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<mods:title id="E13AA0C107266E32D4EBDD86D39D023D">A review of the damselfly genus Megalestes Selys, 1862 (Insecta: Odonata Zygoptera: Synlestidae) using integrative taxonomic methods</mods:title>
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<mods:namePart id="9F33826C8356944E604BF277F3C8F399">Yu, Xin</mods:namePart>
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<mods:namePart id="407DAB7E46FD6E619E58B7FC7368A57B">Xue, Junli</mods:namePart>
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<treatment id="03D487C3FF8DFFBD6EE2F8BFFE9CF9CF" ID-DOI="http://doi.org/10.5281/zenodo.4495344" ID-GBIF-Taxon="177204040" ID-Zenodo-Dep="4495344" LSID="urn:lsid:plazi:treatment:03D487C3FF8DFFBD6EE2F8BFFE9CF9CF" httpUri="http://treatment.plazi.org/id/03D487C3FF8DFFBD6EE2F8BFFE9CF9CF" lastPageId="17" lastPageNumber="261" pageId="16" pageNumber="261">
<subSubSection id="C367655EFF8DFFBC6EE2F8BFFE3EFE36" box="[151,508,1810,1837]" pageId="16" pageNumber="261" type="nomenclature">
<paragraph id="8BC236D5FF8DFFBC6EE2F8BFFE3EFE36" blockId="16.[151,508,1810,1874]" box="[151,508,1810,1837]" pageId="16" pageNumber="261">
<heading id="D08A81B9FF8DFFBC6EE2F8BFFE3EFE36" bold="true" box="[151,508,1810,1837]" fontSize="11" level="1" pageId="16" pageNumber="261" reason="1">
<taxonomicName id="4C7D4D56FF8DFFBC6EE2F8BFFE3EFE36" ID-CoL="72VPL" authority="Selys, 1862" authorityName="Selys" authorityYear="1862" box="[151,508,1810,1837]" class="Insecta" family="Synlestidae" genus="Megalestes" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="261" phylum="Arthropoda" rank="species" species="major">
<emphasis id="B909EAC7FF8DFFBC6EE2F8BFFE3EFE36" bold="true" box="[151,508,1810,1837]" pageId="16" pageNumber="261">
<emphasis id="B909EAC7FF8DFFBC6EE2F8BFFEAEFE37" bold="true" box="[151,364,1810,1837]" italics="true" pageId="16" pageNumber="261">Megalestes major</emphasis>
Selys, 1862
</emphasis>
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<subSubSection id="C367655EFF8DFFBC6EE2F89AFE5AFE48" box="[151,408,1846,1874]" pageId="16" pageNumber="261" type="description">
<paragraph id="8BC236D5FF8DFFBC6EE2F89AFE5AFE48" blockId="16.[151,508,1810,1874]" box="[151,408,1846,1874]" pageId="16" pageNumber="261">
(
<figureCitation id="13462A50FF8DFFBC6EEAF89AFF3CFE4B" box="[159,254,1847,1873]" captionStart="FIGURE 3" captionStartId="14.[151,250,1420,1444]" captionTargetBox="[160,1427,195,1385]" captionTargetId="figure-34@14.[151,1436,181,1396]" captionTargetPageId="14" captionText="FIGURE 3. Caudal appendages of males (dorsal view). a) australis; b) irma, paratype; c) distans, from Yaan, Sichuan; d) distans from Dashahe, Guizhou; e) heros; f) omeiensis, paratype; g) kurahashii; h) major; i) haui; j) “palaceus”; k) micans, bd: basal disk; l) “chengi”; m) riccii; n) “maai”; o) “tuska,” from Damingshan, Guangxi; p) “tuska,” from Jinxiu, Guangxi. (Scale bar = 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.4407489" httpUri="https://zenodo.org/record/4407489/files/figure.png" pageId="16" pageNumber="261">Figs. 3h</figureCitation>
;
<figureCitation id="13462A50FF8DFFBC6F7FF89AFEEBFE4B" box="[266,297,1847,1873]" captionStart="FIGURE 4" captionStartId="15.[151,250,1443,1467]" captionTargetBox="[175,1424,244,1412]" captionTargetId="figure-56@15.[151,1437,237,1419]" captionTargetPageId="15" captionText="FIGURE 4. Caudal appendages of males (lateral view). a) australis; b) irma, paratype; c) distans, from Yaan, Sichuan; d) distans from Qingchengshan, Chengdu, Sichuan; e) heros; f) omeiensis, paratype; g) kurahashii; h) major; i) haui; j) “palaceus”; k) micans; l) “Chengi,” paratype; m) riccii; n) maai; o) tuska, from Damingshan, Guangxi; p) tuska, from Jinxiu, Guangxi. (Scale bar = 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.4407493" httpUri="https://zenodo.org/record/4407493/files/figure.png" pageId="16" pageNumber="261">4h</figureCitation>
;
<figureCitation id="13462A50FF8DFFBC6F41F89BFEABFE4B" box="[308,361,1846,1873]" captionStart="FIGURE 5" captionStartId="18.[151,250,1869,1893]" captionTargetBox="[214,1369,199,1838]" captionTargetId="figure-34@18.[205,1381,181,1845]" captionTargetPageId="18" captionText="FIGURE 5. Genital ligula of males (lateral view) (am) and ventral view (nab). a) australis; b) distans; c) heros; d) omeiensis, paratype; e) major; f) irma, paratype; g) “palaceus”; h) haui; i) “Chengi”; j) maai; k) riccii; l) tuska; m) micans; n) australis, Hr: horn; o) distans, Tlb: terminal lateral lobe; p) heros; q) omeiensis, paratype; r) major; s) irma, paratype; t) kurahashii; u) “palaceus”; v) haui; w) micans, from Jinping, Yunnan; x) “Chengi”; y) maai; z) riccii, Blb: basal lateral lobe; aa) tuska, Mlb: middle lateral lobe; ab) micans, from Fengyangshan, Zhejiang. (Scale bar = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.4407495" httpUri="https://zenodo.org/record/4407495/files/figure.png" pageId="16" pageNumber="261">5e, r</figureCitation>
;
<figureCitation id="13462A50FF8DFFBC6F00F89AFE52FE48" box="[373,400,1847,1874]" captionStart="FIGURE 6" captionStartId="19.[151,250,1319,1343]" captionTargetBox="[168,1412,187,1286]" captionTargetId="figure-17@19.[163,1423,181,1295]" captionTargetPageId="19" captionText="FIGURE 6. Line drawing of male paraprocts (lateral view). a) australis; b) distans; c) gyalsey; d) kurahashii; e) major; f) haui; g) heros; h) omeiensis; i) micans; j) micans; k) riccii; l) irma." figureDoi="http://doi.org/10.5281/zenodo.4407497" httpUri="https://zenodo.org/record/4407497/files/figure.png" pageId="16" pageNumber="261">6e</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C367655EFF8DFFBC6EE2F8DBFAB2FE94" box="[151,1392,1910,1934]" pageId="16" pageNumber="261" type="reference_group">
<paragraph id="8BC236D5FF8DFFBC6EE2F8DBFAB2FE94" blockId="16.[151,1392,1910,1934]" box="[151,1392,1910,1934]" pageId="16" pageNumber="261">
<treatmentCitationGroup id="AB6D11FBFF8DFFBC6EE2F8DBFAB2FE94" box="[151,1392,1910,1934]" pageId="16" pageNumber="261">
<treatmentCitation id="0ADC10C4FF8DFFBC6EE2F8DBFDECFE94" author="Asahina, S." box="[151,558,1910,1934]" page="5" pageId="16" pageNumber="261" year="1985">
<taxonomicName id="4C7D4D56FF8DFFBC6EE2F8DBFDECFE94" ID-CoL="72VPL" authority=": Asahina, 1985 b: 2 - 5" authorityName=": Asahina" authorityPageNumber="2 - 5" authorityYear="1985" box="[151,558,1910,1934]" class="Insecta" family="Synlestidae" genus="Megalestes" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="261" phylum="Arthropoda" rank="species" species="major">
<emphasis id="B909EAC7FF8DFFBC6EE2F8DBFE8EFE97" box="[151,332,1910,1934]" italics="true" pageId="16" pageNumber="261">Megalestes major</emphasis>
:
<bibRefCitation id="EFEC4B24FF8DFFBC6F2CF8DBFDECFE94" author="Asahina, S." box="[345,558,1910,1934]" pageId="16" pageNumber="261" pagination="1 - 18" refId="ref13502" refString="Asahina, S. (1985 b) Contributions to the taxonomic knowledge of the Megalestes species of continental South Asia (Odonata, Synlestidae). Cho-Cho, 8 (10), 1 - 18." type="journal article" year="1985">Asahina, 1985b: 25</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, 1416, figs. 110 [
<collectingCountry id="F36A7645FF8DFFBC6C83F8DBFCF2FE94" box="[758,816,1910,1934]" name="India" pageId="16" pageNumber="261">India</collectingCountry>
;
<collectingCountry id="F36A7645FF8DFFBC6D4FF8DBFCBEFE94" box="[826,892,1910,1934]" name="Nepal" pageId="16" pageNumber="261">Nepal</collectingCountry>
];
<treatmentCitation id="0ADC10C4FF8DFFBC6DF9F8DBFB58FE94" author="Gyeltshen, T. &amp; Kalkman, V. J. &amp; Orr, A. G." box="[908,1178,1910,1934]" page="593" pageId="16" pageNumber="261" year="2017">
<bibRefCitation id="EFEC4B24FF8DFFBC6DF9F8DBFB58FE94" author="Gyeltshen, T. &amp; Kalkman, V. J. &amp; Orr, A. G." box="[908,1178,1910,1934]" pageId="16" pageNumber="261" pagination="588 - 594" refId="ref13981" refString="Gyeltshen, T., Kalkman, V. J. &amp; Orr, A. G. (2017) Honouring His Royal Highness the Crown Prince of Bhutan: Megalestes gyalsey (Odonata: Synlestidae). Zootaxa, 4244 (4), 588 - 594. https: // doi. org / 10.11646 / zootaxa. 4244.4.9" type="journal article" year="2017">
Gyeltshen
<emphasis id="B909EAC7FF8DFFBC6D8CF8DAFBEFFE94" box="[1017,1069,1910,1934]" italics="true" pageId="16" pageNumber="261">et al.</emphasis>
2017: 593
</bibRefCitation>
</treatmentCitation>
, figs. 4cd [
<collectingCountry id="F36A7645FF8DFFBC6B60F8DBFAA4FE94" box="[1301,1382,1910,1934]" name="Bhutan" pageId="16" pageNumber="261">Bhutan</collectingCountry>
].
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</paragraph>
</subSubSection>
<paragraph id="8BC236D5FF8DFFBD6EE2F81AFE9CF9CF" blockId="16.[151,1437,1975,2038]" lastBlockId="17.[151,1436,151,213]" lastPageId="17" lastPageNumber="262" pageId="16" pageNumber="261">
This is the
<typeStatus id="54C68877FF8DFFBC6F64F81AFE83FECB" box="[273,321,1975,2001]" pageId="16" pageNumber="261">type</typeStatus>
species of
<taxonomicName id="4C7D4D56FF8DFFBC6FC8F81AFDF9FECB" ID-CoL="5MR3" authorityName="Selys" authorityYear="1862" box="[445,571,1975,2001]" class="Insecta" family="Synlestidae" genus="Megalestes" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="261" phylum="Arthropoda" rank="genus">
<emphasis id="B909EAC7FF8DFFBC6FC8F81AFDF9FECB" box="[445,571,1975,2001]" italics="true" pageId="16" pageNumber="261">Megalestes</emphasis>
</taxonomicName>
, the only species that has an inner lateral tooth in the middle of the cerci (
<figureCitation id="13462A50FF8DFFBC6B1AF81AFF74FEEC" captionStart="FIGURE 3" captionStartId="14.[151,250,1420,1444]" captionTargetBox="[160,1427,195,1385]" captionTargetId="figure-34@14.[151,1436,181,1396]" captionTargetPageId="14" captionText="FIGURE 3. Caudal appendages of males (dorsal view). a) australis; b) irma, paratype; c) distans, from Yaan, Sichuan; d) distans from Dashahe, Guizhou; e) heros; f) omeiensis, paratype; g) kurahashii; h) major; i) haui; j) “palaceus”; k) micans, bd: basal disk; l) “chengi”; m) riccii; n) “maai”; o) “tuska,” from Damingshan, Guangxi; p) “tuska,” from Jinxiu, Guangxi. (Scale bar = 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.4407489" httpUri="https://zenodo.org/record/4407489/files/figure.png" pageId="16" pageNumber="261">Fig. 3h</figureCitation>
). Its paraprocts are almost spherical, covered with dense setae, with an acute dorsal base tooth pointed directly
<subSubSection id="C367655EFF8CFFBD6EE2FF3AFE9CF9CF" pageId="17" pageNumber="262" type="description">
upwards (
<figureCitation id="13462A50FF8CFFBD6F73FF3AFE9AF9A8" box="[262,344,151,178]" captionStart="FIGURE 6" captionStartId="19.[151,250,1319,1343]" captionTargetBox="[168,1412,187,1286]" captionTargetId="figure-17@19.[163,1423,181,1295]" captionTargetPageId="19" captionText="FIGURE 6. Line drawing of male paraprocts (lateral view). a) australis; b) distans; c) gyalsey; d) kurahashii; e) major; f) haui; g) heros; h) omeiensis; i) micans; j) micans; k) riccii; l) irma." figureDoi="http://doi.org/10.5281/zenodo.4407497" httpUri="https://zenodo.org/record/4407497/files/figure.png" pageId="17" pageNumber="262">Fig. 6e</figureCitation>
).
<taxonomicName id="4C7D4D56FF8CFFBD6F19FF3AFDF6F9AA" authorityName=": Asahina" authorityYear="1985" box="[364,564,151,177]" class="Insecta" family="Synlestidae" genus="Megalestes" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="262" phylum="Arthropoda" rank="species" species="major">
<emphasis id="B909EAC7FF8CFFBD6F19FF3AFDF6F9AA" box="[364,564,151,177]" italics="true" pageId="17" pageNumber="262">Megalestes major</emphasis>
</taxonomicName>
is distributed in
<collectingCountry id="F36A7645FF8CFFBD6C84FF3AFCEFF9AB" box="[753,813,151,177]" name="India" pageId="17" pageNumber="262">India</collectingCountry>
,
<collectingCountry id="F36A7645FF8CFFBD6D4CFF3AFCBDF9AB" box="[825,895,151,177]" name="Nepal" pageId="17" pageNumber="262">Nepal</collectingCountry>
, and
<collectingCountry id="F36A7645FF8CFFBD6DCCFF3AFBCFF9AB" box="[953,1037,151,177]" name="Bhutan" pageId="17" pageNumber="262">Bhutan</collectingCountry>
, and it only occurs on the west side of the Himalayas.
</subSubSection>
</paragraph>
</treatment>
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