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<document id="2D0658864EA82005D961D4C905E7BAF5" ID-DOI="10.5281/zenodo.175790" ID-GBIF-Dataset="2a6512b7-5bd9-4682-b5db-79c72373b68c" ID-ISSN="1175-5326" ID-Zenodo-Dep="175790" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459820893039" checkinUser="plazi" docAuthor="Freitag, Hendrik &amp; Jäch, Manfred A." docDate="2007" docId="03B9F24A7C0EFF80FF305612618F22AF" docLanguage="en" docName="zt01431p044.pdf" docOrigin="Zootaxa 1431" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Hydraena (Hydraenopsis) nielshaggei Freitag &amp; Jach, sp. n." docType="treatment" docVersion="9" lastPageNumber="17" masterDocId="FF808A327C00FF90FFA75435641C2645" masterDocTitle="Revision of the species of Hydraena Kugelann (Coleoptera: Hydraenidae) from Palawan and Busuanga, with descriptions of eleven new species, and redescription of Hydraena (Hydraenopsis) scabra d’Orchymont, 1925" masterLastPageNumber="44" masterPageNumber="1" pageNumber="15" updateTime="1698221773297" updateUser="plazi">
<mods:mods id="0CD1FCD183F06D42A79C040A99E559AB" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="7F9E91AE557E9EE5884F93DA6622E937">
<mods:title id="49D7365520FEB757A772ACD3812CD13D">Revision of the species of Hydraena Kugelann (Coleoptera: Hydraenidae) from Palawan and Busuanga, with descriptions of eleven new species, and redescription of Hydraena (Hydraenopsis) scabra d’Orchymont, 1925</mods:title>
</mods:titleInfo>
<mods:name id="69B96ECDA9CC69ACAD84776A922ACBA2" type="personal">
<mods:role id="DC563FEC43064DF2A5A8E9E1FEF1773B">
<mods:roleTerm id="19B758F7A1DFCD9264198691B4802DB0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="27C6DB6C125AC7946C1DD2517CB3F894">Freitag, Hendrik</mods:namePart>
</mods:name>
<mods:name id="BCD011D123DA46FD41FACED1A33B3FFA" type="personal">
<mods:role id="6BBBEE99662E079B6AA89131231A1300">
<mods:roleTerm id="3D4B7C647B08EFDD1834213FB53BB918">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6B17813F930018B0A7112A06DFE79110">Jäch, Manfred A.</mods:namePart>
</mods:name>
<mods:typeOfResource id="29F976AB689E76649DD5A8AE961C211A">text</mods:typeOfResource>
<mods:relatedItem id="931DA07EEEFCB816D615B2D70D20A159" type="host">
<mods:titleInfo id="C40E7CEF34FF7D2DD25EA1DE2DCF38AD">
<mods:title id="3F4B0066714F92DB0A886CC66E6799CB">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="704E0FDBC66685BC24D0BEB1F0F3ACDE">
<mods:date id="F71574D6117675F1A787641F7986782A">2007</mods:date>
<mods:detail id="088B16640207C669AC40FD602EDE8888" type="volume">
<mods:number id="C26B9A97CB74F22682BB80F31F37B0A6">1431</mods:number>
</mods:detail>
<mods:extent id="CA75B45FFCAF8CAFFF0CFCBA3256145F" unit="page">
<mods:start id="F21349C66E08350763B4313989161BA0">1</mods:start>
<mods:end id="E0C34EF4BFC60CD975A0165F4A0316A4">44</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="F8E0AA7A97D117A24D58156B81CB1389">journal article</mods:classification>
<mods:identifier id="C3AFB7C0A7D2C01CE5C8C8080D92A379" type="DOI">10.5281/zenodo.175790</mods:identifier>
<mods:identifier id="32B304AE8868A42B3E6458AE7C6C13F0" type="GBIF-Dataset">2a6512b7-5bd9-4682-b5db-79c72373b68c</mods:identifier>
<mods:identifier id="EDC3B7C8A16EB20B23A234CE7490EEDE" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="B654A477AA6C938856019D7A700B6AA9" type="Zenodo-Dep">175790</mods:identifier>
</mods:mods>
<treatment id="03B9F24A7C0EFF80FF305612618F22AF" ID-DOI="http://doi.org/10.5281/zenodo.6248071" ID-GBIF-Taxon="119355661" ID-Zenodo-Dep="6248071" LSID="urn:lsid:plazi:treatment:03B9F24A7C0EFF80FF305612618F22AF" httpUri="http://treatment.plazi.org/id/03B9F24A7C0EFF80FF305612618F22AF" lastPageId="16" lastPageNumber="17" pageId="14" pageNumber="15">
<subSubSection id="C30A10D77C0EFF9EFF305612654C2421" pageId="14" pageNumber="15" type="nomenclature">
<paragraph id="8BAF435C7C0EFF9EFF305612676A2404" blockId="14.[151,886,551,612]" box="[151,886,551,577]" pageId="14" pageNumber="15">
<heading id="D0E7F4307C0EFF9EFF305612676A2404" bold="true" box="[151,886,551,577]" fontSize="11" level="1" pageId="14" pageNumber="15" reason="1">
