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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.205.82728" ID-Pensoft-Pub="1314-2003-205-279" ID-Pensoft-UUID="991A7A52D708580E87DF59FCF285C77C" ModsDocID="1314-2003-205-279" checkinTime="1661230117676" checkinUser="pensoft" docAuthor="Tamayo-Cen, Ivan, Torke, Benjamin M., Lopez Contreras, Jose Enrique, Carnevali Fernandez-Concha, German, Ramirez Morillo, Ivon, Can Itza, Lilia Lorena &amp; Duno de Stefano, Rodrigo" docDate="2022" docId="AEBCCA5CA03655D2B77CF42870B7FB8E" docLanguage="en" docName="PhytoKeys 205: 279-298" docOrigin="PhytoKeys 205" docPubDate="2022-08-22" docSource="http://dx.doi.org/10.3897/phytokeys.205.82728" docTitle="Gretheria Duno &amp; Torke 2022, gen. nov." docType="treatment" docVersion="1" id="991A7A52D708580E87DF59FCF285C77C" lastPageNumber="279" masterDocId="991A7A52D708580E87DF59FCF285C77C" masterDocTitle="Revisiting the phylogeny and taxonomy of the Pithecellobium clade (Leguminosae, Caesalpinioideae) with new generic circumscriptions" masterLastPageNumber="298" masterPageNumber="279" pageNumber="279" updateTime="1661230418719" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Revisiting the phylogeny and taxonomy of the Pithecellobium clade (Leguminosae, Caesalpinioideae) with new generic circumscriptions</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tamayo-Cen, Ivan</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-6034-2940</mods:nameIdentifier>
<mods:affiliation>Herbarium CICY, Centro de Investigacion Cientifica de Yucatan, A. C. (CICY), Calle 43 No. 130, Colonia Chuburna de Hidalgo, CP 97200, Merida, Yucatan, Mexico</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Torke, Benjamin M.</mods:namePart>
<mods:affiliation>Institute of Systematic Botany, New York Botanical Garden, 2900 Southern Boulevard, Bronx, New York, 10458 - 5126 USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lopez Contreras, Jose Enrique</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-4994-6465</mods:nameIdentifier>
<mods:affiliation>Herbarium CICY, Centro de Investigacion Cientifica de Yucatan, A. C. (CICY), Calle 43 No. 130, Colonia Chuburna de Hidalgo, CP 97200, Merida, Yucatan, Mexico &amp; Centro de Investigacion de Ciencias Ambientales, Universidad Autonoma del Carmen, C. 56 num, 4 Esquina Avenida Concordia, Colonia Benito Juarez CP 24180, Ciudad del Carmen, Campeche, Mexico</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Carnevali Fernandez-Concha, German</mods:namePart>
<mods:affiliation>Herbarium CICY, Centro de Investigacion Cientifica de Yucatan, A. C. (CICY), Calle 43 No. 130, Colonia Chuburna de Hidalgo, CP 97200, Merida, Yucatan, Mexico</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ramirez Morillo, Ivon</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6288-7984</mods:nameIdentifier>
<mods:affiliation>Herbarium CICY, Centro de Investigacion Cientifica de Yucatan, A. C. (CICY), Calle 43 No. 130, Colonia Chuburna de Hidalgo, CP 97200, Merida, Yucatan, Mexico</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Can Itza, Lilia Lorena</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-6777-9109</mods:nameIdentifier>
<mods:affiliation>Herbarium CICY, Centro de Investigacion Cientifica de Yucatan, A. C. (CICY), Calle 43 No. 130, Colonia Chuburna de Hidalgo, CP 97200, Merida, Yucatan, Mexico</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Duno de Stefano, Rodrigo</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-1707-4121</mods:nameIdentifier>
<mods:affiliation>Herbarium CICY, Centro de Investigacion Cientifica de Yucatan, A. C. (CICY), Calle 43 No. 130, Colonia Chuburna de Hidalgo, CP 97200, Merida, Yucatan, Mexico</mods:affiliation>
<mods:nameIdentifier type="email">roduno@cicy.mx</mods:nameIdentifier>
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<mods:titleInfo>
<mods:title>PhytoKeys</mods:title>
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<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-08-22</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>205</mods:number>
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<mods:extent unit="page">
<mods:start>279</mods:start>
