265 lines
30 KiB
XML
265 lines
30 KiB
XML
<document id="22D6278CC69FC1A837F1C4E1747CB103" ID-CLB-Dataset="24541" ID-DOI="10.11646/zootaxa.4707.1.1" ID-GBIF-Dataset="7130e278-109a-4da2-b2e7-adeca14b5c77" ID-ISSN="1175-5326" ID-Zenodo-Dep="3587155" ID-ZooBank="3C00FB35-708D-4FF0-94C2-B15BD2A1F37A" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1576824974133" checkinUser="plazi" docAuthor="Köhler, Arne" docDate="2019" docId="03D43F593676FFE9FF32FC34346E3A84" docLanguage="en" docName="zootaxa.4707.1.1.pdf" docOrigin="Zootaxa 4707 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Pseudolycoriella jejuna" docType="treatment" docVersion="10" lastPageNumber="30" masterDocId="FFED4721366DFFF4FFA5FF8535443B73" masterDocTitle="The genus Pseudolycoriella Menzel & Mohrig, 1998 (Diptera, Sciaridae) in New Zealand" masterLastPageNumber="69" masterPageNumber="1" pageNumber="28" updateTime="1698775769700" updateUser="ExternalLinkService">
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<mods:title id="6C867BACEC80995CBB13403A35B0719E">The genus Pseudolycoriella Menzel & Mohrig, 1998 (Diptera, Sciaridae) in New Zealand</mods:title>
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<mods:namePart id="F1A4DE398E49BA01DBCDA9BFA7605EBA">Köhler, Arne</mods:namePart>
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<mods:title id="FDCAC53592E4C427CA0C27F737875889">Zootaxa</mods:title>
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<mods:part id="DEFDD449A032BE069DB0607D2D2332A9">
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<mods:date id="4AAE07CA03FAD630B090A1B3E94D0753">2019</mods:date>
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<mods:number id="8E7543305998A55BC083E949967534DE">4707</mods:number>
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<mods:identifier id="3DB1C590B41905EE6725A77058B90D26" type="DOI">10.11646/zootaxa.4707.1.1</mods:identifier>
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<subSubSection id="C367DDC43676FFEFFF32FC3430C53E94" pageId="27" pageNumber="28" type="nomenclature">
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<paragraph id="8BC28E4F3676FFEFFF32FC34376738B8" blockId="27.[151,547,945,972]" box="[151,547,945,972]" pageId="27" pageNumber="28">
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<heading id="D08A39233676FFEFFF32FC34376738B8" bold="true" box="[151,547,945,972]" fontSize="11" level="1" pageId="27" pageNumber="28" reason="1">
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<emphasis id="B909525D3676FFEFFF32FC34376738B8" bold="true" box="[151,547,945,972]" pageId="27" pageNumber="28">
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<taxonomicName id="4C7DF5CC3676FFEFFF32FC3434F038B8" ID-CoL="4P24H" baseAuthorityName="Edwards" baseAuthorityYear="1927" box="[151,436,945,972]" class="Insecta" family="Sciaridae" genus="Pseudolycoriella" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="jejuna">
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<emphasis id="B909525D3676FFEFFF32FC3434F038B8" bold="true" box="[151,436,945,972]" italics="true" pageId="27" pageNumber="28">Pseudolycoriella jejuna</emphasis>
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</taxonomicName>
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complex
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BC28E4F3676FFEFFF32FC7C30C53E94" blockId="27.[151,1437,1017,2015]" pageId="27" pageNumber="28">
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The
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<emphasis id="B909525D3676FFEFFF6CFC7C35A93F60" box="[201,237,1017,1043]" italics="true" pageId="27" pageNumber="28">Psl</emphasis>
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||
.
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<taxonomicName id="4C7DF5CC3676FFEFFF5CFC7F347B3F60" baseAuthorityName="Edwards" baseAuthorityYear="1927" box="[249,319,1018,1043]" class="Insecta" family="Sciaridae" genus="Pseudolycoriella" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="jejuna">
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<emphasis id="B909525D3676FFEFFF5CFC7F347B3F60" box="[249,319,1018,1043]" italics="true" pageId="27" pageNumber="28">jejuna</emphasis>
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</taxonomicName>
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complex is a group of thirteen closely related and cryptic species. These species share a complex and synapomorph structure of their hypopygia. In
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<figureCitation id="134692CA3676FFEFFD39FB9836153F44" box="[668,849,1053,1079]" captionStart="FIGURES 20–24" captionStartId="28.[151,264,1825,1850]" captionTargetBox="[236,1351,167,1806]" captionTargetId="figure@28.[236,1351,166,1806]" captionTargetPageId="28" captionText="FIGURES 20–24. Pseudolycoriella jejuna complex, Pseudolycoriella tewaipounamu sp. n., details of hypopygium. 20. Hypopygium, ventral. 21. Hypopygium, dorsal, tergite IX removed. 22. Hypopygium, ventral. 23. Tegmen, ventral. 24. Tegmen, dorsal. Abbreviations: ac = apical cavity; at = apex of tegmen; be = base of ejaculatory apodeme; be* = base of ejaculatory apodeme covered by the membranous area; dc = dorsal carina; df = dorsal fold; ea = ejaculatory apodeme; gc = gonocoxite; gs = gonostylus; ll = lateral lobe; ma = membranous area; mf = median fold; pa = parameral apodeme; pa* = parameral apodeme covered by integument; sp = gonostylar spine; wh = whip-lash hair." figureDoi="http://doi.org/10.5281/zenodo.3587165" httpUri="https://zenodo.org/record/3587165/files/figure.png" pageId="27" pageNumber="28">Figures 20–22 a</figureCitation>
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hypopygium of
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<emphasis id="B909525D3676FFEFFBA9FB9831743F44" box="[1036,1072,1053,1079]" italics="true" pageId="27" pageNumber="28">Psl</emphasis>
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.
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<taxonomicName id="4C7DF5CC3676FFEFFB9BFB9B31A73F44" authorityName="Köhler" authorityYear="2019" box="[1086,1251,1054,1079]" class="Insecta" family="Sciaridae" genus="Pseudolycoriella" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="tewaipounamu">
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<emphasis id="B909525D3676FFEFFB9BFB9B31A73F44" box="[1086,1251,1054,1079]" italics="true" pageId="27" pageNumber="28">tewaipounamu</emphasis>
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</taxonomicName>
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is depicted. The gonostylus (
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<emphasis id="B909525D3676FFEFFE84FBC7347E3F28" box="[289,314,1090,1115]" italics="true" pageId="27" pageNumber="28">gs</emphasis>
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) is elongated and curved with an inwardly bent apex. The apex possesses one spine (
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<emphasis id="B909525D3676FFEFFB55FBC7304D3F28" box="[1264,1289,1090,1115]" italics="true" pageId="27" pageNumber="28">sp</emphasis>
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) and a whiplash hair (
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<emphasis id="B909525D3676FFEFFEACFBE0346E3F0C" box="[265,298,1125,1151]" italics="true" pageId="27" pageNumber="28">wh</emphasis>
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) and—in most species—on the inward part a cavity (
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<emphasis id="B909525D3676FFEFFC2EFBE336E13F0C" box="[907,933,1126,1151]" italics="true" pageId="27" pageNumber="28">ac</emphasis>
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). The tegmen is highly derivative compared with the tegmen of typical sciarids. Its general shape is onion-shaped: more or less roundish with an apical contraction (
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<emphasis id="B909525D3676FFEFFF77FB2B35AC3FB4" box="[210,232,1198,1223]" italics="true" pageId="27" pageNumber="28">at</emphasis>
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). Adjacent to the apical contraction the tegmen has two lateral lobes (
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<emphasis id="B909525D3676FFEFFC49FB2836B83FB4" box="[1004,1020,1197,1223]" italics="true" pageId="27" pageNumber="28">ll</emphasis>
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) on the ventral side, which can easily be observed in light microscopy. In the middle of the ventral side of the tegmen a membranous area (
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<emphasis id="B909525D3676FFEFFAA3FB57306C3F98" box="[1286,1320,1234,1259]" italics="true" pageId="27" pageNumber="28">ma</emphasis>
