662,908,1493,1546]" gridcol="2" gridrow="12" pageId="3" pageNumber="453">13.2± 1.6 (8.3–15.1); (N=21)</td>
<td id="76F75D920C5E8A9CB49BACC634EFF9E1" box="[965,1253,1493,1546]" gridcol="3" gridrow="12" pageId="3" pageNumber="453">
<emphasis id="B962EABE0C5E7560B49BACC633FBFA00" box="[965,1009,1493,1515]" italics="true" pageId="3" pageNumber="453">43.0</emphasis>
± 
<emphasis id="B962EABE0C5E7560B35BACC6342EFA00" box="[1029,1060,1493,1515]" italics="true" pageId="3" pageNumber="453">2.6</emphasis>
(
<emphasis id="B962EABE0C5E7560B36DACC63492FA00" box="[1075,1176,1493,1515]" italics="true" pageId="3" pageNumber="453">37.0–48.3</emphasis>
); (N=21)
</td>
<td id="76F75D920C5E8A9CB25AACC6352EF9E1" box="[1284,1316,1493,1546]" gridcol="4" gridrow="12" pageId="3" pageNumber="453">1.0</td>
<td id="76F75D920C5E8A9CB215ACC6357DF9E1" box="[1355,1399,1493,1546]" gridcol="5" gridrow="12" pageId="3" pageNumber="453">13.6</td>
</tr>
<tr id="352634EE0C5E8A9CB7C1AF0E357DF9B9" box="[159,1399,1565,1618]" gridrow="13" pageId="3" pageNumber="453">
<th id="76F75D920C5E8A9CB7C1AF0E31CAF9B9" box="[159,448,1565,1618]" gridcol="0" gridrow="13" pageId="3" pageNumber="453">Posterior claw II/III: base</th>
<td id="76F75D920C5E8A9CB6BCAF0E326EF9B9" box="[482,612,1565,1618]" gridcol="1" gridrow="13" pageId="3" pageNumber="453">
14.3 (
<emphasis id="B962EABE0C5E7560B543AF0E3243F9D8" box="[541,585,1565,1587]" italics="true" pageId="3" pageNumber="453">37.6</emphasis>
)
</td>
<td id="76F75D920C5E8A9CB5C8AF0E3386F9B9" box="[662,908,1565,1618]" gridcol="2" gridrow="13" pageId="3" pageNumber="453">11.6± 1.6 (6.3–14.3); (N=22)</td>
<td id="76F75D920C5E8A9CB49BAF0E34EFF9B9" box="[965,1253,1565,1618]" gridcol="3" gridrow="13" pageId="3" pageNumber="453">
<emphasis id="B962EABE0C5E7560B49BAF0E33FBF9D8" box="[965,1009,1565,1587]" italics="true" pageId="3" pageNumber="453">37.6</emphasis>
± 
<emphasis id="B962EABE0C5E7560B35BAF0E342EF9D8" box="[1029,1060,1565,1587]" italics="true" pageId="3" pageNumber="453">2.2</emphasis>
(
<emphasis id="B962EABE0C5E7560B36DAF0E3492F9D8" box="[1075,1176,1565,1587]" italics="true" pageId="3" pageNumber="453">30.9–42.1</emphasis>
); (N=22)
</td>
<td id="76F75D920C5E8A9CB25AAF0E352EF9B9" box="[1284,1316,1565,1618]" gridcol="4" gridrow="13" pageId="3" pageNumber="453">
<emphasis id="B962EABE0C5E7560B25AAF0E3529F9D9" bold="true" box="[1284,1315,1565,1586]" pageId="3" pageNumber="453">1.3</emphasis>
</td>
<td id="76F75D920C5E8A9CB215AF0E357DF9B9" box="[1355,1399,1565,1618]" gridcol="5" gridrow="13" pageId="3" pageNumber="453">11.9</td>
</tr>
<tr id="352634EE0C5E8A9CB7C1AF76357DF971" box="[159,1399,1637,1690]" gridrow="14" pageId="3" pageNumber="453">
<th id="76F75D920C5E8A9CB7C1AF7631CAF971" box="[159,448,1637,1690]" gridcol="0" gridrow="14" pageId="3" pageNumber="453">Internal claw II/III length</th>
<td id="76F75D920C5E8A9CB6BCAF76326EF971" box="[482,612,1637,1690]" gridcol="1" gridrow="14" pageId="3" pageNumber="453">
13.8 (
<emphasis id="B962EABE0C5E7560B543AF763243F990" box="[541,585,1637,1659]" italics="true" pageId="3" pageNumber="453">36.3</emphasis>
)
</td>
<td id="76F75D920C5E8A9CB5C8AF763386F971" box="[662,908,1637,1690]" gridcol="2" gridrow="14" pageId="3" pageNumber="453">11.9± 1.6 (6.8–13.8); (N=17)</td>
<td id="76F75D920C5E8A9CB49BAF7634EFF971" box="[965,1253,1637,1690]" gridcol="3" gridrow="14" pageId="3" pageNumber="453">
<emphasis id="B962EABE0C5E7560B49BAF7633FBF990" box="[965,1009,1637,1659]" italics="true" pageId="3" pageNumber="453">38.5</emphasis>
± 
<emphasis id="B962EABE0C5E7560B35BAF76342EF990" box="[1029,1060,1637,1659]" italics="true" pageId="3" pageNumber="453">2.5</emphasis>
(
<emphasis id="B962EABE0C5E7560B36DAF763492F990" box="[1075,1176,1637,1659]" italics="true" pageId="3" pageNumber="453">33.3–45.3</emphasis>
); (N=17)
</td>
<td id="76F75D920C5E8A9CB25AAF76352EF971" box="[1284,1316,1637,1690]" gridcol="4" gridrow="14" pageId="3" pageNumber="453">1.1</td>
<td id="76F75D920C5E8A9CB215AF76357DF971" box="[1355,1399,1637,1690]" gridcol="5" gridrow="14" pageId="3" pageNumber="453">12.4</td>
</tr>
<tr id="352634EE0C5E8A9CB7C1AFBE357DF909" box="[159,1399,1709,1762]" gridrow="15" pageId="3" pageNumber="453">
<th id="76F75D920C5E8A9CB7C1AFBE31CAF909" box="[159,448,1709,1762]" gridcol="0" gridrow="15" pageId="3" pageNumber="453">External claw I: primary branch</th>
<td id="76F75D920C5E8A9CB6BCAFBE326EF909" box="[482,612,1709,1762]" gridcol="1" gridrow="15" pageId="3" pageNumber="453">
17.6 (
<emphasis id="B962EABE0C5E7560B543AFBE3243F928" box="[541,585,1709,1731]" italics="true" pageId="3" pageNumber="453">46.3</emphasis>
)
</td>
<td id="76F75D920C5E8A9CB5C8AFBE3386F909" box="[662,908,1709,1762]" gridcol="2" gridrow="15" pageId="3" pageNumber="453">15.2± 1.8 (10.4–17.6); (N=15)</td>
<td id="76F75D920C5E8A9CB49BAFBE34EFF909" box="[965,1253,1709,1762]" gridcol="3" gridrow="15" pageId="3" pageNumber="453">
<emphasis id="B962EABE0C5E7560B49BAFBE33FBF928" box="[965,1009,1709,1731]" italics="true" pageId="3" pageNumber="453">51.0</emphasis>
± 
<emphasis id="B962EABE0C5E7560B35BAFBE342EF928" box="[1029,1060,1709,1731]" italics="true" pageId="3" pageNumber="453">3.2</emphasis>
(
<emphasis id="B962EABE0C5E7560B36AAFBE3492F928" box="[1076,1176,1709,1731]" italics="true" pageId="3" pageNumber="453">46.3–53.0</emphasis>
); (N=15)
</td>
<td id="76F75D920C5E8A9CB25AAFBE352EF909" box="[1284,1316,1709,1762]" gridcol="4" gridrow="15" pageId="3" pageNumber="453">0.9</td>
<td id="76F75D920C5E8A9CB215AFBE357DF909" box="[1355,1399,1709,1762]" gridcol="5" gridrow="15" pageId="3" pageNumber="453">16.0</td>
</tr>
<tr id="352634EE0C5E8A9CB7C1AFE6357DF8C1" box="[159,1399,1781,1834]" gridrow="16" pageId="3" pageNumber="453">
<th id="76F75D920C5E8A9CB7C1AFE631CAF8C1" box="[159,448,1781,1834]" gridcol="0" gridrow="16" pageId="3" pageNumber="453">Posterior claw IV: base</th>
<td id="76F75D920C5E8A9CB6BCAFE6326EF8C1" box="[482,612,1781,1834]" gridcol="1" gridrow="16" pageId="3" pageNumber="453">
14.7 (
<emphasis id="B962EABE0C5E7560B543AFE63243F8E0" box="[541,585,1781,1803]" italics="true" pageId="3" pageNumber="453">38.7</emphasis>
