181 lines
19 KiB
XML
181 lines
19 KiB
XML
<document id="257DF4576C8714D9DDD919E33DF1F212" ID-DOI="10.11646/phytotaxa.257.3.2" ID-ISSN="1179-3163" ID-Zenodo-Dep="13670670" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725398280652" checkinUser="felipe" docAuthor="Doucette, Alfonso, Portilla, Jose & Cameron, Kenneth M." docDate="2016" docId="03B987A5BA5D346AA7963CB154DFF84D" docLanguage="en" docName="phytotaxa.257.3.2.pdf" docOrigin="Phytotaxa 257 (3)" docSource="http://dx.doi.org/10.11646/phytotaxa.257.3.2" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Pleurothallis papillingua A. Doucette & J. Portilla 2016, sp. nov." docType="treatment" docVersion="2" lastPageNumber="239" masterDocId="FF80FFDDBA553463A71E3A395174FFE3" masterDocTitle="Ten new taxa in the orchid subtribe Pleurothallidinae (Epidendroideae, Epidendreae) from Ecuador" masterLastPageNumber="248" masterPageNumber="230" pageNumber="238" updateTime="1725420285918" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="B19C6E0D4BCBAB145AAE9EE189EC8719">
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<mods:title id="969A8948F1B4987FD97A305846D1AB6D">Ten new taxa in the orchid subtribe Pleurothallidinae (Epidendroideae, Epidendreae) from Ecuador</mods:title>
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<mods:name id="3237FB98E1C98BDF978B9412947DE695" type="personal">
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<mods:namePart id="BDE1FFF6E2ECC10C3B6DA3E02AA957B1">Doucette, Alfonso</mods:namePart>
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<mods:namePart id="5510CADDFEC62C78C75E27E2283E6BDB">Portilla, Jose</mods:namePart>
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<mods:affiliation id="FF66CB6013E61EC1DA4CC018EDFBE626">Km. 2 1 / 1 Vía a Cuenca Sector Llampasay, Ecuagenera, Gualaceo, Ecuador</mods:affiliation>
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<mods:roleTerm id="3EBECFEC2BDCA8AED45CC97CD484E02E">Author</mods:roleTerm>
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<mods:namePart id="E9835522D9DA380EFCA8DDC2A7A8BCB9">Cameron, Kenneth M.</mods:namePart>
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<mods:typeOfResource id="BA0A561160516FCE425A675FB1B7419C">text</mods:typeOfResource>
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<mods:title id="B8FDE5B46F04891DA63986026A710C21">Phytotaxa</mods:title>
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<mods:part id="5869C868EDF810DD9FE1AF90280C232A">
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<mods:date id="FDA4B659CB239233521D8A821C0B75C9">2016</mods:date>
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<mods:number id="5389661B8DFAC320775B99DC499DBC24">2016-04-20</mods:number>
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<mods:number id="7BCBE0507B598823EB093DFCDAC21322">257</mods:number>
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<mods:start id="2819A96F2B5421B04D3004ADE2A718B1">230</mods:start>
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<mods:url id="D41E7C914CD6531B21D4EF7AA3280EF9">http://dx.doi.org/10.11646/phytotaxa.257.3.2</mods:url>
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<mods:identifier id="CC30A2F2AAE960421C819F9EB7E388D5" type="ISSN">1179-3163</mods:identifier>
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<mods:identifier id="D5F0AFDB24BA3E3334A1FFB30821DC50" type="Zenodo-Dep">13670670</mods:identifier>
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<treatment id="03B987A5BA5D346AA7963CB154DFF84D" LSID="urn:lsid:plazi:treatment:03B987A5BA5D346AA7963CB154DFF84D" httpUri="http://treatment.plazi.org/id/03B987A5BA5D346AA7963CB154DFF84D" lastPageId="9" lastPageNumber="239" pageId="8" pageNumber="238">
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<subSubSection id="C30A6538BA5D346BA7963CB152CEF942" box="[136,954,1672,1697]" pageId="8" pageNumber="238" type="nomenclature">
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<paragraph id="8BAF36B3BA5D346BA7963CB152CEF942" blockId="8.[136,954,1672,1697]" box="[136,954,1672,1697]" pageId="8" pageNumber="238">
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<heading id="D0E781DFBA5D346BA7963CB152CEF942" box="[136,954,1672,1697]" fontSize="11" level="2" pageId="8" pageNumber="238" reason="3">
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<taxonomicName id="4C104D30BA5D346BA7963CB153CFF942" ID-CoL="4KD9V" authority="A.Doucette & J.Portilla" authorityName="A. Doucette & J. Portilla" authorityYear="2016" box="[136,699,1672,1697]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="8" pageNumber="238" phylum="Tracheophyta" rank="species" species="papillingua" status="sp. nov.">