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) bordered by two median folds (
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<emphasis id="B909525D3676FFEFFE28FB7034ED3E7C" box="[397,425,1269,1295]" italics="true" pageId="27" pageNumber="28">mf</emphasis>
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) can easily be observed by using scanning electron microscopy. In light microscopy these delicate structures are hardly visible. At this membranous area a bulge of the base of the ejaculatory apodeme (
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<emphasis id="B909525D3676FFEFFADDFA9C30D63E40" box="[1400,1426,1305,1331]" italics="true" pageId="27" pageNumber="28">be</emphasis>
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) can be observed. An area of teeth is absent. In light microscopy, the parameral apodemes (
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<emphasis id="B909525D3676FFEFFB23FABB31E63E24" box="[1158,1186,1342,1367]" italics="true" pageId="27" pageNumber="28">pa</emphasis>
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) do not fuse medially. Instead they are stretched to the apex of the tegmen and form the apical contraction (
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<emphasis id="B909525D3676FFEFFBFDFAE7312A3E08" box="[1112,1134,1378,1403]" italics="true" pageId="27" pageNumber="28">at</emphasis>
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). On the dorsal side of the tegmen the parameral apodemes protrude and form a dorsal carina (
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<emphasis id="B909525D3676FFEFFC26FA0036D93EEC" box="[899,925,1413,1439]" italics="true" pageId="27" pageNumber="28">dc</emphasis>
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). This carina is bent around the apex and ends subapically on the ventral side of the tegmen. The bulges of the parameral apodemes are accompanied by a dorsal fold (
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<emphasis id="B909525D3676FFEFFF70FA4835AF3E94" box="[213,235,1485,1511]" italics="true" pageId="27" pageNumber="28">df</emphasis>
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), which can also be observed in light microscopy. The phylogeny of this complex is shown in
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<figureCitation id="134692CA3676FFEFFAABFA4830393E94" box="[1294,1405,1485,1511]" captionStart="FIGURE 61" captionStartId="63.[151,250,1859,1884]" captionTargetBox="[218,1376,171,1833]" captionTargetId="figure@63.[210,1377,166,1839]" captionTargetPageId="63" captionText="FIGURE 61. Phylogenetic tree of the New Zealand Pseudolycoriella species inferred from Maximum Likelihood analysis based on three concatenated genes (COI, 16S, and 28S; total length 3,053 bp) applying the model GTR+F+R4. Ultrafast bootstrap values (UFBoot; 1,000 replicates) are given if these values are greater than or equal to 95%. Intraspecific UFBoot values are not specified. Pseudolycoriella species without available genetic data are also inserted and assigned to a species complex according to the morphological results." figureDoi="http://doi.org/10.5281/zenodo.3587177" httpUri="https://zenodo.org/record/3587177/files/figure.png" pageId="27" pageNumber="28">Figure 61</figureCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C367DDC43676FFE9FF62FA75346E3A84" lastPageId="29" lastPageNumber="30" pageId="27" pageNumber="28" type="discussion">
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<paragraph id="8BC28E4F3676FFE9FF62FA7537153BA4" blockId="27.[151,1437,1017,2015]" lastBlockId="29.[151,1437,153,503]" lastPageId="29" lastPageNumber="30" pageId="27" pageNumber="28">
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<emphasis id="B909525D3676FFEFFF62FA7534083D78" bold="true" box="[199,332,1520,1547]" pageId="27" pageNumber="28">Discussion.</emphasis>
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The species of the
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<emphasis id="B909525D3676FFEFFDBAFA7437073D78" box="[543,579,1521,1547]" italics="true" pageId="27" pageNumber="28">Psl</emphasis>
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.
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<taxonomicName id="4C7DF5CC3676FFEFFDF5FA7737D23D78" baseAuthorityName="Edwards" baseAuthorityYear="1927" box="[592,662,1522,1547]" class="Insecta" family="Sciaridae" genus="Pseudolycoriella" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="jejuna">
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<emphasis id="B909525D3676FFEFFDF5FA7737D23D78" box="[592,662,1522,1547]" italics="true" pageId="27" pageNumber="28">jejuna</emphasis>
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</taxonomicName>
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complex can be assigned to the
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<emphasis id="B909525D3676FFEFFC5FFA74315A3D78" box="[1018,1054,1521,1547]" italics="true" pageId="27" pageNumber="28">Psl</emphasis>
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.
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<taxonomicName id="4C7DF5CC3676FFEFFB8EFA74313E3D78" baseAuthorityName="Winnertz" baseAuthorityYear="1867" box="[1067,1146,1521,1547]" class="Insecta" family="Sciaridae" genus="Pseudolycoriella" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="bruckii">
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<emphasis id="B909525D3676FFEFFB8EFA74313E3D78" box="[1067,1146,1521,1547]" italics="true" pageId="27" pageNumber="28">bruckii</emphasis>
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</taxonomicName>
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group. The monophyly of this complex is strongly supported by genetics and can also be demonstrated by some autapomorphic morphological characters such as the extraordinary tegmen, the reduction of the number of gonostylar spines, and the secondary absence of an area of teeth. Due to this unique character set, all species of the
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<emphasis id="B909525D3676FFEFFC52F9D8315F3D04" box="[1015,1051,1629,1655]" italics="true" pageId="27" pageNumber="28">Psl</emphasis>
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.
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<taxonomicName id="4C7DF5CC3676FFEFFB8DF9DB312A3D04" baseAuthorityName="Edwards" baseAuthorityYear="1927" box="[1064,1134,1630,1655]" class="Insecta" family="Sciaridae" genus="Pseudolycoriella" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="jejuna">
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<emphasis id="B909525D3676FFEFFB8DF9DB312A3D04" box="[1064,1134,1630,1655]" italics="true" pageId="27" pageNumber="28">jejuna</emphasis>
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</taxonomicName>
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complex can easily be recognized. Only
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<emphasis id="B909525D3676FFEFFE99F90434243DE8" box="[316,352,1665,1691]" italics="true" pageId="27" pageNumber="28">Psl</emphasis>
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.
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<emphasis id="B909525D3676FFEFFEC8F90437503DE8" box="[365,532,1665,1691]" italics="true" pageId="27" pageNumber="28">fuscorubroides</emphasis>
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(Mohrig & Blasco-Zumeta) from
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<collectingCountry id="F36ACEDF3676FFEFFC31F90436903DE8" box="[916,980,1665,1691]" name="Spain" pageId="27" pageNumber="28">Spain</collectingCountry>
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has a similar tegmen (Mohrig & Blasco- Zumeta 1996). It also possesses a dorsal carina that derives from the parameral apodemes. However, there are differences: the parameral apodemes are fused and not separated by a gap, as in the
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<emphasis id="B909525D3676FFEFFBB3F94C317E3D90" box="[1046,1082,1737,1763]" italics="true" pageId="27" pageNumber="28">Psl</emphasis>
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.
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<taxonomicName id="4C7DF5CC3676FFEFFBE2F94F31C93D90" baseAuthorityName="Edwards" baseAuthorityYear="1927" box="[1095,1165,1738,1763]" class="Insecta" family="Sciaridae" genus="Pseudolycoriella" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="jejuna">
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<emphasis id="B909525D3676FFEFFBE2F94F31C93D90" box="[1095,1165,1738,1763]" italics="true" pageId="27" pageNumber="28">jejuna</emphasis>
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</taxonomicName>
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complex, and the carina in
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<emphasis id="B909525D3676FFEFFF16F96835933C74" box="[179,215,1773,1799]" italics="true" pageId="27" pageNumber="28">Psl</emphasis>
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.
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<emphasis id="B909525D3676FFEFFF41F96834CF3C74" box="[228,395,1773,1799]" italics="true" pageId="27" pageNumber="28">fuscorubroides</emphasis>
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arises vertically in the middle base of the tegmen. Furthermore,
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<emphasis id="B909525D3676FFEFFBF6F96831333C74" box="[1107,1143,1773,1799]" italics="true" pageId="27" pageNumber="28">Psl</emphasis>
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.
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<emphasis id="B909525D3676FFEFFB21F968306F3C74" box="[1156,1323,1773,1799]" italics="true" pageId="27" pageNumber="28">fuscorubroides</emphasis>
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possesses a large area of conspicous teeth (
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<bibRefCitation id="EFECF3BE3676FFEFFDAEF89436313C5F" author="Mohrig, W. & Blasco-Zumeta, J." box="[523,885,1809,1836]" pageId="27" pageNumber="28" pagination="99 - 116" refId="ref59254" refString="Mohrig, W. & Blasco-Zumeta, J. (1996) The sciarid fauna (Diptera, Sciaridae) of a Juniperus thurifera L. forest of the Monegros region (Zaragoza, Spain) with description of ten new species. Miscellanea Zoologica, 18, 99 - 116." type="journal article" year="1996">Mohrig & Blasco-Zumeta 1996</bibRefCitation>
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). Several other structures of the hypopygium are also disimilar. For instance, the gonostylus of
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<emphasis id="B909525D3676FFEFFD37F8B037F23C3C" box="[658,694,1845,1871]" italics="true" pageId="27" pageNumber="28">Psl</emphasis>
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.
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<emphasis id="B909525D3676FFEFFD67F8B0362D3C3C" box="[706,873,1845,1871]" italics="true" pageId="27" pageNumber="28">fuscorubroides</emphasis>
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is armed with two short spines and has a conspicuous lobe on the dorsal margin and therefore differs significantly from the gonostyli of the
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<emphasis id="B909525D3676FFEFFB21F8DC31EC3C00" box="[1156,1192,1881,1907]" italics="true" pageId="27" pageNumber="28">Psl</emphasis>
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.
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<taxonomicName id="4C7DF5CC3676FFEFFB13F8DF31B83C00" baseAuthorityName="Edwards" baseAuthorityYear="1927" box="[1206,1276,1882,1907]" class="Insecta" family="Sciaridae" genus="Pseudolycoriella" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="jejuna">
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<emphasis id="B909525D3676FFEFFB13F8DF31B83C00" box="[1206,1276,1882,1907]" italics="true" pageId="27" pageNumber="28">jejuna</emphasis>
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</taxonomicName>
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complex. It is not clear whether the dorsal carina of the tegmen is a convergent or a homologous character. Maybe
|
||
<emphasis id="B909525D3676FFEFFAA9F8F830743CE4" box="[1292,1328,1917,1943]" italics="true" pageId="27" pageNumber="28">Psl</emphasis>
|
||
.
|
||
<emphasis id="B909525D3676FFEFFA9AF8F835AE3CC8" italics="true" pageId="27" pageNumber="28">fuscorubroides</emphasis>
|
||
is closely related to the
|
||
<emphasis id="B909525D3676FFEFFE59F82437643CC8" box="[508,544,1953,1979]" italics="true" pageId="27" pageNumber="28">Psl</emphasis>
|
||
.
|
||
<taxonomicName id="4C7DF5CC3676FFEFFD8BF82737303CC8" baseAuthorityName="Edwards" baseAuthorityYear="1927" box="[558,628,1954,1979]" class="Insecta" family="Sciaridae" genus="Pseudolycoriella" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="jejuna">
|
||
<emphasis id="B909525D3676FFEFFD8BF82737303CC8" box="[558,628,1954,1979]" italics="true" pageId="27" pageNumber="28">jejuna</emphasis>
|
||
</taxonomicName>
|
||
complex, but it certainly does not belong to the crown group of the
|
||
<emphasis id="B909525D3676FFEFFAD4F82430D13CC8" box="[1393,1429,1953,1979]" italics="true" pageId="27" pageNumber="28">Psl</emphasis>
|
||
.
|
||
<taxonomicName id="4C7DF5CC3676FFEFFF32F84335993CAC" baseAuthorityName="Edwards" baseAuthorityYear="1927" box="[151,221,1990,2015]" class="Insecta" family="Sciaridae" genus="Pseudolycoriella" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="jejuna">
|
||
<emphasis id="B909525D3676FFEFFF32F84335993CAC" box="[151,221,1990,2015]" italics="true" pageId="27" pageNumber="28">jejuna</emphasis>
|
||
</taxonomicName>
|
||
complex, since the absence of an area of teeth can be regarded as a synapomorpy of the
|
||
<collectingCountry id="F36ACEDF3676FFEFFB77F840302D3CAC" box="[1234,1385,1989,2015]" name="New Zealand" pageId="27" pageNumber="28">New Zealand</collectingCountry>
|
||
<emphasis id="B909525D3676FFEFFAD4F84030D13CAC" box="[1393,1429,1989,2015]" italics="true" pageId="27" pageNumber="28">Psl</emphasis>
|
||
.
|
||
<taxonomicName id="4C7DF5CC3670FFE9FF32FF1F35993BC0" baseAuthorityName="Edwards" baseAuthorityYear="1927" box="[151,221,154,179]" class="Insecta" family="Sciaridae" genus="Pseudolycoriella" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="jejuna">
|
||
<emphasis id="B909525D3670FFE9FF32FF1F35993BC0" box="[151,221,154,179]" italics="true" pageId="29" pageNumber="30">jejuna</emphasis>
|
||
</taxonomicName>
|
||
complex members. In other words, the hypothesis of a monophyly of the
|
||
<collectingCountry id="F36ACEDF3670FFE9FBB7FF1C31E33BC0" box="[1042,1191,153,179]" name="New Zealand" pageId="29" pageNumber="30">New Zealand</collectingCountry>
|
||
representatives of the
|
||
<emphasis id="B909525D3670FFE9FF32FF3835FF3BA4" box="[151,187,189,215]" italics="true" pageId="29" pageNumber="30">Psl</emphasis>
|
||
.
|
||
<taxonomicName id="4C7DF5CC3670FFE9FF6CFF3B344B3BA4" baseAuthorityName="Edwards" baseAuthorityYear="1927" box="[201,271,190,215]" class="Insecta" family="Sciaridae" genus="Pseudolycoriella" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="jejuna">
|
||
<emphasis id="B909525D3670FFE9FF6CFF3B344B3BA4" box="[201,271,190,215]" italics="true" pageId="29" pageNumber="30">jejuna</emphasis>
|
||
</taxonomicName>
|
||
complex cannot be rejected.
|
||
</paragraph>
|
||
<caption id="DF02DEC73671FFE8FF32F8A436003C9D" ID-DOI="http://doi.org/10.5281/zenodo.3587165" ID-Zenodo-Dep="3587165" httpUri="https://zenodo.org/record/3587165/files/figure.png" pageId="28" pageNumber="29" startId="28.[151,264,1825,1850]" targetBox="[236,1351,167,1806]" targetPageId="28">
|
||
<paragraph id="8BC28E4F3671FFE8FF32F8A436003C9D" blockId="28.[151,1437,1825,2030]" pageId="28" pageNumber="29">
|
||
<emphasis id="B909525D3671FFE8FF32F8A434123C49" bold="true" box="[151,342,1825,1850]" pageId="28" pageNumber="29">FIGURES 20–24.</emphasis>
|
||
<taxonomicName id="4C7DF5CC3671FFE8FEF8F8A737143C49" baseAuthorityName="Edwards" baseAuthorityYear="1927" box="[349,592,1826,1850]" class="Insecta" family="Sciaridae" genus="Pseudolycoriella" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="jejuna">
|
||
<emphasis id="B909525D3671FFE8FEF8F8A737143C49" box="[349,592,1826,1850]" italics="true" pageId="28" pageNumber="29">Pseudolycoriella jejuna</emphasis>
|
||
</taxonomicName>
|
||
complex,
|
||
<taxonomicName id="4C7DF5CC3671FFE8FD1BF8A731433C49" authority="Köhler, 2019" authorityName="Köhler" authorityYear="2019" box="[702,1031,1826,1850]" class="Insecta" family="Sciaridae" genus="Pseudolycoriella" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="tewaipounamu" status="sp. nov.">
|
||
<emphasis id="B909525D3671FFE8FD1BF8A731433C49" box="[702,1031,1826,1850]" italics="true" pageId="28" pageNumber="29">Pseudolycoriella tewaipounamu</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B909525D3671FFE8FBAAF8A7310D3C49" bold="true" box="[1039,1097,1826,1850]" pageId="28" pageNumber="29">
|
||
<taxonomicNameLabel id="A23AEF263671FFE8FBAAF8A7310D3C49" box="[1039,1097,1826,1850]" pageId="28" pageNumber="29" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, details of hypopygium.