<mods:end>298</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/phytokeys.205.82728</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.205.82728</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-205-279</mods:identifier>
<mods:identifier type="Pensoft-UUID">991A7A52D708580E87DF59FCF285C77C</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:AEBCCA5CA03655D2B77CF42870B7FB8E" httpUri="http://treatment.plazi.org/id/AEBCCA5CA03655D2B77CF42870B7FB8E" lastPageNumber="279" pageId="0" pageNumber="279">
<subSubSection pageId="0" pageNumber="279" type="nomenclature">
<paragraph pageId="0" pageNumber="279">
<taxonomicName LSID="AEBCCA5C-A036-55D2-B77C-F42870B7FB8E" authority="Duno &amp; Torke" authorityName="Duno &amp; Torke" authorityYear="2022" family="Fabaceae" genus="Gretheria" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="Gretheria" order="Fabales" pageId="0" pageNumber="279" rank="genus" status="gen. nov.">Gretheria Duno &amp; Torke</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="279">gen. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="279" type="diagnosis">
<paragraph pageId="0" pageNumber="279">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="279">
Similar to
<taxonomicName authorityName="J.K.Small" authorityYear="1901" class="Magnoliopsida" family="Fabaceae" genus="Havardia" kingdom="Plantae" lsidName="Havardia" order="Fabales" pageId="0" pageNumber="279" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="279">Havardia</emphasis>
</taxonomicName>
in arboreal or shrubby habit, vegetative branches arising both proleptically and sylleptically, leaves microphyllous, inflorescence arising on long shoots, pod flattened-compressed and seed plumply disciform to orbicular, but differing in the pinnae distally accrescent (vs. decrescent at each end in
<taxonomicName authorityName="J.K.Small" authorityYear="1901" class="Magnoliopsida" family="Fabaceae" genus="Havardia" kingdom="Plantae" lsidName="Havardia" order="Fabales" pageId="0" pageNumber="279" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="279">Havardia</emphasis>
</taxonomicName>
), the calyx longer and deeply campanulate (vs. shorter, and shallowly campanulate), and the corolla lobes erect-ascending at anthesis (vs. recurved).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="279" type="materials_examined">
<paragraph pageId="0" pageNumber="279">
<typeStatus>Type</typeStatus>
.
</paragraph>
<paragraph pageId="0" pageNumber="279">
<taxonomicName authorityName="Duno &amp; Torke" authorityYear="2022" baseAuthorityName="Duno &amp; Torke" baseAuthorityYear="2022" family="Fabaceae" genus="Gretheria" kingdom="Plantae" lsidName="Gretheria sonorae" order="Fabales" pageId="0" pageNumber="279" rank="species" species="sonorae">
<emphasis italics="true" pageId="0" pageNumber="279">Gretheria sonorae</emphasis>
</taxonomicName>
(S. Watson) Duno &amp; Torke.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="279" type="description">
<paragraph pageId="0" pageNumber="279">Description.</paragraph>
<paragraph pageId="0" pageNumber="279">
Xerophytic, microphyllous arborescent shrubs and small trees, 2-14 m tall, commonly armed with stout recurved, lignescent stipules on the trunk and at each node of long-shoots, indumented with minute whitish trichomes on new growth.
<emphasis bold="true" italics="true" pageId="0" pageNumber="279">Leaves</emphasis>
bipinnate, with 1-6 (13) pairs of pinnae; leaflets 10-31 per pinna; principal leaf axis typically 2-15 cm long, with the petiole 2-24 mm long, bearing a nectary at or below the mid-point of the petiole, the nectary sessile, shallow-cupular, thick-rimmed or plane and dimpled, pinnae axes sometimes with similar but smaller nectaries between 1-2 (3) distal-most pinna pairs and a minute one at the tip of some pinna-rachises; leaflets opposite, the blade oblong-elliptic to linear-oblong, subcordate at base, obtuse or shortly apiculate at apex; pilosulous or glabrous and marginally ciliolate; venation pinnate, immersed above, prominulous beneath, the mid-rib slightly displaced, giving rise on each side to 2-5 weak secondary veins expiring submarginally or faintly brochidodromous.