<emphasis id="B9649F4E7C0EFF9EFF305612676A2404" bold="true" box="[151,886,551,577]" pageId="14" pageNumber="15">
<taxonomicName id="4C1038DF7C0EFF9EFF305612673A2404" ID-CoL="6MJCK" authority="Freitag &amp; Jach" authorityName="Freitag &amp; Jach" box="[151,806,551,577]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="nielshaggei" status="sp. nov." subGenus="Hydraenopsis">
<emphasis id="B9649F4E7C0EFF9EFF305612650C2404" bold="true" box="[151,272,551,577]" italics="true" pageId="14" pageNumber="15">Hydraena</emphasis>
(
<emphasis id="B9649F4E7C0EFF9EFE87561265DA2404" bold="true" box="[288,454,551,577]" italics="true" pageId="14" pageNumber="15">Hydraenopsis</emphasis>
)
<emphasis id="B9649F4E7C0EFF9EFE705612667E2404" bold="true" box="[471,610,551,577]" italics="true" pageId="14" pageNumber="15">nielshaggei</emphasis>
Freitag &amp; Jäch
</taxonomicName>
,
<taxonomicNameLabel id="A25722357C0EFF9EFC935612676A2404" box="[820,886,551,577]" pageId="14" pageNumber="15" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF435C7C0EFF9EFF30567F654C2421" blockId="14.[151,886,551,612]" box="[151,336,586,612]" pageId="14" pageNumber="15">(Figs. 5, 17a–h)</paragraph>
</subSubSection>
<subSubSection id="C30A10D77C0EFF9EFF3056AF654C23C1" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="8BAF435C7C0EFF9EFF3056AF61732499" blockId="14.[151,1437,666,2012]" pageId="14" pageNumber="15">
<emphasis id="B9649F4E7C0EFF9EFF3056AF652024F1" bold="true" box="[151,316,666,692]" pageId="14" pageNumber="15">
<typeStatus id="54ABFDFE7C0EFF9EFF3056AF64C824F1" box="[151,212,666,692]" pageId="14" pageNumber="15">Type</typeStatus>
locality.
</emphasis>
Cabayugan River, near Nagsatayan Creek, south of Martarpi;
<geoCoordinate id="EE24259B7C0EFF9EFBB156AF608024F1" box="[1046,1180,666,692]" direction="north" orientation="latitude" pageId="14" pageNumber="15" precision="9" value="10.163">10°09.78'N</geoCoordinate>
<geoCoordinate id="EE24259B7C0EFF9EFB0456AF612A24F1" box="[1187,1334,666,692]" direction="east" orientation="longitude" pageId="14" pageNumber="15" precision="9" value="118.8435">118°50.61'E</geoCoordinate>
; St. Paul Subterranean River National Park, Cabayugan district, Puerto Princesa City, central Palawan,
<collectingCountry id="F30703CC7C0EFF9EFB4356F761702499" box="[1252,1388,706,732]" name="Philippines" pageId="14" pageNumber="15">Philippines</collectingCountry>
.
</paragraph>
<paragraph id="8BAF435C7C0EFF9EFF6056DF654C23C1" blockId="14.[151,1437,666,2012]" pageId="14" pageNumber="15">
<emphasis id="B9649F4E7C0EFF9EFF6056DF660E2541" bold="true" box="[199,530,746,772]" pageId="14" pageNumber="15">
<typeStatus id="54ABFDFE7C0EFF9EFF6056DF65182541" box="[199,260,746,772]" pageId="14" pageNumber="15">Type</typeStatus>
material.
<typeStatus id="54ABFDFE7C0EFF9EFE0756DF660E2541" box="[416,530,746,772]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
</emphasis>
ɗ (
<collectionCode id="ED01DB997C0EFF9EFDFE56DF66B72541" box="[601,683,746,772]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="14" pageNumber="15">NMW</collectionCode>
): “
<collectionCode id="ED01DB997C0EFF9EFD7F56DF673F2541" box="[728,803,746,772]" pageId="14" pageNumber="15">PHIL</collectionCode>
.: Palawan, P. Princesa S Maratarpi, Cabayugan R.
<geoCoordinate id="EE24259B7C0EFF9EFF085727655A2569" box="[175,326,786,812]" direction="north" orientation="latitude" pageId="14" pageNumber="15" precision="1" value="10.163">10°09.780'N</geoCoordinate>
<geoCoordinate id="EE24259B7C0EFF9EFEF7572765F22569" box="[336,494,786,812]" direction="east" orientation="longitude" pageId="14" pageNumber="15" precision="1" value="118.26017">
118°15.610'
<geoCoordinate id="EE24259B7C0EFF9EFE7C572765F22569" box="[475,494,786,812]" direction="east" orientation="longitude" pageId="14" pageNumber="15" precision="5555" value="16.5">E</geoCoordinate>
</geoCoordinate>
<date id="FFAE659C7C0EFF9EFE5F5727666C2569" box="[504,624,786,812]" pageId="14" pageNumber="15" value="2001-05-16">16.5.2001</date>
, leg. Freitag (a21)”, abdomen and aedeagus glued separately, some parts of legs lacking.
<emphasis id="B9649F4E7C0EFF9EFE31570F66012511" bold="true" box="[406,541,826,852]" pageId="14" pageNumber="15">
<typeStatus id="54ABFDFE7C0EFF9EFE31570F66042511" box="[406,536,826,852]" pageId="14" pageNumber="15" type="paratype">Paratypes</typeStatus>
:
</emphasis>
1 Ψ (
<collectionCode id="ED01DB997C0EFF9EFDC7570F66B12511" box="[608,685,826,852]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="14" pageNumber="15">NMW</collectionCode>
, terminal part of abdomen lacking): same locality data as
<typeStatus id="54ABFDFE7C0EFF9EFAF9570F64D12539" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
; 1 ɗ (
<collectionCode id="ED01DB997C0EFF9EFE825757656B2539" box="[293,375,866,892]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="14" pageNumber="15">NMW</collectionCode>
): “
<collectionCode id="ED01DB997C0EFF9EFE3B575765FB2539" box="[412,487,866,892]" pageId="14" pageNumber="15">PHIL</collectionCode>
.: Palawan, P. Princesa S Maratarpi, Cabayugan R.
<geoCoordinate id="EE24259B7C0EFF9EFBFE575760EC2539" box="[1113,1264,866,892]" direction="north" orientation="latitude" pageId="14" pageNumber="15" precision="1" value="10.163">10°09.780'N</geoCoordinate>
<geoCoordinate id="EE24259B7C0EFF9EFB5A575761802539" box="[1277,1436,866,892]" direction="east" orientation="longitude" pageId="14" pageNumber="15" precision="1" value="118.26017">
118°15.610'
<geoCoordinate id="EE24259B7C0EFF9EFA2E575761802539" box="[1417,1436,866,892]" direction="east" orientation="longitude" pageId="14" pageNumber="15" precision="5555" value="16.5">E</geoCoordinate>
</geoCoordinate>
<date id="FFAE659C7C0EFF9EFF3057BF651325E1" box="[151,271,906,932]" pageId="14" pageNumber="15" value="2001-05-16">16.5.2001</date>
, leg. Freitag (a2)”; 1 ɗ (WPU, aedeagus lacking): “
<collectionCode id="ED01DB997C0EFF9EFCCD57BF67A925E1" box="[874,949,906,932]" pageId="14" pageNumber="15">PHIL</collectionCode>
.: Palawan, P. Princesa S Maratarpi, Cabayugan R.
<geoCoordinate id="EE24259B7C0EFF9EFEA6578765842589" box="[257,408,946,972]" direction="north" orientation="latitude" pageId="14" pageNumber="15" precision="1" value="10.163">10°09.780'N</geoCoordinate>
<geoCoordinate id="EE24259B7C0EFF9EFE055787665C2589" box="[418,576,946,972]" direction="east" orientation="longitude" pageId="14" pageNumber="15" precision="1" value="118.26017">
118°15.610'
<geoCoordinate id="EE24259B7C0EFF9EFD8A5787665C2589" box="[557,576,946,972]" direction="east" orientation="longitude" pageId="14" pageNumber="15" precision="5555" value="16.5">E</geoCoordinate>
</geoCoordinate>
<date id="FFAE659C7C0EFF9EFDED578766DE2589" box="[586,706,946,972]" pageId="14" pageNumber="15" value="2001-05-16">16.5.2001</date>
, leg. Freitag (a19)”; 2 ɗɗ (
<collectionCode id="ED01DB997C0EFF9EFBB5578760432589" box="[1042,1119,946,972]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="14" pageNumber="15">NMW</collectionCode>
, 1 ex. mutilated): “
<collectionCode id="ED01DB997C0EFF9EFAEE578761882589" box="[1353,1428,946,972]" pageId="14" pageNumber="15">PHIL</collectionCode>
.: Palawan, P. Princesa S Maratarpi, Cabayugan R.
<geoCoordinate id="EE24259B7C0EFF9EFD6F57EF677C25B1" box="[712,864,986,1012]" direction="north" orientation="latitude" pageId="14" pageNumber="15" precision="1" value="10.163">10°09.780'N</geoCoordinate>
<geoCoordinate id="EE24259B7C0EFF9EFCC157EF601825B1" box="[870,1028,986,1012]" direction="east" orientation="longitude" pageId="14" pageNumber="15" precision="1" value="118.26017">
118°15.610'
<geoCoordinate id="EE24259B7C0EFF9EFC5657EF601825B1" box="[1009,1028,986,1012]" direction="east" orientation="longitude" pageId="14" pageNumber="15" precision="5555" value="16.5">E</geoCoordinate>
</geoCoordinate>
<date id="FFAE659C7C0EFF9EFBAC57EF609F25B1" box="[1035,1155,986,1012]" pageId="14" pageNumber="15" value="2001-05-16">16.5.2001</date>
, leg. Freitag (a20)”; 1 ɗ (
<collectionCode id="ED01DB997C0EFF9EFF38503764EE2259" box="[159,242,1026,1052]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="14" pageNumber="15">NMW</collectionCode>
): “
<collectionCode id="ED01DB997C0EFF9EFEB6503765402259" box="[273,348,1026,1052]" pageId="14" pageNumber="15">PHIL</collectionCode>
.: Palawan, P. Princesa S Maratarpi, Cabayugan R.