)
</td>
<td id="76F75D920C5E8A9CB5C8AFE63386F8C1" box="[662,908,1781,1834]" gridcol="2" gridrow="16" pageId="3" pageNumber="453">12.1± 1.9 (7.2–14.7); (N=15)</td>
<td id="76F75D920C5E8A9CB49BAFE634EFF8C1" box="[965,1253,1781,1834]" gridcol="3" gridrow="16" pageId="3" pageNumber="453">
<emphasis id="B962EABE0C5E7560B49BAFE633FBF8E0" box="[965,1009,1781,1803]" italics="true" pageId="3" pageNumber="453">39.4</emphasis>
± 
<emphasis id="B962EABE0C5E7560B35BAFE6342EF8E0" box="[1029,1060,1781,1803]" italics="true" pageId="3" pageNumber="453">3.1</emphasis>
(
<emphasis id="B962EABE0C5E7560B36AAFE63492F8E0" box="[1076,1176,1781,1803]" italics="true" pageId="3" pageNumber="453">34.5–46.9</emphasis>
); (N=15)
</td>
<td id="76F75D920C5E8A9CB25AAFE6352EF8C1" box="[1284,1316,1781,1834]" gridcol="4" gridrow="16" pageId="3" pageNumber="453">1.1</td>
<td id="76F75D920C5E8A9CB215AFE6357DF8C1" box="[1355,1399,1781,1834]" gridcol="5" gridrow="16" pageId="3" pageNumber="453">12.4</td>
</tr>
<tr id="352634EE0C5E8A9CB7C1AE2E357DF899" box="[159,1399,1853,1906]" gridrow="17" pageId="3" pageNumber="453">
<th id="76F75D920C5E8A9CB7C1AE2E31CAF899" box="[159,448,1853,1906]" gridcol="0" gridrow="17" pageId="3" pageNumber="453">Anterior claw IV length</th>
<td id="76F75D920C5E8A9CB6BCAE2E326EF899" box="[482,612,1853,1906]" gridcol="1" gridrow="17" pageId="3" pageNumber="453">
14.4 (
<emphasis id="B962EABE0C5E7560B543AE2E3243F8B8" box="[541,585,1853,1875]" italics="true" pageId="3" pageNumber="453">37.9</emphasis>
)
</td>
<td id="76F75D920C5E8A9CB5C8AE2E3386F899" box="[662,908,1853,1906]" gridcol="2" gridrow="17" pageId="3" pageNumber="453">12.7± 1.8 (7.4–14.4); (N=13)</td>
<td id="76F75D920C5E8A9CB49BAE2E34EFF899" box="[965,1253,1853,1906]" gridcol="3" gridrow="17" pageId="3" pageNumber="453">
<emphasis id="B962EABE0C5E7560B49BAE2E33FBF8B8" box="[965,1009,1853,1875]" italics="true" pageId="3" pageNumber="453">41.1</emphasis>
± 
<emphasis id="B962EABE0C5E7560B35BAE2E342EF8B8" box="[1029,1060,1853,1875]" italics="true" pageId="3" pageNumber="453">2.5</emphasis>
(
<emphasis id="B962EABE0C5E7560B36AAE2E3492F8B8" box="[1076,1176,1853,1875]" italics="true" pageId="3" pageNumber="453">36.3–44.7</emphasis>
); (N=13)
</td>
<td id="76F75D920C5E8A9CB25AAE2E352EF899" box="[1284,1316,1853,1906]" gridcol="4" gridrow="17" pageId="3" pageNumber="453">1.1</td>
<td id="76F75D920C5E8A9CB215AE2E357DF899" box="[1355,1399,1853,1906]" gridcol="5" gridrow="17" pageId="3" pageNumber="453">13.2</td>
</tr>
</table>
</paragraph>
<caption id="DF6966240C597567B7C9AF8A30C5F906" httpUri="https://zenodo.org/record/573632/files/figure.png" pageId="4" pageNumber="454" targetBox="[441,1155,608,1658]" targetPageId="4">
<paragraph id="8BA936AC0C597567B7C9AF8A30C5F906" blockId="4.[151,1436,1689,1773]" pageId="4" pageNumber="454">
<emphasis id="B962EABE0C597567B7C9AF8A311CF945" bold="true" box="[151,278,1689,1711]" pageId="4" pageNumber="454">FIGURE 2.</emphasis>
Schematic representation of muscle attachment points in 
<taxonomicName id="4C164D2F0C597567B424AF893455F944" box="[890,1119,1689,1711]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="4" pageNumber="454" phylum="Tardigrada" rank="species" species="littoreus" status="sp. nov.">
<emphasis id="B962EABE0C597567B424AF893455F944" box="[890,1119,1689,1711]" italics="true" pageId="4" pageNumber="454">Ramazzottius littoreus</emphasis>
</taxonomicName>
<emphasis id="B962EABE0C597567B336AF8934B6F944" bold="true" box="[1128,1212,1690,1711]" pageId="4" pageNumber="454">
<taxonomicNameLabel id="A25157C50C597567B336AF8934B6F944" box="[1128,1212,1690,1711]" pageId="4" pageNumber="454" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(A: ventral view; B: dorsal view) and 
<taxonomicName id="4C164D2F0C597567B610AFAB3249F925" box="[334,579,1720,1742]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="4" pageNumber="454" phylum="Tardigrada" rank="species" species="halophilus" status="sp. nov.">
<emphasis id="B962EABE0C597567B610AFAB3249F925" box="[334,579,1720,1742]" italics="true" pageId="4" pageNumber="454">Macrobiotus halophilus</emphasis>
</taxonomicName>
<emphasis id="B962EABE0C597567B512AFAB32AAF926" bold="true" box="[588,672,1720,1741]" pageId="4" pageNumber="454">
<taxonomicNameLabel id="A25157C50C597567B512AFAB32AAF926" box="[588,672,1720,1741]" pageId="4" pageNumber="454" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(C: ventral view; D: dorsal view). Numerals (I–XV) identify transversal rows.
</paragraph>
</caption>
<caption id="DF6966240C587566B7C9AC2A30D5FA21" httpUri="https://zenodo.org/record/573633/files/figure.png" pageId="5" pageNumber="455" targetBox="[151,1436,195,1316]" targetPageId="5">
<paragraph id="8BA936AC0C587566B7C9AC2A30D5FA21" blockId="5.[151,1436,1337,1482]" pageId="5" pageNumber="455">
<emphasis id="B962EABE0C587566B7C9AC2A311EFAA5" bold="true" box="[151,276,1337,1359]" pageId="5" pageNumber="455">FIGURE 3.</emphasis>
<taxonomicName id="4C164D2F0C587566B642AC29320AFAA4" box="[284,512,1337,1359]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="455" phylum="Tardigrada" rank="species" species="littoreus" status="sp. nov.">
<emphasis id="B962EABE0C587566B642AC29320AFAA4" box="[284,512,1337,1359]" italics="true" pageId="5" pageNumber="455">Ramazzottius littoreus</emphasis>
</taxonomicName>
<emphasis id="B962EABE0C587566B559AC293250FAA4" bold="true" box="[519,602,1338,1359]" pageId="5" pageNumber="455">
<taxonomicNameLabel id="A25157C50C587566B559AC293250FAA4" box="[519,602,1338,1359]" pageId="5" pageNumber="455" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(holotype). Bucco-pharyngeal apparatus in dorsal view (holotype); arrow shows posterior band with four dorsal teeth (A). Bucco-pharyngeal apparatus in ventral view (paratype); arrow shows posterior band with four ventral teeth (only three visible) (B). Claws III (holotype); arrow shows main branch of external claw with robust accessory points, and arrowhead shows cuticle granulation (C). Claws IV (holotype); arrows show lunules (D). All scale bars = 10 µm.
</paragraph>
</caption>
<paragraph id="8BA936AC0C587566B799AF033316F8CE" blockId="5.[151,1436,1552,1932]" pageId="5" pageNumber="455">
Bucco-pharyngeal apparatus of 
<taxonomicName id="4C164D2F0C587566B571AF0232CEF9C3" box="[559,708,1553,1576]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="455" phylum="Tardigrada" rank="genus">
<emphasis id="B962EABE0C587566B571AF0232CEF9C3" box="[559,708,1553,1576]" italics="true" pageId="5" pageNumber="455">Ramazzottius</emphasis>
</taxonomicName>
<typeStatus id="54AD880E0C587566B593AF02330AF9C2" box="[717,768,1553,1577]" pageId="5" pageNumber="455">type</typeStatus>
, with a rigid buccal tube without ventral lamina (
<figureCitation id="132D2A290C587566B26DAF03357CF9C3" box="[1331,1398,1552,1577]" captionStart="FIGURE 3" captionStartId="5.[151,250,1337,1359]" captionTargetBox="[151,1436,195,1316]" captionTargetId="figure@5.[151,1436,195,1316]" captionTargetPageId="5" captionText="FIGURE 3. Ramazzottius littoreus sp. nov. (holotype). Bucco-pharyngeal apparatus in dorsal view (holotype); arrow shows posterior band with four dorsal teeth (A). Bucco-pharyngeal apparatus in ventral view (paratype); arrow shows posterior band with four ventral teeth (only three visible) (B). Claws III (holotype); arrow shows main branch of external claw with robust accessory points, and arrowhead shows cuticle granulation (C). Claws IV (holotype); arrows show lunules (D). All scale bars = 10 µm." httpUri="https://zenodo.org/record/573633/files/figure.png" pageId="5" pageNumber="455">Fig. 3</figureCitation>
A); asymmetrical dorsal and ventral blunt hook shaped apophyses for the insertion of stylet muscles. Antero-ventral mouth opening, without peribuccal 
<emphasis id="B962EABE0C587566B579AF4B328BF99B" box="[551,641,1624,1648]" italics="true" pageId="5" pageNumber="455">papulae</emphasis>
and 
<emphasis id="B962EABE0C587566B5E7AF4B3313F99B" box="[697,793,1624,1648]" italics="true" pageId="5" pageNumber="455">lamellae</emphasis>
. Oral cavity with an anterior band of teeth poorly visible and a posterior band with four evident dorsal and four ventral teeth (
<figureCitation id="132D2A290C587566B4F9AF6F33E4F97E" box="[935,1006,1660,1685]" captionStart="FIGURE 3" captionStartId="5.[151,250,1337,1359]" captionTargetBox="[151,1436,195,1316]" captionTargetId="figure@5.[151,1436,195,1316]" captionTargetPageId="5" captionText="FIGURE 3. Ramazzottius littoreus sp. nov. (holotype). Bucco-pharyngeal apparatus in dorsal view (holotype); arrow shows posterior band with four dorsal teeth (A). Bucco-pharyngeal apparatus in ventral view (paratype); arrow shows posterior band with four ventral teeth (only three visible) (B). Claws III (holotype); arrow shows main branch of external claw with robust accessory points, and arrowhead shows cuticle granulation (C). Claws IV (holotype); arrows show lunules (D). All scale bars = 10 µm." httpUri="https://zenodo.org/record/573633/files/figure.png" pageId="5" pageNumber="455">Figs 3</figureCitation>
A, B, arrows). Buccal tube with thick wall, particularly posterior to the insertion of stylet supports (
<figureCitation id="132D2A290C587566B410AFB3339BF953" box="[846,913,1696,1721]" captionStart="FIGURE 3" captionStartId="5.[151,250,1337,1359]" captionTargetBox="[151,1436,195,1316]" captionTargetId="figure@5.[151,1436,195,1316]" captionTargetPageId="5" captionText="FIGURE 3. Ramazzottius littoreus sp. nov. (holotype). Bucco-pharyngeal apparatus in dorsal view (holotype); arrow shows posterior band with four dorsal teeth (A). Bucco-pharyngeal apparatus in ventral view (paratype); arrow shows posterior band with four ventral teeth (only three visible) (B). Claws III (holotype); arrow shows main branch of external claw with robust accessory points, and arrowhead shows cuticle granulation (C). Claws IV (holotype); arrows show lunules (D). All scale bars = 10 µm." httpUri="https://zenodo.org/record/573633/files/figure.png" pageId="5" pageNumber="455">Fig. 3</figureCitation>
A). Stylet supports inserted on the buccal tube at 60.8% of its length. Ovoid pharyngeal bulb with large apophyses and two grain-like macroplacoids without constrictions. First macroplacoid slightly longer than the second (respectively: 3.4 and 3.3 µm long). Microplacoid and septulum absent (
<figureCitation id="132D2A290C587566B6D5AE1F31C7F8CE" box="[395,461,1804,1829]" captionStart="FIGURE 3" captionStartId="5.[151,250,1337,1359]" captionTargetBox="[151,1436,195,1316]" captionTargetId="figure@5.[151,1436,195,1316]" captionTargetPageId="5" captionText="FIGURE 3. Ramazzottius littoreus sp. nov. (holotype). Bucco-pharyngeal apparatus in dorsal view (holotype); arrow shows posterior band with four dorsal teeth (A). Bucco-pharyngeal apparatus in ventral view (paratype); arrow shows posterior band with four ventral teeth (only three visible) (B). Claws III (holotype); arrow shows main branch of external claw with robust accessory points, and arrowhead shows cuticle granulation (C). Claws IV (holotype); arrows show lunules (D). All scale bars = 10 µm." httpUri="https://zenodo.org/record/573633/files/figure.png" pageId="5" pageNumber="455">Fig. 3</figureCitation>
A). Placoid row, 8.1 µm long.
</paragraph>
<paragraph id="8BA936AC0C587566B799AE2333F5F867" blockId="5.[151,1436,1552,1932]" pageId="5" pageNumber="455">
Claws of 
<taxonomicName id="4C164D2F0C587566B66CAE2231CDF8A3" box="[306,455,1841,1864]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="455" phylum="Tardigrada" rank="genus">
<emphasis id="B962EABE0C587566B66CAE2231CDF8A3" box="[306,455,1841,1864]" italics="true" pageId="5" pageNumber="455">Ramazzottius</emphasis>
</taxonomicName>
<typeStatus id="54AD880E0C587566B690AE22320BF8A2" box="[462,513,1841,1865]" pageId="5" pageNumber="455">type</typeStatus>
, well developed (
<emphasis id="B962EABE0C587566B598AE2232D6F8A3" box="[710,732,1841,1864]" italics="true" pageId="5" pageNumber="455">pt</emphasis>
of posterior claw IV 
<emphasis id="B962EABE0C587566B495AE2233E6F8A3" box="[971,1004,1841,1864]" italics="true" pageId="5" pageNumber="455">ca.</emphasis>
85%), long main branches on external claws with robust accessory points (
<figureCitation id="132D2A290C587566B576AE423260F881" box="[552,618,1873,1898]" captionStart="FIGURE 3" captionStartId="5.[151,250,1337,1359]" captionTargetBox="[151,1436,195,1316]" captionTargetId="figure@5.[151,1436,195,1316]" captionTargetPageId="5" captionText="FIGURE 3. Ramazzottius littoreus sp. nov. (holotype). Bucco-pharyngeal apparatus in dorsal view (holotype); arrow shows posterior band with four dorsal teeth (A). Bucco-pharyngeal apparatus in ventral view (paratype); arrow shows posterior band with four ventral teeth (only three visible) (B). Claws III (holotype); arrow shows main branch of external claw with robust accessory points, and arrowhead shows cuticle granulation (C). Claws IV (holotype); arrows show lunules (D). All scale bars = 10 µm." httpUri="https://zenodo.org/record/573633/files/figure.png" pageId="5" pageNumber="455">Fig. 3</figureCitation>
C, arrow). Poorly visible lunules on legs I–III and well developed lunules on the hind legs (
<figureCitation id="132D2A290C587566B606AE603190F860" box="[344,410,1907,1932]" captionStart="FIGURE 3" captionStartId="5.[151,250,1337,1359]" captionTargetBox="[151,1436,195,1316]" captionTargetId="figure@5.[151,1436,195,1316]" captionTargetPageId="5" captionText="FIGURE 3. Ramazzottius littoreus sp. nov. (holotype). Bucco-pharyngeal apparatus in dorsal view (holotype); arrow shows posterior band with four dorsal teeth (A). Bucco-pharyngeal apparatus in ventral view (paratype); arrow shows posterior band with four ventral teeth (only three visible) (B). Claws III (holotype); arrow shows main branch of external claw with robust accessory points, and arrowhead shows cuticle granulation (C). Claws IV (holotype); arrows show lunules (D). All scale bars = 10 µm." httpUri="https://zenodo.org/record/573633/files/figure.png" pageId="5" pageNumber="455">Fig. 3</figureCitation>
D, arrows). Other cuticular thickenings on legs absent.
</paragraph>
<caption id="DF6966240C5B7565B7C9ADC834D7FAA6" httpUri="https://zenodo.org/record/573634/files/figure.png" pageId="6" pageNumber="456" targetBox="[151,1436,193,1222]" targetPageId="6">
<paragraph id="8BA936AC0C5B7565B7C9ADC834D7FAA6" blockId="6.[151,1436,1243,1357]" pageId="6" pageNumber="456">
<emphasis id="B962EABE0C5B7565B7C9ADC8311FFB1B" bold="true" box="[151,277,1243,1265]" pageId="6" pageNumber="456">FIGURE 4.</emphasis>
<taxonomicName id="4C164D2F0C5B7565B640ADCF320BFB1A" box="[286,513,1243,1265]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="456" phylum="Tardigrada" rank="species" species="littoreus" status="sp. nov.">
<emphasis id="B962EABE0C5B7565B640ADCF320BFB1A" box="[286,513,1243,1265]" italics="true" pageId="6" pageNumber="456">Ramazzottius littoreus</emphasis>
</taxonomicName>
<emphasis id="B962EABE0C5B7565B554ADCF3256FB1A" bold="true" box="[522,604,1244,1265]" pageId="6" pageNumber="456">
<taxonomicNameLabel id="A25157C50C5B7565B554ADCF3256FB1A" box="[522,604,1244,1265]" pageId="6" pageNumber="456" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Egg with low density of short processes; arrowheads show reticulated bases of processes (A). Egg with high density of processes; arrowheads show long funnel-shaped processes with enlarged tips (B). Egg with conical processes; arrows show reticulation on entire processes (C). Egg with conical processes; arrow shows reticulation only in the base of the process and arrowhead shows a process without reticulation (D). Scale bars = 10 µm.
</paragraph>
</caption>
<paragraph id="8BA936AC0C5B7565B799AC8B30D2F8F2" blockId="6.[151,1437,1432,1817]" pageId="6" pageNumber="456">
<emphasis id="B962EABE0C5B7565B799AC8B310BFA5A" bold="true" box="[199,257,1432,1457]" pageId="6" pageNumber="456">Eggs</emphasis>
(
<figureCitation id="132D2A290C5B7565B64CAC8B3157FA5A" box="[274,349,1432,1457]" captionStart="FIGURE 4" captionStartId="6.[151,250,1243,1265]" captionTargetBox="[151,1436,193,1222]" captionTargetId="figure@6.[151,1436,193,1222]" captionTargetPageId="6" captionText="FIGURE 4. Ramazzottius littoreus sp. nov. Egg with low density of short processes; arrowheads show reticulated bases of processes (A). Egg with high density of processes; arrowheads show long funnel-shaped processes with enlarged tips (B). Egg with conical processes; arrows show reticulation on entire processes (C). Egg with conical processes; arrow shows reticulation only in the base of the process and arrowhead shows a process without reticulation (D). Scale bars = 10 µm." httpUri="https://zenodo.org/record/573634/files/figure.png" pageId="6" pageNumber="456">Figs 4</figureCitation>
A, D; table 2): free laid, whitish, spherical (diameter 51.3–68.2 and 61.3–84.1µm without and with egg processes respectively). Egg processes are well spaced (24–28 processes on the circumference) and coneshaped with a bulbous base and a thinner apical point. The size and shape of the egg processes are variable between eggs and, sometimes also within the same egg (some egg processes have a longer, thin apical point, 
<figureCitation id="132D2A290C5B7565B21CAF173582F9F6" box="[1346,1416,1540,1565]" captionStart="FIGURE 4" captionStartId="6.[151,250,1243,1265]" captionTargetBox="[151,1436,193,1222]" captionTargetId="figure@6.[151,1436,193,1222]" captionTargetPageId="6" captionText="FIGURE 4. Ramazzottius littoreus sp. nov. Egg with low density of short processes; arrowheads show reticulated bases of processes (A). Egg with high density of processes; arrowheads show long funnel-shaped processes with enlarged tips (B). Egg with conical processes; arrows show reticulation on entire processes (C). Egg with conical processes; arrow shows reticulation only in the base of the process and arrowhead shows a process without reticulation (D). Scale bars = 10 µm." httpUri="https://zenodo.org/record/573634/files/figure.png" pageId="6" pageNumber="456">Fig. 4</figureCitation>
A arrowheads, and a few are funnel-shaped with enlarged tips, 
<figureCitation id="132D2A290C5B7565B42BAF3B33B7F9AA" box="[885,957,1576,1601]" captionStart="FIGURE 4" captionStartId="6.[151,250,1243,1265]" captionTargetBox="[151,1436,193,1222]" captionTargetId="figure@6.[151,1436,193,1222]" captionTargetPageId="6" captionText="FIGURE 4. Ramazzottius littoreus sp. nov. Egg with low density of short processes; arrowheads show reticulated bases of processes (A). Egg with high density of processes; arrowheads show long funnel-shaped processes with enlarged tips (B). Egg with conical processes; arrows show reticulation on entire processes (C). Egg with conical processes; arrow shows reticulation only in the base of the process and arrowhead shows a process without reticulation (D). Scale bars = 10 µm." httpUri="https://zenodo.org/record/573634/files/figure.png" pageId="6" pageNumber="456">Fig. 4</figureCitation>
B arrowheads). Length of egg processes 5.4–16.3 µm and with base diameter 1.8–6.2 µm. The bulbous base of egg processes is clearly reticulated (
<figureCitation id="132D2A290C5B7565B265AF5F3588F98E" box="[1339,1410,1612,1637]" captionStart="FIGURE 4" captionStartId="6.[151,250,1243,1265]" captionTargetBox="[151,1436,193,1222]" captionTargetId="figure@6.[151,1436,193,1222]" captionTargetPageId="6" captionText="FIGURE 4. Ramazzottius littoreus sp. nov. Egg with low density of short processes; arrowheads show reticulated bases of processes (A). Egg with high density of processes; arrowheads show long funnel-shaped processes with enlarged tips (B). Egg with conical processes; arrows show reticulation on entire processes (C). Egg with conical processes; arrow shows reticulation only in the base of the process and arrowhead shows a process without reticulation (D). Scale bars = 10 µm." httpUri="https://zenodo.org/record/573634/files/figure.png" pageId="6" pageNumber="456">Figs 4</figureCitation>
A, arrowheads, C, arrows). However, the egg process coverage of this reticulation is variable, and can be restricted to the base (
<figureCitation id="132D2A290C5B7565B65FAF87314EF946" box="[257,324,1684,1709]" captionStart="FIGURE 4" captionStartId="6.[151,250,1243,1265]" captionTargetBox="[151,1436,193,1222]" captionTargetId="figure@6.[151,1436,193,1222]" captionTargetPageId="6" captionText="FIGURE 4. Ramazzottius littoreus sp. nov. Egg with low density of short processes; arrowheads show reticulated bases of processes (A). Egg with high density of processes; arrowheads show long funnel-shaped processes with enlarged tips (B). Egg with conical processes; arrows show reticulation on entire processes (C). Egg with conical processes; arrow shows reticulation only in the base of the process and arrowhead shows a process without reticulation (D). Scale bars = 10 µm." httpUri="https://zenodo.org/record/573634/files/figure.png" pageId="6" pageNumber="456">Fig. 4</figureCitation>
D, arrow), in some egg processes is not visible (
<figureCitation id="132D2A290C5B7565B400AF8733ABF946" box="[862,929,1684,1709]" captionStart="FIGURE 4" captionStartId="6.[151,250,1243,1265]" captionTargetBox="[151,1436,193,1222]" captionTargetId="figure@6.[151,1436,193,1222]" captionTargetPageId="6" captionText="FIGURE 4. Ramazzottius littoreus sp. nov. Egg with low density of short processes; arrowheads show reticulated bases of processes (A). Egg with high density of processes; arrowheads show long funnel-shaped processes with enlarged tips (B). Egg with conical processes; arrows show reticulation on entire processes (C). Egg with conical processes; arrow shows reticulation only in the base of the process and arrowhead shows a process without reticulation (D). Scale bars = 10 µm." httpUri="https://zenodo.org/record/573634/files/figure.png" pageId="6" pageNumber="456">Fig. 4</figureCitation>
D, arrowhead), or can cover almost the entire processes, tip excluded (
<figureCitation id="132D2A290C5B7565B6F5AFAB31E5F93A" box="[427,495,1720,1745]" captionStart="FIGURE 4" captionStartId="6.[151,250,1243,1265]" captionTargetBox="[151,1436,193,1222]" captionTargetId="figure@6.[151,1436,193,1222]" captionTargetPageId="6" captionText="FIGURE 4. Ramazzottius littoreus sp. nov. Egg with low density of short processes; arrowheads show reticulated bases of processes (A). Egg with high density of processes; arrowheads show long funnel-shaped processes with enlarged tips (B). Egg with conical processes; arrows show reticulation on entire processes (C). Egg with conical processes; arrow shows reticulation only in the base of the process and arrowhead shows a process without reticulation (D). Scale bars = 10 µm." httpUri="https://zenodo.org/record/573634/files/figure.png" pageId="6" pageNumber="456">Fig. 4</figureCitation>
C, arrows). The inter-process egg surface is faintly dotted/granulated (in some eggs the inter-process surface appears smooth when observed with PCM but it is dotted/granulated when observed with DIC).
</paragraph>
<caption id="DF6966240C5A7564B7C9A98B310EFF04" pageId="7" pageNumber="457">
<paragraph id="8BA936AC0C5A7564B7C9A98B310EFF04" blockId="7.[151,1436,151,239]" pageId="7" pageNumber="457">
<emphasis id="B962EABE0C5A7564B7C9A98B3104FF44" bold="true" box="[151,270,152,175]" pageId="7" pageNumber="457">TABLE 2.</emphasis>
Measurements (in µm) of selected morphological structures for eggs of 
<taxonomicName id="4C164D2F0C5A7564B31BA98B353DFF44" box="[1093,1335,151,175]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="457" phylum="Tardigrada" rank="species" species="littoreus" status="sp. nov.">
<emphasis id="B962EABE0C5A7564B31BA98B353DFF44" box="[1093,1335,151,175]" italics="true" pageId="7" pageNumber="457">Ramazzottius littoreus</emphasis>
</taxonomicName>
<emphasis id="B962EABE0C5A7564B21CA98B3596FF44" bold="true" box="[1346,1436,152,175]" pageId="7" pageNumber="457">
<taxonomicNameLabel id="A25157C50C5A7564B21CA98B3596FF44" box="[1346,1436,152,175]" pageId="7" pageNumber="457" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(N—number of eggs/structures measured; SD—Standard deviation; Range refers to the smallest and largest measured structure).
</paragraph>
</caption>
<paragraph id="8BA936AC0C5A7564B7C1A819340AFE77" blockId="7.[159,1062,266,495]" pageId="7" pageNumber="457">Characteristics of the egg Mean ± SD (Range; N) Diameter without processes 59.7± 4.8 (51.3–68.2; 9) Diameter with processes 74.2 ± 6.7 (61.3–84.1; 9) Number processes on circumference 24.9 ± 1.5 (24–28; 9)</paragraph>
<paragraph id="8BA936AC0C5A7564B7C1A8A23311FD9B" blockId="7.[159,1062,266,495]" lastBlockId="7.[151,1436,564,2029]" pageId="7" pageNumber="457">
Processes base 4.2 ± 1.1 (1.8–6.2; 29) Processes length 9.9 ± 2.9 (5.4–16.3; 26) 
<emphasis id="B962EABE0C5A7564B799AB273131FDA6" bold="true" box="[199,315,564,589]" pageId="7" pageNumber="457">Remarks.</emphasis>
Measurements and statistics of structures obtained for specimens of 
<taxonomicName id="4C164D2F0C5A7564B323AB2634F5FDA7" box="[1149,1279,564,588]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="457" phylum="Tardigrada" rank="species" species="littoreus">
<emphasis id="B962EABE0C5A7564B323AB2634F5FDA7" box="[1149,1279,564,588]" italics="true" pageId="7" pageNumber="457">R. littoreus</emphasis>
</taxonomicName>
<emphasis id="B962EABE0C5A7564B253AB273553FDA6" bold="true" box="[1293,1369,564,589]" pageId="7" pageNumber="457">sp. no</emphasis>
<taxonomicName id="4C164D2F0C5A7564B207AB263596FDA7" box="[1369,1436,565,588]" pageId="7" pageNumber="457" rank="variety" variety="are">v. are</taxonomicName>
provided in table 1, and raw data as Supplementary Files.
</paragraph>
<paragraph id="8BA936AC0C5A7564B799AB6F31F8FD37" blockId="7.[151,1436,564,2029]" pageId="7" pageNumber="457">
Although not visible in the 
<typeStatus id="54AD880E0C5A7564B6A4AB6F3255FD7E" box="[506,607,636,661]" pageId="7" pageNumber="457" type="holotype">holotype</typeStatus>
, in 
<typeStatus id="54AD880E0C5A7564B5D9AB6E32F8FD7E" box="[647,754,637,661]" pageId="7" pageNumber="457" type="paratype">paratypes</typeStatus>
mounted in lateral position the anterior portion of the buccal tube is bent ventrally. A very slight second bend is also visible in the posterior part of the buccal tube, posterior to the insertion of stylet supports.
</paragraph>
<paragraph id="8BA936AC0C5A7564B799ABF4336EFC33" blockId="7.[151,1436,564,2029]" pageId="7" pageNumber="457">
According to 
<bibRefCitation id="EF874B5D0C5A7564B63CABFB3223FCEB" author="Baumman" box="[354,553,743,768]" pageId="7" pageNumber="457" refString="Baumman, H. (1966) Der Lebenslauf von Hypsibius (H.) oberhaeuseri Doyere (Tardigrada). Veroeffentlichungen aus den Ubersee-Museum, Bremen, Series A, 3, 245 - 258." type="journal article" year="1966">Baumman (1966)</bibRefCitation>
, 
<bibRefCitation id="EF874B5D0C5A7564B568ABF432D5FCEB" author="Biserov" box="[566,735,743,768]" pageId="7" pageNumber="457" refString="Biserov V. I. (1985) Hypsibius subanomalus sp. n. (Eutardigrada, Hypsibiidae) from the Astrakhan District. Zoologicheskii Zhurnal, 64 (1), 131 - 135." type="journal article" year="1985">Biserov (1985)</bibRefCitation>
and Rebecchi &amp; Bertolani (1988), some 
<taxonomicName id="4C164D2F0C5A7564B3F0ABFB3549FD14" box="[1198,1347,744,767]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="457" phylum="Tardigrada" rank="genus">
<emphasis id="B962EABE0C5A7564B3F0ABFB3549FD14" box="[1198,1347,744,767]" italics="true" pageId="7" pageNumber="457">Ramazzottius</emphasis>
</taxonomicName>
species can have lateral gibbosities on hind legs that are flattened in males and roundish or absent in females. Sex dimorphism in 
<taxonomicName id="4C164D2F0C5A7564B617AA2331D4FCAC" box="[329,478,816,839]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="457" phylum="Tardigrada" rank="genus">
<emphasis id="B962EABE0C5A7564B617AA2331D4FCAC" box="[329,478,816,839]" italics="true" pageId="7" pageNumber="457">Ramazzottius</emphasis>
</taxonomicName>
species can indicate an amphimictic mode of reproduction (Rebecchi &amp; Bertolani 1994; 
<bibRefCitation id="EF874B5D0C5A7564B7BBAA473180FC87" author="Dastych" box="[229,394,852,877]" pageId="7" pageNumber="457" refString="Dastych, H. (2011) Ramazzottius agannae sp. nov., a new tardigrade species from the nival zone of the Austrian Central Alps (Tardigrada). Entomologisch Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut, 15, 184, 237 - 253." type="journal article" year="2011">Dastych 2011</bibRefCitation>
). In the new species, lateral gibbosities on hind legs were exhibited by all examined specimens. However, distinctly formed gonads with male germ cells were not observed in the examined aceticorcein stained specimens of 
<taxonomicName id="4C164D2F0C5A7564B684AA8E325CFC5F" box="[474,598,924,948]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="457" phylum="Tardigrada" rank="species" species="littoreus" status="sp. nov.">
<emphasis id="B962EABE0C5A7564B684AA8E325CFC5F" box="[474,598,924,948]" italics="true" pageId="7" pageNumber="457">R. littoreus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25157C50C5A7564B501AA8F32B0FC5F" box="[607,698,924,949]" pageId="7" pageNumber="457" rank="species">
<emphasis id="B962EABE0C5A7564B501AA8F32B9FC5E" bold="true" box="[607,691,924,949]" pageId="7" pageNumber="457">sp. nov</emphasis>
.
</taxonomicNameLabel>
We were, therefore, unable to determine gender of specimens of the new species and thus the reproductive mode is inconclusive.
</paragraph>
<paragraph id="8BA936AC0C5A7564B799AAF63497FC17" blockId="7.[151,1436,564,2029]" box="[199,1181,996,1021]" pageId="7" pageNumber="457">
In the same sample 
<taxonomicName id="4C164D2F0C5A7564B6FBAAF6322AFC17" box="[421,544,996,1020]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="457" phylum="Tardigrada" rank="species" species="littoreus" status="sp. nov.">
<emphasis id="B962EABE0C5A7564B6FBAAF6322AFC17" box="[421,544,996,1020]" italics="true" pageId="7" pageNumber="457">R. littoreus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25157C50C5A7564B579AAF7328BFC17" box="[551,641,996,1021]" pageId="7" pageNumber="457" rank="species">
<emphasis id="B962EABE0C5A7564B579AAF73270FC16" bold="true" box="[551,634,996,1021]" pageId="7" pageNumber="457">sp. nov</emphasis>
.
</taxonomicNameLabel>
was found jointly with 
<emphasis id="B962EABE0C5A7564B4D2AAF73497FC17" box="[908,1181,996,1020]" italics="true" pageId="7" pageNumber="457">
<taxonomicName id="4C164D2F0C5A7564B4D2AAF73493FC17" box="[908,1177,996,1020]" class="Polychaeta" family="Aphroditidae" genus="Milnesium" kingdom="Animalia" order="Phyllodocida" pageId="7" pageNumber="457" phylum="Annelida" rank="species" species="tardigradum">Milnesium tardigradum</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C30C65270C5A7564B799AD14351EFBAE" pageId="7" pageNumber="457" type="etymology">
<paragraph id="8BA936AC0C5A7564B799AD14351EFBAE" blockId="7.[151,1436,564,2029]" pageId="7" pageNumber="457">
<emphasis id="B962EABE0C5A7564B799AD143144FBCB" bold="true" box="[199,334,1031,1056]" pageId="7" pageNumber="457">Etymology.</emphasis>
The name 
<taxonomicName id="4C164D2F0C5A7564B697AD1B322FFBCB" box="[457,549,1032,1056]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="457" phylum="Tardigrada" rank="species" species="littoreus">
<emphasis id="B962EABE0C5A7564B697AD1B322FFBCB" box="[457,549,1032,1056]" italics="true" pageId="7" pageNumber="457">littoreus</emphasis>
</taxonomicName>
refers to the environment where the species was found, the littoral, a derivation from the Latin noun 
<emphasis id="B962EABE0C5A7564B620AD3F31A4FBAF" box="[382,430,1068,1092]" italics="true" pageId="7" pageNumber="457">litus</emphasis>
(genitive 
<emphasis id="B962EABE0C5A7564B57EAD3F3269FBAF" box="[544,611,1068,1092]" italics="true" pageId="7" pageNumber="457">litoris</emphasis>
) meaning the shore; 
<emphasis id="B962EABE0C5A7564B411AD3F33CCFBAF" box="[847,966,1068,1092]" italics="true" pageId="7" pageNumber="457">
<taxonomicName id="4C164D2F0C5A7564B411AD3F33A1FBAF" box="[847,939,1068,1092]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="457" phylum="Tardigrada" rank="species" species="littoreus">littoreus</taxonomicName>
=
</emphasis>
inhabiting near the shoreline.
</paragraph>
</subSubSection>
<subSubSection id="C30C65270C5A756BB799AD5C323CFE8F" lastPageId="8" lastPageNumber="458" pageId="7" pageNumber="457" type="diagnosis">
<paragraph id="8BA936AC0C5A7564B799AD5C331DFB66" blockId="7.[151,1436,564,2029]" pageId="7" pageNumber="457">
<emphasis id="B962EABE0C5A7564B799AD5C31DAFB83" bold="true" box="[199,464,1103,1128]" pageId="7" pageNumber="457">Differential diagnosis.</emphasis>
Currently, 27 
<taxonomicName id="4C164D2F0C5A7564B527AD433304FB8C" box="[633,782,1104,1127]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="457" phylum="Tardigrada" rank="genus">
<emphasis id="B962EABE0C5A7564B527AD433304FB8C" box="[633,782,1104,1127]" italics="true" pageId="7" pageNumber="457">Ramazzottius</emphasis>
</taxonomicName>
species have been described. Some are very similar and can only be distinguished by a different egg morphology.
</paragraph>
<paragraph id="8BA936AC0C5A7564B799AD8432A7FAF6" blockId="7.[151,1436,564,2029]" pageId="7" pageNumber="457">
Of these 27, there are seven 
<taxonomicName id="4C164D2F0C5A7564B55AAD8B3293FB44" box="[516,665,1176,1199]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="457" phylum="Tardigrada" rank="genus">
<emphasis id="B962EABE0C5A7564B55AAD8B3293FB44" box="[516,665,1176,1199]" italics="true" pageId="7" pageNumber="457">Ramazzottius</emphasis>
</taxonomicName>
sp. that share the 
<taxonomicName id="4C164D2F0C5A7564B43AAD8A33D5FB5B" box="[868,991,1176,1200]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="457" phylum="Tardigrada" rank="species" species="littoreus" status="sp. nov.">
<emphasis id="B962EABE0C5A7564B43AAD8A33D5FB5B" box="[868,991,1176,1200]" italics="true" pageId="7" pageNumber="457">R. littoreus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25157C50C5A7564B4B8AD84344AFB5B" box="[998,1088,1175,1200]" pageId="7" pageNumber="457" rank="species">
<emphasis id="B962EABE0C5A7564B4B8AD843433FB5B" bold="true" box="[998,1081,1175,1200]" pageId="7" pageNumber="457">sp. nov</emphasis>
.
</taxonomicNameLabel>
characters of: absence of eyes, narrow buccal tube (less than 2.0 µm inner diameter), the dorsal cuticle without dorsal gibbosities and entirely sculptured with small tubercles (diameter about 2–3 µm), and by having medium sized claws (
<emphasis id="B962EABE0C5A7564B3E5ADF234DBFB13" box="[1211,1233,1249,1272]" italics="true" pageId="7" pageNumber="457">pt</emphasis>
posterior claw IV &lt;100) with accessory points on main branches.
</paragraph>
<paragraph id="8BA936AC0C5A7564B799AC3433ECFA46" blockId="7.[151,1436,564,2029]" pageId="7" pageNumber="457">
Of these seven, three species are clearly differentiate from those, like 
<taxonomicName id="4C164D2F0C5A7564B4BAAC3A346AFAAB" box="[996,1120,1320,1344]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="457" phylum="Tardigrada" rank="species" species="littoreus" status="sp. nov.">
<emphasis id="B962EABE0C5A7564B4BAAC3A346AFAAB" box="[996,1120,1320,1344]" italics="true" pageId="7" pageNumber="457">R. littoreus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25157C50C5A7564B337AC3434C9FAAB" box="[1129,1219,1319,1344]" pageId="7" pageNumber="457" rank="species">
<emphasis id="B962EABE0C5A7564B337AC3434B7FAAB" bold="true" box="[1129,1213,1319,1344]" pageId="7" pageNumber="457">sp. nov</emphasis>
.
</taxonomicNameLabel>
, that produce eggs with well separated, conical processes. 
<taxonomicName id="4C164D2F0C5A7564B53FAC5E339EFA8F" box="[609,916,1356,1380]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="457" phylum="Tardigrada" rank="species" species="oberhaeuseri">
<emphasis id="B962EABE0C5A7564B53FAC5E339EFA8F" box="[609,916,1356,1380]" italics="true" pageId="7" pageNumber="457">Ramazzottius oberhaeuseri</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C164D2F0C5A7564B48CAC5E3592FA8F" authority="Pilato, D'Urso &amp; Lisi, 2013" authorityName="Pilato, D'Urso &amp; Lisi" authorityYear="2013" box="[978,1432,1356,1381]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="457" phylum="Tardigrada" rank="species" species="lybicus">
<emphasis id="B962EABE0C5A7564B48CAC5E3449FA8F" box="[978,1091,1356,1380]" italics="true" pageId="7" pageNumber="457">R. lybicus</emphasis>
Pilato, D’Urso &amp; 
<bibRefCitation id="EF874B5D0C5A7564B27EAC5F3592FA8F" author="Lisi" box="[1312,1432,1356,1381]" pageId="7" pageNumber="457" refString="Pilato, G., D'Urso, V. &amp; Lisi, O. (2013) Ramazzottius thulini (Pilato, 1970) bona species and description of Ramazzottius lybicus sp. nov. (Eutardigrada, Ramazzottidae). Zootaxa, 3681 (3), 270 - 280." type="journal article" year="2013">Lisi, 2013</bibRefCitation>
</taxonomicName>
, produce eggs with hemispheric processes, and 
<taxonomicName id="4C164D2F0C5A7564B5FDAC62354AFA63" authority="Kaczmarek, Michalczyk &amp; Diduszko, 2006" authorityName="Kaczmarek, Michalczyk &amp; Diduszko" authorityYear="2006" box="[675,1344,1391,1416]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="457" phylum="Tardigrada" rank="species" species="bunikowskae">
<emphasis id="B962EABE0C5A7564B5FDAC623359FA63" box="[675,851,1392,1416]" italics="true" pageId="7" pageNumber="457">R. bunikowskae</emphasis>
<bibRefCitation id="EF874B5D0C5A7564B404AC7C354AFA63" author="Kaczmarek" box="[858,1344,1391,1416]" pageId="7" pageNumber="457" refString="Kaczmarek, L., Michalczyk, L. &amp; Diduszko, D. (2006) Ramazzottius bunikowskae, a new species of Tardigrada (Eutardigrada, Hypsibiidae) from Russia. Zootaxa, 1229, 49 - 57." type="journal article" year="2006">Kaczmarek, Michalczyk &amp; Diduszko, 2006</bibRefCitation>
</taxonomicName>
has egg processes with low enlarged bases (almost in contact) and long slender tips.
</paragraph>
<paragraph id="8BA936AC0C5A7564B799ACA43307F926" blockId="7.[151,1436,564,2029]" pageId="7" pageNumber="457">
The remaining four species have eggs with similar shaped egg processes but, contrary to the new species, 
<taxonomicName id="4C164D2F0C5A7564B2DAACAA325BFA1F" authority="Bertolani &amp; Rebecchi, 1988" authorityName="Bertolani &amp; Rebecchi" authorityYear="1988" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="457" phylum="Tardigrada" rank="species" species="tribulosus">
<emphasis id="B962EABE0C5A7564B2DAACAA310DFA1F" italics="true" pageId="7" pageNumber="457">R. tribulosus</emphasis>
Bertolani &amp; Rebecchi, 1988
</taxonomicName>
, 
<taxonomicName id="4C164D2F0C5A7564B503ACCE338EFA1F" authority="Biserov, 1997" authorityName="Biserov" authorityYear="1997" box="[605,900,1500,1525]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="457" phylum="Tardigrada" rank="species" species="andreevi">
<emphasis id="B962EABE0C5A7564B503ACCE32D5FA1F" box="[605,735,1500,1524]" italics="true" pageId="7" pageNumber="457">R. andreevi</emphasis>
<bibRefCitation id="EF874B5D0C5A7564B5B6ACCF338EFA1F" author="Biserov" box="[744,900,1500,1525]" pageId="7" pageNumber="457" refString="Biserov, V. I. (1997 / 98) Tardigrades of the Caucasus with a taxonomic analysis of the genus Ramazzottius (Parachela: Hypsibiidae). Zoologischer Anzeiger, 236, 139 - 159." type="book chapter" year="1997">Biserov, 1997</bibRefCitation>
</taxonomicName>
/98 and 
<taxonomicName id="4C164D2F0C5A7564B4BCACCE34FDFA1F" authority="Biserov, 1999" authorityName="Biserov" authorityYear="1999" box="[994,1271,1500,1525]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="457" phylum="Tardigrada" rank="species" species="rupeus">
<emphasis id="B962EABE0C5A7564B4BCACCE3444FA1F" box="[994,1102,1501,1524]" italics="true" pageId="7" pageNumber="457">R. rupeus</emphasis>
<bibRefCitation id="EF874B5D0C5A7564B309ACCF34FDFA1F" author="Biserov" box="[1111,1271,1500,1525]" pageId="7" pageNumber="457" refString="Biserov, V. I. (1999) A review of the Tardigrada from Novaya Zemlya, with descriptions of three new species, and an evaluation of the environment in this region. Zoologischer Anzeiger, 238, 169 - 182." type="book chapter" year="1999">Biserov, 1999</bibRefCitation>
</taxonomicName>
, lay eggs with smooth egg shells. Only 
<taxonomicName id="4C164D2F0C5A7564B6F1AF123322F9F3" authority="Binda &amp; Pilato, 1994" authorityName="Binda &amp; Pilato" authorityYear="1994" box="[431,808,1535,1560]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="457" phylum="Tardigrada" rank="species" species="horningi">
<emphasis id="B962EABE0C5A7564B6F1AF123225F9F3" box="[431,559,1536,1560]" italics="true" pageId="7" pageNumber="457">R. horningi</emphasis>
<bibRefCitation id="EF874B5D0C5A7564B569ACEC3322F9F3" author="Binda" box="[567,808,1535,1560]" pageId="7" pageNumber="457" refString="Binda, M. G. &amp; Pilato, G. (1994) Notizie sui Tardigradi delle Isole Hawaii con descrizione di due specie nuove, Animalia, 21 (1 / 3), 57 - 62." type="journal article" year="1994">Binda &amp; Pilato, 1994</bibRefCitation>
</taxonomicName>
lay eggs with dotted shells but, unlike the new species, the base of the egg processes is not sculptured. Adult specimens of 
<taxonomicName id="4C164D2F0C5A7564B4CCAF363418F9D7" box="[914,1042,1572,1596]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="457" phylum="Tardigrada" rank="species" species="horningi">
<emphasis id="B962EABE0C5A7564B4CCAF363418F9D7" box="[914,1042,1572,1596]" italics="true" pageId="7" pageNumber="457">R. horningi</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="4C164D2F0C5A7564B3F6AF363529F9D7" box="[1192,1315,1572,1596]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="457" phylum="Tardigrada" rank="species" species="littoreus">
<emphasis id="B962EABE0C5A7564B3F6AF363529F9D7" box="[1192,1315,1572,1596]" italics="true" pageId="7" pageNumber="457">R. littoreus</emphasis>
</taxonomicName>
<emphasis id="B962EABE0C5A7564B272AF373572F9D6" bold="true" box="[1324,1400,1572,1597]" pageId="7" pageNumber="457">sp nov</emphasis>
. in having unsculptured bands on dorsal side of the body, the cuticular granules are smaller (the largest have a diameter less than 1.9 µm; but up to 3.0 µm in the new species), and the placoid row and macroplacoids, particularly the first, are longer (
<emphasis id="B962EABE0C5A7564B609AF823167F943" box="[343,365,1681,1704]" italics="true" pageId="7" pageNumber="457">pt</emphasis>
of placoid row and first macroplacoid 
<emphasis id="B962EABE0C5A7564B475AF833356F943" box="[811,860,1680,1704]" italics="true" pageId="7" pageNumber="457">23.5</emphasis>
and 
<emphasis id="B962EABE0C5A7564B4CBAF8333CCF943" box="[917,966,1680,1704]" italics="true" pageId="7" pageNumber="457">12.9</emphasis>
respectively in a 233 µm specimen of 
<taxonomicName id="4C164D2F0C5A7564B2DAAF8230FDF927" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="457" phylum="Tardigrada" rank="species" species="horningi">
<emphasis id="B962EABE0C5A7564B2DAAF8230FDF927" italics="true" pageId="7" pageNumber="457">R. horningi</emphasis>
</taxonomicName>
; 
<emphasis id="B962EABE0C5A7564B658AFA63199F927" box="[262,403,1716,1740]" italics="true" pageId="7" pageNumber="457">pt 19.1–23.5</emphasis>
and 
<quantity id="4CEE9B490C5A7564B697AFA7324CF926" box="[457,582,1716,1741]" metricMagnitude="-1" metricUnit="m" metricValue="2.5654" metricValueMax="3.0226" metricValueMin="2.1082" pageId="7" pageNumber="457" unit="in" value="10.1" valueMax="11.9" valueMin="8.3">
<emphasis id="B962EABE0C5A7564B697AFA73223F927" box="[457,553,1716,1740]" italics="true" pageId="7" pageNumber="457">8.3–11.9</emphasis>
in
</quantity>
the new species).
</paragraph>
<paragraph id="8BA936AC0C5A756BB799AFCB3255FF3E" blockId="7.[151,1436,564,2029]" lastBlockId="8.[151,1436,151,213]" lastPageId="8" lastPageNumber="458" pageId="7" pageNumber="457">
As the dotted/granulated egg surface is faint and appears smooth in some 
<taxonomicName id="4C164D2F0C5A7564B351AFCA3486F91B" box="[1039,1164,1752,1776]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="457" phylum="Tardigrada" rank="species" species="littoreus" status="sp. nov.">
<emphasis id="B962EABE0C5A7564B351AFCA3486F91B" box="[1039,1164,1752,1776]" italics="true" pageId="7" pageNumber="457">R. littoreus</emphasis>
</taxonomicName>
<emphasis id="B962EABE0C5A7564B3CAAFC434E5F91B" bold="true" box="[1172,1263,1751,1776]" pageId="7" pageNumber="457">
<taxonomicNameLabel id="A25157C50C5A7564B3CAAFC434E5F91B" box="[1172,1263,1751,1776]" pageId="7" pageNumber="457" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
eggs, we think opportune to distinguish the new species from 
<emphasis id="B962EABE0C5A7564B5E7AFEE3358F8FF" box="[697,850,1788,1812]" italics="true" pageId="7" pageNumber="457">
<taxonomicName id="4C164D2F0C5A7564B5E7AFEE3345F8FF" box="[697,847,1788,1812]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="457" phylum="Tardigrada" rank="species" species="tribulosus">R. tribulosus</taxonomicName>
.
</emphasis>
The presence of conical egg processes with bases sculptured with a reticular pattern, puts 
<taxonomicName id="4C164D2F0C5A7564B530AE323308F8D3" box="[622,770,1824,1848]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="457" phylum="Tardigrada" rank="species" species="tribulosus">
<emphasis id="B962EABE0C5A7564B530AE323308F8D3" box="[622,770,1824,1848]" italics="true" pageId="7" pageNumber="457">R. tribulosus</emphasis>
</taxonomicName>
as the most similar species to 
<taxonomicName id="4C164D2F0C5A7564B328AE3234FCF8D3" box="[1142,1270,1824,1848]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="457" phylum="Tardigrada" rank="species" species="littoreus" status="sp. nov.">
<emphasis id="B962EABE0C5A7564B328AE3234FCF8D3" box="[1142,1270,1824,1848]" italics="true" pageId="7" pageNumber="457">R. littoreus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25157C50C5A7564B25FAE0C3555F8DC" box="[1281,1375,1823,1848]" pageId="7" pageNumber="457" rank="species">
<emphasis id="B962EABE0C5A7564B25FAE0C3552F8D3" bold="true" box="[1281,1368,1823,1848]" pageId="7" pageNumber="457">sp. nov</emphasis>
.
</taxonomicNameLabel>
This character was not referred to in the original description of 
<taxonomicName id="4C164D2F0C5A7564B440AE5633A7F8B7" box="[798,941,1860,1884]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="457" phylum="Tardigrada" rank="species" species="tribulosus">
<emphasis id="B962EABE0C5A7564B440AE5633A7F8B7" box="[798,941,1860,1884]" italics="true" pageId="7" pageNumber="457">R. tribulosus</emphasis>
</taxonomicName>
, despite being obvious in the figure (see fig. 4, page 368, 
<emphasis id="B962EABE0C5A7564B66AAE7B3140F894" box="[308,330,1896,1919]" italics="true" pageId="7" pageNumber="457">in</emphasis>
Bertolani &amp; Rebecchi 1988). Furthermore, 
<bibRefCitation id="EF874B5D0C5A7564B40EAE74340EF86B" author="Dastych" box="[848,1028,1895,1920]" pageId="7" pageNumber="457" refString="Dastych, H. (1993) A new genus and four new species of semiterrestrial water-bears from South Africa (Tardigrada). Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut, 90, 175 - 186." type="journal article" year="1993">Dastych (1993)</bibRefCitation>
, noted in his description of the 
<taxonomicName id="4C164D2F0C5A7564B2DAAE7A310DF84F" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="457" phylum="Tardigrada" rank="species" species="tribulosus">
<emphasis id="B962EABE0C5A7564B2DAAE7A310DF84F" italics="true" pageId="7" pageNumber="457">R. tribulosus</emphasis>
</taxonomicName>
egg processes as having “irregular sculptures internally”. However, adult specimens of the two species can be distinguished by the different buccal armature (posterior band with four ventral and four dorsal teeth in the new species, and six ventral and six dorsal teeth in 
<taxonomicName id="4C164D2F0C5A7564B58AAEC6336EF807" box="[724,868,2004,2028]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="457" phylum="Tardigrada" rank="species" species="tribulosus">
<emphasis id="B962EABE0C5A7564B58AAEC6336EF807" box="[724,868,2004,2028]" italics="true" pageId="7" pageNumber="457">R. tribulosus</emphasis>
</taxonomicName>
), and by the unconstricted first macroplacoids and undivided elliptical organs in the new species (
<taxonomicName id="4C164D2F0C55756BB5E1A98A3358FF5B" box="[703,850,152,176]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="458" phylum="Tardigrada" rank="species" species="tribulosus">
<emphasis id="B962EABE0C55756BB5E1A98A3358FF5B" box="[703,850,152,176]" italics="true" pageId="8" pageNumber="458">R. tribulosus</emphasis>
</taxonomicName>
has macroplacoids with constriction and elliptical organs divided by a longitudinal sulcus).
</paragraph>
<paragraph id="8BA936AC0C55756BB7C9A834323CFE8F" blockId="8.[151,885,295,392]" pageId="8" pageNumber="458">
<emphasis id="B962EABE0C55756BB7C9A834313EFEAB" bold="true" box="[151,308,295,320]" pageId="8" pageNumber="458">Superfamily:</emphasis>
Macrobiotoidea 
<bibRefCitation id="EF874B5D0C55756BB6ADA834328EFEAB" author="Thulin" box="[499,644,295,320]" pageId="8" pageNumber="458" refString="Thulin, G. (1928) Uber die Phylogenie und das System der Tardigraden. Hereditas, 11, 207 - 266. https: // doi. org / 10.1111 / j. 1601 - 5223.1928. tb 02488. x" type="journal article" year="1928">
Thulin, 
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</bibRefCitation>
in 
<bibRefCitation id="EF874B5D0C55756BB5F6A834337FFEAB" author="Marley" box="[680,885,295,320]" pageId="8" pageNumber="458" refString="Marley, N. J., McInnes, S. &amp; Sands, C. J. (2011) Phylum Tardigrada: A re-evaluation of the Parachela. Zootaxa, 2819, 51 - 64." type="journal article" year="2011">
Marley 
<emphasis id="B962EABE0C55756BB5A1A83A333BFEAB" box="[767,817,296,320]" italics="true" pageId="8" pageNumber="458">et al</emphasis>
. 2011
</bibRefCitation>
<emphasis id="B962EABE0C55756BB7C9A85F30FEFE8E" bold="true" box="[151,244,332,357]" pageId="8" pageNumber="458">Family:</emphasis>
<taxonomicName id="4C164D2F0C55756BB7A5A85F323CFE8F" authority="Thulin, 1928" authorityName="Thulin" authorityYear="1928" box="[251,566,332,357]" class="Eutardigrada" family="Macrobiotidae" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="458" phylum="Tardigrada" rank="family">
Macrobiotidae 
<bibRefCitation id="EF874B5D0C55756BB6F8A85F323CFE8F" author="Thulin" box="[422,566,332,357]" pageId="8" pageNumber="458" refString="Thulin, G. (1928) Uber die Phylogenie und das System der Tardigraden. Hereditas, 11, 207 - 266. https: // doi. org / 10.1111 / j. 1601 - 5223.1928. tb 02488. x" type="journal article" year="1928">Thulin, 1928</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C30C65270C55756BB7C9A87C328BFE63" box="[151,641,367,392]" pageId="8" pageNumber="458" type="description">
<paragraph id="8BA936AC0C55756BB7C9A87C328BFE63" blockId="8.[151,885,295,392]" box="[151,641,367,392]" pageId="8" pageNumber="458">
<emphasis id="B962EABE0C55756BB7C9A87C30E6FE63" bold="true" box="[151,236,367,392]" pageId="8" pageNumber="458">Genus:</emphasis>
<taxonomicName id="4C164D2F0C55756BB7ADA863328BFE63" authority="C.A.S. Schultze, 1834" authorityName="C.A.S. Schultze" authorityYear="1834" box="[243,641,367,392]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="458" phylum="Tardigrada" rank="genus">
<emphasis id="B962EABE0C55756BB7ADA863318BFE63" box="[243,385,368,392]" italics="true" pageId="8" pageNumber="458">Macrobiotus</emphasis>
<bibRefCitation id="EF874B5D0C55756BB6D6A863328BFE63" author="Schultze" box="[392,641,367,392]" pageId="8" pageNumber="458" refString="Schultze, C. A. S. (1834) Macrobiotus hufelandi animal e crustaceorum classe novum reviviscendi post diuturnam asphyxiam et ariditatem potens, etc. 8 Seiten. C. Curths, Berlin, 6 pp." type="book" year="1834">C.A.S. Schultze, 1834</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>