<emphasis bold="true" italics="true" pageId="0" pageNumber="279">Inflorescence</emphasis>
capituliform racemes arising from leaf axils of long shoots and coeval with or preceding the leaf and/or arising from brachyblasts; peduncles (1.3) 2 cm long; capitula 10-37-flowered, receptacle clavate, 1.5-2.5 mm diameter; bracts ovate, minute, less than 1 mm long, sessile, persistent into anthesis.
<emphasis bold="true" italics="true" pageId="0" pageNumber="279">Flowers</emphasis>
sessile, homomorphic, the perianth 5-merous; calyx deeply campanulate, glabrous, teeth deltate-ovate, ciliolate and sometimes distally puberulent or strigose; corolla subcylindrical, lobes erect, white-silky strigose dorsally; androecium 40-52-merous, 9-13 mm long, tube 3.6-5 mm long, nectar disc simple callosities or 5-lobed, 0.2 - 0.35 mm tall; ovary subsessile, slenderly ellipsoid, stipe 0.1-0.25 mm long; style about as long as stamens, the stigma poriform.
<emphasis bold="true" italics="true" pageId="0" pageNumber="279">Pods</emphasis>
1-3 per capitulum, oblong in profile, contracted at base into a pseudostipe
<normalizedToken originalValue="±">+/-</normalizedToken>
5-14 mm long and abruptly so at apex into an erect cusp 1.5-8 mm long; body straight or almost straight, 6.5-13
<normalizedToken originalValue="×">x</normalizedToken>
(1.3) 1.2-2.4 (2.6) cm, plano-compressed, the valves bluntly framed by longitudinally 3-ridged sutures
<normalizedToken originalValue="±">+/-</normalizedToken>
1.5-2 mm wide, stiff, somewhat brittle, brownish-green, externally veinless, glabrous, red-granular or both granular and puberulent outside, the cavity continuous; funicle dilated, sigmoid.
<emphasis bold="true" italics="true" pageId="0" pageNumber="279">Seeds</emphasis>
transverse, 8-13, plumply disciform to orbicular in outline, 9-12
<normalizedToken originalValue="×">x</normalizedToken>
7-10 mm, the pleurogram U-shaped.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="279" type="distribution">
<paragraph pageId="0" pageNumber="279">Geographic distribution.</paragraph>
<paragraph pageId="0" pageNumber="279">
<taxonomicName authorityName="Duno &amp; Torke" authorityYear="2022" family="Fabaceae" genus="Gretheria" kingdom="Plantae" lsidName="Gretheria" order="Fabales" pageId="0" pageNumber="279" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="279">Gretheria</emphasis>
</taxonomicName>
comprises two species in United States (Texas), Mexico and Central America (Honduras and Nicaragua).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="279" type="habitat">
<paragraph pageId="0" pageNumber="279">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="279">
<taxonomicName authorityName="Duno &amp; Torke" authorityYear="2022" family="Fabaceae" genus="Gretheria" kingdom="Plantae" lsidName="Gretheria" order="Fabales" pageId="0" pageNumber="279" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="279">Gretheria</emphasis>
</taxonomicName>
grows in tropical deciduous dry forests, thorn scrubs and brush-woodlands, between sea level and 400 m elevation, occasionally ascending to 700 m.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="279" type="etymology">
<paragraph pageId="0" pageNumber="279">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="279">
The generic name honours Rosaura Grether
<normalizedToken originalValue="González">Gonzalez</normalizedToken>
, an extraordinary and prolific Mexican botanist, with whom we had the pleasure of sharing her experience as a botanist and colleague. Her profound dedication and perseverant commitment to botanical research over decades has contributed importantly to our knowledge and understanding of
<taxonomicName authorityName="Duno &amp; Torke" authorityYear="2022" family="Leguminosae" genus="Gretheria" kingdom="Plantae" lsidName="" order="Fabales" pageId="0" pageNumber="279" rank="family">Leguminosae</taxonomicName>
, especially of the genus
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Mimosa" kingdom="Plantae" lsidName="Mimosa" order="Fabales" pageId="0" pageNumber="279" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="279">Mimosa</emphasis>
</taxonomicName>
L.
</paragraph>
</subSubSection>
</treatment>
</document>