<geoCoordinate id="EE24259B7C0EFF9EFC00503760222259" box="[935,1086,1026,1052]" direction="north" orientation="latitude" pageId="14" pageNumber="15" precision="1" value="10.163">10°09.780'N</geoCoordinate>
<geoCoordinate id="EE24259B7C0EFF9EFBE1503760F82259" box="[1094,1252,1026,1052]" direction="east" orientation="longitude" pageId="14" pageNumber="15" precision="1" value="118.26017">
118°15.610'
<geoCoordinate id="EE24259B7C0EFF9EFB76503760F82259" box="[1233,1252,1026,1052]" direction="east" orientation="longitude" pageId="14" pageNumber="15" precision="5555" value="16.5">E</geoCoordinate>
</geoCoordinate>
<date id="FFAE659C7C0EFF9EFB4B503761782259" box="[1260,1380,1026,1052]" pageId="14" pageNumber="15" value="2001-05-16">16.5.2001</date>
, leg. Freitag (b8)”; 2 ΨΨ (
<collectionCode id="ED01DB997C0EFF9EFE2D501F65C62201" box="[394,474,1066,1092]" httpUri="http://grbio.org/cool/134h-pm2f" name="Museum of Natural History, University of the Philippines" pageId="14" pageNumber="15">UPLB</collectionCode>
; mutilated): “
<collectionCode id="ED01DB997C0EFF9EFDDF501F66DF2201" box="[632,707,1066,1092]" pageId="14" pageNumber="15">PHIL</collectionCode>
.: Palawan, P. Princesa S Maratarpi, Cabayugan R.
<geoCoordinate id="EE24259B7C0EFF9EFAA1501F61802201" box="[1286,1436,1066,1092]" direction="north" orientation="latitude" pageId="14" pageNumber="15" precision="1" value="10.163">10°09.780'N</geoCoordinate>
<geoCoordinate id="EE24259B7C0EFF9EFF30506765292229" box="[151,309,1106,1132]" direction="east" orientation="longitude" pageId="14" pageNumber="15" precision="1" value="118.26017">
118°15.610'
<geoCoordinate id="EE24259B7C0EFF9EFE85506765292229" box="[290,309,1106,1132]" direction="east" orientation="longitude" pageId="14" pageNumber="15" precision="5555" value="16.5">E</geoCoordinate>
</geoCoordinate>
<date id="FFAE659C7C0EFF9EFE98506765AB2229" box="[319,439,1106,1132]" pageId="14" pageNumber="15" value="2001-05-16">16.5.2001</date>
, leg. Freitag (a30)”; 1 Ψ (WPU, mutilated): “
<collectionCode id="ED01DB997C0EFF9EFC7A506760342229" box="[989,1064,1106,1132]" pageId="14" pageNumber="15">PHIL</collectionCode>
.: Palawan, P. Princesa S Maratarpi, Cabayugan R.
<geoCoordinate id="EE24259B7C0EFF9EFE26504F660422D1" box="[385,536,1146,1172]" direction="north" orientation="latitude" pageId="14" pageNumber="15" precision="1" value="10.163">10°09.780'N</geoCoordinate>
<geoCoordinate id="EE24259B7C0EFF9EFD87504F66A222D1" box="[544,702,1146,1172]" direction="east" orientation="longitude" pageId="14" pageNumber="15" precision="1" value="118.26017">
118°15.610'
<geoCoordinate id="EE24259B7C0EFF9EFD0C504F66A222D1" box="[683,702,1146,1172]" direction="east" orientation="longitude" pageId="14" pageNumber="15" precision="5555" value="16.5">E</geoCoordinate>
</geoCoordinate>
<date id="FFAE659C7C0EFF9EFD60504F672322D1" box="[711,831,1146,1172]" pageId="14" pageNumber="15" value="2001-05-16">16.5.2001</date>
, leg. Freitag (d1)”; 1 ɗ (
<collectionCode id="ED01DB997C0EFF9EFBC2504F60AB22D1" box="[1125,1207,1146,1172]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="14" pageNumber="15">NMW</collectionCode>
): “
<collectionCode id="ED01DB997C0EFF9EFB71504F613D22D1" box="[1238,1313,1146,1172]" pageId="14" pageNumber="15">PHIL</collectionCode>
.: Palawan, P. Princesa SSW Martarpi Cabayugan R1
<geoCoordinate id="EE24259B7C0EFF9EFD0A5097672922F9" box="[685,821,1186,1212]" direction="north" orientation="latitude" pageId="14" pageNumber="15" precision="15" value="10.162777">10°09'46''N</geoCoordinate>
<geoCoordinate id="EE24259B7C0EFF9EFCE3509767CA22F9" box="[836,982,1186,1212]" direction="east" orientation="longitude" pageId="14" pageNumber="15" precision="15" value="118.82472">
118°49'29''
<geoCoordinate id="EE24259B7C0EFF9EFC64509767CA22F9" box="[963,982,1186,1212]" direction="east" orientation="longitude" pageId="14" pageNumber="15" precision="5555" value="16.5">E</geoCoordinate>
</geoCoordinate>
<date id="FFAE659C7C0EFF9EFC425097604122F9" box="[997,1117,1186,1212]" pageId="14" pageNumber="15" value="2000-05-16">16.5.2000</date>
, leg. Freitag (a24)”; 1 ɗ (
<collectionCode id="ED01DB997C0EFF9EFF3850FF64EE22A1" box="[159,242,1226,1252]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="14" pageNumber="15">NMW</collectionCode>
): “
<collectionCode id="ED01DB997C0EFF9EFEB450FF654222A1" box="[275,350,1226,1252]" pageId="14" pageNumber="15">PHIL</collectionCode>
.: Palawan, P. Princesa SSW Martarpi Cabayugan R.1
<geoCoordinate id="EE24259B7C0EFF9EFC4850FF606B22A1" box="[1007,1143,1226,1252]" direction="north" orientation="latitude" pageId="14" pageNumber="15" precision="15" value="10.162777">10°09'46''N</geoCoordinate>
<geoCoordinate id="EE24259B7C0EFF9EFB2550FF610822A1" box="[1154,1300,1226,1252]" direction="east" orientation="longitude" pageId="14" pageNumber="15" precision="15" value="118.82472">
118°49'29''
<geoCoordinate id="EE24259B7C0EFF9EFAA650FF610822A1" box="[1281,1300,1226,1252]" direction="east" orientation="longitude" pageId="14" pageNumber="15" precision="5555" value="16.5">E</geoCoordinate>
</geoCoordinate>
<date id="FFAE659C7C0EFF9EFAB850FF618B22A1" box="[1311,1431,1226,1252]" pageId="14" pageNumber="15" value="2001-05-16">16.5.2001</date>
, leg. Freitag (8)”; 1 ɗ (
<collectionCode id="ED01DB997C0EFF9EFE1550C766182349" box="[434,516,1266,1292]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="14" pageNumber="15">NMW</collectionCode>
): “
<collectionCode id="ED01DB997C0EFF9EFD8050C7666E2349" box="[551,626,1266,1292]" pageId="14" pageNumber="15">PHIL</collectionCode>
.: Palawan, P. Princesa SSW Martarpi Cabayugan R.1,
<geoCoordinate id="EE24259B7C0EFF9EFAB450C761802349" box="[1299,1436,1266,1292]" direction="north" orientation="latitude" pageId="14" pageNumber="15" precision="15" value="10.162777">10°09'46''N</geoCoordinate>
<geoCoordinate id="EE24259B7C0EFF9EFF30512F65352371" box="[151,297,1306,1332]" direction="east" orientation="longitude" pageId="14" pageNumber="15" precision="15" value="118.82472">118°49'29''E</geoCoordinate>
<date id="FFAE659C7C0EFF9EFE96512F65DE2371" box="[305,450,1306,1332]" pageId="14" pageNumber="15" value="2000-12-11">11.XII.2000</date>
, leg. Freitag (
<date id="FFAE659C7C0EFF9EFDC4512F66912371" box="[611,653,1306,1332]" pageId="14" pageNumber="15" value="1944-01">i44</date>
)E”; 1 ex. (WPU, terminal part of abdomen lacking): “
<collectionCode id="ED01DB997C0EFF9EFAA1512F614D2371" box="[1286,1361,1306,1332]" pageId="14" pageNumber="15">PHIL</collectionCode>
.: Palawan, P.Princesa; mountain stream
<quantity id="4CE8EEB97C0EFF9EFD8E517766422319" box="[553,606,1346,1372]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="15" unit="km" value="2.0">2km</quantity>
SE Laptay/Napsan, semi-prim.forest,
<geoCoordinate id="EE24259B7C0EFF9EFB875177606F2319" box="[1056,1139,1346,1372]" direction="north" orientation="latitude" pageId="14" pageNumber="15" precision="925" value="9.716666">9°43'N</geoCoordinate>
<geoCoordinate id="EE24259B7C0EFF9EFBDD517760F82319" box="[1146,1252,1346,1372]" direction="east" orientation="longitude" pageId="14" pageNumber="15" precision="925" value="118.46667">
118°28'
<geoCoordinate id="EE24259B7C0EFF9EFB76517760F82319" box="[1233,1252,1346,1372]" direction="east" orientation="longitude" pageId="14" pageNumber="15" precision="5555" value="29.3">E</geoCoordinate>
</geoCoordinate>
<date id="FFAE659C7C0EFF9EFB4A517761792319" box="[1261,1381,1346,1372]" pageId="14" pageNumber="15" value="1995-03-29">29.3.1995</date>
, leg. Freitag (23)M”.
</paragraph>
</subSubSection>
<subSubSection id="C30A10D77C0EFF80FF6051A7600025BF" lastPageId="16" lastPageNumber="17" pageId="14" pageNumber="15" type="description">
<paragraph id="8BAF435C7C0EFF9EFF6051A7674A23B9" blockId="14.[151,1437,666,2012]" pageId="14" pageNumber="15">
<emphasis id="B9649F4E7C0EFF9EFF6051A7654323E9" bold="true" box="[199,351,1426,1452]" pageId="14" pageNumber="15">Description.</emphasis>
Body
<quantity id="4CE8EEB97C0EFF9EFE0951A7664523E9" box="[430,601,1426,1452]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" metricValueMax="1.15" metricValueMin="1.05" pageId="14" pageNumber="15" unit="mm" value="1.1" valueMax="1.15" valueMin="1.05">1.05–1.15 mm</quantity>
long. Habitus as in Fig. 5. Elytra and rectangular spot – if present – in the middle of pronotal disc dark brown, otherwise pronotal disc pale grey-brown; frons darker, in most specimens reddish-brown; pronotal margins and legs much paler.
</paragraph>
<paragraph id="8BAF435C7C0EFF9EFF60523F605D20A9" blockId="14.[151,1437,666,2012]" pageId="14" pageNumber="15">Pronotum wider than long, broadest at posterior 0.3–0.5; entire pronotum densely punctate; punctures moderately large and moderately deeply impressed, interstices glabrous; foveae shallowly impressed and hardly perceptible; anterior margin slightly concave; anterior angles rounded; lateral rim denticulate; lateral margins anteriorly convergent, posteriorly sinuously convergent (inflexed) forming a distinct pronoto-elytral angle; posterior margin straight to slightly bisinuate; lateral portions moderately deflexed; lateral hypomeron approximately as broad as profemora, mesal hypomeron narrow, inconspicuous.</paragraph>
<paragraph id="8BAF435C7C0EFF9EFF6052CF652C2199" blockId="14.[151,1437,666,2012]" pageId="14" pageNumber="15">Elytra elongately oval, conjointly rounded apically; with ca. 15 longitudinal, quite irregular, not impressed rows of punctures (10 between suture and shoulder); punctures small and shallowly impressed; interstices and intervals almost flat, glabrous; inflexed lateral portion of elytra moderately explanate, not reaching elytral apex, distinctly narrower than metafemora; rim inconspicuously denticulate posteriorly; epipleuron inconspicuous; pseudepipleuron distinct, reaching ca. posterior 0.25, glabrous, with an indistinct row of punctures.</paragraph>
<caption id="DF6F13D47C0FFF9FFF3052B367772085" httpUri="https://zenodo.org/record/175795/files/figure.png" pageId="15" pageNumber="16" targetBox="[271,1314,196,1635]" targetPageId="15">
<paragraph id="8BAF435C7C0FFF9FFF3052B367772085" blockId="15.[151,1436,1670,1728]" pageId="15" pageNumber="16">
<emphasis id="B9649F4E7C0FFF9FFF3052B3653A20DB" bold="true" box="[151,294,1670,1694]" pageId="15" pageNumber="16">FIGURE 17.</emphasis>
<taxonomicName id="4C1038DF7C0FFF9FFE8B52B2660B20DB" box="[300,535,1671,1694]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="nielshaggei" status="sp. nov.">
<emphasis id="B9649F4E7C0FFF9FFE8B52B2660B20DB" box="[300,535,1671,1694]" italics="true" pageId="15" pageNumber="16">Hydraena nielshaggei</emphasis>
</taxonomicName>
<emphasis id="B9649F4E7C0FFF9FFDBA52B3664420DB" bold="true" box="[541,600,1670,1694]" pageId="15" pageNumber="16">
<taxonomicNameLabel id="A25722357C0FFF9FFDBA52B3664420DB" box="[541,600,1670,1694]" pageId="15" pageNumber="16" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
a–c) aedeagus (holotype) in ventral, dorsal and lateral view; d) male sternite X and spiculum; e) gonocoxite; f–g) spermatheca; h) female tergite X.
</paragraph>
</caption>
<paragraph id="8BAF435C7C0FFF9FFF6052C1667521EB" blockId="15.[151,1437,1780,2006]" pageId="15" pageNumber="16">Mesoventrite with well developed longitudinal ridges; mesoventral intercoxal process short, almost as broad as pseudepipleuron; metaventral disc deeply impressed; metaventral plaques distinct, widely separated (distance wider than inflexed lateral portion of elytra), very narrow, glabrous; intercoxal sternite very large, approximately as broad as metafemur and as long as broad, sub-triangular; metaventrite and abdominal sternites moderately densely pubescent.</paragraph>
<paragraph id="8BAF435C7C0FFF9FFF60538960B02193" blockId="15.[151,1437,1780,2006]" box="[199,1196,1980,2006]" pageId="15" pageNumber="16">Femora with inner margins straight to concave, outer margins conspicuously convex.</paragraph>
<paragraph id="8BAF435C7C10FF80FF6054AD66C527E7" blockId="16.[151,1437,152,1258]" pageId="16" pageNumber="17">
Aedeagus (
<figureCitation id="132B5FD97C10FF80FEEC54AD65B626F7" box="[331,426,152,178]" captionStart="FIGURE 17" captionStartId="15.[151,255,1670,1694]" captionTargetBox="[271,1314,196,1635]" captionTargetId="figure@15.[264,1323,194,1646]" captionTargetPageId="15" captionText="FIGURE 17. Hydraena nielshaggei sp. n. a – c) aedeagus (holotype) in ventral, dorsal and lateral view; d) male sternite X and spiculum; e) gonocoxite; f – g) spermatheca; h) female tergite X." httpUri="https://zenodo.org/record/175795/files/figure.png" pageId="16" pageNumber="17">Figs. 17</figureCitation>
a–c) rather simple, elongate and slender. Main piece rather straight, slightly curved to right side (ventral/dorsal view), without setae, with five micropores; phallobase asymmetric. Distal lobe small, with few short amorphic appendages, not clearly delimited from main piece. Parameres very short, almost completely reduced, not articulately connected with main piece; left paramere inserted at about basal 0.55 of aedeagus, with ca. eight moderately long setae (two dorsal apical, one ventral apical, five ventral lateral, arranged in a longitudinal row); right paramere inserted at about basal 0.6 of aedeagus, with ca. eight moderately long setae (five dorsal, three ventral).
</paragraph>
<paragraph id="8BAF435C7C10FF80FF605585673E2407" blockId="16.[151,1437,152,1258]" pageId="16" pageNumber="17">
Gonocoxite (
<figureCitation id="132B5FD97C10FF80FEC3558565A1278F" box="[356,445,432,458]" captionStart="FIGURE 17" captionStartId="15.[151,255,1670,1694]" captionTargetBox="[271,1314,196,1635]" captionTargetId="figure@15.[264,1323,194,1646]" captionTargetPageId="15" captionText="FIGURE 17. Hydraena nielshaggei sp. n. a – c) aedeagus (holotype) in ventral, dorsal and lateral view; d) male sternite X and spiculum; e) gonocoxite; f – g) spermatheca; h) female tergite X." httpUri="https://zenodo.org/record/175795/files/figure.png" pageId="16" pageNumber="17">Fig. 17</figureCitation>
e) subtriangular, rather short; lateral margins strongly convex subbasally, slightly emarginate medially; apical area large; margin between apical and basal area well developed, sinuous; subapical tuft setae curved; basal area without distinct pubescence; condyles prominent; dorsal plate simple, slightly surpassing ventral plate, without distinct cavea.
</paragraph>
<paragraph id="8BAF435C7C10FF80FF6056656003242F" blockId="16.[151,1437,152,1258]" box="[199,1055,592,618]" pageId="16" pageNumber="17">
Spermatheca (
<figureCitation id="132B5FD97C10FF80FEC9566565D7242F" box="[366,459,592,618]" captionStart="FIGURE 17" captionStartId="15.[151,255,1670,1694]" captionTargetBox="[271,1314,196,1635]" captionTargetId="figure@15.[264,1323,194,1646]" captionTargetPageId="15" captionText="FIGURE 17. Hydraena nielshaggei sp. n. a – c) aedeagus (holotype) in ventral, dorsal and lateral view; d) male sternite X and spiculum; e) gonocoxite; f – g) spermatheca; h) female tergite X." httpUri="https://zenodo.org/record/175795/files/figure.png" pageId="16" pageNumber="17">Figs. 17</figureCitation>
f–g) very similar to that of
<taxonomicName id="4C1038DF7C10FF80FCB556656004242C" box="[786,1048,592,617]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="jojoorculloi">
<emphasis id="B9649F4E7C10FF80FCB556656004242C" box="[786,1048,592,617]" italics="true" pageId="16" pageNumber="17">Hydraena jojoorculloi</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BAF435C7C10FF80FF60564D67C225C7" blockId="16.[151,1437,152,1258]" pageId="16" pageNumber="17">
Secondary sexual characters. Male profemur with tiny denticle near base of ventral face. Metaventral plaques narrower in male; meso- and metatibia in male slightly curved. Male terminal sternite and spiculum gastrale (
<figureCitation id="132B5FD97C10FF80FEA456FD654524A7" box="[259,345,712,738]" captionStart="FIGURE 17" captionStartId="15.[151,255,1670,1694]" captionTargetBox="[271,1314,196,1635]" captionTargetId="figure@15.[264,1323,194,1646]" captionTargetPageId="15" captionText="FIGURE 17. Hydraena nielshaggei sp. n. a – c) aedeagus (holotype) in ventral, dorsal and lateral view; d) male sternite X and spiculum; e) gonocoxite; f – g) spermatheca; h) female tergite X." httpUri="https://zenodo.org/record/175795/files/figure.png" pageId="16" pageNumber="17">Fig. 17</figureCitation>
d) very similar to
<taxonomicName id="4C1038DF7C10FF80FD9756FD672B24A4" box="[560,823,712,737]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="jojoorculloi">
<emphasis id="B9649F4E7C10FF80FD9756FD672B24A4" box="[560,823,712,737]" italics="true" pageId="16" pageNumber="17">Hydraena jojoorculloi</emphasis>
</taxonomicName>
, but micropores at mediolateral portion of terminal sternite less conspicuous, and spiculum gastrale comparatively longer. Male tergite X distinctly excised apically. Female tergite X (
<figureCitation id="132B5FD97C10FF80FE46572D66242577" box="[481,568,792,818]" captionStart="FIGURE 17" captionStartId="15.[151,255,1670,1694]" captionTargetBox="[271,1314,196,1635]" captionTargetId="figure@15.[264,1323,194,1646]" captionTargetPageId="15" captionText="FIGURE 17. Hydraena nielshaggei sp. n. a – c) aedeagus (holotype) in ventral, dorsal and lateral view; d) male sternite X and spiculum; e) gonocoxite; f – g) spermatheca; h) female tergite X." httpUri="https://zenodo.org/record/175795/files/figure.png" pageId="16" pageNumber="17">Fig. 17</figureCitation>
h) subtrapezoidal; anterior and posterior corners widely rounded; lateral margins evenly rounded; disc sparsely covered with setae; subapical fringe of trichoid setae widely interrupted medially; hyaline apical margin distinctly notched medially.
</paragraph>
<paragraph id="8BAF435C7C10FF80FF6057BA600025BF" blockId="16.[151,1437,152,1258]" pageId="16" pageNumber="17">
<emphasis id="B9649F4E7C10FF80FF6057BA65C525EC" bold="true" box="[199,473,911,937]" pageId="16" pageNumber="17">Differential diagnosis.</emphasis>
General habitus and colour resemble
<taxonomicName id="4C1038DF7C10FF80FC3257A5608725EC" box="[917,1179,912,937]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="jojoorculloi">
<emphasis id="B9649F4E7C10FF80FC3257A5608725EC" box="[917,1179,912,937]" italics="true" pageId="16" pageNumber="17">Hydraena jojoorculloi</emphasis>
</taxonomicName>
, from which it can be distinguished by its slightly smaller size, the slender bent aedeagus with characteristic short amorphic appendages, and by the subapical fringe of trichoid setae of female tergite X.
</paragraph>
</subSubSection>
<subSubSection id="C30A10D77C10FF80FF60503264D02287" pageId="16" pageNumber="17" type="distribution">
<paragraph id="8BAF435C7C10FF80FF60503266A7220F" blockId="16.[151,1437,152,1258]" pageId="16" pageNumber="17">
<emphasis id="B9649F4E7C10FF80FF605032657C2264" bold="true" box="[199,352,1031,1057]" pageId="16" pageNumber="17">Distribution</emphasis>
(
<figureCitation id="132B5FD97C10FF80FED7503D65D42267" box="[368,456,1032,1058]" captionStart="FIGURE 30" captionStartId="41.[151,255,1142,1166]" captionTargetBox="[267,1314,183,1127]" captionTargetId="figure@41.[264,1323,183,1135]" captionTargetPageId="41" captionText="FIGURE 30. Collecting sites and geographical distribution of Hydraena species in Palawan and Busuanga, including enlarged map of St. Paul National Park: Hydraena scabra d’Orchymont, H. jojoorculloi sp. n., H. nielshaggei sp. n., H. zetteli sp. n., H. castanescens sp. n., H. claudia sp. n., H. kodadai sp. n." httpUri="https://zenodo.org/record/175808/files/figure.png" pageId="16" pageNumber="17">Fig. 30</figureCitation>
). This species is known from the Cabayugan River and the Ulangoan River, central Palawan. It is most likely endemic to Palawan.
</paragraph>
<paragraph id="8BAF435C7C10FF80FF60506264D02287" blockId="16.[151,1437,152,1258]" pageId="16" pageNumber="17">
<emphasis id="B9649F4E7C10FF80FF605062652C2234" bold="true" box="[199,304,1111,1137]" pageId="16" pageNumber="17">Ecology.</emphasis>
The occurrence in headwaters of rather undisturbed, forest-covered areas suggests an affinity to pristine habitats. Most specimens were collected in drift traps. Two specimens were caught in an emergence trap.
</paragraph>
</subSubSection>
<subSubSection id="C30A10D77C10FF80FF6050FA618F22AF" box="[199,1427,1231,1258]" pageId="16" pageNumber="17" type="etymology">
<paragraph id="8BAF435C7C10FF80FF6050FA618F22AF" blockId="16.[151,1437,152,1258]" box="[199,1427,1231,1258]" pageId="16" pageNumber="17">
<emphasis id="B9649F4E7C10FF80FF6050FA654822AC" bold="true" box="[199,340,1231,1257]" pageId="16" pageNumber="17">Etymology.</emphasis>
Named for Niels Hagge (Hamburg), zoologist, for his support during field work in Palawan.
</paragraph>
</subSubSection>
</treatment>
</document>
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