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<emphasis id="B964EAA1BA5D346BA7963CB150DCF942" bold="true" box="[136,424,1672,1697]" italics="true" pageId="8" pageNumber="238">Pleurothallis papillingua</emphasis>
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A.Doucette & J.Portilla
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</taxonomicName>
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,
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<emphasis id="B964EAA1BA5D346BA5D83CB1526FF942" box="[710,795,1672,1697]" italics="true" pageId="8" pageNumber="238">
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<taxonomicNameLabel id="A25757DABA5D346BA5D83CB1526FF942" box="[710,795,1672,1697]" pageId="8" pageNumber="238" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
(
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<figureCitation id="132B2A36BA5D346BA4343CB1520DF942" box="[810,889,1672,1697]" captionStart="FIGURE 7" captionStartId="10.[136,229,1580,1601]" captionTargetBox="[304,1288,200,1551]" captionTargetId="figure-14@10.[285,1302,175,1563]" captionTargetPageId="10" captionText="FIGURE 7. Pleurothallis papillingua. A. Habit, drawn to 5 cm scale. B. Flower. C. Dorsal sepal. D. Lateral sepals. E. Petal. F. Lower surface of lip. G. Upper surface of lip. H. Flower with petals and sepals removed. All drawn to 1 mm scale." figureDoi="http://doi.org/10.5281/zenodo.13670694" httpUri="https://zenodo.org/record/13670694/files/figure.png" pageId="8" pageNumber="238">
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Figs.
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<geoCoordinate id="EE245074BA5D346BA4773CB1520DF942" box="[873,889,1672,1697]" degrees="7, 11" direction="east" orientation="longitude" pageId="8" pageNumber="238" precision="555" value="7.11">7</geoCoordinate>
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</figureCitation>
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,
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<figureCitation id="132B2A36BA5D346BA49B3CB152C6F942" box="[901,946,1672,1697]" captionStart="FIGURE 11" captionStartId="15.[136,229,1891,1912]" captionTargetBox="[208,1380,202,1851]" captionTargetId="figure-14@15.[197,1391,190,1866]" captionTargetPageId="15" captionText="FIGURE 11. A. Masdevallia constricta f. glabra. B. Masdevallia pachyura f. pardinella. C. Masdevallia posadae f. rugosepala. D. Acianthera nikoleana. E. Pleurothallis papillingua. F. Pleurothallis gigiportillae. G. Pleurothallis cardiostola f. magnidraba. H. Zootrophion machaqway. I. Stelis glaucus. J. Trisetella portillae." figureDoi="http://doi.org/10.5281/zenodo.13670708" httpUri="https://zenodo.org/record/13670708/files/figure.png" pageId="8" pageNumber="238">11E</figureCitation>
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)
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30A6538BA5D346AA7963CEB5479F935" lastPageId="9" lastPageNumber="239" pageId="8" pageNumber="238" type="materials_examined">
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<paragraph id="8BAF36B3BA5D346BA7963CEB502CF8E8" blockId="8.[136,1451,1746,1875]" pageId="8" pageNumber="238">
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Type:—
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<collectingCountry id="F3077623BA5D346BA7CB3CEB5039F904" box="[213,333,1746,1767]" name="Ecuador" pageId="8" pageNumber="238">ECUADOR</collectingCountry>
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. Without a specific locality, flowered in cultivation at Ecuagenera Orchid Nursery,
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<date id="FFAE1073BA5D346BA3783CEB5462F904" box="[1126,1302,1746,1767]" pageId="8" pageNumber="238" value="2015-09-04">4 September 2015</date>
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,
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<emphasis id="B964EAA1BA5D346BA2013CEB54DFF904" box="[1311,1451,1746,1767]" italics="true" pageId="8" pageNumber="238">Doucette 0314</emphasis>
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(
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<typeStatus id="54AB8811BA5D346BA7DA3CCF5069F8E8" box="[196,285,1782,1803]" pageId="8" pageNumber="238" type="holotype">holotype</typeStatus>
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:
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<collectionCode id="ED01AE76BA5D346BA6383CCF5039F8E8" box="[294,333,1782,1803]" country="Ecuador" lsid="urn:lsid:biocol.org:col:14290" name="Universidad del Azuay" pageId="8" pageNumber="238" type="Herbarium">HA</collectionCode>
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).
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</paragraph>
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<paragraph id="8BAF36B3BA5D346BA7963D235304F8B0" blockId="8.[136,1451,1746,1875]" pageId="8" pageNumber="238">
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Diagnosis: The species is most similar to
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<taxonomicName id="4C104D30BA5D346BA5273D2352E7F8CC" authority="Luer & Hirtz" authorityName="Luer & Hirtz" box="[569,915,1818,1839]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="8" pageNumber="238" phylum="Tracheophyta" rank="species" species="tiarata">
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<emphasis id="B964EAA1BA5D346BA5273D235275F8CC" box="[569,769,1818,1839]" italics="true" pageId="8" pageNumber="238">Pleurothallis tiarata</emphasis>
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Luer & Hirtz
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</taxonomicName>
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in
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<bibRefCitation id="EF814B42BA5D346BA4A53D23551DF8CC" author="Luer, C." box="[955,1129,1818,1839]" pageId="8" pageNumber="238" pagination="1 - 265" refId="ref8472" refString="Luer, C. (2004) Icones pleurothallidinarum XXVI: Pleurothallis subgenus Acianthera and three allied subgenera; a second century of new species of Stelis of Ecuador; Epibator, Ophidion, Zootrophion. Pleurothallis. Monographs in Systematic Botany 95: 1 - 265." type="journal article" year="2004">Luer (2004: 238)</bibRefCitation>
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differing in the fewer-flowered inflorescence, longer petals and papillose lip.
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</paragraph>
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<caption id="DF6F663BBA5C346AA7963F4255FAFA57" ID-DOI="http://doi.org/10.5281/zenodo.13670692" ID-Zenodo-Dep="13670692" httpUri="https://zenodo.org/record/13670692/files/figure.png" pageId="9" pageNumber="239" startId="9.[136,229,1403,1424]" targetBox="[376,1214,212,1367]" targetPageId="9" targetType="figure">
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<paragraph id="8BAF36B3BA5C346AA7963F4255FAFA57" blockId="9.[136,1452,1403,1460]" pageId="9" pageNumber="239">
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<emphasis id="B964EAA1BA5C346AA7963F42518AFA73" bold="true" box="[136,254,1403,1424]" pageId="9" pageNumber="239">FIGURE 6.</emphasis>
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<taxonomicName id="4C104D30BA5C346AA61B3F425375FA73" authorityName="A. Doucette & J. Portilla" authorityYear="2016" box="[261,513,1403,1424]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="9" pageNumber="239" phylum="Tracheophyta" rank="species" species="gigiportillae">
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<emphasis id="B964EAA1BA5C346AA61B3F425375FA73" box="[261,513,1403,1424]" italics="true" pageId="9" pageNumber="239">Pleurothallis gigiportillae</emphasis>
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</taxonomicName>
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. A. Habit, drawn to 5 cm scale. B. Flower. C. Dorsal sepal. D. Lateral sepals. E. Petal. F. Lower surface of lip. G. Upper surface of lip. H. Flower with petals and sepals removed. All drawn to 1 mm scale.
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</paragraph>
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</caption>
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<paragraph id="8BAF36B3BA5C346AA7963C305479F935" blockId="9.[136,1452,1545,1966]" pageId="9" pageNumber="239">
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Epiphytic, caespitose herb; roots
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<quantity id="4CE89B56BA5C346AA51C3C305331F9C1" box="[514,581,1545,1570]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="239" unit="mm" value="1.0">1 mm</quantity>
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in diam. The stem 120–335 5
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<quantity id="4CE89B56BA5C346AA4BD3C30529CF9C1" box="[931,1000,1545,1570]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="239" unit="mm" value="3.0">3 mm</quantity>
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, and enveloped by two to three tubular, papery sheaths 13–55 5
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<quantity id="4CE89B56BA5C346AA68D3C1450A2F9A5" box="[403,470,1581,1606]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="239" unit="mm" value="3.0">3 mm</quantity>
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. Leaves sessile, lanceolate,
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<quantity id="4CE89B56BA5C346AA4173C1452FBF9A5" box="[777,911,1581,1606]" metricMagnitude="-1" metricUnit="m" metricValue="1.2" metricValueMax="1.5" metricValueMin="0.9" pageId="9" pageNumber="239" unit="mm" value="120.0" valueMax="150.0" valueMin="90.0">90–150 mm</quantity>
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5
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<quantity id="4CE89B56BA5C346AA4B63C145557F9A5" box="[936,1059,1581,1606]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" metricValueMax="2.0" metricValueMin="1.6" pageId="9" pageNumber="239" unit="mm" value="18.0" valueMax="20.0" valueMin="16.0">16–20 mm</quantity>
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. Inflorescence with a single flower, rarely a two flowered raceme, protected by a papery sheath at the base,
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<quantity id="4CE89B56BA5C346AA4BD3C685285F989" box="[931,1009,1617,1642]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="9" pageNumber="239" unit="mm" value="18.0">18 mm</quantity>
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long; the peduncle 24–27 5
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<quantity id="4CE89B56BA5C346AA2323C68541BF989" box="[1324,1391,1617,1642]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="239" unit="mm" value="1.0">1 mm</quantity>
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, with a tubular bract before the pedicel, 7 5
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||
<quantity id="4CE89B56BA5C346AA5333C4C5304F96D" box="[557,624,1653,1678]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="239" unit="mm" value="2.0">2 mm</quantity>
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||
; the pedicel 6 5
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||
<quantity id="4CE89B56BA5C346AA43A3C4C521CF96D" box="[804,872,1653,1678]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="239" unit="mm" value="1.0">1 mm</quantity>
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; ovary 5.0 5
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||
<quantity id="4CE89B56BA5C346AA4E83C4C553AF96D" box="[1014,1102,1653,1678]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="239" unit="mm" value="1.5">1.5 mm</quantity>
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, flowers non-resupinate, dorsal sepal ovate, beige, 7 5
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||
<quantity id="4CE89B56BA5C346AA6963CA050BFF951" box="[392,459,1689,1714]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="9" pageNumber="239" unit="mm" value="6.0">6 mm</quantity>
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; synsepalum broadly ovate, similar in color to the dorsal sepal, 7 5
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||
<quantity id="4CE89B56BA5C346AA3A33CA05475F951" box="[1213,1281,1689,1714]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="9" pageNumber="239" unit="mm" value="8.0">8 mm</quantity>
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; petal sigmoid, darker beige than the sepals, 10 5
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||
<quantity id="4CE89B56BA5C346AA51B3C84533DF935" box="[517,585,1725,1750]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="239" unit="mm" value="2.0">2 mm</quantity>
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; lip reniform, papillose, pink, 4 5
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<quantity id="4CE89B56BA5C346AA4D83C84557EF935" box="[966,1034,1725,1750]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="239" unit="mm" value="4.0">4 mm</quantity>
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||
; column 3.0 5
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||
<quantity id="4CE89B56BA5C346AA3B13C84547CF935" box="[1199,1288,1725,1750]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="239" unit="mm" value="1.5">1.5 mm</quantity>
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||
.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30A6538BA5C346AA7A33CD854DFF84D" pageId="9" pageNumber="239" type="etymology">
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<paragraph id="8BAF36B3BA5C346AA7A33CD8524EF8FD" blockId="9.[136,1452,1545,1966]" pageId="9" pageNumber="239">
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<emphasis id="B964EAA1BA5C346AA7A33CD85010F919" bold="true" box="[189,356,1761,1786]" pageId="9" pageNumber="239">Etymology:—</emphasis>
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From the Latin
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<emphasis id="B964EAA1BA5C346AA50D3CD85315F919" box="[531,609,1761,1786]" italics="true" pageId="9" pageNumber="239">papula</emphasis>
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, a small protuberance, and
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<emphasis id="B964EAA1BA5C346AA48B3CD852A9F919" box="[917,989,1761,1786]" italics="true" pageId="9" pageNumber="239">lingua</emphasis>
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, tongue, in reference to the texture of the lip, which distinguishes the new species from similar species.
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</paragraph>
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<paragraph id="8BAF36B3BA5C346AA7A33D1054DFF84D" blockId="9.[136,1452,1545,1966]" pageId="9" pageNumber="239">
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<emphasis id="B964EAA1BA5C346AA7A33D105052F8A1" bold="true" box="[189,294,1833,1858]" pageId="9" pageNumber="239">Notes:—</emphasis>
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The species belongs to
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<taxonomicName id="4C104D30BA5C346AA5393D1052CEF8A1" box="[551,954,1833,1858]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="9" pageNumber="239" phylum="Tracheophyta" rank="subSection" subSection="Macrophyllae">
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<emphasis id="B964EAA1BA5C346AA5393D1053CCF8A1" box="[551,696,1833,1858]" italics="true" pageId="9" pageNumber="239">Pleurothallis</emphasis>
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subsect.
|
||
<emphasis id="B964EAA1BA5C346AA4003D1052CEF8A1" box="[798,954,1833,1858]" italics="true" pageId="9" pageNumber="239">Macrophyllae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B964EAA1BA5C346AA4A43D105530F8A1" box="[954,1092,1833,1858]" italics="true" pageId="9" pageNumber="239">-Racemosae</emphasis>
|
||
<bibRefCitation id="EF814B42BA5C346AA3573D10558DF8A1" author="Luer, C." box="[1097,1273,1833,1858]" pageId="9" pageNumber="239" pagination="1 - 109" refId="ref8171" refString="Luer, C. (1986) Icones pleurothallidinarum III: Systematics of Pleurothallis. Monographs in Systematic Botany 20: 1 - 109." type="journal article" year="1986">Luer (1986: 61)</bibRefCitation>
|
||
and is excluded from the key (
|
||
<bibRefCitation id="EF814B42BA5C346AA6313D7450DCF885" author="Luer, C." box="[303,424,1869,1894]" pageId="9" pageNumber="239" pagination="1 - 182" refId="ref8335" refString="Luer, C. (1999) Icones pleurothallidinarum XVIII: aystematics of Pleurothallis subgen. Pleurothallis, sect. Pleurothallis, subsect. Antenniferae, subsect. Longiracemosae, subsect. Macrophyllae-Racemosae, subsect. Perplexae, subgen. Pseudostelis, subgen. Acuminatia. Monographs in Systematic Botany 76: 1 - 182." type="journal article" year="1999">Luer 1999</bibRefCitation>
|
||
) at couplet 17 because it lacks both calli and broadly rounded lateral lobes. The species is unique in the subsection in having a densely papillose lip. No new species attributable to
|
||
<taxonomicName id="4C104D30BA5C346AA3A93D48543CF869" box="[1207,1352,1905,1930]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="9" pageNumber="239" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B964EAA1BA5C346AA3A93D48543CF869" box="[1207,1352,1905,1930]" italics="true" pageId="9" pageNumber="239">Pleurothallis</emphasis>
|
||
</taxonomicName>
|
||
subsect.
|
||
<emphasis id="B964EAA1BA5C346AA7963DAC50D9F84D" box="[136,429,1941,1966]" italics="true" pageId="9" pageNumber="239">Macrophyllae-Racemosae</emphasis>
|
||
is reported by
|
||
<emphasis id="B964EAA1BA5C346AA54B3DAC52B1F84D" box="[597,965,1941,1966]" italics="true" pageId="9" pageNumber="239">
|
||
International Plant
|
||
<taxonomicName id="4C104D30BA5C346AA42D3DAC52B1F84D" box="[819,965,1941,1966]" class="Liliopsida" family="Orchidaceae" genus="Names" kingdom="Plantae" order="Asparagales" pageId="9" pageNumber="239" phylum="Tracheophyta" rank="species" species="index">Names Index</taxonomicName>
|
||
</emphasis>
|
||
(2016) to have been published 2005–2015.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |