treatments-xml/data/03/8F/FF/038FFF5A3843FFA2FF2CFC8943A7FE71.xml
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<document id="6530DDAC7D8FEA09EA49A1BB7FC99196" ID-CLB-Dataset="55057" ID-DOI="10.11646/zootaxa.5244.3.2" ID-GBIF-Dataset="512007ce-c83f-4a27-adcb-cb8b6325085f" ID-ISSN="1175-5326" ID-Zenodo-Dep="7656249" ID-ZooBank="E5A4C931-F8F4-41A0-9270-209C09272B43" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1676882713289" checkinUser="plazi" docAuthor="Hoskin, Conrad J." docDate="2023" docId="038FFF5A3843FFA2FF2CFC8943A7FE71" docLanguage="en" docName="zootaxa.5244.3.2.pdf" docOrigin="Zootaxa 5244 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5244.3.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Phyllurus fimbriatus Hoskin 2023, sp. nov." docType="treatment" docVersion="7" lastPageNumber="241" masterDocId="FFB687223841FFAAFFBBFFF64211FFA0" masterDocTitle="A new species of Phyllurus leaf-tailed gecko (Lacertilia: Carphodactylidae) from Scawfell Island, mid-east Queensland, Australia" masterLastPageNumber="243" masterPageNumber="233" pageNumber="235" updateTime="1699475451312" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="BE1A4B859BC5B9FAF690C8BADC75D573">A new species of Phyllurus leaf-tailed gecko (Lacertilia: Carphodactylidae) from Scawfell Island, mid-east Queensland, Australia</mods:title>
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<mods:namePart id="527DC80B00161365FE656306FC787580">Hoskin, Conrad J.</mods:namePart>
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<mods:date id="98A4D1C862F0BB8B4901F3D9A6586750">2023</mods:date>
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<mods:number id="CCD4B1E0E5C9D233D16FCE1215E8348D">2023-02-20</mods:number>
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<mods:identifier id="BEF6248BFA73967E238B753DA0591B07" type="DOI">10.11646/zootaxa.5244.3.2</mods:identifier>
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<treatment id="038FFF5A3843FFA2FF2CFC8943A7FE71" ID-DOI="http://doi.org/10.5281/zenodo.7671266" ID-GBIF-Taxon="209342051" ID-Zenodo-Dep="7671266" LSID="urn:lsid:plazi:treatment:038FFF5A3843FFA2FF2CFC8943A7FE71" httpUri="http://treatment.plazi.org/id/038FFF5A3843FFA2FF2CFC8943A7FE71" lastPageId="8" lastPageNumber="241" pageId="2" pageNumber="235">
<subSubSection id="C33C1DC73843FFA8FF2CFC8943E9FC3A" box="[151,504,895,922]" pageId="2" pageNumber="235" type="nomenclature">
<paragraph id="8B994E4C3843FFA8FF2CFC8943E9FC3A" blockId="2.[151,534,895,994]" box="[151,504,895,922]" pageId="2" pageNumber="235">
<heading id="D0D1F9203843FFA8FF2CFC8943E9FC3A" bold="true" box="[151,504,895,922]" fontSize="11" level="1" pageId="2" pageNumber="235" reason="1">
<emphasis id="B952925E3843FFA8FF2CFC8943E9FC3A" bold="true" box="[151,504,895,922]" pageId="2" pageNumber="235">
<taxonomicName id="4C2635CF3843FFA8FF2CFC894385FC3A" authority="Hoskin, 2023" authorityName="Hoskin" authorityYear="2023" box="[151,404,895,922]" class="Reptilia" family="Carphodactylidae" genus="Phyllurus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="235" phylum="Chordata" rank="species" species="fimbriatus" status="sp. nov.">
<emphasis id="B952925E3843FFA8FF2CFC894385FC3A" bold="true" box="[151,404,895,922]" italics="true" pageId="2" pageNumber="235">Phyllurus fimbriatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2612F253843FFA8FE21FC7643E9FC3A" box="[410,504,896,922]" pageId="2" pageNumber="235" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C33C1DC73843FFA8FF2CFC524007FC1E" box="[151,534,932,958]" pageId="2" pageNumber="235" type="vernacular_names">
<paragraph id="8B994E4C3843FFA8FF2CFC524007FC1E" blockId="2.[151,534,895,994]" box="[151,534,932,958]" pageId="2" pageNumber="235">Scawfell Island Leaf-tailed Gecko</paragraph>
</subSubSection>
<subSubSection id="C33C1DC73843FFA8FF2CFC3E433EFC42" box="[151,303,968,994]" pageId="2" pageNumber="235" type="description">
<paragraph id="8B994E4C3843FFA8FF2CFC3E433EFC42" blockId="2.[151,534,895,994]" box="[151,303,968,994]" pageId="2" pageNumber="235">
(
<figureCitation id="131D52C93843FFA8FF24FC3E431DFC42" box="[159,268,968,994]" captionStart="FIGURE 2" captionStartId="4.[151,250,1722,1746]" captionTargetBox="[151,1436,489,1698]" captionTargetId="figure-180@4.[151,1436,489,1698]" captionTargetPageId="4" captionText="FIGURE 2. Holotype of P. fimbriatus sp. nov. (QM J97578). Photos: (A) Edward Evans; (B) Conrad Hoskin." figureDoi="http://doi.org/10.5281/zenodo.7656253" httpUri="https://zenodo.org/record/7656253/files/figure.png" pageId="2" pageNumber="235">Figures 2</figureCitation>
,
<figureCitation id="131D52C93843FFA8FEA3FC3E4337FC42" box="[280,294,968,994]" captionStart="FIGURE 3" captionStartId="5.[151,250,1236,1260]" captionTargetBox="[151,1435,381,1212]" captionTargetId="figure-123@5.[151,1435,381,1212]" captionTargetPageId="5" captionText="FIGURE 3. Phyllurus fimbriatus sp. nov. in life: (A) adult female with original tail; (B) juvenile with original tail; (C) adult female with regenerated tail; (D) adult male of unusual colouration, with regenerated tail. Photos: Conrad Hoskin." figureDoi="http://doi.org/10.5281/zenodo.7656255" httpUri="https://zenodo.org/record/7656255/files/figure.png" pageId="2" pageNumber="235">3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C33C1DC73843FFA8FF2CFBFC4034FB31" pageId="2" pageNumber="235" type="materials_examined">
<paragraph id="8B994E4C3843FFA8FF2CFBFC4034FB31" blockId="2.[151,1437,1034,2033]" pageId="2" pageNumber="235">
<emphasis id="B952925E3843FFA8FF2CFBFC4392FB84" bold="true" box="[151,387,1034,1060]" pageId="2" pageNumber="235">Material examined.</emphasis>
<materialsCitation id="3B4E44113843FFA8FE37FBFD4667FBE9" ID-GBIF-Occurrence="4036166304" collectingDate="2021-11-18" collectionCode="QM" collectorName="C. J. Hoskin; E. Carmichael &amp; E. Evans" elevation="200" latitude="-20.8743" location="Scawfell Island" longLatPrecision="7" longitude="149.6102" pageId="2" pageNumber="235" specimenCode="J97578" specimenCount="1" typeStatus="holotype">
<emphasis id="B952925E3843FFA8FE37FBFD4011FB85" bold="true" box="[396,512,1035,1061]" pageId="2" pageNumber="235">
<typeStatus id="549DF0EE3843FFA8FE37FBFD43EAFB85" box="[396,507,1035,1061]" pageId="2" pageNumber="235" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="ED37D6893843FFA8FDB2FBFD4024FB85" box="[521,565,1035,1061]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="2" pageNumber="235" type="Museum">QM</collectionCode>
<specimenCode id="DB80E6373843FFA8FD84FBFC4081FB85" box="[575,656,1034,1061]" collectionCode="QM" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="2" pageNumber="235" type="Museum">J97578</specimenCode>
, adult female, original tail,
<location id="8EF918973843FFA8FC7CFBFD4667FB85" LSID="urn:lsid:plazi:treatment:038FFF5A3843FFA2FF2CFC8943A7FE71:8EF918973843FFA8FC7CFBFD4667FB85" box="[967,1142,1035,1061]" latitude="-20.8743" longLatPrecision="7" longitude="149.6102" name="Scawfell Island" pageId="2" pageNumber="235">Scawfell Island</location>
(
<geoCoordinate id="EE12288B3843FFA8FB33FBFD4710FB85" box="[1160,1281,1035,1061]" degrees="20.8743" direction="south" orientation="latitude" pageId="2" pageNumber="235" precision="5" value="-20.8743">20.8743°S</geoCoordinate>
,
<geoCoordinate id="EE12288B3843FFA8FAB4FBFD4789FB85" box="[1295,1432,1035,1061]" degrees="149.6102" direction="east" orientation="longitude" pageId="2" pageNumber="235" precision="5" value="149.6102">149.6102°E</geoCoordinate>
;
<elevation id="000BA97F3843FFA8FF2CFBD94331FBE9" box="[151,288,1071,1097]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="235" unit="m" value="200.0">
<quantity id="4CDEE3A93843FFA8FF2CFBD942F0FBE9" box="[151,225,1071,1097]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="235" unit="m" value="200.0">200 m</quantity>
a.s.l.
</elevation>
),
<date id="FF98688C3843FFA8FE81FBD9400AFBE9" box="[314,539,1071,1097]" pageId="2" pageNumber="235" value="2021-11-18">
<collectingDate id="EFDC91643843FFA8FE81FBD9400AFBE9" box="[314,539,1071,1097]" pageId="2" pageNumber="235" value="2021-11-18">18 November 2021</collectingDate>
</date>
,
<collectorName id="26D32B9A3843FFA8FD91FBD940ACFBE9" box="[554,701,1071,1097]" pageId="2" pageNumber="235">C. J. Hoskin</collectorName>
,
<collectorName id="26D32B9A3843FFA8FD77FBD9417EFBE9" box="[716,879,1071,1097]" pageId="2" pageNumber="235">E. Carmichael</collectorName>
&amp;
<collectorName id="26D32B9A3843FFA8FC22FBD94612FBE9" box="[921,1027,1071,1097]" pageId="2" pageNumber="235">E. Evans</collectorName>
;
<figureCitation id="131D52C93843FFA8FBA9FBD94667FBE9" box="[1042,1142,1071,1097]" captionStart="FIGURE 2" captionStartId="4.[151,250,1722,1746]" captionTargetBox="[151,1436,489,1698]" captionTargetId="figure-180@4.[151,1436,489,1698]" captionTargetPageId="4" captionText="FIGURE 2. Holotype of P. fimbriatus sp. nov. (QM J97578). Photos: (A) Edward Evans; (B) Conrad Hoskin." figureDoi="http://doi.org/10.5281/zenodo.7656253" httpUri="https://zenodo.org/record/7656253/files/figure.png" pageId="2" pageNumber="235">Figure 2</figureCitation>
</materialsCitation>
.
<materialsCitation id="3B4E44113843FFA8FB3FFBD943CFFBCD" ID-GBIF-Occurrence="4036166303" collectionCode="QM" pageId="2" pageNumber="235" specimenCode="J97576" specimenCount="1" typeStatus="paratype">
<emphasis id="B952925E3843FFA8FB3FFBD94712FBE9" bold="true" box="[1156,1283,1071,1097]" pageId="2" pageNumber="235">
<typeStatus id="549DF0EE3843FFA8FB3FFBD946EFFBE9" box="[1156,1278,1071,1097]" pageId="2" pageNumber="235" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectionCode id="ED37D6893843FFA8FAB6FBD9472BFBE9" box="[1293,1338,1071,1097]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="2" pageNumber="235" type="Museum">QM</collectionCode>
<specimenCode id="DB80E6373843FFA8FAFFFBD84789FBE9" box="[1348,1432,1070,1097]" collectionCode="QM" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="2" pageNumber="235" type="Museum">J97576</specimenCode>
, subadult female, original tail
</materialsCitation>
;
<materialsCitation id="3B4E44113843FFA8FE51FBA541BDFBCD" ID-GBIF-Occurrence="4036166302" box="[490,940,1106,1133]" collectionCode="QM" pageId="2" pageNumber="235" specimenCode="J97577" specimenCount="1" typeStatus="paratype">
<collectionCode id="ED37D6893843FFA8FE51FBA54006FBCD" box="[490,535,1107,1133]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="2" pageNumber="235" type="Museum">QM</collectionCode>
<specimenCode id="DB80E6373843FFA8FDA5FBA4407EFBCD" box="[542,623,1106,1133]" collectionCode="QM" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="2" pageNumber="235" type="Museum">J97577</specimenCode>
adult male, regenerated tail
</materialsCitation>
;
<materialsCitation id="3B4E44113843FFA8FC03FBA54033FB31" ID-GBIF-Occurrence="4036166301" collectionCode="QM" pageId="2" pageNumber="235" specimenCode="J97579" specimenCount="1" typeStatus="holotype">
<collectionCode id="ED37D6893843FFA8FC03FBA541F4FBCD" box="[952,997,1107,1133]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="2" pageNumber="235" type="Museum">QM</collectionCode>
<specimenCode id="DB80E6373843FFA8FC56FBA44650FBCD" box="[1005,1089,1106,1133]" collectionCode="QM" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="2" pageNumber="235" type="Museum">J97579</specimenCode>
, adult female, regenerated tail; collection details same as
<typeStatus id="549DF0EE3843FFA8FE07FB814033FB31" box="[444,546,1143,1169]" pageId="2" pageNumber="235" type="holotype">holotype</typeStatus>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33C1DC73843FFA8FF7CFB6C46CBFA81" pageId="2" pageNumber="235" type="diagnosis">
<paragraph id="8B994E4C3843FFA8FF7CFB6C46CBFA81" blockId="2.[151,1437,1034,2033]" pageId="2" pageNumber="235">
<emphasis id="B952925E3843FFA8FF7CFB6C4353FB14" bold="true" box="[199,322,1178,1204]" pageId="2" pageNumber="235">Diagnosis.</emphasis>
Distinguished from congeners by the following characters: large size (SVL to
<quantity id="4CDEE3A93843FFA8FB02FB6D470BFB15" box="[1209,1306,1179,1205]" metricMagnitude="-1" metricUnit="m" metricValue="1.13" pageId="2" pageNumber="235" unit="mm" value="113.0">113 mm</quantity>
); obviously flared original and regenerated tail; rostral scale partially divided by a single groove; body and tail surfaces with small tubercles, with larger spinose tubercles largely restricted to margins of original tail; anterior-most white band on original tail V-shaped or notched, and spanning full width of tail; ventral surfaces immaculate.
</paragraph>
</subSubSection>
<subSubSection id="C33C1DC73843FFA8FF7CFADD4040FAC9" pageId="2" pageNumber="235" type="etymology">
<paragraph id="8B994E4C3843FFA8FF7CFADD4040FAC9" blockId="2.[151,1437,1034,2033]" pageId="2" pageNumber="235">
<emphasis id="B952925E3843FFA8FF7CFADD435FFAE5" bold="true" box="[199,334,1323,1349]" pageId="2" pageNumber="235">Etymology.</emphasis>
The species name
<taxonomicName id="4C2635CF3843FFA8FDA5FADD4080FAE5" authorityName="Hoskin" authorityYear="2023" box="[542,657,1323,1349]" class="Reptilia" family="Carphodactylidae" genus="Phyllurus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="235" phylum="Chordata" rank="species" species="fimbriatus">
<emphasis id="B952925E3843FFA8FDA5FADD4080FAE5" box="[542,657,1323,1349]" italics="true" pageId="2" pageNumber="235">fimbriatus</emphasis>
</taxonomicName>
is Latin for fringed, referring to the restriction of spines on the original tail to a fringe along the outer edge.
</paragraph>
</subSubSection>
<subSubSection id="C33C1DC73843FFAEFF7CFA8543B1FE71" lastPageId="4" lastPageNumber="237" pageId="2" pageNumber="235" type="description">
<paragraph id="8B994E4C3843FFA9FF7CFA8546BBFA39" blockId="2.[151,1437,1034,2033]" lastBlockId="3.[151,1437,150,2043]" lastPageId="3" lastPageNumber="236" pageId="2" pageNumber="235">
<emphasis id="B952925E3843FFA8FF7CFA854022FA2D" bold="true" box="[199,563,1394,1421]" italics="true" pageId="2" pageNumber="235">Measurements and scale counts.</emphasis>
All measurements and scale counts for the
<typeStatus id="549DF0EE3843FFA8FBB1FA854678FA2D" box="[1034,1129,1395,1421]" pageId="2" pageNumber="235" type="holotype">holotype</typeStatus>
and
<typeStatus id="549DF0EE3843FFA8FB25FA854717FA2D" box="[1182,1286,1395,1421]" pageId="2" pageNumber="235" type="paratype">paratypes</typeStatus>
are presented in
<tableCitation id="C6A47BF73843FFA8FF0DFA61431AFA11" box="[182,267,1431,1457]" captionStart="TABLE 1" captionStartId="6.[152,249,150,176]" captionTargetPageId="6" captionText="TABLE 1. Morphological measurements, proportions, and scale counts for the type specimens, and SVL data from individuals measured in the field. All measurements are in mm. Field data is only for individuals deemed to be mature adults (see text), is split by sex, and is presented as average followed by range in brackets." httpUri="http://table.plazi.org/id/DF591EC43847FFACFF23FF6046ACFF59" pageId="2" pageNumber="235" tableUuid="DF591EC43847FFACFF23FF6046ACFF59">Table 1</tableCitation>
.
<emphasis id="B952925E3843FFA8FEA0FA614054FA11" bold="true" box="[283,581,1430,1457]" italics="true" pageId="2" pageNumber="235">Description of type series.</emphasis>
SVL = average
<quantity id="4CDEE3A93843FFA8FCBEFA614166FA11" box="[773,887,1431,1457]" metricMagnitude="-1" metricUnit="m" metricValue="1.017" pageId="2" pageNumber="235" unit="mm" value="101.7">101.7 mm</quantity>
(90.7110.7). Proportions (average, followed by range in brackets): AG/SVL = 0.47 (0.450.48), FL/SVL = 0.20 (0.190.20), LHL/SVL = 0.23 (0.220.24), L1/SVL = 0.46 (0.440.48), L2/SVL = 0.53 (0.510.54), NL/SVL = 0.19 (0.190.20), NW/SVL = 0.11 (0.100.12), HL/SVL = 0.27 (0.270.28), HW/SVL = 0.22 (0.220.23), HD/SVL = 0.110 (
<geoCoordinate id="EE12288B3843FFA8FCD6F9F441BDF9BD" box="[877,940,1538,1565]" degrees="0.107" direction="north" orientation="latitude" pageId="2" pageNumber="235" precision="55" value="0.107">0.107</geoCoordinate>
<geoCoordinate id="EE12288B3843FFA8FC16F9F441EBF9BD" box="[941,1018,1538,1565]" degrees="0.115" direction="west" orientation="longitude" pageId="2" pageNumber="235" precision="55" value="-0.115">0.115</geoCoordinate>
), SL/SVL = 0.124 (
<geoCoordinate id="EE12288B3843FFA8FB6BF9F54701F9BD" box="[1232,1296,1539,1565]" degrees="0.118" direction="north" orientation="latitude" pageId="2" pageNumber="235" precision="55" value="0.118">0.118</geoCoordinate>
<geoCoordinate id="EE12288B3843FFA8FAABF9F5474CF9BD" box="[1296,1373,1539,1565]" degrees="0.131" direction="west" orientation="longitude" pageId="2" pageNumber="235" precision="55" value="-0.131">0.131</geoCoordinate>
), ED/ SVL = 0.061 (
<geoCoordinate id="EE12288B3843FFA8FE8CF9D04367F9E1" box="[311,374,1574,1601]" degrees="0.057" direction="north" orientation="latitude" pageId="2" pageNumber="235" precision="55" value="0.057">0.057</geoCoordinate>
<geoCoordinate id="EE12288B3843FFA8FECDF9D043D4F9E1" box="[374,453,1574,1601]" degrees="0.064" direction="west" orientation="longitude" pageId="2" pageNumber="235" precision="55" value="-0.064">0.064</geoCoordinate>
), TL/SVL (original) = 0.77 (0.760.78), TT/SVL (original) = 0.41 (0.410.42), TW/SVL (original) = 0.20 (0.180.21), TD/SVL (original) = 0.066 (
<geoCoordinate id="EE12288B3843FFA8FC9FF9BC4175F9C5" box="[804,868,1610,1637]" degrees="0.059" direction="north" orientation="latitude" pageId="2" pageNumber="235" precision="55" value="0.059">0.059</geoCoordinate>
<geoCoordinate id="EE12288B3843FFA8FCDFF9BC41A3F9C5" box="[868,946,1610,1637]" degrees="0.073" direction="west" orientation="longitude" pageId="2" pageNumber="235" precision="55" value="-0.073">0.073</geoCoordinate>
), TL/SVL (regenerated) = 0.49 (0.470.50), TT/SVL (regenerated) = 0.24 (0.220.25), TW/SVL (regenerated) = 0.23 (0.210.25), TD/SVL (regenerated) = 0.079 (
<geoCoordinate id="EE12288B3843FFA8FF24F96542CEF90D" box="[159,223,1683,1709]" degrees="0.077" direction="north" orientation="latitude" pageId="2" pageNumber="235" precision="55" value="0.077">0.077</geoCoordinate>
<geoCoordinate id="EE12288B3843FFA8FF65F965433CF90D" box="[222,301,1683,1709]" degrees="0.082" direction="west" orientation="longitude" pageId="2" pageNumber="235" precision="55" value="-0.082">0.082</geoCoordinate>
). Scale counts (average, followed by range in brackets): internasals = 6.8 (67), supralabials = 15.0 (1416), infralabials = 14.5 (1316), subdigital lamellae under 4
<superScript id="7C53E3043843FFA8FCD1F9404166F964" attach="left" box="[874,887,1718,1732]" fontSize="6" pageId="2" pageNumber="235">th</superScript>
finger = 16.3 (1518), subdigital lamellae under 4
<superScript id="7C53E3043843FFA8FF1EF92C42A3F948" attach="left" box="[165,178,1754,1768]" fontSize="6" pageId="2" pageNumber="235">th</superScript>
toe = 19.0 (1820).
<emphasis id="B952925E3843FFA8FE2DF92C43CBF955" bold="true" box="[406,474,1754,1781]" pageId="2" pageNumber="235">
<emphasis id="B952925E3843FFA8FE2DF92C43C2F955" bold="true" box="[406,467,1754,1781]" italics="true" pageId="2" pageNumber="235">Head</emphasis>
.
</emphasis>
Large, depressed, triangular; covered in very small granules with scattered larger, pale, conical tubercles on sides of head; top of snout and canthul region have fine granular scales and no enlarged tubercles; canthul region moderately steep, with a moderately defined bony ridge extending forward from upper anterior margin of each eye towards naris; skin of head co-ossified with skull; deep, vertical groove partially (2050%) dividing rostral scale (
<tableCitation id="C6A47BF73843FFA8FF60F89D433CF825" box="[219,301,1899,1925]" captionStart="TABLE 1" captionStartId="6.[152,249,150,176]" captionTargetPageId="6" captionText="TABLE 1. Morphological measurements, proportions, and scale counts for the type specimens, and SVL data from individuals measured in the field. All measurements are in mm. Field data is only for individuals deemed to be mature adults (see text), is split by sex, and is presented as average followed by range in brackets." httpUri="http://table.plazi.org/id/DF591EC43847FFACFF23FF6046ACFF59" pageId="2" pageNumber="235" tableUuid="DF591EC43847FFACFF23FF6046ACFF59">Table 1</tableCitation>
); rostral not in contact with nostril; 67 scales along the upper margin of rostral shield (i.e., internasals); first supralabial scale taller than wide, remaining supralabials broad and steadily decreasing in size; first infralabial scale taller than wide, remaining infralabials broad and steadily decreasing in size; granular scales of chin and throat homogenously minute granules, with the exception of larger scales along the edge of the mental and infralabials (particularly abutting infralabials 13); ear opening conspicuous, vertical, much less than half as large as eye.
<emphasis id="B952925E3842FFA9FAAFFF604745FF11" bold="true" box="[1300,1364,150,177]" italics="true" pageId="3" pageNumber="236">Neck.</emphasis>
Fairly broad; covered in small granules that are intermixed with evenly scattered larger pale conical tubercles the sides and top.
<emphasis id="B952925E3842FFA9FF7CFF284317FF59" bold="true" box="[199,262,222,249]" italics="true" pageId="3" pageNumber="236">Body.</emphasis>
Depressed, covered in small granules; flanks and back evenly covered in scattered, small, conical tubercles; those on the back and side evenly small and scattered (as for neck); all ventral surfaces (even under limbs) covered in even, smooth, fine granules, with no enlarged tubercles; single male has a large post-cloacal bulge, with conspicuous cloacal spurs, consisting of a cluster of white triangular scales, on the anterior-lateral margin; no pre-cloacal pores on any individuals; axilla deeply invaginated.
<emphasis id="B952925E3842FFA9FDC8FE9840D1FE29" bold="true" box="[627,704,366,393]" italics="true" pageId="3" pageNumber="236">Limbs.</emphasis>
Long and very slender, covered in small granular scales and with scattered, small, pointed tubercles dorsally; lacking tubercles on ventral surface; digits strongly compressed distally; dorsal surface of hands, feet and digits without enlarged conical tubercles.
<emphasis id="B952925E3842FFA9FC69FE40464CFE71" bold="true" box="[978,1117,438,465]" italics="true" pageId="3" pageNumber="236">Original tail</emphasis>
(
<figureCitation id="131D52C93842FFA9FBD6FE4146A0FE71" box="[1133,1201,439,465]" captionStart="FIGURE 2" captionStartId="4.[151,250,1722,1746]" captionTargetBox="[151,1436,489,1698]" captionTargetId="figure-180@4.[151,1436,489,1698]" captionTargetPageId="4" captionText="FIGURE 2. Holotype of P. fimbriatus sp. nov. (QM J97578). Photos: (A) Edward Evans; (B) Conrad Hoskin." figureDoi="http://doi.org/10.5281/zenodo.7656253" httpUri="https://zenodo.org/record/7656253/files/figure.png" pageId="3" pageNumber="236">Fig. 2</figureCitation>
). Anterior half flared and leaf-shaped, then tapers abruptly into a long attenuated tip (tip 54% of total tail length for both specimens;
<tableCitation id="C6A47BF73842FFA9FADBFE2D42B5FDB9" captionStart="TABLE 1" captionStartId="6.[152,249,150,176]" captionTargetPageId="6" captionText="TABLE 1. Morphological measurements, proportions, and scale counts for the type specimens, and SVL data from individuals measured in the field. All measurements are in mm. Field data is only for individuals deemed to be mature adults (see text), is split by sex, and is presented as average followed by range in brackets." httpUri="http://table.plazi.org/id/DF591EC43847FFACFF23FF6046ACFF59" pageId="3" pageNumber="236" tableUuid="DF591EC43847FFACFF23FF6046ACFF59">Table 1</tableCitation>
); flared section strongly flattened; distal section rounded, terminating with a minute, rounded, white knob; dorsal surface covered in minute granules, with sparsely scattered enlarged spinose tubercles; margin of flared portion of tail undulating, with the peaks each having several obvious, large spinose tubercles, making the tail distinctly spiny on the edge but smooth on the top; ventral surface smooth with a mosaic of small rounded to square or hexagonal scales.
<emphasis id="B952925E3842FFA9FF2CFD784346FD09" bold="true" box="[151,343,654,681]" italics="true" pageId="3" pageNumber="236">Regenerated tail.</emphasis>
Disc-like, rounded shape, with thin, tapering distal portion (tip 47% and 50% of total tail length in the
<specimenCount id="9D2085C53842FFA9FF05FD454373FD6D" box="[190,354,691,717]" pageId="3" pageNumber="236" type="generic">two specimens</specimenCount>
;
<tableCitation id="C6A47BF73842FFA9FED0FD4543ABFD6D" box="[363,442,691,717]" captionStart="TABLE 1" captionStartId="6.[152,249,150,176]" captionTargetPageId="6" captionText="TABLE 1. Morphological measurements, proportions, and scale counts for the type specimens, and SVL data from individuals measured in the field. All measurements are in mm. Field data is only for individuals deemed to be mature adults (see text), is split by sex, and is presented as average followed by range in brackets." httpUri="http://table.plazi.org/id/DF591EC43847FFACFF23FF6046ACFF59" pageId="3" pageNumber="236" tableUuid="DF591EC43847FFACFF23FF6046ACFF59">Table 1</tableCitation>
); dorsal and lateral surfaces of disc and attenuated portion covered in fine, pointed granules; small, rounded knob on the end; ventral surface covered in mosaic of block like scales.
<emphasis id="B952925E3842FFA9FBE2FD20474FFD51" bold="true" box="[1113,1374,726,753]" italics="true" pageId="3" pageNumber="236">Colour pattern in spirit</emphasis>
(e.g.,
<figureCitation id="131D52C93842FFA9FF2CFD0D42CBFCB5" box="[151,218,763,789]" captionStart="FIGURE 2" captionStartId="4.[151,250,1722,1746]" captionTargetBox="[151,1436,489,1698]" captionTargetId="figure-180@4.[151,1436,489,1698]" captionTargetPageId="4" captionText="FIGURE 2. Holotype of P. fimbriatus sp. nov. (QM J97578). Photos: (A) Edward Evans; (B) Conrad Hoskin." figureDoi="http://doi.org/10.5281/zenodo.7656253" httpUri="https://zenodo.org/record/7656253/files/figure.png" pageId="3" pageNumber="236">Fig. 2</figureCitation>
). Dorsal base colour grey to tan, with irregular large dark brown blotches on neck and body, and smaller, dark brown blotches on head and limbs; dark blotches on neck and back connected to be arranged as irregular, dark, transverse bars; dark, longitudinal markings extend from above hips to base of tail; top of head and snout has less pattern but W-shaped mark across top of snout, connecting the anterior margin of the eyes; diffuse pale band from lower anterior margin of eye diagonally to midway along jaw, and more obvious similar pale band from lower posterior margin of eye diagonally to just behind jaw; white spot (from cluster of minute tubercles) above the ear; all dorsal and lateral surface of neck, body and limbs covered in fine white flecks (that are generally the small conical tubercles); ventral surfaces of chin, throat, chest, abdomen and limbs fairly even cream colour, except for dark smudging under the knees, feet and hands; original tail has pale grey-brown background with extensive dark brown-black mottling, and white bands; the white bands consist of a band at the base of the flared portion that is distinctly V-shaped, and unbroken across the width of the tail, then a very thin white band three-quarters of the way along the flared portion, then a thick white band at the start of the attenuated section, then two white bands along the attenuated section, and a white tip at the end of the tail; underside of flared portion of original tail has a pale background, with heavy, dark brown flecked and scattered white spots; ventral surface of attenuated portion of original tail starting with V-shaped white marking, then dark brown and white corresponding to stripes on dorsal surface; dorsal surface of regenerated tail dark brown with a few irregular pale markings (QM J97579) or light brown background with fairly even dark mottling (QM J97577); ventral surfaces have a pale background that is heavily (QM J97579) or moderately covered in dark brown mottling (QM J97577).
<emphasis id="B952925E3842FFA9FAF6FAC143DFFAD6" bold="true" italics="true" pageId="3" pageNumber="236">Colour pattern of type series in life.</emphasis>
As above but base colour of body generally lighter (whitish grey), white and black markings (e.g., particularly on original tail) more contrasting, and ventral surfaces immaculate white.
</paragraph>
<paragraph id="8B994E4C3842FFAEFF7CFA5443B1FE71" blockId="3.[151,1437,150,2043]" lastBlockId="4.[151,1437,150,465]" lastPageId="4" lastPageNumber="237" pageId="3" pageNumber="236">
<emphasis id="B952925E3842FFA9FF7CFA544047FA1D" bold="true" box="[199,598,1442,1469]" pageId="3" pageNumber="236">
Field data.
<emphasis id="B952925E3842FFA9FEF7FA544047FA1D" bold="true" box="[332,598,1442,1469]" italics="true" pageId="3" pageNumber="236">Body size and maturity.</emphasis>
</emphasis>
SVL and tail data were collected from 23 individuals in the field, and these individuals were photographed to assess consistency in colour pattern traits.
<collectorName id="26D32B9A3842FFA9FC6BFA3041EAFA40" box="[976,1019,1478,1504]" pageId="3" pageNumber="236">The</collectorName>
majority of individuals observed were subadults of approximately
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SVL.
<collectorName id="26D32B9A3842FFA9FDD9FA1C409CF9A4" box="[610,653,1514,1540]" pageId="3" pageNumber="236">The</collectorName>
smallest male with a pronounced post-cloacal bulge was
<quantity id="4CDEE3A93842FFA9FAB0FA1C474BF9A5" box="[1291,1370,1514,1541]" metricMagnitude="-2" metricUnit="m" metricValue="9.6" pageId="3" pageNumber="236" unit="mm" value="96.0">96 mm</quantity>
SVL.
<collectorName id="26D32B9A3842FFA9FF2CF9F84300F988" box="[151,273,1550,1576]" pageId="3" pageNumber="236">Individuals</collectorName>
deemed male below that SVL (up to
<quantity id="4CDEE3A93842FFA9FD08F9F84113F988" box="[691,770,1550,1577]" metricMagnitude="-2" metricUnit="m" metricValue="8.5" pageId="3" pageNumber="236" unit="mm" value="85.0">85 mm</quantity>
) did not have a pronounced post-cloacal bulge.
<collectorName id="26D32B9A3842FFA9FAADF9F8475FF988" box="[1302,1358,1550,1576]" pageId="3" pageNumber="236">Male</collectorName>
sexual maturity was therefore taken to be
<quantity id="4CDEE3A93842FFA9FDAAF9C7404FF9EC" box="[529,606,1585,1612]" metricMagnitude="-2" metricUnit="m" metricValue="9.6" pageId="3" pageNumber="236" unit="mm" value="96.0">96 mm</quantity>
SVL for the field data and, for the
<specimenCount id="9D2085C53842FFA9FC61F9C74646F9EB" box="[986,1111,1585,1611]" pageId="3" pageNumber="236" type="male">eight males</specimenCount>
that were this size and above, the average SVL was
<quantity id="4CDEE3A93842FFA9FE34F9A343FDF9D0" box="[399,492,1621,1648]" metricMagnitude="-1" metricUnit="m" metricValue="1.02" pageId="3" pageNumber="236" unit="mm" value="102.0">102 mm</quantity>
(with a range between
<quantity id="4CDEE3A93842FFA9FD4EF9A3416FF9D0" box="[757,894,1621,1648]" metricMagnitude="-1" metricUnit="m" metricValue="1.005" metricValueMax="1.05" metricValueMin="0.96" pageId="3" pageNumber="236" unit="mm" value="100.5" valueMax="105.0" valueMin="96.0">96105 mm</quantity>
;
<tableCitation id="C6A47BF73842FFA9FC30F9A341CCF9CF" box="[907,989,1621,1647]" captionStart="TABLE 1" captionStartId="6.[152,249,150,176]" captionTargetPageId="6" captionText="TABLE 1. Morphological measurements, proportions, and scale counts for the type specimens, and SVL data from individuals measured in the field. All measurements are in mm. Field data is only for individuals deemed to be mature adults (see text), is split by sex, and is presented as average followed by range in brackets." httpUri="http://table.plazi.org/id/DF591EC43847FFACFF23FF6046ACFF59" pageId="3" pageNumber="236" tableUuid="DF591EC43847FFACFF23FF6046ACFF59">Table 1</tableCitation>
).
<collectorName id="26D32B9A3842FFA9FC4DF9A34630F9CF" box="[1014,1057,1621,1647]" pageId="3" pageNumber="236">The</collectorName>
smallest female with eggs visible through the body wall of the abdomen was
<quantity id="4CDEE3A93842FFA9FD3AF98F40CCF934" box="[641,733,1657,1684]" metricMagnitude="-1" metricUnit="m" metricValue="1.03" pageId="3" pageNumber="236" unit="mm" value="103.0">103 mm</quantity>
SVL. None of the females inspected below that size (up to
<quantity id="4CDEE3A93842FFA9FA3BF98F42D3F917" metricMagnitude="-2" metricUnit="m" metricValue="9.6" pageId="3" pageNumber="236" unit="mm" value="96.0">96 mm</quantity>
SVL) had visible eggs, whereas six of the
<specimenCount id="9D2085C53842FFA9FD26F96B4121F917" box="[669,816,1693,1719]" pageId="3" pageNumber="236" type="female">eight females</specimenCount>
inspected above
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SVL had visible eggs.
<collectorName id="26D32B9A3842FFA9FAF7F96B478CF917" box="[1356,1437,1693,1719]" pageId="3" pageNumber="236">Female</collectorName>
sexual maturity was therefore taken to be
<quantity id="4CDEE3A93842FFA9FDDAF93740ADF97B" box="[609,700,1729,1755]" metricMagnitude="-1" metricUnit="m" metricValue="1.03" pageId="3" pageNumber="236" unit="mm" value="103.0">103 mm</quantity>
SVL for the field data and, for the
<specimenCount id="9D2085C53842FFA9FB85F93646C0F97A" box="[1086,1233,1728,1754]" pageId="3" pageNumber="236" type="female">eight females</specimenCount>
that were this size and above, the average SVL was
<quantity id="4CDEE3A93842FFA9FE44F9134048F95F" box="[511,601,1765,1791]" metricMagnitude="-1" metricUnit="m" metricValue="1.09" pageId="3" pageNumber="236" unit="mm" value="109.0">109 mm</quantity>
(with a range between
<quantity id="4CDEE3A93842FFA9FCEEF91341F9F95F" box="[853,1000,1765,1791]" metricMagnitude="-1" metricUnit="m" metricValue="1.08" metricValueMax="1.13" metricValueMin="1.03" pageId="3" pageNumber="236" unit="mm" value="108.0" valueMax="113.0" valueMin="103.0">103113 mm</quantity>
;
<tableCitation id="C6A47BF73842FFA9FC49F9124650F95F" box="[1010,1089,1764,1791]" captionStart="TABLE 1" captionStartId="6.[152,249,150,176]" captionTargetPageId="6" captionText="TABLE 1. Morphological measurements, proportions, and scale counts for the type specimens, and SVL data from individuals measured in the field. All measurements are in mm. Field data is only for individuals deemed to be mature adults (see text), is split by sex, and is presented as average followed by range in brackets." httpUri="http://table.plazi.org/id/DF591EC43847FFACFF23FF6046ACFF59" pageId="3" pageNumber="236" tableUuid="DF591EC43847FFACFF23FF6046ACFF59">Table 1</tableCitation>
).
<emphasis id="B952925E3842FFA9FBECF91246F0F95F" bold="true" box="[1111,1249,1764,1791]" italics="true" pageId="3" pageNumber="236">
<collectorName id="26D32B9A3842FFA9FBECF91246A7F95F" box="[1111,1206,1764,1791]" pageId="3" pageNumber="236">Original</collectorName>
tail
</emphasis>
(e.g.,
<figureCitation id="131D52C93842FFA9FA9FF9124779F95F" box="[1316,1384,1764,1791]" captionStart="FIGURE 2" captionStartId="4.[151,250,1722,1746]" captionTargetBox="[151,1436,489,1698]" captionTargetId="figure-180@4.[151,1436,489,1698]" captionTargetPageId="4" captionText="FIGURE 2. Holotype of P. fimbriatus sp. nov. (QM J97578). Photos: (A) Edward Evans; (B) Conrad Hoskin." figureDoi="http://doi.org/10.5281/zenodo.7656253" httpUri="https://zenodo.org/record/7656253/files/figure.png" pageId="3" pageNumber="236">Figs 2</figureCitation>
,
<figureCitation id="131D52C93842FFA9FACFF91342ABF883" captionStart="FIGURE 3" captionStartId="5.[151,250,1236,1260]" captionTargetBox="[151,1435,381,1212]" captionTargetId="figure-123@5.[151,1435,381,1212]" captionTargetPageId="5" captionText="FIGURE 3. Phyllurus fimbriatus sp. nov. in life: (A) adult female with original tail; (B) juvenile with original tail; (C) adult female with regenerated tail; (D) adult male of unusual colouration, with regenerated tail. Photos: Conrad Hoskin." figureDoi="http://doi.org/10.5281/zenodo.7656255" httpUri="https://zenodo.org/record/7656255/files/figure.png" pageId="3" pageNumber="236">3A, 3B</figureCitation>
).
<collectorName id="26D32B9A3842FFA9FF74F8FE42FDF882" box="[207,236,1800,1826]" pageId="3" pageNumber="236">Of</collectorName>
the 23 individuals for which tail status was recorded in the field, 8 (i.e., 35%) had an original tail.
<collectorName id="26D32B9A3842FFA9FA89F8FF4789F883" box="[1330,1432,1801,1827]" pageId="3" pageNumber="236">However</collectorName>
, there was a clear relationship with body size: all 7 individuals &lt;
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SVL had an original tail, whereas only 1 out of 16 individuals (i.e., 6%) that were
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or longer had an original tail.
<collectorName id="26D32B9A3842FFA9FC7EF8A7460EF8CB" box="[965,1055,1873,1899]" pageId="3" pageNumber="236">Average</collectorName>
TL/SVL for the 8 individuals with an original tail was 0.77 (range 0.740.81). Shape as described for
<typeStatus id="549DF0EE3842FFA9FCC5F88341BCF82F" box="[894,941,1909,1935]" pageId="3" pageNumber="236">type</typeStatus>
specimens.
<collectorName id="26D32B9A3842FFA9FB8EF882464EF82E" box="[1077,1119,1908,1934]" pageId="3" pageNumber="236">Tail</collectorName>
colouration as described but with the following variation.
<collectorName id="26D32B9A3842FFA9FE66F86E4019F812" box="[477,520,1944,1970]" pageId="3" pageNumber="236">The</collectorName>
white band halfway along the flared portion varies from thin but distinct in some individuals to a single, central white dot in others.
<collectorName id="26D32B9A3842FFA9FD00F84A40F7F876" box="[699,742,1980,2006]" pageId="3" pageNumber="236">The</collectorName>
white band at the end of the flared portion is often V-shaped, or sometimes a diagonal line, rather than straight and transverse.
<collectorName id="26D32B9A3842FFA9FC80F8164177F85A" box="[827,870,2016,2042]" pageId="3" pageNumber="236">The</collectorName>
degree of dark brown mottling on the flared portion of the tail varies from light to heavy.
<emphasis id="B952925E3845FFAEFDA4FF6040C4FF11" bold="true" box="[543,725,150,177]" italics="true" pageId="4" pageNumber="237">
<collectorName id="26D32B9A3845FFAEFDA4FF6040BDFF11" box="[543,684,150,177]" pageId="4" pageNumber="237">Regenerated</collectorName>
tail
</emphasis>
(e.g.,
<figureCitation id="131D52C93845FFAEFCACFF61418AFF11" box="[791,923,151,177]" captionStart="FIGURE 3" captionStartId="5.[151,250,1236,1260]" captionTargetBox="[151,1435,381,1212]" captionTargetId="figure-123@5.[151,1435,381,1212]" captionTargetPageId="5" captionText="FIGURE 3. Phyllurus fimbriatus sp. nov. in life: (A) adult female with original tail; (B) juvenile with original tail; (C) adult female with regenerated tail; (D) adult male of unusual colouration, with regenerated tail. Photos: Conrad Hoskin." figureDoi="http://doi.org/10.5281/zenodo.7656255" httpUri="https://zenodo.org/record/7656255/files/figure.png" pageId="4" pageNumber="237">Figs 3C, 3D</figureCitation>
).
<collectorName id="26D32B9A3845FFAEFC17FF61463DFF11" box="[940,1068,151,177]" pageId="4" pageNumber="237">Average TL</collectorName>
/SVL for three individuals that appeared to have a full regenerated tail was 0.53 (range 0.460.60). Shape as described for
<typeStatus id="549DF0EE3845FFAEFBE9FF4D4690FF75" box="[1106,1153,187,213]" pageId="4" pageNumber="237">type</typeStatus>
specimens, but varying in the degree to which the tail has regenerated, with the attenuated portion appearing to get relatively longer with tail size.
<collectorName id="26D32B9A3845FFAEFF2CFEF542F2FEBD" box="[151,227,259,285]" pageId="4" pageNumber="237">Colour</collectorName>
also appears to vary as the tail regenerates, with freshly regenerating tails being almost black and those more fully regenerated being paler and more similar to the background colouration of the body.
<emphasis id="B952925E3845FFAEFBE5FED0471BFEE1" bold="true" box="[1118,1290,294,321]" italics="true" pageId="4" pageNumber="237">
<collectorName id="26D32B9A3845FFAEFBE5FED046BCFEE1" box="[1118,1197,294,321]" pageId="4" pageNumber="237">Colour</collectorName>
pattern.
</emphasis>
<collectorName id="26D32B9A3845FFAEFAB4FED1473CFEE1" box="[1295,1325,295,321]" pageId="4" pageNumber="237">As</collectorName>
described for the
<typeStatus id="549DF0EE3845FFAEFF5FFEBD4302FEC5" box="[228,275,331,357]" pageId="4" pageNumber="237">type</typeStatus>
series, noting the following variation.
<collectorName id="26D32B9A3845FFAEFD08FEBD40F9FEC5" box="[691,744,331,357]" pageId="4" pageNumber="237">Base</collectorName>
colour varies from light grey to brown, giving some individuals a much darker appearance (e.g.,
<figureCitation id="131D52C93845FFAEFE42FE99405CFE29" box="[505,589,367,393]" captionStart="FIGURE 3" captionStartId="5.[151,250,1236,1260]" captionTargetBox="[151,1435,381,1212]" captionTargetId="figure-123@5.[151,1435,381,1212]" captionTargetPageId="5" captionText="FIGURE 3. Phyllurus fimbriatus sp. nov. in life: (A) adult female with original tail; (B) juvenile with original tail; (C) adult female with regenerated tail; (D) adult male of unusual colouration, with regenerated tail. Photos: Conrad Hoskin." figureDoi="http://doi.org/10.5281/zenodo.7656255" httpUri="https://zenodo.org/record/7656255/files/figure.png" pageId="4" pageNumber="237">Fig. 3C</figureCitation>
).
<collectorName id="26D32B9A3845FFAEFDDAFE99409EFE29" box="[609,655,367,393]" pageId="4" pageNumber="237">One</collectorName>
individual (
<figureCitation id="131D52C93845FFAEFCA9FE994179FE29" box="[786,872,367,393]" captionStart="FIGURE 3" captionStartId="5.[151,250,1236,1260]" captionTargetBox="[151,1435,381,1212]" captionTargetId="figure-123@5.[151,1435,381,1212]" captionTargetPageId="5" captionText="FIGURE 3. Phyllurus fimbriatus sp. nov. in life: (A) adult female with original tail; (B) juvenile with original tail; (C) adult female with regenerated tail; (D) adult male of unusual colouration, with regenerated tail. Photos: Conrad Hoskin." figureDoi="http://doi.org/10.5281/zenodo.7656255" httpUri="https://zenodo.org/record/7656255/files/figure.png" pageId="4" pageNumber="237">Fig. 3D</figureCitation>
) was obviously different in colour pattern, being an even, soft brown colour with four longitudinal lines of dark blotches down the vertebral zone and a dorsolateral line of enlarged, pale tubercles.
</paragraph>
</subSubSection>
<caption id="DF591EC43845FFAEFF2CF94C46E1F972" ID-DOI="http://doi.org/10.5281/zenodo.7656253" ID-Zenodo-Dep="7656253" box="[151,1264,1722,1746]" httpUri="https://zenodo.org/record/7656253/files/figure.png" pageId="4" pageNumber="237" startId="4.[151,250,1722,1746]" targetBox="[151,1436,489,1698]" targetPageId="4">
<paragraph id="8B994E4C3845FFAEFF2CF94C46E1F972" blockId="4.[151,1264,1722,1746]" box="[151,1264,1722,1746]" pageId="4" pageNumber="237">
<emphasis id="B952925E3845FFAEFF2CF94C431CF972" bold="true" box="[151,269,1722,1746]" pageId="4" pageNumber="237">FIGURE 2</emphasis>
. Holotype of
<taxonomicName id="4C2635CF3845FFAEFE22F94D400BF972" authority="Hoskin, 2023" authorityName="Hoskin" authorityYear="2023" box="[409,538,1722,1746]" class="Reptilia" family="Carphodactylidae" genus="Phyllurus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="237" phylum="Chordata" rank="species" species="fimbriatus" status="sp. nov.">
<emphasis id="B952925E3845FFAEFE22F94D400BF972" box="[409,538,1722,1746]" italics="true" pageId="4" pageNumber="237">P. fimbriatus</emphasis>
</taxonomicName>
<emphasis id="B952925E3845FFAEFD9AF94C4060F972" bold="true" box="[545,625,1722,1746]" pageId="4" pageNumber="237">
<taxonomicNameLabel id="A2612F253845FFAEFD9AF94C4060F972" box="[545,625,1722,1746]" pageId="4" pageNumber="237" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(QM J97578). Photos: (A) Edward Evans; (B) Conrad Hoskin.
</paragraph>
</caption>
<subSubSection id="C33C1DC73845FFAFFF7CF8F6470FF918" lastPageId="5" lastPageNumber="238" pageId="4" pageNumber="237" type="discussion">
<paragraph id="8B994E4C3845FFAFFF7CF8F640DBFEC5" blockId="4.[151,1436,1792,2035]" lastBlockId="5.[151,1437,151,357]" lastPageId="5" lastPageNumber="238" pageId="4" pageNumber="237">
<emphasis id="B952925E3845FFAEFF7CF8F6437CF8BA" bold="true" box="[199,365,1792,1818]" pageId="4" pageNumber="237">Comparisons.</emphasis>
<taxonomicName id="4C2635CF3845FFAEFECCF8F64073F8BA" authority="Hoskin, 2023" authorityName="Hoskin" authorityYear="2023" box="[375,610,1792,1818]" class="Reptilia" family="Carphodactylidae" genus="Phyllurus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="237" phylum="Chordata" rank="species" species="fimbriatus" status="sp. nov.">
<emphasis id="B952925E3845FFAEFECCF8F64073F8BA" box="[375,610,1792,1818]" italics="true" pageId="4" pageNumber="237">Phyllurus fimbriatus</emphasis>
</taxonomicName>
<emphasis id="B952925E3845FFAEFDD7F8F640D8F8BA" bold="true" box="[620,713,1792,1818]" pageId="4" pageNumber="237">
<taxonomicNameLabel id="A2612F253845FFAEFDD7F8F640D8F8BA" box="[620,713,1792,1818]" pageId="4" pageNumber="237" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
could only be confused with congeners. The obviously flared original and regenerated tail readily distinguishes it from the following congeners that have a cylindrical, tapering tail:
<emphasis id="B952925E3845FFAEFF72F8BF4000F8C2" box="[201,529,1864,1890]" italics="true" pageId="4" pageNumber="237">
<taxonomicName id="4C2635CF3845FFAEFF72F8BF4381F8C2" authorityName="Covacevich" authorityYear="1975" box="[201,400,1864,1890]" class="Reptilia" family="Carphodactylidae" genus="Phyllurus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="237" phylum="Chordata" rank="species" species="caudiannulatus">P. caudiannulatus</taxonomicName>
<taxonomicName id="4C2635CF3845FFAEFE2DF8BF4000F8C2" authorityName="Couper, Hamley &amp; Hoskin" authorityYear="2008" box="[406,529,1864,1890]" class="Reptilia" family="Carphodactylidae" genus="Phyllurus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="237" phylum="Chordata" rank="species" species="kabikabi">P. kabikabi</taxonomicName>
</emphasis>
,
<taxonomicName id="4C2635CF3845FFAEFDA5F8BF4083F8C2" authorityName="Hoskin, Couper &amp; Schneider" authorityYear="2003" box="[542,658,1864,1890]" class="Reptilia" family="Carphodactylidae" genus="Phyllurus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="237" phylum="Chordata" rank="species" species="gulbaru">
<emphasis id="B952925E3845FFAEFDA5F8BF4083F8C2" box="[542,658,1864,1890]" italics="true" pageId="4" pageNumber="237">P. gulbaru</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2635CF3845FFAEFD7DF8BF417CF8C2" authorityName="Hoskin, Bertola &amp; Higgie" authorityYear="2019" box="[710,877,1864,1890]" class="Reptilia" family="Carphodactylidae" genus="Phyllurus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="237" phylum="Chordata" rank="species" species="pinnaclensis">
<emphasis id="B952925E3845FFAEFD7DF8BF417CF8C2" box="[710,877,1864,1890]" italics="true" pageId="4" pageNumber="237">P. pinnaclensis</emphasis>
</taxonomicName>
.
<taxonomicName id="4C2635CF3845FFAEFCC1F8BE4671F8C2" authority="Hoskin, 2023" authorityName="Hoskin" authorityYear="2023" box="[890,1120,1864,1890]" class="Reptilia" family="Carphodactylidae" genus="Phyllurus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="237" phylum="Chordata" rank="species" species="fimbriatus" status="sp. nov.">
<emphasis id="B952925E3845FFAEFCC1F8BE4671F8C2" box="[890,1120,1864,1890]" italics="true" pageId="4" pageNumber="237">Phyllurus fimbriatus</emphasis>
</taxonomicName>
<emphasis id="B952925E3845FFAEFBDDF8BE46AEF8C2" bold="true" box="[1126,1215,1864,1890]" pageId="4" pageNumber="237">
<taxonomicNameLabel id="A2612F253845FFAEFBDDF8BE46AEF8C2" box="[1126,1215,1864,1890]" pageId="4" pageNumber="237" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from congeners with flared tails in the following ways (comparative data from
<bibRefCitation id="EFB733BD3845FFAEFC3BF89B4615F826" author="Bauer, A. M." box="[896,1028,1900,1927]" pageId="4" pageNumber="237" pagination="1 - 217" refId="ref6124" refString="Bauer, A. M. (1990) Phylogenetic systematics and biogeography of the Carphodactylini (Reptilia: Gekkonidae). Bonner Zoologische Monographien, 30, 1 - 217." type="journal article" year="1990">Bauer 1990</bibRefCitation>
;
<bibRefCitation id="EFB733BD3845FFAEFBB5F89A46CFF826" author="Couper, P. J. &amp; Covacevich, J. A. &amp; Moritz, C." box="[1038,1246,1900,1926]" pageId="4" pageNumber="237" pagination="95 - 124" refId="ref6280" refString="Couper, P. J., Covacevich, J. A. &amp; Moritz, C. (1993) A review of the leaf-tailed geckos endemic to eastern Australia: a new genus, four new species, and other new data. Memoirs of the Queensland Museum, 34, 95 - 124." type="journal article" year="1993">
Couper
<emphasis id="B952925E3845FFAEFBDCF89B468FF826" box="[1127,1182,1900,1926]" italics="true" pageId="4" pageNumber="237">et al.</emphasis>
1993
</bibRefCitation>
, 2000; Couper &amp;
<bibRefCitation id="EFB733BD3845FFAEFF2CF866433DF80A" author="Hoskin, C. J. &amp; Couper, P. J." box="[151,300,1936,1962]" pageId="4" pageNumber="237" pagination="543 - 558" refId="ref6957" refString="Hoskin, C. J. &amp; Couper, P. J. (2013) A spectacular new leaf-tailed gecko (Carphodactylidae: Saltuarius) from the Melville Range, north-east Australia. Zootaxa, 3717 (4), 543 - 558. https: // doi. org / 10.11646 / zootaxa. 3717.4.6" type="journal article" year="2013">Hoskin 2013</bibRefCitation>
). The V-shaped anterior-most white band on the original tail separates it from all other species. Large body size (av. SVL
<quantity id="4CDEE3A93845FFAEFEC8F84243C3F86F" box="[371,466,1972,1999]" metricMagnitude="-1" metricUnit="m" metricValue="1.05" pageId="4" pageNumber="237" unit="mm" value="105.0">105 mm</quantity>
, max.
<quantity id="4CDEE3A93845FFAEFDA7F842406AF86F" box="[540,635,1972,1999]" metricMagnitude="-1" metricUnit="m" metricValue="1.13" pageId="4" pageNumber="237" unit="mm" value="113.0">113 mm</quantity>
; including field measurements) readily separates it from
<taxonomicName id="4C2635CF3845FFAEFB43F843478DF86E" authorityName="Schneider, Couper, Hoskin &amp; Covacevich" authorityYear="2000" box="[1272,1436,1972,1998]" class="Reptilia" family="Carphodactylidae" genus="Phyllurus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="237" phylum="Chordata" rank="species" species="championae">
<emphasis id="B952925E3845FFAEFB43F843478DF86E" box="[1272,1436,1972,1998]" italics="true" pageId="4" pageNumber="237">P. championae</emphasis>
</taxonomicName>
(av.
<quantity id="4CDEE3A93845FFAEFF73F82E4308F853" box="[200,281,2008,2035]" metricMagnitude="-2" metricUnit="m" metricValue="6.2" pageId="4" pageNumber="237" unit="mm" value="62.0">62 mm</quantity>
, max.
<quantity id="4CDEE3A93845FFAEFEDFF82E43A6F852" box="[356,439,2008,2034]" metricMagnitude="-2" metricUnit="m" metricValue="8.1" pageId="4" pageNumber="237" unit="mm" value="81.0">81 mm</quantity>
),
<taxonomicName id="4C2635CF3845FFAEFE71F82F401AF852" authorityName="Couper, Covacevich &amp; Moritz" authorityYear="1993" box="[458,523,2009,2034]" class="Reptilia" family="Carphodactylidae" genus="Phyllurus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="237" phylum="Chordata" rank="species" species="isis">
<emphasis id="B952925E3845FFAEFE71F82F401AF852" box="[458,523,2009,2034]" italics="true" pageId="4" pageNumber="237">P. isis</emphasis>
</taxonomicName>
(av.
<quantity id="4CDEE3A93845FFAEFDFFF82E4084F853" box="[580,661,2008,2035]" metricMagnitude="-2" metricUnit="m" metricValue="6.9" pageId="4" pageNumber="237" unit="mm" value="69.0">69 mm</quantity>
, max.
<quantity id="4CDEE3A93845FFAEFD5BF82E4122F852" box="[736,819,2008,2034]" metricMagnitude="-2" metricUnit="m" metricValue="7.6" pageId="4" pageNumber="237" unit="mm" value="76.0">76 mm</quantity>
),
<taxonomicName id="4C2635CF3845FFAEFCFCF82F4184F852" authorityName="Couper, Covacevich &amp; Moritz" authorityYear="1993" box="[839,917,2009,2034]" class="Reptilia" family="Carphodactylidae" genus="Phyllurus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="237" phylum="Chordata" rank="species" species="ossa">
<emphasis id="B952925E3845FFAEFCFCF82F4184F852" box="[839,917,2009,2034]" italics="true" pageId="4" pageNumber="237">P. ossa</emphasis>
</taxonomicName>
(av.
<quantity id="4CDEE3A93845FFAEFC77F82E460FF853" box="[972,1054,2008,2035]" metricMagnitude="-2" metricUnit="m" metricValue="7.4" pageId="4" pageNumber="237" unit="mm" value="74.0">74 mm</quantity>
, max.
<quantity id="4CDEE3A93845FFAEFBD3F82E46ADF852" box="[1128,1212,2008,2034]" metricMagnitude="-2" metricUnit="m" metricValue="8.9" pageId="4" pageNumber="237" unit="mm" value="89.0">89 mm</quantity>
),
<taxonomicName id="4C2635CF3845FFAEFB74F82F4759F852" baseAuthorityName="White" baseAuthorityYear="1790" box="[1231,1352,2008,2034]" class="Reptilia" family="Carphodactylidae" genus="Phyllurus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="237" phylum="Chordata" rank="species" species="platurus">
<emphasis id="B952925E3845FFAEFB74F82F4759F852" box="[1231,1352,2008,2034]" italics="true" pageId="4" pageNumber="237">P. platurus</emphasis>
</taxonomicName>
(av.
<quantity id="4CDEE3A93845FFAFFA3BF82E42D4FF11" lastPageId="5" lastPageNumber="238" metricMagnitude="-2" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="237" unit="mm" value="80.0">80 mm</quantity>
, max.
<quantity id="4CDEE3A93844FFAFFEB6FF61434EFF11" box="[269,351,151,177]" metricMagnitude="-2" metricUnit="m" metricValue="9.9" pageId="5" pageNumber="238" unit="mm" value="99.0">99 mm</quantity>
), partially from
<taxonomicName id="4C2635CF3844FFAFFDA9FF614093FF11" authorityName="Couper, Covacevich &amp; Moritz" authorityYear="1993" box="[530,642,151,177]" class="Reptilia" family="Carphodactylidae" genus="Phyllurus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="238" phylum="Chordata" rank="species" species="nepthys">
<emphasis id="B952925E3844FFAFFDA9FF614093FF11" box="[530,642,151,177]" italics="true" pageId="5" pageNumber="238">P. nepthys</emphasis>
</taxonomicName>
(av.
<quantity id="4CDEE3A93844FFAFFD0CFF614116FF11" box="[695,775,151,177]" metricMagnitude="-2" metricUnit="m" metricValue="9.1" pageId="5" pageNumber="238" unit="mm" value="91.0">91 mm</quantity>
, max.
<quantity id="4CDEE3A93844FFAFFCF4FF6141BEFF11" box="[847,943,151,177]" metricMagnitude="-1" metricUnit="m" metricValue="1.03" pageId="5" pageNumber="238" unit="mm" value="103.0">103 mm</quantity>
), but not from
<taxonomicName id="4C2635CF3844FFAFFBE9FF6146C4FF11" authorityName="Hoskin, Couper, Schneider &amp; Covacevich" authorityYear="2000" box="[1106,1237,151,177]" class="Reptilia" family="Carphodactylidae" genus="Phyllurus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="238" phylum="Chordata" rank="species" species="amnicola">
<emphasis id="B952925E3844FFAFFBE9FF6146C4FF11" box="[1106,1237,151,177]" italics="true" pageId="5" pageNumber="238">P. amnicola</emphasis>
</taxonomicName>
(av.
<quantity id="4CDEE3A93844FFAFFAB1FF61474BFF11" box="[1290,1370,151,177]" metricMagnitude="-2" metricUnit="m" metricValue="9.7" pageId="5" pageNumber="238" unit="mm" value="97.0">97 mm</quantity>
, max.
<quantity id="4CDEE3A93844FFAFFF2CFF4D42E7FF75" box="[151,246,187,213]" metricMagnitude="-1" metricUnit="m" metricValue="1.13" pageId="5" pageNumber="238" unit="mm" value="113.0">113 mm</quantity>
; including field data). Neck, body, and dorsal surface of original tail relatively smooth, versus moderately to extremely spinose in
<taxonomicName id="4C2635CF3844FFAFFE17FF294042FF59" authorityName="Schneider, Couper, Hoskin &amp; Covacevich" authorityYear="2000" box="[428,595,223,249]" class="Reptilia" family="Carphodactylidae" genus="Phyllurus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="238" phylum="Chordata" rank="species" species="championae">
<emphasis id="B952925E3844FFAFFE17FF294042FF59" box="[428,595,223,249]" italics="true" pageId="5" pageNumber="238">P. championae</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2635CF3844FFAFFDD8FF2940A2FF58" authorityName="Couper, Covacevich &amp; Moritz" authorityYear="1993" box="[611,691,223,248]" class="Reptilia" family="Carphodactylidae" genus="Phyllurus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="238" phylum="Chordata" rank="species" species="ossa">
<emphasis id="B952925E3844FFAFFDD8FF2940A2FF58" box="[611,691,223,248]" italics="true" pageId="5" pageNumber="238">P. ossa</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2635CF3844FFAFFD55FF294173FF59" authorityName="Couper, Covacevich &amp; Moritz" authorityYear="1993" box="[750,866,223,249]" class="Reptilia" family="Carphodactylidae" genus="Phyllurus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="238" phylum="Chordata" rank="species" species="nepthys">
<emphasis id="B952925E3844FFAFFD55FF294173FF59" box="[750,866,223,249]" italics="true" pageId="5" pageNumber="238">P. nepthys</emphasis>
</taxonomicName>
. Single groove partially dividing the rostral scale separates it from
<taxonomicName id="4C2635CF3844FFAFFEE0FEF54011FEBD" authorityName="Schneider, Couper, Hoskin &amp; Covacevich" authorityYear="2000" box="[347,512,259,285]" class="Reptilia" family="Carphodactylidae" genus="Phyllurus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="238" phylum="Chordata" rank="species" species="championae">
<emphasis id="B952925E3844FFAFFEE0FEF54011FEBD" box="[347,512,259,285]" italics="true" pageId="5" pageNumber="238">P. championae</emphasis>
</taxonomicName>
(single groove completely divides rostral) and
<taxonomicName id="4C2635CF3844FFAFFBAEFEF54675FEBC" authorityName="Couper, Covacevich &amp; Moritz" authorityYear="1993" box="[1045,1124,259,284]" class="Reptilia" family="Carphodactylidae" genus="Phyllurus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="238" phylum="Chordata" rank="species" species="ossa">
<emphasis id="B952925E3844FFAFFBAEFEF54675FEBC" box="[1045,1124,259,284]" italics="true" pageId="5" pageNumber="238">P. ossa</emphasis>
</taxonomicName>
(typically three, sometimes two, partial grooves). Further differs from
<taxonomicName id="4C2635CF3844FFAFFDD4FED140BEFEE0" authorityName="Couper, Covacevich &amp; Moritz" authorityYear="1993" box="[623,687,295,320]" class="Reptilia" family="Carphodactylidae" genus="Phyllurus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="238" phylum="Chordata" rank="species" species="isis">
<emphasis id="B952925E3844FFAFFDD4FED140BEFEE0" box="[623,687,295,320]" italics="true" pageId="5" pageNumber="238">P. isis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2635CF3844FFAFFD5FFED14120FEE0" authorityName="Couper, Covacevich &amp; Moritz" authorityYear="1993" box="[740,817,295,320]" class="Reptilia" family="Carphodactylidae" genus="Phyllurus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="238" phylum="Chordata" rank="species" species="ossa">
<emphasis id="B952925E3844FFAFFD5FFED14120FEE0" box="[740,817,295,320]" italics="true" pageId="5" pageNumber="238">P. ossa</emphasis>
</taxonomicName>
in having 6 or 7 scales along posterior margin of rostral scale (i.e., internasals) (
<taxonomicName id="4C2635CF3844FFAFFE1AFEBD43F3FEC4" authorityName="Couper, Covacevich &amp; Moritz" authorityYear="1993" box="[417,482,331,356]" class="Reptilia" family="Carphodactylidae" genus="Phyllurus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="238" phylum="Chordata" rank="species" species="isis">
<emphasis id="B952925E3844FFAFFE1AFEBD43F3FEC4" box="[417,482,331,356]" italics="true" pageId="5" pageNumber="238">P. isis</emphasis>
</taxonomicName>
911;
<taxonomicName id="4C2635CF3844FFAFFD94FEBD406DFEC4" authorityName="Couper, Covacevich &amp; Moritz" authorityYear="1993" box="[559,636,331,356]" class="Reptilia" family="Carphodactylidae" genus="Phyllurus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="238" phylum="Chordata" rank="species" species="ossa">
<emphasis id="B952925E3844FFAFFD94FEBD406DFEC4" box="[559,636,331,356]" italics="true" pageId="5" pageNumber="238">P. ossa</emphasis>
</taxonomicName>
811).
</paragraph>
<caption id="DF591EC43844FFAFFF2CFB22470AFAB0" ID-DOI="http://doi.org/10.5281/zenodo.7656255" ID-Zenodo-Dep="7656255" httpUri="https://zenodo.org/record/7656255/files/figure.png" pageId="5" pageNumber="238" startId="5.[151,250,1236,1260]" targetBox="[151,1435,381,1212]" targetPageId="5">
<paragraph id="8B994E4C3844FFAFFF2CFB22470AFAB0" blockId="5.[151,1437,1236,1296]" pageId="5" pageNumber="238">
<emphasis id="B952925E3844FFAFFF2CFB22431FFB4C" bold="true" box="[151,270,1236,1260]" pageId="5" pageNumber="238">FIGURE 3</emphasis>
.
<taxonomicName id="4C2635CF3844FFAFFEA7FB2243FFFB4C" authority="Hoskin, 2023" authorityName="Hoskin" authorityYear="2023" box="[284,494,1236,1260]" class="Reptilia" family="Carphodactylidae" genus="Phyllurus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="238" phylum="Chordata" rank="species" species="fimbriatus" status="sp. nov.">
<emphasis id="B952925E3844FFAFFEA7FB2243FFFB4C" box="[284,494,1236,1260]" italics="true" pageId="5" pageNumber="238">Phyllurus fimbriatus</emphasis>
</taxonomicName>
<emphasis id="B952925E3844FFAFFE4EFB224056FB4C" bold="true" box="[501,583,1236,1260]" pageId="5" pageNumber="238">
<taxonomicNameLabel id="A2612F253844FFAFFE4EFB224056FB4C" box="[501,583,1236,1260]" pageId="5" pageNumber="238" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in life: (A) adult female with original tail; (B) juvenile with original tail; (C) adult female with regenerated tail; (D) adult male of unusual colouration, with regenerated tail. Photos: Conrad Hoskin.
</paragraph>
</caption>
<paragraph id="8B994E4C3844FFAFFF7CFACF470FF918" blockId="5.[151,1437,1337,2043]" pageId="5" pageNumber="238">
Broadly, most similar to
<taxonomicName id="4C2635CF3844FFAFFE62FACF404AFAF3" authorityName="Hoskin, Couper, Schneider &amp; Covacevich" authorityYear="2000" box="[473,603,1337,1363]" class="Reptilia" family="Carphodactylidae" genus="Phyllurus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="238" phylum="Chordata" rank="species" species="amnicola">
<emphasis id="B952925E3844FFAFFE62FACF404AFAF3" box="[473,603,1337,1363]" italics="true" pageId="5" pageNumber="238">P. amnicola</emphasis>
</taxonomicName>
(
<figureCitation id="131D52C93844FFAFFDD2FACF40E3FAF3" box="[617,754,1337,1363]" captionStart="FIGURE 4" captionStartId="7.[151,250,961,985]" captionTargetBox="[151,1435,181,937]" captionTargetId="figure-15@7.[151,1435,181,937]" captionTargetPageId="7" captionText="FIGURE 4. Phyllurus amnicola with original (A) and regenerated tail (B), and P. nepthys with original (C) and regenerated tail (D). Photos: (A, B) Conrad Hoskin; (C, D) Anders Zimny." figureDoi="http://doi.org/10.5281/zenodo.7656257" httpUri="https://zenodo.org/record/7656257/files/figure.png" pageId="5" pageNumber="238">Figs 4A, 4B</figureCitation>
) and
<taxonomicName id="4C2635CF3844FFAFFC95FACF418EFAF3" authorityName="Couper, Covacevich &amp; Moritz" authorityYear="1993" box="[814,927,1337,1363]" class="Reptilia" family="Carphodactylidae" genus="Phyllurus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="238" phylum="Chordata" rank="species" species="nepthys">
<emphasis id="B952925E3844FFAFFC95FACF418EFAF3" box="[814,927,1337,1363]" italics="true" pageId="5" pageNumber="238">P. nepthys</emphasis>
</taxonomicName>
(
<figureCitation id="131D52C93844FFAFFC17FACF4626FAF3" box="[940,1079,1337,1363]" captionStart="FIGURE 4" captionStartId="7.[151,250,961,985]" captionTargetBox="[151,1435,181,937]" captionTargetId="figure-15@7.[151,1435,181,937]" captionTargetPageId="7" captionText="FIGURE 4. Phyllurus amnicola with original (A) and regenerated tail (B), and P. nepthys with original (C) and regenerated tail (D). Photos: (A, B) Conrad Hoskin; (C, D) Anders Zimny." figureDoi="http://doi.org/10.5281/zenodo.7656257" httpUri="https://zenodo.org/record/7656257/files/figure.png" pageId="5" pageNumber="238">Figs 4C, 4D</figureCitation>
), due to large body size and partial division of rostral scale. Differs from
<taxonomicName id="4C2635CF3844FFAFFDD3FAAB40FDFAD7" authorityName="Hoskin, Couper, Schneider &amp; Covacevich" authorityYear="2000" box="[616,748,1373,1399]" class="Reptilia" family="Carphodactylidae" genus="Phyllurus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="238" phylum="Chordata" rank="species" species="amnicola">
<emphasis id="B952925E3844FFAFFDD3FAAB40FDFAD7" box="[616,748,1373,1399]" italics="true" pageId="5" pageNumber="238">P. amnicola</emphasis>
</taxonomicName>
in having a wider (HW/SVL =
<geoCoordinate id="EE12288B3844FFAFFBF6FAAB469AFAD7" box="[1101,1163,1373,1399]" degrees="0.217" direction="north" orientation="latitude" pageId="5" pageNumber="238" precision="55" value="0.217">0.217</geoCoordinate>
<geoCoordinate id="EE12288B3844FFAFFB37FAAB46C8FAD7" box="[1164,1241,1373,1399]" degrees="0.232" direction="west" orientation="longitude" pageId="5" pageNumber="238" precision="55" value="-0.232">0.232</geoCoordinate>
vs.
<geoCoordinate id="EE12288B3844FFAFFABCFAAB4757FAD7" box="[1287,1350,1373,1399]" degrees="0.207" direction="north" orientation="latitude" pageId="5" pageNumber="238" precision="55" value="0.207">0.207</geoCoordinate>
<geoCoordinate id="EE12288B3844FFAFFAFDFAAB4784FAD7" box="[1350,1429,1373,1399]" degrees="0.214" direction="west" orientation="longitude" pageId="5" pageNumber="238" precision="55" value="-0.214">0.214</geoCoordinate>
) and shorter head (HL/SVL =
<geoCoordinate id="EE12288B3844FFAFFE55FA76403CFA3A" box="[494,557,1408,1434]" degrees="0.266" direction="north" orientation="latitude" pageId="5" pageNumber="238" precision="55" value="0.266">0.266</geoCoordinate>
<geoCoordinate id="EE12288B3844FFAFFD96FA76406BFA3A" box="[557,634,1408,1434]" degrees="0.276" direction="west" orientation="longitude" pageId="5" pageNumber="238" precision="55" value="-0.276">0.276</geoCoordinate>
vs.
<geoCoordinate id="EE12288B3844FFAFFD15FA7740E1FA3B" box="[686,752,1409,1435]" degrees="0.277" direction="north" orientation="latitude" pageId="5" pageNumber="238" precision="55" value="0.277">0.277</geoCoordinate>
<geoCoordinate id="EE12288B3844FFAFFD4BFA77412EFA3B" box="[752,831,1409,1435]" degrees="0.291" direction="west" orientation="longitude" pageId="5" pageNumber="238" precision="55" value="-0.291">0.291</geoCoordinate>
); a shorter neck (NL/SVL =
<geoCoordinate id="EE12288B3844FFAFFB34FA7746DFFA3B" box="[1167,1230,1409,1435]" degrees="0.188" direction="north" orientation="latitude" pageId="5" pageNumber="238" precision="55" value="0.188">0.188</geoCoordinate>
<geoCoordinate id="EE12288B3844FFAFFB75FA77470DFA3B" box="[1230,1308,1409,1435]" degrees="0.203" direction="west" orientation="longitude" pageId="5" pageNumber="238" precision="55" value="-0.203">0.203</geoCoordinate>
vs.
<geoCoordinate id="EE12288B3844FFAFFAF4FA77479FFA3B" box="[1359,1422,1409,1435]" degrees="0.207" direction="north" orientation="latitude" pageId="5" pageNumber="238" precision="55" value="0.207">0.207</geoCoordinate>
<geoCoordinate id="EE12288B3844FFAFFA35FA7742CAFA1E" degrees="0.233" direction="west" orientation="longitude" pageId="5" pageNumber="238" precision="55" value="-0.233"> 0.233</geoCoordinate>
); a less flared original tail; a V-shaped, unbroken anterior-most white band on the original tail (vs. straight and usually not continuous); and by the restriction of prominent spines on the original tail to the outer margin (vs. prominent spines also on anterior one-third of the dorsal surface of the tail in
<taxonomicName id="4C2635CF3844FFAFFBAEFA1A468BF9A6" authorityName="Hoskin, Couper, Schneider &amp; Covacevich" authorityYear="2000" box="[1045,1178,1516,1542]" class="Reptilia" family="Carphodactylidae" genus="Phyllurus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="238" phylum="Chordata" rank="species" species="amnicola">
<emphasis id="B952925E3844FFAFFBAEFA1A468BF9A6" box="[1045,1178,1516,1542]" italics="true" pageId="5" pageNumber="238">P. amnicola</emphasis>
</taxonomicName>
).
<taxonomicName id="4C2635CF3844FFAFFB08FA1A478DF9A6" authority="Hoskin, 2023" authorityName="Hoskin" authorityYear="2023" box="[1203,1436,1516,1542]" class="Reptilia" family="Carphodactylidae" genus="Phyllurus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="238" phylum="Chordata" rank="species" species="fimbriatus" status="sp. nov.">
<emphasis id="B952925E3844FFAFFB08FA1A478DF9A6" box="[1203,1436,1516,1542]" italics="true" pageId="5" pageNumber="238">Phyllurus fimbriatus</emphasis>
</taxonomicName>
<emphasis id="B952925E3844FFAFFF2CF9F942E1F989" bold="true" box="[151,240,1551,1577]" pageId="5" pageNumber="238">
<taxonomicNameLabel id="A2612F253844FFAFFF2CF9F942E1F989" box="[151,240,1551,1577]" pageId="5" pageNumber="238" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="4C2635CF3844FFAFFE38F9E643E5F989" authorityName="Couper, Covacevich &amp; Moritz" authorityYear="1993" box="[387,500,1551,1577]" class="Reptilia" family="Carphodactylidae" genus="Phyllurus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="238" phylum="Chordata" rank="species" species="nepthys">
<emphasis id="B952925E3844FFAFFE38F9E643E5F989" box="[387,500,1551,1577]" italics="true" pageId="5" pageNumber="238">P. nepthys</emphasis>
</taxonomicName>
by larger body size (SVL av.
<quantity id="4CDEE3A93844FFAFFC84F9F9418FF98A" box="[831,926,1551,1578]" metricMagnitude="-1" metricUnit="m" metricValue="1.05" pageId="5" pageNumber="238" unit="mm" value="105.0">105 mm</quantity>
, max.
<quantity id="4CDEE3A93844FFAFFC5CF9F94653F98A" box="[999,1090,1551,1578]" metricMagnitude="-1" metricUnit="m" metricValue="1.13" pageId="5" pageNumber="238" unit="mm" value="113.0">113 mm</quantity>
vs. av.
<quantity id="4CDEE3A93844FFAFFB2EF9F946F7F98A" box="[1173,1254,1551,1578]" metricMagnitude="-2" metricUnit="m" metricValue="9.1" pageId="5" pageNumber="238" unit="mm" value="91.0">91 mm</quantity>
, max.
<quantity id="4CDEE3A93844FFAFFA94F9F9479EF989" box="[1327,1423,1551,1577]" metricMagnitude="-1" metricUnit="m" metricValue="1.03" pageId="5" pageNumber="238" unit="mm" value="103.0">103 mm</quantity>
); more flared original and regenerated tail; V-shaped anterior-most white band on the original tail (vs. straight); much less spinose dorsal surfaces of head, body and limbs (vs. extremely spinose); spines restricted to margin of flared portion of original tail (vs. prominent spines over dorsal and marginal surfaces); no spines on regenerated tail (vs. covered in spines); and immaculate white ventral surfaces (vs. ventral surfaces peppered with brown).
</paragraph>
</subSubSection>
<subSubSection id="C33C1DC73844FFAFFF7CF9344112F883" pageId="5" pageNumber="238" type="distribution">
<paragraph id="8B994E4C3844FFAFFF7CF9344112F883" blockId="5.[151,1437,1337,2043]" pageId="5" pageNumber="238">
<emphasis id="B952925E3844FFAFFF7CF9344370F97C" bold="true" box="[199,353,1730,1756]" pageId="5" pageNumber="238">Distribution.</emphasis>
Appears to be restricted to Scawfell Island, approximately
<quantity id="4CDEE3A93844FFAFFBBAF9344658F97C" box="[1025,1097,1730,1757]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="238" unit="km" value="50.0">50 km</quantity>
north-east of Mackay in mideastern
<collectingRegion id="49E280AE3844FFAFFF55F9104363F8A0" box="[238,370,1766,1792]" country="Australia" name="Queensland" pageId="5" pageNumber="238">Queensland</collectingRegion>
(
<figureCitation id="131D52C93844FFAFFE38F91043D4F8A0" box="[387,453,1766,1792]" captionStart="FIGURE 1" captionStartId="1.[151,250,1574,1598]" captionTargetBox="[173,1415,306,1548]" captionTargetId="figure-77@1.[173,1415,306,1548]" captionTargetPageId="1" captionText="FIGURE 1. Map showing the distributions of Phyllurus species and subspecies in mid-east Queensland." figureDoi="http://doi.org/10.5281/zenodo.7656251" httpUri="https://zenodo.org/record/7656251/files/figure.png" pageId="5" pageNumber="238">Fig. 1</figureCitation>
). On Scawfell Island, known from deeply-piled boulders in two gullies but other areas of similar habitat on the island have not been surveyed.
</paragraph>
</subSubSection>
<subSubSection id="C33C1DC73844FFA2FF7CF8DB43A7FE71" lastPageId="8" lastPageNumber="241" pageId="5" pageNumber="238" type="biology_ecology">
<paragraph id="8B994E4C3844FFAFFF7CF8DB404FF85A" blockId="5.[151,1437,1337,2043]" pageId="5" pageNumber="238">
<emphasis id="B952925E3844FFAFFF7CF8DB43A2F8E7" bold="true" box="[199,435,1837,1863]" pageId="5" pageNumber="238">Habitat and habits.</emphasis>
Deeply layered rock with associated rainforest vegetation. Both known sites consist of deeply-piled granite boulders in a gully (
<figureCitation id="131D52C93844FFAFFDE7F8A7408EF8CC" box="[604,671,1873,1900]" captionStart="FIGURE 5" captionStartId="7.[151,250,1900,1924]" captionTargetBox="[151,1435,1045,1876]" captionTargetId="figure-49@7.[151,1435,1045,1876]" captionTargetPageId="7" captionText="FIGURE 5. Habitat of P. fimbriatus sp. nov. on Scawfell Island. Photo: Conrad Hoskin." figureDoi="http://doi.org/10.5281/zenodo.7656259" httpUri="https://zenodo.org/record/7656259/files/figure.png" pageId="5" pageNumber="238">Fig. 5</figureCitation>
). The boulders are covered in ferns (e.g.,
<taxonomicName id="4C2635CF3844FFAFFBD6F8A746F2F8CB" box="[1133,1251,1873,1899]" class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" kingdom="Plantae" order="Polypodiales" pageId="5" pageNumber="238" phylum="Tracheophyta" rank="genus">
<emphasis id="B952925E3844FFAFFBD6F8A746F2F8CB" box="[1133,1251,1873,1899]" italics="true" pageId="5" pageNumber="238">Asplenium</emphasis>
</taxonomicName>
) and vines (e.g.,
<taxonomicName id="4C2635CF3844FFAFFF2CF88342C5F82E" box="[151,212,1909,1934]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="5" pageNumber="238" phylum="Tracheophyta" rank="genus">
<emphasis id="B952925E3844FFAFFF2CF88342C5F82E" box="[151,212,1909,1934]" italics="true" pageId="5" pageNumber="238">Hoya</emphasis>
</taxonomicName>
) and have a canopy of
<taxonomicName id="4C2635CF3844FFAFFE6DF8834005F82E" authorityName="Roding" authorityYear="1798" box="[470,532,1909,1934]" class="Gastropoda" family="Ficidae" genus="Ficus" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="238" phylum="Mollusca" rank="genus">
<emphasis id="B952925E3844FFAFFE6DF8834005F82E" box="[470,532,1909,1934]" italics="true" pageId="5" pageNumber="238">Ficus</emphasis>
</taxonomicName>
and other rainforest trees (
<figureCitation id="131D52C93844FFAFFCFBF8834195F82F" box="[832,900,1908,1935]" captionStart="FIGURE 5" captionStartId="7.[151,250,1900,1924]" captionTargetBox="[151,1435,1045,1876]" captionTargetId="figure-49@7.[151,1435,1045,1876]" captionTargetPageId="7" captionText="FIGURE 5. Habitat of P. fimbriatus sp. nov. on Scawfell Island. Photo: Conrad Hoskin." figureDoi="http://doi.org/10.5281/zenodo.7656259" httpUri="https://zenodo.org/record/7656259/files/figure.png" pageId="5" pageNumber="238">Fig. 5</figureCitation>
).
<taxonomicName id="4C2635CF3844FFAFFC22F883466EF82F" authority="Hoskin, 2023" authorityName="Hoskin" authorityYear="2023" box="[921,1151,1909,1935]" class="Reptilia" family="Carphodactylidae" genus="Phyllurus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="238" phylum="Chordata" rank="species" species="fimbriatus" status="sp. nov.">
<emphasis id="B952925E3844FFAFFC22F883466EF82F" box="[921,1151,1909,1935]" italics="true" pageId="5" pageNumber="238">Phyllurus fimbriatus</emphasis>
</taxonomicName>
<emphasis id="B952925E3844FFAFFB3DF88346CEF82F" bold="true" box="[1158,1247,1909,1935]" pageId="5" pageNumber="238">
<taxonomicNameLabel id="A2612F253844FFAFFB3DF88346CEF82F" box="[1158,1247,1909,1935]" pageId="5" pageNumber="238" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was restricted to areas with deeply-piled boulders—surveys in adjacent areas, including other areas of rainforest vegetation along the gullies and slopes did not find the species. Individuals were found on rock surfaces at night, and presumably retreat deep among the boulders during the day.
</paragraph>
<caption id="DF591EC43847FFACFF23FF6046ACFF59" ID-Table-UUID="DF591EC43847FFACFF23FF6046ACFF59" httpUri="http://table.plazi.org/id/DF591EC43847FFACFF23FF6046ACFF59" pageId="6" pageNumber="239" startId="6.[152,249,150,176]" targetBox="[166,1410,264,1966]" targetIsTable="true" targetPageId="6">
<paragraph id="8B994E4C3847FFACFF23FF6046ACFF59" blockId="6.[151,1437,150,249]" pageId="6" pageNumber="239">
<emphasis id="B952925E3847FFACFF23FF604304FF10" bold="true" box="[152,277,150,176]" pageId="6" pageNumber="239">TABLE 1.</emphasis>
Morphological measurements, proportions, and scale counts for the type specimens, and SVL data from individuals measured in the field. All measurements are in mm. Field data is only for individuals deemed to be mature adults (see text), is split by sex, and is presented as average followed by range in brackets.
</paragraph>
</caption>
<paragraph id="8B994E4C3847FFACFE7DFEFE41F6F80E" pageId="6" pageNumber="239">
<table id="F926BCEC38470055FF1DFEFE4793F80E" box="[166,1410,264,1966]" gridcols="7" gridrows="40" pageId="6" pageNumber="239">
<tr id="35164C0E38470055FF1DFEFE4793FEC8" box="[166,1410,264,360]" gridrow="0" pageId="6" pageNumber="239" rowspan-0="1">
<th id="76C7257238470055FE7DFEFE4054FEC8" box="[454,581,264,360]" gridcol="1" gridrow="0" pageId="6" pageNumber="239">
<emphasis id="B952925E3847FFACFE7DFEFE4054FE80" bold="true" box="[454,581,264,288]" pageId="6" pageNumber="239">QM J97578</emphasis>
Holotype Female
</th>
<th id="76C7257238470055FDC8FEFE40E3FEC8" box="[627,754,264,360]" gridcol="2" gridrow="0" pageId="6" pageNumber="239">
<emphasis id="B952925E3847FFACFDC8FEFE40E3FE80" bold="true" box="[627,754,264,288]" pageId="6" pageNumber="239">QM J97576</emphasis>
Paratype Female
</th>
<th id="76C7257238470055FC9AFEFE418EFEC8" box="[801,927,264,360]" gridcol="3" gridrow="0" pageId="6" pageNumber="239">
<emphasis id="B952925E3847FFACFC9AFEFE418EFE80" bold="true" box="[801,927,264,288]" pageId="6" pageNumber="239">QM J97577</emphasis>
Paratype Male
</th>
<th id="76C7257238470055FC75FEFE465DFEC8" box="[974,1100,264,360]" gridcol="4" gridrow="0" pageId="6" pageNumber="239">
<emphasis id="B952925E3847FFACFC75FEFE465DFE80" bold="true" box="[974,1100,264,288]" pageId="6" pageNumber="239">QM J97579</emphasis>
Paratype Female
</th>
<th id="76C7257238470055FBC0FEFE46F9FEC8" box="[1147,1256,264,360]" gridcol="5" gridrow="0" pageId="6" pageNumber="239">
<emphasis id="B952925E3847FFACFBC0FEFE46F9FE80" bold="true" box="[1147,1256,264,288]" pageId="6" pageNumber="239">Field data</emphasis>
Adult females
</th>
<th id="76C7257238470055FAAEFEFE4793FEC8" box="[1301,1410,264,360]" gridcol="6" gridrow="0" pageId="6" pageNumber="239">
<emphasis id="B952925E3847FFACFAAEFEFE4793FE80" bold="true" box="[1301,1410,264,288]" pageId="6" pageNumber="239">Field data</emphasis>
Adult males
</th>
</tr>
<tr id="35164C0E38470055FF1DFE8E4793FE31" box="[166,1410,376,401]" gridrow="1" pageId="6" pageNumber="239" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="76C7257238470055FF1DFE8E4383FE31" box="[166,402,376,401]" gridcol="0" gridrow="1" pageId="6" pageNumber="239">
<emphasis id="B952925E3847FFACFF1DFE8E4354FE30" bold="true" box="[166,325,376,400]" pageId="6" pageNumber="239">Measurements</emphasis>
</th>
<td id="76C7257238470055FBC0FE8E46F9FE31" box="[1147,1256,376,401]" gridcol="5" gridrow="1" pageId="6" pageNumber="239">N = 8</td>
<td id="76C7257238470055FAAEFE8E4793FE31" box="[1301,1410,376,401]" gridcol="6" gridrow="1" pageId="6" pageNumber="239">N = 8</td>
</tr>
<tr id="35164C0E38470055FF1DFE574793FE7D" box="[166,1410,417,477]" gridrow="2" pageId="6" pageNumber="239">
<th id="76C7257238470055FF1DFE574383FE7D" box="[166,402,417,477]" gridcol="0" gridrow="2" pageId="6" pageNumber="239">Snout to vent (SVL)</th>
<td id="76C7257238470055FE7DFE574054FE7D" box="[454,581,417,477]" gridcol="1" gridrow="2" pageId="6" pageNumber="239">104.2</td>
<td id="76C7257238470055FDC8FE5740E3FE7D" box="[627,754,417,477]" gridcol="2" gridrow="2" pageId="6" pageNumber="239">90.7</td>
<td id="76C7257238470055FC9AFE57418EFE7D" box="[801,927,417,477]" gridcol="3" gridrow="2" pageId="6" pageNumber="239">101.4</td>
<td id="76C7257238470055FC75FE57465DFE7D" box="[974,1100,417,477]" gridcol="4" gridrow="2" pageId="6" pageNumber="239">110.7</td>
<td id="76C7257238470055FBC0FE5746F9FE7D" box="[1147,1256,417,477]" gridcol="5" gridrow="2" pageId="6" pageNumber="239">109 (103113)</td>
<td id="76C7257238470055FAAEFE574793FE7D" box="[1301,1410,417,477]" gridcol="6" gridrow="2" pageId="6" pageNumber="239">102 (96105)</td>
</tr>
<tr id="35164C0E38470055FF1DFE1B4793FDA5" box="[166,1410,493,517]" gridrow="3" pageId="6" pageNumber="239" rowspan-5="1" rowspan-6="1">
<th id="76C7257238470055FF1DFE1B4383FDA5" box="[166,402,493,517]" gridcol="0" gridrow="3" pageId="6" pageNumber="239">Interlimb length (AG)</th>
<td id="76C7257238470055FE7DFE1B4054FDA5" box="[454,581,493,517]" gridcol="1" gridrow="3" pageId="6" pageNumber="239">50.3</td>
<td id="76C7257238470055FDC8FE1B40E3FDA5" box="[627,754,493,517]" gridcol="2" gridrow="3" pageId="6" pageNumber="239">42.7</td>
<td id="76C7257238470055FC9AFE1B418EFDA5" box="[801,927,493,517]" gridcol="3" gridrow="3" pageId="6" pageNumber="239">45.4</td>
<td id="76C7257238470055FC75FE1B465DFDA5" box="[974,1100,493,517]" gridcol="4" gridrow="3" pageId="6" pageNumber="239">51.0</td>
</tr>
<tr id="35164C0E38470055FF1DFDE34793FD8D" box="[166,1410,533,557]" gridrow="4" pageId="6" pageNumber="239" rowspan-5="1" rowspan-6="1">
<th id="76C7257238470055FF1DFDE34383FD8D" box="[166,402,533,557]" gridcol="0" gridrow="4" pageId="6" pageNumber="239">Forearm length (FL)</th>
<td id="76C7257238470055FE7DFDE34054FD8D" box="[454,581,533,557]" gridcol="1" gridrow="4" pageId="6" pageNumber="239">20.7</td>
<td id="76C7257238470055FDC8FDE340E3FD8D" box="[627,754,533,557]" gridcol="2" gridrow="4" pageId="6" pageNumber="239">18.3</td>
<td id="76C7257238470055FC9AFDE3418EFD8D" box="[801,927,533,557]" gridcol="3" gridrow="4" pageId="6" pageNumber="239">20.2</td>
<td id="76C7257238470055FC75FDE3465DFD8D" box="[974,1100,533,557]" gridcol="4" gridrow="4" pageId="6" pageNumber="239">21.2</td>
</tr>
<tr id="35164C0E38470055FF1DFDCB4793FDF6" box="[166,1410,573,598]" gridrow="5" pageId="6" pageNumber="239" rowspan-5="1" rowspan-6="1">
<th id="76C7257238470055FF1DFDCB4383FDF6" box="[166,402,573,598]" gridcol="0" gridrow="5" pageId="6" pageNumber="239">Lower hindlimb (LHL)</th>
<td id="76C7257238470055FE7DFDCB4054FDF6" box="[454,581,573,598]" gridcol="1" gridrow="5" pageId="6" pageNumber="239">23.2</td>
<td id="76C7257238470055FDC8FDCB40E3FDF6" box="[627,754,573,598]" gridcol="2" gridrow="5" pageId="6" pageNumber="239">21.4</td>
<td id="76C7257238470055FC9AFDCB418EFDF6" box="[801,927,573,598]" gridcol="3" gridrow="5" pageId="6" pageNumber="239">23.1</td>
<td id="76C7257238470055FC75FDCB465DFDF6" box="[974,1100,573,598]" gridcol="4" gridrow="5" pageId="6" pageNumber="239">24.8</td>
</tr>
<tr id="35164C0E38470055FF1DFD904793FDDE" box="[166,1410,614,638]" gridrow="6" pageId="6" pageNumber="239" rowspan-5="1" rowspan-6="1">
<th id="76C7257238470055FF1DFD904383FDDE" box="[166,402,614,638]" gridcol="0" gridrow="6" pageId="6" pageNumber="239">Full forelimb (L1)</th>
<td id="76C7257238470055FE7DFD904054FDDE" box="[454,581,614,638]" gridcol="1" gridrow="6" pageId="6" pageNumber="239">46.5</td>
<td id="76C7257238470055FDC8FD9040E3FDDE" box="[627,754,614,638]" gridcol="2" gridrow="6" pageId="6" pageNumber="239">43.3</td>
<td id="76C7257238470055FC9AFD90418EFDDE" box="[801,927,614,638]" gridcol="3" gridrow="6" pageId="6" pageNumber="239">44.2</td>
<td id="76C7257238470055FC75FD90465DFDDE" box="[974,1100,614,638]" gridcol="4" gridrow="6" pageId="6" pageNumber="239">51.9</td>
</tr>
<tr id="35164C0E38470055FF1DFD784793FD06" box="[166,1410,654,678]" gridrow="7" pageId="6" pageNumber="239" rowspan-5="1" rowspan-6="1">
<th id="76C7257238470055FF1DFD784383FD06" box="[166,402,654,678]" gridcol="0" gridrow="7" pageId="6" pageNumber="239">Full hindlimb (L2)</th>
<td id="76C7257238470055FE7DFD784054FD06" box="[454,581,654,678]" gridcol="1" gridrow="7" pageId="6" pageNumber="239">55.4</td>
<td id="76C7257238470055FDC8FD7840E3FD06" box="[627,754,654,678]" gridcol="2" gridrow="7" pageId="6" pageNumber="239">48.8</td>
<td id="76C7257238470055FC9AFD78418EFD06" box="[801,927,654,678]" gridcol="3" gridrow="7" pageId="6" pageNumber="239">51.9</td>
<td id="76C7257238470055FC75FD78465DFD06" box="[974,1100,654,678]" gridcol="4" gridrow="7" pageId="6" pageNumber="239">58.5</td>
</tr>
<tr id="35164C0E38470055FF1DFD404793FD6E" box="[166,1410,694,718]" gridrow="8" pageId="6" pageNumber="239" rowspan-5="1" rowspan-6="1">
<th id="76C7257238470055FF1DFD404383FD6E" box="[166,402,694,718]" gridcol="0" gridrow="8" pageId="6" pageNumber="239">Neck length (NL)</th>
<td id="76C7257238470055FE7DFD404054FD6E" box="[454,581,694,718]" gridcol="1" gridrow="8" pageId="6" pageNumber="239">19.5</td>
<td id="76C7257238470055FDC8FD4040E3FD6E" box="[627,754,694,718]" gridcol="2" gridrow="8" pageId="6" pageNumber="239">18.1</td>
<td id="76C7257238470055FC9AFD40418EFD6E" box="[801,927,694,718]" gridcol="3" gridrow="8" pageId="6" pageNumber="239">20.6</td>
<td id="76C7257238470055FC75FD40465DFD6E" box="[974,1100,694,718]" gridcol="4" gridrow="8" pageId="6" pageNumber="239">20.9</td>
</tr>
<tr id="35164C0E38470055FF1DFD284793FD57" box="[166,1410,734,759]" gridrow="9" pageId="6" pageNumber="239" rowspan-5="1" rowspan-6="1">
<th id="76C7257238470055FF1DFD284383FD57" box="[166,402,734,759]" gridcol="0" gridrow="9" pageId="6" pageNumber="239">Neck width (NW)</th>
<td id="76C7257238470055FE7DFD284054FD57" box="[454,581,734,759]" gridcol="1" gridrow="9" pageId="6" pageNumber="239">10.7</td>
<td id="76C7257238470055FDC8FD2840E3FD57" box="[627,754,734,759]" gridcol="2" gridrow="9" pageId="6" pageNumber="239">10.1</td>
<td id="76C7257238470055FC9AFD28418EFD57" box="[801,927,734,759]" gridcol="3" gridrow="9" pageId="6" pageNumber="239">12.1</td>
<td id="76C7257238470055FC75FD28465DFD57" box="[974,1100,734,759]" gridcol="4" gridrow="9" pageId="6" pageNumber="239">11.5</td>
</tr>
<tr id="35164C0E38470055FF1DFCF14793FCBF" box="[166,1410,775,799]" gridrow="10" pageId="6" pageNumber="239" rowspan-5="1" rowspan-6="1">
<th id="76C7257238470055FF1DFCF14383FCBF" box="[166,402,775,799]" gridcol="0" gridrow="10" pageId="6" pageNumber="239">Head length (HL)</th>
<td id="76C7257238470055FE7DFCF14054FCBF" box="[454,581,775,799]" gridcol="1" gridrow="10" pageId="6" pageNumber="239">27.7</td>
<td id="76C7257238470055FDC8FCF140E3FCBF" box="[627,754,775,799]" gridcol="2" gridrow="10" pageId="6" pageNumber="239">25.1</td>
<td id="76C7257238470055FC9AFCF1418EFCBF" box="[801,927,775,799]" gridcol="3" gridrow="10" pageId="6" pageNumber="239">27.4</td>
<td id="76C7257238470055FC75FCF1465DFCBF" box="[974,1100,775,799]" gridcol="4" gridrow="10" pageId="6" pageNumber="239">30.0</td>
</tr>
<tr id="35164C0E38470055FF1DFCD94793FCE7" box="[166,1410,815,839]" gridrow="11" pageId="6" pageNumber="239" rowspan-5="1" rowspan-6="1">
<th id="76C7257238470055FF1DFCD94383FCE7" box="[166,402,815,839]" gridcol="0" gridrow="11" pageId="6" pageNumber="239">Head width (HW)</th>
<td id="76C7257238470055FE7DFCD94054FCE7" box="[454,581,815,839]" gridcol="1" gridrow="11" pageId="6" pageNumber="239">22.6</td>
<td id="76C7257238470055FDC8FCD940E3FCE7" box="[627,754,815,839]" gridcol="2" gridrow="11" pageId="6" pageNumber="239">21.0</td>
<td id="76C7257238470055FC9AFCD9418EFCE7" box="[801,927,815,839]" gridcol="3" gridrow="11" pageId="6" pageNumber="239">23.4</td>
<td id="76C7257238470055FC75FCD9465DFCE7" box="[974,1100,815,839]" gridcol="4" gridrow="11" pageId="6" pageNumber="239">24.3</td>
</tr>
<tr id="35164C0E38470055FF1DFCA14793FCCF" box="[166,1410,855,879]" gridrow="12" pageId="6" pageNumber="239" rowspan-5="1" rowspan-6="1">
<th id="76C7257238470055FF1DFCA14383FCCF" box="[166,402,855,879]" gridcol="0" gridrow="12" pageId="6" pageNumber="239">Head depth (HD)</th>
<td id="76C7257238470055FE7DFCA14054FCCF" box="[454,581,855,879]" gridcol="1" gridrow="12" pageId="6" pageNumber="239">11.3</td>
<td id="76C7257238470055FDC8FCA140E3FCCF" box="[627,754,855,879]" gridcol="2" gridrow="12" pageId="6" pageNumber="239">10.4</td>
<td id="76C7257238470055FC9AFCA1418EFCCF" box="[801,927,855,879]" gridcol="3" gridrow="12" pageId="6" pageNumber="239">11.1</td>
<td id="76C7257238470055FC75FCA1465DFCCF" box="[974,1100,855,879]" gridcol="4" gridrow="12" pageId="6" pageNumber="239">11.9</td>
</tr>
<tr id="35164C0E38470055FF1DFC894793FC38" box="[166,1410,895,920]" gridrow="13" pageId="6" pageNumber="239" rowspan-5="1" rowspan-6="1">
<th id="76C7257238470055FF1DFC894383FC38" box="[166,402,895,920]" gridcol="0" gridrow="13" pageId="6" pageNumber="239">Snout length (SL)</th>
<td id="76C7257238470055FE7DFC894054FC38" box="[454,581,895,920]" gridcol="1" gridrow="13" pageId="6" pageNumber="239">12.9</td>
<td id="76C7257238470055FDC8FC8940E3FC38" box="[627,754,895,920]" gridcol="2" gridrow="13" pageId="6" pageNumber="239">11.3</td>
<td id="76C7257238470055FC9AFC89418EFC38" box="[801,927,895,920]" gridcol="3" gridrow="13" pageId="6" pageNumber="239">12.0</td>
<td id="76C7257238470055FC75FC89465DFC38" box="[974,1100,895,920]" gridcol="4" gridrow="13" pageId="6" pageNumber="239">14.5</td>
</tr>
<tr id="35164C0E38470055FF1DFC514793FC60" box="[166,1410,935,960]" gridrow="14" pageId="6" pageNumber="239" rowspan-5="1" rowspan-6="1">
<th id="76C7257238470055FF1DFC514383FC60" box="[166,402,935,960]" gridcol="0" gridrow="14" pageId="6" pageNumber="239">Eye diameter (ED)</th>
<td id="76C7257238470055FE7DFC514054FC60" box="[454,581,935,960]" gridcol="1" gridrow="14" pageId="6" pageNumber="239">6.1</td>
<td id="76C7257238470055FDC8FC5140E3FC60" box="[627,754,935,960]" gridcol="2" gridrow="14" pageId="6" pageNumber="239">5.8</td>
<td id="76C7257238470055FC9AFC51418EFC60" box="[801,927,935,960]" gridcol="3" gridrow="14" pageId="6" pageNumber="239">6.5</td>
<td id="76C7257238470055FC75FC51465DFC60" box="[974,1100,935,960]" gridcol="4" gridrow="14" pageId="6" pageNumber="239">6.3</td>
</tr>
<tr id="35164C0E38470055FF1DFC264793FC48" box="[166,1410,976,1000]" gridrow="15" pageId="6" pageNumber="239">
<th id="76C7257238470055FF1DFC264793FC48" box="[166,1410,976,1000]" colspan="7" colspanRight="6" gridcol="0" gridrow="15" pageId="6" pageNumber="239">Original Tail</th>
</tr>
<tr id="35164C0E38470055FF1DFC0E4793FBB0" box="[166,1410,1016,1040]" gridrow="16" pageId="6" pageNumber="239" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<th id="76C7257238470055FF1DFC0E4383FBB0" box="[166,402,1016,1040]" gridcol="0" gridrow="16" pageId="6" pageNumber="239">Tail Length (TL)</th>
<td id="76C7257238470055FE7DFC0E4054FBB0" box="[454,581,1016,1040]" gridcol="1" gridrow="16" pageId="6" pageNumber="239">78.9</td>
<td id="76C7257238470055FDC8FC0E40E3FBB0" box="[627,754,1016,1040]" gridcol="2" gridrow="16" pageId="6" pageNumber="239">70.8</td>
</tr>
<tr id="35164C0E38470055FF1DFBD64793FB99" box="[166,1410,1056,1081]" gridrow="17" pageId="6" pageNumber="239" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<th id="76C7257238470055FF1DFBD64383FB99" box="[166,402,1056,1081]" gridcol="0" gridrow="17" pageId="6" pageNumber="239">Tail tip length (TT)</th>
<td id="76C7257238470055FE7DFBD64054FB99" box="[454,581,1056,1081]" gridcol="1" gridrow="17" pageId="6" pageNumber="239">42.3</td>
<td id="76C7257238470055FDC8FBD640E3FB99" box="[627,754,1056,1081]" gridcol="2" gridrow="17" pageId="6" pageNumber="239">38.1</td>
</tr>
<tr id="35164C0E38470055FF1DFBBF4793FBC1" box="[166,1410,1097,1121]" gridrow="18" pageId="6" pageNumber="239" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<th id="76C7257238470055FF1DFBBF4383FBC1" box="[166,402,1097,1121]" gridcol="0" gridrow="18" pageId="6" pageNumber="239">Tail Width (TW)</th>
<td id="76C7257238470055FE7DFBBF4054FBC1" box="[454,581,1097,1121]" gridcol="1" gridrow="18" pageId="6" pageNumber="239">18.6</td>
<td id="76C7257238470055FDC8FBBF40E3FBC1" box="[627,754,1097,1121]" gridcol="2" gridrow="18" pageId="6" pageNumber="239">19.4</td>
</tr>
<tr id="35164C0E38470055FF1DFB874793FB29" box="[166,1410,1137,1161]" gridrow="19" pageId="6" pageNumber="239" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<th id="76C7257238470055FF1DFB874383FB29" box="[166,402,1137,1161]" gridcol="0" gridrow="19" pageId="6" pageNumber="239">Tail Depth (TD)</th>
<td id="76C7257238470055FE7DFB874054FB29" box="[454,581,1137,1161]" gridcol="1" gridrow="19" pageId="6" pageNumber="239">6.2</td>
<td id="76C7257238470055FDC8FB8740E3FB29" box="[627,754,1137,1161]" gridcol="2" gridrow="19" pageId="6" pageNumber="239">6.6</td>
</tr>
<tr id="35164C0E38470055FF1DFB6F4793FB11" box="[166,1410,1177,1201]" gridrow="20" pageId="6" pageNumber="239">
<th id="76C7257238470055FF1DFB6F4793FB11" box="[166,1410,1177,1201]" colspan="7" colspanRight="6" gridcol="0" gridrow="20" pageId="6" pageNumber="239">Regenerated Tail</th>
</tr>
<tr id="35164C0E38470055FF1DFB374793FB79" box="[166,1410,1217,1241]" gridrow="21" pageId="6" pageNumber="239" rowspan-1="1" rowspan-2="1" rowspan-5="1" rowspan-6="1">
<th id="76C7257238470055FF1DFB374383FB79" box="[166,402,1217,1241]" gridcol="0" gridrow="21" pageId="6" pageNumber="239">Tail Length (TL)</th>
<td id="76C7257238470055FC9AFB37418EFB79" box="[801,927,1217,1241]" gridcol="3" gridrow="21" pageId="6" pageNumber="239">50.5</td>
<td id="76C7257238470055FC75FB37465DFB79" box="[974,1100,1217,1241]" gridcol="4" gridrow="21" pageId="6" pageNumber="239">52.4</td>
</tr>
<tr id="35164C0E38470055FF1DFB1F4793FAA2" box="[166,1410,1257,1282]" gridrow="22" pageId="6" pageNumber="239" rowspan-1="1" rowspan-2="1" rowspan-5="1" rowspan-6="1">
<th id="76C7257238470055FF1DFB1F4383FAA2" box="[166,402,1257,1282]" gridcol="0" gridrow="22" pageId="6" pageNumber="239">Tail tip length (TT)</th>
<td id="76C7257238470055FC9AFB1F418EFAA2" box="[801,927,1257,1282]" gridcol="3" gridrow="22" pageId="6" pageNumber="239">25.4</td>
<td id="76C7257238470055FC75FB1F465DFAA2" box="[974,1100,1257,1282]" gridcol="4" gridrow="22" pageId="6" pageNumber="239">24.9</td>
</tr>
<tr id="35164C0E38470055FF1DFAE44793FA8A" box="[166,1410,1298,1322]" gridrow="23" pageId="6" pageNumber="239" rowspan-1="1" rowspan-2="1" rowspan-5="1" rowspan-6="1">
<th id="76C7257238470055FF1DFAE44383FA8A" box="[166,402,1298,1322]" gridcol="0" gridrow="23" pageId="6" pageNumber="239">Tail Width (TW)</th>
<td id="76C7257238470055FC9AFAE4418EFA8A" box="[801,927,1298,1322]" gridcol="3" gridrow="23" pageId="6" pageNumber="239">25.0</td>
<td id="76C7257238470055FC75FAE4465DFA8A" box="[974,1100,1298,1322]" gridcol="4" gridrow="23" pageId="6" pageNumber="239">23.4</td>
</tr>
<tr id="35164C0E38470055FF1DFACC4793FAF2" box="[166,1410,1338,1362]" gridrow="24" pageId="6" pageNumber="239" rowspan-1="1" rowspan-2="1" rowspan-5="1" rowspan-6="1">
<th id="76C7257238470055FF1DFACC4383FAF2" box="[166,402,1338,1362]" gridcol="0" gridrow="24" pageId="6" pageNumber="239">Tail Depth (TD)</th>
<td id="76C7257238470055FC9AFACC418EFAF2" box="[801,927,1338,1362]" gridcol="3" gridrow="24" pageId="6" pageNumber="239">8.3</td>
<td id="76C7257238470055FC75FACC465DFAF2" box="[974,1100,1338,1362]" gridcol="4" gridrow="24" pageId="6" pageNumber="239">8.5</td>
</tr>
<tr id="35164C0E38470055FF1DFA944793FADA" box="[166,1410,1378,1402]" gridrow="25" pageId="6" pageNumber="239">
<th id="76C7257238470055FF1DFA944793FADA" box="[166,1410,1378,1402]" colspan="7" colspanRight="6" gridcol="0" gridrow="25" pageId="6" pageNumber="239">Scale counts</th>
</tr>
<tr id="35164C0E38470055FF1DFA7C4793FA03" box="[166,1410,1418,1443]" gridrow="26" pageId="6" pageNumber="239" rowspan-5="1" rowspan-6="1">
<th id="76C7257238470055FF1DFA7C4383FA03" box="[166,402,1418,1443]" gridcol="0" gridrow="26" pageId="6" pageNumber="239">Rostral groove %</th>
<td id="76C7257238470055FE7DFA7C4054FA03" box="[454,581,1418,1443]" gridcol="1" gridrow="26" pageId="6" pageNumber="239">50</td>
<td id="76C7257238470055FDC8FA7C40E3FA03" box="[627,754,1418,1443]" gridcol="2" gridrow="26" pageId="6" pageNumber="239">20</td>
<td id="76C7257238470055FC9AFA7C418EFA03" box="[801,927,1418,1443]" gridcol="3" gridrow="26" pageId="6" pageNumber="239">40</td>
<td id="76C7257238470055FC75FA7C465DFA03" box="[974,1100,1418,1443]" gridcol="4" gridrow="26" pageId="6" pageNumber="239">40</td>
</tr>
<tr id="35164C0E38470055FF1DFA454793FA6B" box="[166,1410,1459,1483]" gridrow="27" pageId="6" pageNumber="239" rowspan-5="1" rowspan-6="1">
<th id="76C7257238470055FF1DFA454383FA6B" box="[166,402,1459,1483]" gridcol="0" gridrow="27" pageId="6" pageNumber="239">Internasals</th>
<td id="76C7257238470055FE7DFA454054FA6B" box="[454,581,1459,1483]" gridcol="1" gridrow="27" pageId="6" pageNumber="239">7</td>
<td id="76C7257238470055FDC8FA4540E3FA6B" box="[627,754,1459,1483]" gridcol="2" gridrow="27" pageId="6" pageNumber="239">6</td>
<td id="76C7257238470055FC9AFA45418EFA6B" box="[801,927,1459,1483]" gridcol="3" gridrow="27" pageId="6" pageNumber="239">7</td>
<td id="76C7257238470055FC75FA45465DFA6B" box="[974,1100,1459,1483]" gridcol="4" gridrow="27" pageId="6" pageNumber="239">7</td>
</tr>
<tr id="35164C0E38470055FF1DFA2D4793FA53" box="[166,1410,1499,1523]" gridrow="28" pageId="6" pageNumber="239" rowspan-5="1" rowspan-6="1">
<th id="76C7257238470055FF1DFA2D4383FA53" box="[166,402,1499,1523]" gridcol="0" gridrow="28" pageId="6" pageNumber="239">Supralabials</th>
<td id="76C7257238470055FE7DFA2D4054FA53" box="[454,581,1499,1523]" gridcol="1" gridrow="28" pageId="6" pageNumber="239">15</td>
<td id="76C7257238470055FDC8FA2D40E3FA53" box="[627,754,1499,1523]" gridcol="2" gridrow="28" pageId="6" pageNumber="239">14</td>
<td id="76C7257238470055FC9AFA2D418EFA53" box="[801,927,1499,1523]" gridcol="3" gridrow="28" pageId="6" pageNumber="239">16</td>
<td id="76C7257238470055FC75FA2D465DFA53" box="[974,1100,1499,1523]" gridcol="4" gridrow="28" pageId="6" pageNumber="239">15</td>
</tr>
<tr id="35164C0E38470055FF1DF9F54793F9BB" box="[166,1410,1539,1563]" gridrow="29" pageId="6" pageNumber="239" rowspan-5="1" rowspan-6="1">
<th id="76C7257238470055FF1DF9F54383F9BB" box="[166,402,1539,1563]" gridcol="0" gridrow="29" pageId="6" pageNumber="239">Infralabials</th>
<td id="76C7257238470055FE7DF9F54054F9BB" box="[454,581,1539,1563]" gridcol="1" gridrow="29" pageId="6" pageNumber="239">16</td>
<td id="76C7257238470055FDC8F9F540E3F9BB" box="[627,754,1539,1563]" gridcol="2" gridrow="29" pageId="6" pageNumber="239">13</td>
<td id="76C7257238470055FC9AF9F5418EF9BB" box="[801,927,1539,1563]" gridcol="3" gridrow="29" pageId="6" pageNumber="239">14</td>
<td id="76C7257238470055FC75F9F5465DF9BB" box="[974,1100,1539,1563]" gridcol="4" gridrow="29" pageId="6" pageNumber="239">15</td>
</tr>
<tr id="35164C0E38470055FF1DF9DD4793F9E4" box="[166,1410,1579,1604]" gridrow="30" pageId="6" pageNumber="239" rowspan-5="1" rowspan-6="1">
<th id="76C7257238470055FF1DF9DD4383F9E4" box="[166,402,1579,1604]" gridcol="0" gridrow="30" pageId="6" pageNumber="239">1st finger lamellae</th>
<td id="76C7257238470055FE7DF9DD4054F9E4" box="[454,581,1579,1604]" gridcol="1" gridrow="30" pageId="6" pageNumber="239">12</td>
<td id="76C7257238470055FDC8F9DD40E3F9E4" box="[627,754,1579,1604]" gridcol="2" gridrow="30" pageId="6" pageNumber="239">12</td>
<td id="76C7257238470055FC9AF9DD418EF9E4" box="[801,927,1579,1604]" gridcol="3" gridrow="30" pageId="6" pageNumber="239">12</td>
<td id="76C7257238470055FC75F9DD465DF9E4" box="[974,1100,1579,1604]" gridcol="4" gridrow="30" pageId="6" pageNumber="239">13</td>
</tr>
<tr id="35164C0E38470055FF1DF9A24793F9CC" box="[166,1410,1620,1644]" gridrow="31" pageId="6" pageNumber="239" rowspan-5="1" rowspan-6="1">
<th id="76C7257238470055FF1DF9A24383F9CC" box="[166,402,1620,1644]" gridcol="0" gridrow="31" pageId="6" pageNumber="239">2nd finger lamellae</th>
<td id="76C7257238470055FE7DF9A24054F9CC" box="[454,581,1620,1644]" gridcol="1" gridrow="31" pageId="6" pageNumber="239">15</td>
<td id="76C7257238470055FDC8F9A240E3F9CC" box="[627,754,1620,1644]" gridcol="2" gridrow="31" pageId="6" pageNumber="239">17</td>
<td id="76C7257238470055FC9AF9A2418EF9CC" box="[801,927,1620,1644]" gridcol="3" gridrow="31" pageId="6" pageNumber="239">15</td>
<td id="76C7257238470055FC75F9A2465DF9CC" box="[974,1100,1620,1644]" gridcol="4" gridrow="31" pageId="6" pageNumber="239">16</td>
</tr>
<tr id="35164C0E38470055FF1DF98A4793F934" box="[166,1410,1660,1684]" gridrow="32" pageId="6" pageNumber="239" rowspan-5="1" rowspan-6="1">
<th id="76C7257238470055FF1DF98A4383F934" box="[166,402,1660,1684]" gridcol="0" gridrow="32" pageId="6" pageNumber="239">3rd finger lamellae</th>
<td id="76C7257238470055FE7DF98A4054F934" box="[454,581,1660,1684]" gridcol="1" gridrow="32" pageId="6" pageNumber="239">16</td>
<td id="76C7257238470055FDC8F98A40E3F934" box="[627,754,1660,1684]" gridcol="2" gridrow="32" pageId="6" pageNumber="239">18</td>
<td id="76C7257238470055FC9AF98A418EF934" box="[801,927,1660,1684]" gridcol="3" gridrow="32" pageId="6" pageNumber="239">16</td>
<td id="76C7257238470055FC75F98A465DF934" box="[974,1100,1660,1684]" gridcol="4" gridrow="32" pageId="6" pageNumber="239">18</td>
</tr>
<tr id="35164C0E38470055FF1DF9524793F91C" box="[166,1410,1700,1724]" gridrow="33" pageId="6" pageNumber="239" rowspan-5="1" rowspan-6="1">
<th id="76C7257238470055FF1DF9524383F91C" box="[166,402,1700,1724]" gridcol="0" gridrow="33" pageId="6" pageNumber="239">4th finger lamellae</th>
<td id="76C7257238470055FE7DF9524054F91C" box="[454,581,1700,1724]" gridcol="1" gridrow="33" pageId="6" pageNumber="239">15</td>
<td id="76C7257238470055FDC8F95240E3F91C" box="[627,754,1700,1724]" gridcol="2" gridrow="33" pageId="6" pageNumber="239">18</td>
<td id="76C7257238470055FC9AF952418EF91C" box="[801,927,1700,1724]" gridcol="3" gridrow="33" pageId="6" pageNumber="239">15</td>
<td id="76C7257238470055FC75F952465DF91C" box="[974,1100,1700,1724]" gridcol="4" gridrow="33" pageId="6" pageNumber="239">17</td>
</tr>
<tr id="35164C0E38470055FF1DF93A4793F945" box="[166,1410,1740,1765]" gridrow="34" pageId="6" pageNumber="239" rowspan-5="1" rowspan-6="1">
<th id="76C7257238470055FF1DF93A4383F945" box="[166,402,1740,1765]" gridcol="0" gridrow="34" pageId="6" pageNumber="239">5th finger lamellae</th>
<td id="76C7257238470055FE7DF93A4054F945" box="[454,581,1740,1765]" gridcol="1" gridrow="34" pageId="6" pageNumber="239">18</td>
<td id="76C7257238470055FDC8F93A40E3F945" box="[627,754,1740,1765]" gridcol="2" gridrow="34" pageId="6" pageNumber="239">15</td>
<td id="76C7257238470055FC9AF93A418EF945" box="[801,927,1740,1765]" gridcol="3" gridrow="34" pageId="6" pageNumber="239">14</td>
<td id="76C7257238470055FC75F93A465DF945" box="[974,1100,1740,1765]" gridcol="4" gridrow="34" pageId="6" pageNumber="239">16</td>
</tr>
<tr id="35164C0E38470055FF1DF9024793F8AD" box="[166,1410,1780,1805]" gridrow="35" pageId="6" pageNumber="239" rowspan-5="1" rowspan-6="1">
<th id="76C7257238470055FF1DF9024383F8AD" box="[166,402,1780,1805]" gridcol="0" gridrow="35" pageId="6" pageNumber="239">1st toe lamellae</th>
<td id="76C7257238470055FE7DF9024054F8AD" box="[454,581,1780,1805]" gridcol="1" gridrow="35" pageId="6" pageNumber="239">12</td>
<td id="76C7257238470055FDC8F90240E3F8AD" box="[627,754,1780,1805]" gridcol="2" gridrow="35" pageId="6" pageNumber="239">12</td>
<td id="76C7257238470055FC9AF902418EF8AD" box="[801,927,1780,1805]" gridcol="3" gridrow="35" pageId="6" pageNumber="239">12</td>
<td id="76C7257238470055FC75F902465DF8AD" box="[974,1100,1780,1805]" gridcol="4" gridrow="35" pageId="6" pageNumber="239">12</td>
</tr>
<tr id="35164C0E38470055FF1DF8EB4793F895" box="[166,1410,1821,1845]" gridrow="36" pageId="6" pageNumber="239" rowspan-5="1" rowspan-6="1">
<th id="76C7257238470055FF1DF8EB4383F895" box="[166,402,1821,1845]" gridcol="0" gridrow="36" pageId="6" pageNumber="239">2nd toe lamellae</th>
<td id="76C7257238470055FE7DF8EB4054F895" box="[454,581,1821,1845]" gridcol="1" gridrow="36" pageId="6" pageNumber="239">17</td>
<td id="76C7257238470055FDC8F8EB40E3F895" box="[627,754,1821,1845]" gridcol="2" gridrow="36" pageId="6" pageNumber="239">17</td>
<td id="76C7257238470055FC9AF8EB418EF895" box="[801,927,1821,1845]" gridcol="3" gridrow="36" pageId="6" pageNumber="239">17</td>
<td id="76C7257238470055FC75F8EB465DF895" box="[974,1100,1821,1845]" gridcol="4" gridrow="36" pageId="6" pageNumber="239">17</td>
</tr>
<tr id="35164C0E38470055FF1DF8B34793F8FD" box="[166,1410,1861,1885]" gridrow="37" pageId="6" pageNumber="239" rowspan-5="1" rowspan-6="1">
<th id="76C7257238470055FF1DF8B34383F8FD" box="[166,402,1861,1885]" gridcol="0" gridrow="37" pageId="6" pageNumber="239">3rd toe lamellae</th>
<td id="76C7257238470055FE7DF8B34054F8FD" box="[454,581,1861,1885]" gridcol="1" gridrow="37" pageId="6" pageNumber="239">18</td>
<td id="76C7257238470055FDC8F8B340E3F8FD" box="[627,754,1861,1885]" gridcol="2" gridrow="37" pageId="6" pageNumber="239">18</td>
<td id="76C7257238470055FC9AF8B3418EF8FD" box="[801,927,1861,1885]" gridcol="3" gridrow="37" pageId="6" pageNumber="239">18</td>
<td id="76C7257238470055FC75F8B3465DF8FD" box="[974,1100,1861,1885]" gridcol="4" gridrow="37" pageId="6" pageNumber="239">20</td>
</tr>
<tr id="35164C0E38470055FF1DF89B4793F826" box="[166,1410,1901,1926]" gridrow="38" pageId="6" pageNumber="239" rowspan-5="1" rowspan-6="1">
<th id="76C7257238470055FF1DF89B4383F826" box="[166,402,1901,1926]" gridcol="0" gridrow="38" pageId="6" pageNumber="239">4th toe lamellae</th>
<td id="76C7257238470055FE7DF89B4054F826" box="[454,581,1901,1926]" gridcol="1" gridrow="38" pageId="6" pageNumber="239">18</td>
<td id="76C7257238470055FDC8F89B40E3F826" box="[627,754,1901,1926]" gridcol="2" gridrow="38" pageId="6" pageNumber="239">19</td>
<td id="76C7257238470055FC9AF89B418EF826" box="[801,927,1901,1926]" gridcol="3" gridrow="38" pageId="6" pageNumber="239">20</td>
<td id="76C7257238470055FC75F89B465DF826" box="[974,1100,1901,1926]" gridcol="4" gridrow="38" pageId="6" pageNumber="239">19</td>
</tr>
<tr id="35164C0E38470055FF1DF8604793F80E" box="[166,1410,1942,1966]" gridrow="39" pageId="6" pageNumber="239" rowspan-5="1" rowspan-6="1">
<th id="76C7257238470055FF1DF8604383F80E" box="[166,402,1942,1966]" gridcol="0" gridrow="39" pageId="6" pageNumber="239">5th toe lamellae</th>
<td id="76C7257238470055FE7DF8604054F80E" box="[454,581,1942,1966]" gridcol="1" gridrow="39" pageId="6" pageNumber="239">20</td>
<td id="76C7257238470055FDC8F86040E3F80E" box="[627,754,1942,1966]" gridcol="2" gridrow="39" pageId="6" pageNumber="239">19</td>
<td id="76C7257238470055FC9AF860418EF80E" box="[801,927,1942,1966]" gridcol="3" gridrow="39" pageId="6" pageNumber="239">18</td>
<td id="76C7257238470055FC75F860465DF80E" box="[974,1100,1942,1966]" gridcol="4" gridrow="39" pageId="6" pageNumber="239">19</td>
</tr>
</table>
</paragraph>
<caption id="DF591EC43846FFADFF2CFC3740F7FC5D" ID-DOI="http://doi.org/10.5281/zenodo.7656257" ID-Zenodo-Dep="7656257" httpUri="https://zenodo.org/record/7656257/files/figure.png" pageId="7" pageNumber="240" startId="7.[151,250,961,985]" targetBox="[151,1435,181,937]" targetPageId="7">
<paragraph id="8B994E4C3846FFADFF2CFC3740F7FC5D" blockId="7.[151,1436,961,1021]" pageId="7" pageNumber="240">
<emphasis id="B952925E3846FFADFF2CFC37431CFC79" bold="true" box="[151,269,961,985]" pageId="7" pageNumber="240">FIGURE 4</emphasis>
.
<taxonomicName id="4C2635CF3846FFADFEA3FC3743CEFC79" authorityName="Hoskin, Couper, Schneider &amp; Covacevich" authorityYear="2000" box="[280,479,961,985]" class="Reptilia" family="Carphodactylidae" genus="Phyllurus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="240" phylum="Chordata" rank="species" species="amnicola">
<emphasis id="B952925E3846FFADFEA3FC3743CEFC79" box="[280,479,961,985]" italics="true" pageId="7" pageNumber="240">Phyllurus amnicola</emphasis>
</taxonomicName>
with original (A) and regenerated tail (B), and
<taxonomicName id="4C2635CF3846FFADFC00FC344631FC79" authorityName="Couper, Covacevich &amp; Moritz" authorityYear="1993" box="[955,1056,961,985]" class="Reptilia" family="Carphodactylidae" genus="Phyllurus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="240" phylum="Chordata" rank="species" species="nepthys">
<emphasis id="B952925E3846FFADFC00FC344631FC79" box="[955,1056,961,985]" italics="true" pageId="7" pageNumber="240">P. nepthys</emphasis>
</taxonomicName>
with original (C) and regenerated tail (D). Photos: (A, B) Conrad Hoskin; (C, D) Anders Zimny.
</paragraph>
</caption>
<caption id="DF591EC43846FFADFF2CF89A4609F824" ID-DOI="http://doi.org/10.5281/zenodo.7656259" ID-Zenodo-Dep="7656259" box="[151,1048,1900,1924]" httpUri="https://zenodo.org/record/7656259/files/figure.png" pageId="7" pageNumber="240" startId="7.[151,250,1900,1924]" targetBox="[151,1435,1045,1876]" targetPageId="7">
<paragraph id="8B994E4C3846FFADFF2CF89A4609F824" blockId="7.[151,1048,1900,1924]" box="[151,1048,1900,1924]" pageId="7" pageNumber="240">
<emphasis id="B952925E3846FFADFF2CF89A4302F824" bold="true" box="[151,275,1900,1924]" pageId="7" pageNumber="240">FIGURE 5.</emphasis>
Habitat of
<taxonomicName id="4C2635CF3846FFADFE3DF89B4016F824" authority="Hoskin, 2023" authorityName="Hoskin" authorityYear="2023" box="[390,519,1900,1924]" class="Reptilia" family="Carphodactylidae" genus="Phyllurus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="240" phylum="Chordata" rank="species" species="fimbriatus" status="sp. nov.">
<emphasis id="B952925E3846FFADFE3DF89B4016F824" box="[390,519,1900,1924]" italics="true" pageId="7" pageNumber="240">P. fimbriatus</emphasis>
</taxonomicName>
<emphasis id="B952925E3846FFADFDB6F89A404FF824" bold="true" box="[525,606,1900,1924]" pageId="7" pageNumber="240">
<taxonomicNameLabel id="A2612F253846FFADFDB6F89A404FF824" box="[525,606,1900,1924]" pageId="7" pageNumber="240" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
on Scawfell Island. Photo: Conrad Hoskin.
</paragraph>
</caption>
<paragraph id="8B994E4C3849FFA2FF7CFF614600FEC5" blockId="8.[151,1437,151,465]" pageId="8" pageNumber="241">
Density was high within suitable habitat, with at least fifteen individuals found within approximately three hours search time at each of the two sites. During the survey period (
<date id="FF98688C3849FFA2FC11FF4D46D1FF75" box="[938,1216,186,213]" pageId="8" pageNumber="241" value="2021-11-16" valueMax="2021-11-19" valueMin="2021-11-16">
1619
<superScript id="7C53E3043849FFA2FC4AFF4C41EFFF68" attach="left" box="[1009,1022,186,200]" fontSize="6" pageId="8" pageNumber="241">th</superScript>
November 2021
</date>
), at least half of all individuals found were subadults of small size (approximately
<quantity id="4CDEE3A93849FFA2FCE0FF2841BFFF59" box="[859,942,222,249]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" pageId="8" pageNumber="241" unit="mm" value="65.0">65 mm</quantity>
). Of
<specimenCount id="9D2085C53849FFA2FC5DFF29465CFF59" box="[998,1101,223,249]" pageId="8" pageNumber="241" type="adult">16 adults</specimenCount>
inspected in the hand, 8 were males and 8 were females. Six of the
<specimenCount id="9D2085C53849FFA2FDFFFEF540ADFEBD" box="[580,700,259,285]" pageId="8" pageNumber="241" type="adult">eight adult</specimenCount>
females had two large eggs visible through the skin on the body wall, and two had no eggs visible. For estimates of size at sexual maturity for each sex, and data on original versus regenerated tail status in subadults and adults, see the Field Data section above.
</paragraph>
<paragraph id="8B994E4C3849FFA2FF7CFE9943A7FE71" blockId="8.[151,1437,151,465]" pageId="8" pageNumber="241">
Other gecko species observed at the two sites were
<taxonomicName id="4C2635CF3849FFA2FCAFFE99469FFE29" authority="(Gray, 1845)" baseAuthorityName="Gray" baseAuthorityYear="1845" box="[788,1166,366,393]" class="Reptilia" family="Diplodactylidae" genus="Amalosia" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="241" phylum="Chordata" rank="species" species="rhombifer">
<emphasis id="B952925E3849FFA2FCAFFE9941E4FE29" box="[788,1013,367,393]" italics="true" pageId="8" pageNumber="241">Amalosia rhombifer</emphasis>
(
<bibRefCitation id="EFB733BD3849FFA2FBBDFE994697FE29" author="Gray, J. E." box="[1030,1158,366,393]" pageId="8" pageNumber="241" refId="ref6628" refString="Gray, J. E. (1845) Catalogue of the Specimens of Lizards in the Collection of the British Museum. British Museum, London, xxviii + 289 pp." type="book" year="1845">Gray, 1845</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C2635CF3849FFA2FB73FE99433DFE0D" authority="(Gray, 1845)" baseAuthorityName="Gray" baseAuthorityYear="1845" class="Reptilia" family="Gekkonidae" genus="Heteronotia" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="241" phylum="Chordata" rank="species" species="binoei">
<emphasis id="B952925E3849FFA2FB73FE99478DFE29" box="[1224,1436,367,393]" italics="true" pageId="8" pageNumber="241">Heteronotia binoei</emphasis>
(
<bibRefCitation id="EFB733BD3849FFA2FF24FE654335FE0D" author="Gray, J. E." box="[159,292,402,429]" pageId="8" pageNumber="241" refId="ref6628" refString="Gray, J. E. (1845) Catalogue of the Specimens of Lizards in the Collection of the British Museum. British Museum, London, xxviii + 289 pp." type="book" year="1845">Gray, 1845</bibRefCitation>
)
</taxonomicName>
. The introduced Asian House Gecko (
<taxonomicName id="4C2635CF3849FFA2FD50FE6541FBFE0D" authorityName="Dumeril &amp; Bibron" authorityYear="1836" box="[747,1002,403,429]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="241" phylum="Chordata" rank="species" species="frenatus">
<emphasis id="B952925E3849FFA2FD50FE6541FBFE0D" box="[747,1002,403,429]" italics="true" pageId="8" pageNumber="241">Hemidactylus frenatus</emphasis>
</taxonomicName>
) was seen on granite rocks and trees within
<quantity id="4CDEE3A93849FFA2FF5DFE41430EFE71" box="[230,287,439,465]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="241" unit="km" value="1.0">1 km</quantity>
of both sites.
</paragraph>
</subSubSection>
</treatment>
</document>