|
||
<emphasis id="B909525D3671FFE8FAE8F8A430293C49" bold="true" box="[1357,1389,1825,1850]" pageId="28" pageNumber="29">20.</emphasis>
|
||
Hypopygium, ventral.
|
||
<emphasis id="B909525D3671FFE8FEC5F8C0343B3C2D" bold="true" box="[352,383,1861,1886]" pageId="28" pageNumber="29">21.</emphasis>
|
||
Hypopygium, dorsal, tergite IX removed.
|
||
<emphasis id="B909525D3671FFE8FC93F8C036123C2D" bold="true" box="[822,854,1861,1886]" pageId="28" pageNumber="29">22.</emphasis>
|
||
Hypopygium, ventral.
|
||
<emphasis id="B909525D3671FFE8FBE1F8C031203C2D" bold="true" box="[1092,1124,1861,1886]" pageId="28" pageNumber="29">23.</emphasis>
|
||
Tegmen, ventral.
|
||
<emphasis id="B909525D3671FFE8FABBF8C0307A3C2D" bold="true" box="[1310,1342,1861,1886]" pageId="28" pageNumber="29">24.</emphasis>
|
||
Tegmen, dorsal. Abbreviations: ac = apical cavity; at = apex of tegmen; be = base of ejaculatory apodeme; be* = base of ejaculatory apodeme covered by the membranous area; dc = dorsal carina; df = dorsal fold; ea = ejaculatory apodeme; gc = gonocoxite; gs = gonostylus; ll = lateral lobe; ma = membranous area; mf = median fold; pa = parameral apodeme; pa* = parameral apodeme covered by integument; sp = gonostylar spine; wh = whip-lash hair.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC28E4F3670FFE9FF62FF64346E3A84" blockId="29.[151,1437,153,503]" pageId="29" pageNumber="30">
|
||
The occurrence of two atavisms among the specimens of the
|
||
<emphasis id="B909525D3670FFE9FCD7FF6436D23B88" box="[882,918,225,251]" italics="true" pageId="29" pageNumber="30">Psl</emphasis>
|
||
.
|
||
<taxonomicName id="4C7DF5CC3670FFE9FC01FF6736AE3B88" baseAuthorityName="Edwards" baseAuthorityYear="1927" box="[932,1002,226,251]" class="Insecta" family="Sciaridae" genus="Pseudolycoriella" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="jejuna">
|
||
<emphasis id="B909525D3670FFE9FC01FF6736AE3B88" box="[932,1002,226,251]" italics="true" pageId="29" pageNumber="30">jejuna</emphasis>
|
||
</taxonomicName>
|
||
complex confirms the above character polarisation regarding the single gonostylar tooth and the absence of the area of teeth. One
|
||
<typeStatus id="54C630ED3670FFE9FB3BFE8031BA3A6C" box="[1182,1278,261,287]" pageId="29" pageNumber="30" type="paratype">paratype</typeStatus>
|
||
of
|
||
<emphasis id="B909525D3670FFE9FA80FE80300D3A6C" box="[1317,1353,261,287]" italics="true" pageId="29" pageNumber="30">Psl</emphasis>
|
||
.
|
||
<taxonomicName id="4C7DF5CC3670FFE9FAFDFE8030D83A6C" authorityName="Köhler" authorityYear="2019" box="[1368,1436,261,287]" class="Insecta" family="Sciaridae" genus="Pseudolycoriella" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="dagae">
|
||
<emphasis id="B909525D3670FFE9FAFDFE8030D83A6C" box="[1368,1436,261,287]" italics="true" pageId="29" pageNumber="30">dagae</emphasis>
|
||
</taxonomicName>
|
||
has two spines on the left gonostylus while the right one only bears one. This atavistic state is unique among all 233 investigated specimens belonging to the
|
||
<emphasis id="B909525D3670FFE9FD3CFEC837F93A14" box="[665,701,333,359]" italics="true" pageId="29" pageNumber="30">Psl</emphasis>
|
||
.
|
||
<taxonomicName id="4C7DF5CC3670FFE9FD69FECB36563A14" baseAuthorityName="Edwards" baseAuthorityYear="1927" box="[716,786,334,359]" class="Insecta" family="Sciaridae" genus="Pseudolycoriella" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="jejuna">
|
||
<emphasis id="B909525D3670FFE9FD69FECB36563A14" box="[716,786,334,359]" italics="true" pageId="29" pageNumber="30">jejuna</emphasis>
|
||
</taxonomicName>
|
||
complex and indicates that the species of the
|
||
<emphasis id="B909525D3670FFE9FA86FEC830033A14" box="[1315,1351,333,359]" italics="true" pageId="29" pageNumber="30">Psl</emphasis>
|
||
.
|
||
<taxonomicName id="4C7DF5CC3670FFE9FAF3FECB30D83A14" baseAuthorityName="Edwards" baseAuthorityYear="1927" box="[1366,1436,334,359]" class="Insecta" family="Sciaridae" genus="Pseudolycoriella" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="jejuna">
|
||
<emphasis id="B909525D3670FFE9FAF3FECB30D83A14" box="[1366,1436,334,359]" italics="true" pageId="29" pageNumber="30">jejuna</emphasis>
|
||
</taxonomicName>
|
||
complex derivate from an ancestor that bears two spines at the gonostylus, like the extant species of the
|
||
<emphasis id="B909525D3670FFE9FABCFEF430793AF8" box="[1305,1341,369,395]" italics="true" pageId="29" pageNumber="30">Psl</emphasis>
|
||
.
|
||
<taxonomicName id="4C7DF5CC3670FFE9FAEFFEF430D83AF8" authorityName="in Mohrig & Jaschhof" authorityYear="1999" box="[1354,1436,369,395]" class="Insecta" family="Sciaridae" genus="Pseudolycoriella" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="bispina">
|
||
<emphasis id="B909525D3670FFE9FAEFFEF430D83AF8" box="[1354,1436,369,395]" italics="true" pageId="29" pageNumber="30">bispina</emphasis>
|
||
</taxonomicName>
|
||
complex or the
|
||
<emphasis id="B909525D3670FFE9FEEBFE1034363ADC" box="[334,370,405,431]" italics="true" pageId="29" pageNumber="30">Psl</emphasis>
|
||
.
|
||
<taxonomicName id="4C7DF5CC3670FFE9FE26FE1034B83ADC" baseAuthorityName="Edwards" baseAuthorityYear="1927" box="[387,508,405,431]" class="Insecta" family="Sciaridae" genus="Pseudolycoriella" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="zealandica">
|
||
<emphasis id="B909525D3670FFE9FE26FE1034B83ADC" box="[387,508,405,431]" italics="true" pageId="29" pageNumber="30">zealandica</emphasis>
|
||
</taxonomicName>
|
||
complex. The second detected atavism occured in two
|
||
<typeStatus id="54C630ED3670FFE9FB24FE1031A83ADC" box="[1153,1260,405,431]" pageId="29" pageNumber="30" type="paratype">paratypes</typeStatus>
|
||
of
|
||
<emphasis id="B909525D3670FFE9FABDFE1030783ADC" box="[1304,1340,405,431]" italics="true" pageId="29" pageNumber="30">Psl</emphasis>
|
||
.
|
||
<taxonomicName id="4C7DF5CC3670FFE9FAE9FE1030D13ADC" authorityName="Köhler" authorityYear="2019" box="[1356,1429,405,431]" class="Insecta" family="Sciaridae" genus="Pseudolycoriella" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="whena">
|
||
<emphasis id="B909525D3670FFE9FAE9FE1030D13ADC" box="[1356,1429,405,431]" italics="true" pageId="29" pageNumber="30">whena</emphasis>
|
||
</taxonomicName>
|
||
, where an area of the teeth is present, although to a varying degree—one
|
||
<typeStatus id="54C630ED3670FFE9FC7CFE3C317D3AA0" box="[985,1081,441,467]" pageId="29" pageNumber="30" type="paratype">paratype</typeStatus>
|
||
shows only five tiny teeth, the other twelve.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |