401 lines
35 KiB
XML
401 lines
35 KiB
XML
<document id="77D33729F1EE99DA5C8ABCFF7EADD38E" ID-CLB-Dataset="29764" ID-DOI="10.11646/zootaxa.4442.2.5" ID-GBIF-Dataset="0fea0081-d093-4611-89c2-316f9baf6ca4" ID-ISSN="1175-5326" ID-Zenodo-Dep="1303125" ID-ZooBank="8EB2105E-3EAA-444A-A20E-819AD215B3C3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1530533589605" checkinUser="plazi" docAuthor="Khaustov, Alexander A. & Frolov, Andrey V." docDate="2018" docId="A1192263FE51BD3AC7E1038F426BFF1A" docLanguage="en" docName="zootaxa.4442.2.5.pdf" docOrigin="Zootaxa 4442 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nasutidania Khaustov & Frolov 2018, gen. nov." docType="treatment" docVersion="7" lastPageNumber="285" masterDocId="5D205A1BFE57BD32C776016A461FFFCF" masterDocTitle="New taxa of pygmephoroid mites (Acari: Pygmephoroidea: Pygmephoridae, Neopygmephoridae) phoretic on Enoplotrupes sharpi (Coleoptera: Geotrupidae) from Thailand" masterLastPageNumber="292" masterPageNumber="277" pageNumber="283" updateTime="1698552142957" updateUser="ExternalLinkService">
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<mods:mods id="02022A8DAA886D57557504FB29500EED" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="4F89178F74F03E2493742F9FDE7CCA3C">
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<mods:title id="3117DED9D1A455F31694E970AAFF7BF6">New taxa of pygmephoroid mites (Acari: Pygmephoroidea: Pygmephoridae, Neopygmephoridae) phoretic on Enoplotrupes sharpi (Coleoptera: Geotrupidae) from Thailand</mods:title>
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</mods:titleInfo>
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<mods:name id="A43BEF7CA3C32E081CBA1B081B31E15B" type="personal">
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<mods:roleTerm id="FE077A2D6B51635DD7CF7F3845B1900D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="3F0E3E1464872F9EA39BEC46EB8011C8">Khaustov, Alexander A.</mods:namePart>
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</mods:name>
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<mods:name id="DDF945554A438D85F638E536F4E6EE7C" type="personal">
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<mods:role id="8A47BF215F1204706843951A596DEE3D">
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<mods:roleTerm id="B04658E92273F2BF21FBF82382EB22E2">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="CA376A6ECF34D39E5BC76567B3B03AF8">Frolov, Andrey V.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="D4D43A17E27B0F6E4FDA51BC53A19D39">text</mods:typeOfResource>
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<mods:titleInfo id="49ED7FB77941EC7B2F6A40BC3A0EE93D">
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<mods:title id="ED60B7996FD7DBB0E48A50FD4B02762B">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="470A3AF7FF27D2364EB0CC1352CF440D">
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<mods:date id="CE0AC4739D4F49773F6A90E0C8DB11D8">2018</mods:date>
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<mods:detail id="48F40CC023DF45DEFFA750783EE6F9FC" type="pubDate">
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<mods:number id="D66E04F487CDFFEBD4B51AD96D9E3D1D">2018-07-02</mods:number>
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</mods:detail>
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<mods:detail id="F45C9E938B93E2C6965746D9660D99A6" type="volume">
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<mods:number id="4A4723744FB1FBD410982436321A342A">4442</mods:number>
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</mods:detail>
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<mods:detail id="B3A0B8928079B51B202945050BA037AE" type="issue">
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<mods:number id="4CB7FEFA1C4763CDB7B130298A68F4F9">2</mods:number>
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<mods:classification id="C2488892E50FD9162ED11038B7448C05">journal article</mods:classification>
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<mods:identifier id="8E991950882519F9722954F29EE3C017" type="CLB-Dataset">29764</mods:identifier>
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<mods:identifier id="FCF709420E47F606FDC74617E6E282E9" type="DOI">10.11646/zootaxa.4442.2.5</mods:identifier>
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<mods:identifier id="D83DD7A1838605905BB6FFCB61A87D14" type="GBIF-Dataset">0fea0081-d093-4611-89c2-316f9baf6ca4</mods:identifier>
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<mods:identifier id="61EC1D3D7713349B3044CFB9839B260F" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
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<treatment id="A1192263FE51BD3AC7E1038F426BFF1A" ID-DOI="http://doi.org/10.5281/zenodo.5949611" ID-GBIF-Taxon="146044082" ID-Zenodo-Dep="5949611" LSID="urn:lsid:plazi:treatment:A1192263FE51BD3AC7E1038F426BFF1A" httpUri="http://treatment.plazi.org/id/A1192263FE51BD3AC7E1038F426BFF1A" lastPageId="8" lastPageNumber="285" pageId="6" pageNumber="283">
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<subSubSection id="61AAC0FEFE51BD34C7E1038F47E8FD30" box="[151,503,741,767]" pageId="6" pageNumber="283" type="nomenclature">
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<paragraph id="290F9375FE51BD34C7E1038F47E8FD30" blockId="6.[151,503,741,767]" box="[151,503,741,767]" pageId="6" pageNumber="283">
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<heading id="72472419FE51BD34C7E1038F47E8FD30" bold="true" box="[151,503,741,767]" fontSize="11" level="1" pageId="6" pageNumber="283" reason="1">
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<emphasis id="1BC44F67FE51BD34C7E1038F47E8FD30" bold="true" box="[151,503,741,767]" pageId="6" pageNumber="283">
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Genus
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<taxonomicName id="EEB0E8F6FE51BD34C798038F479CFD30" authority="Khaustov & Frolov, 2018" authorityName="Khaustov & Frolov" authorityYear="2018" box="[238,387,741,767]" class="Arachnida" family="Pygmephoridae" genus="Nasutidania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="283" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="1BC44F67FE51BD34C798038F479CFD30" bold="true" box="[238,387,741,767]" italics="true" pageId="6" pageNumber="283">Nasutidania</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="00F7F21CFE51BD34C6FF038F47E8FD30" box="[393,503,741,767]" pageId="6" pageNumber="283" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="61AAC0FEFE51BD34C7E10246447FFC8F" box="[151,608,811,833]" pageId="6" pageNumber="283" type="materials_examined">
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<paragraph id="290F9375FE51BD34C7E10246447FFC8F" blockId="6.[151,608,811,833]" box="[151,608,811,833]" pageId="6" pageNumber="283">
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Type species:
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<taxonomicName id="EEB0E8F6FE51BD34C65002414416FC8E" authority="Khaustov & Frolov, 2018" authorityName="Khaustov & Frolov" authorityYear="2018" box="[294,521,811,833]" class="Arachnida" family="Pygmephoridae" genus="Nasutidania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="283" phylum="Arthropoda" rank="species" species="orientalis" status="sp. nov.">
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<emphasis id="1BC44F67FE51BD34C65002414416FC8E" box="[294,521,811,833]" italics="true" pageId="6" pageNumber="283">Nasutidania orientalis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="00F7F21CFE51BD34C5790241447FFC8F" box="[527,608,811,832]" pageId="6" pageNumber="283" rank="species">
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<emphasis id="1BC44F67FE51BD34C5790241447FFC8F" bold="true" box="[527,608,811,832]" pageId="6" pageNumber="283">sp. nov.</emphasis>
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</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="61AAC0FEFE51BD35C7E102074263F86B" lastPageId="7" lastPageNumber="284" pageId="6" pageNumber="283" type="diagnosis">
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<paragraph id="290F9375FE51BD34C7E102074737F8D7" blockId="6.[151,1437,877,2032]" pageId="6" pageNumber="283">
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<emphasis id="1BC44F67FE51BD34C7E102074715FC49" bold="true" box="[151,266,877,902]" pageId="6" pageNumber="283">Diagnosis</emphasis>
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. FEMALE. Body oval. Gnathosomal capsule elongate, beak-like, its length almost three times longer than width, dorsally with two pairs of cheliceral setae (
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<emphasis id="1BC44F67FE51BD34C46402FB4525FC65" box="[786,826,913,938]" italics="true" pageId="6" pageNumber="283">сha</emphasis>
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,
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<emphasis id="1BC44F67FE51BD34C43C02FB456DFC65" box="[842,882,913,938]" italics="true" pageId="6" pageNumber="283">сhb</emphasis>
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); postpalpal setae (
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<emphasis id="1BC44F67FE51BD34C32202F9426FFC65" box="[1108,1136,915,938]" italics="true" pageId="6" pageNumber="283">pp</emphasis>
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) present; palps very long, with two pairs of dorsal setae (
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<emphasis id="1BC44F67FE51BD34C68D02DF4438FC01" box="[507,551,949,974]" italics="true" pageId="6" pageNumber="283">dFe</emphasis>
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,
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<emphasis id="1BC44F67FE51BD34C54002DF447AFC01" box="[566,613,949,974]" italics="true" pageId="6" pageNumber="283">dGe</emphasis>
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). Palpal tibiotarsus without tibial claw and seta. Subcapitulum with one pair of setae
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<emphasis id="1BC44F67FE51BD34C64C02B14751FC3D" box="[314,334,987,1010]" italics="true" pageId="6" pageNumber="283">m</emphasis>
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and round pits in posterior part; palps ventrally without accessory setigenous structure and solenidion; ventrolaterally with unusual oval depressions (dep). Rostrum of subcapitulum with several lateral short spiniform projections in distal half. Pharyngeal pumps tripartite, striated, situated closely to each other; pharyngeal pump 1 small, butterfly-like, pharyngeal pump 2 large, subrectangular and about three times longer than oval pump 3. Posterior margin of prodorsum slightly covered by anterior part of tergite C. Prodorsum with two pairs of simple setae (
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<date id="5D0EB5B5FE51BD34C7A905E546EBFB64" box="[223,244,1167,1195]" pageId="6" pageNumber="283">
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<emphasis id="1BC44F67FE51BD34C7A905E546F4FB69" box="[223,235,1167,1190]" italics="true" pageId="6" pageNumber="283">v</emphasis>
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2
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</date>
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,
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<emphasis id="1BC44F67FE51BD34C67705E54707FB69" box="[257,280,1167,1190]" italics="true" pageId="6" pageNumber="283">sс</emphasis>
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2), pair of clavate trichobothria (
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<emphasis id="1BC44F67FE51BD34C5F605E54488FB69" box="[640,663,1167,1190]" italics="true" pageId="6" pageNumber="283">sс</emphasis>
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1) and small, round stigmata located just anteriorly to setae
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<date id="5D0EB5B5FE51BD34C25905E54355FB69" box="[1327,1354,1167,1195]" pageId="6" pageNumber="283">
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<emphasis id="1BC44F67FE51BD34C25905E54324FB69" box="[1327,1339,1167,1190]" italics="true" pageId="6" pageNumber="283">v</emphasis>
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2.
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</date>
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Dorsal idiosomal setae not modified. Tergite C with two pairs of setae (
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<emphasis id="1BC44F67FE51BD34C4F405DC4591FB02" box="[898,910,1206,1229]" italics="true" pageId="6" pageNumber="283">с</emphasis>
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1,
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<emphasis id="1BC44F67FE51BD34C4D105DC45ACFB02" box="[935,947,1206,1229]" italics="true" pageId="6" pageNumber="283">с</emphasis>
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2); tergite D with one pair of setae (
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<emphasis id="1BC44F67FE51BD34C22505DE437EFB02" box="[1363,1377,1204,1229]" italics="true" pageId="6" pageNumber="283">d</emphasis>
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) and round cupules
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<emphasis id="1BC44F67FE51BD34C63505B64746FB3C" box="[323,345,1244,1267]" italics="true" pageId="6" pageNumber="283">ia</emphasis>
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; tergite EF with two pairs of setae (
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<emphasis id="1BC44F67FE51BD34C47005B6450DFB3C" box="[774,786,1244,1267]" italics="true" pageId="6" pageNumber="283">e</emphasis>
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,
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<emphasis id="1BC44F67FE51BD34C45205B64533FB3B" box="[804,812,1244,1268]" italics="true" pageId="6" pageNumber="283">f</emphasis>
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); tergite H with two pairs of setae (
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<emphasis id="1BC44F67FE51BD34C3A105B142FAFB3B" box="[1239,1253,1243,1268]" italics="true" pageId="6" pageNumber="283">h</emphasis>
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1,
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<emphasis id="1BC44F67FE51BD34C38805B14313FB3B" box="[1278,1292,1243,1268]" italics="true" pageId="6" pageNumber="283">h</emphasis>
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2) and round cupules
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<emphasis id="1BC44F67FE51BD34C78304684714FAD4" box="[245,267,1282,1307]" italics="true" pageId="6" pageNumber="283">ih</emphasis>
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. Coxal fields I with two pairs of setae (1
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<emphasis id="1BC44F67FE51BD34C591046944EAFAD5" box="[743,757,1283,1306]" italics="true" pageId="6" pageNumber="283">a</emphasis>
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, 1
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<emphasis id="1BC44F67FE51BD34C4620468453DFAD4" box="[788,802,1282,1307]" italics="true" pageId="6" pageNumber="283">b</emphasis>
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); coxal fields II with two pairs of setae (2
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<emphasis id="1BC44F67FE51BD34C27D04694306FAD5" box="[1291,1305,1283,1306]" italics="true" pageId="6" pageNumber="283">a</emphasis>
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, 2
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<emphasis id="1BC44F67FE51BD34C2410468435AFAD4" box="[1335,1349,1282,1307]" italics="true" pageId="6" pageNumber="283">b</emphasis>
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); coxal fields III with three pairs of setae (3
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<emphasis id="1BC44F67FE51BD34C54A044D4455FAF1" box="[572,586,1319,1342]" italics="true" pageId="6" pageNumber="283">a</emphasis>
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, 3
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<emphasis id="1BC44F67FE51BD34C51F044C4468FAF0" box="[617,631,1318,1343]" italics="true" pageId="6" pageNumber="283">b</emphasis>
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, 3
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<emphasis id="1BC44F67FE51BD34C5E0044D44BDFAF1" box="[662,674,1319,1342]" italics="true" pageId="6" pageNumber="283">с</emphasis>
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); coxal fields IV with three pairs of setae (4
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<emphasis id="1BC44F67FE51BD34C3DC044D42A7FAF1" box="[1194,1208,1319,1342]" italics="true" pageId="6" pageNumber="283">a</emphasis>
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, 4
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<emphasis id="1BC44F67FE51BD34C3A1044C42FAFAF0" box="[1239,1253,1318,1343]" italics="true" pageId="6" pageNumber="283">b</emphasis>
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, 4
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<emphasis id="1BC44F67FE51BD34C272044D430FFAF1" box="[1284,1296,1319,1342]" italics="true" pageId="6" pageNumber="283">с</emphasis>
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). Pseudanal segment with three pairs of setae (
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<emphasis id="1BC44F67FE51BD34C56E0421442FFAAD" box="[536,560,1355,1378]" italics="true" pageId="6" pageNumber="283">ps</emphasis>
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1-3). Ap1 and ap2 well developed and joined together and with appr, apsej well developed and joined with appr; ap3 very weak, linear, ap4 long, reaching distinctly beyond bases of setae 3
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<emphasis id="1BC44F67FE51BD34C2F1041B438AFA45" box="[1415,1429,1393,1418]" italics="true" pageId="6" pageNumber="283">b</emphasis>
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. Apodemes 5 vestigial, located near the bases of trochanters IV. Secondary transverse apodeme absent. Posterior margin of posterior sternal plate tripartite. Ags bell-like, pgs triangular, median genital sclerite (mgs) present. Ventral idiosomal setae not modified. Legs I and II subequal in length. Tibiotarsus I slightly thickened, with distinct claw directed laterally, situated on pretarsus; ventrodistal part of pretarsus with short and thick projection (fused unguinal setae); seta
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<emphasis id="1BC44F67FE51BD34C6A8074F47F3F9F1" box="[478,492,1573,1598]" italics="true" pageId="6" pageNumber="283">d</emphasis>
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of femur I hook-like, seta
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<emphasis id="1BC44F67FE51BD34C458074F4525F9F1" box="[814,826,1573,1598]" italics="true" pageId="6" pageNumber="283">k</emphasis>
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long, pointed, seta-like. Claws on tarsus IV simple, distinctly thinner than padded claws on tarsi II and III. Tarsus IV not strongly elongate. Femora III–IV divided into basi- and telofemur. Leg setation: leg I; Tr 1 (
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<emphasis id="1BC44F67FE51BD34C5E3070444B4F94A" box="[661,683,1646,1669]" italics="true" pageId="6" pageNumber="283">v’</emphasis>
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), Fe 3 (
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<emphasis id="1BC44F67FE51BD34C4720707450DF949" box="[772,786,1645,1670]" italics="true" pageId="6" pageNumber="283">d</emphasis>
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,
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<emphasis id="1BC44F67FE51BD34C4690707452FF949" box="[799,816,1645,1670]" italics="true" pageId="6" pageNumber="283">l’</emphasis>
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||
,
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<emphasis id="1BC44F67FE51BD34C44807044545F94A" box="[830,858,1646,1669]" italics="true" pageId="6" pageNumber="283">v”</emphasis>
|
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), Ge 4 (
|
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<emphasis id="1BC44F67FE51BD34C4CE070745D6F949" box="[952,969,1645,1670]" italics="true" pageId="6" pageNumber="283">l’</emphasis>
|
||
,
|
||
<emphasis id="1BC44F67FE51BD34C4A1070745F1F949" box="[983,1006,1645,1670]" italics="true" pageId="6" pageNumber="283">l”</emphasis>
|
||
,
|
||
<emphasis id="1BC44F67FE51BD34C48A0704420EF94A" box="[1020,1041,1646,1669]" italics="true" pageId="6" pageNumber="283">v’</emphasis>
|
||
,
|
||
<emphasis id="1BC44F67FE51BD34C35607044224F94A" box="[1056,1083,1646,1669]" italics="true" pageId="6" pageNumber="283">v”</emphasis>
|
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), TiTa 16(4) (
|
||
<emphasis id="1BC44F67FE51BD34C3AC070742F7F949" box="[1242,1256,1645,1670]" italics="true" pageId="6" pageNumber="283">d</emphasis>
|
||
,
|
||
<emphasis id="1BC44F67FE51BD34C38007074318F949" box="[1270,1287,1645,1670]" italics="true" pageId="6" pageNumber="283">l’</emphasis>
|
||
,
|
||
<emphasis id="1BC44F67FE51BD34C26307074332F949" box="[1301,1325,1645,1670]" italics="true" pageId="6" pageNumber="283">l”</emphasis>
|
||
,
|
||
<emphasis id="1BC44F67FE51BD34C24D0704434FF94A" box="[1339,1360,1646,1669]" italics="true" pageId="6" pageNumber="283">v’</emphasis>
|
||
,
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||
<emphasis id="1BC44F67FE51BD34C22807044365F94A" box="[1374,1402,1646,1669]" italics="true" pageId="6" pageNumber="283">v”</emphasis>
|
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,
|
||
<emphasis id="1BC44F67FE51BD34C2FE0707438BF949" box="[1416,1428,1645,1670]" italics="true" pageId="6" pageNumber="283">k</emphasis>
|
||
,,
|
||
<emphasis id="1BC44F67FE51BD34C7D307F846DDF966" box="[165,194,1682,1705]" italics="true" pageId="6" pageNumber="283">tс’</emphasis>
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,
|
||
<emphasis id="1BC44F67FE51BD34C7A607F846EBF966" box="[208,244,1682,1705]" italics="true" pageId="6" pageNumber="283">tс”</emphasis>
|
||
,
|
||
<emphasis id="1BC44F67FE51BD34C67407F84700F966" box="[258,287,1682,1705]" italics="true" pageId="6" pageNumber="283">p”</emphasis>
|
||
,
|
||
<emphasis id="1BC44F67FE51BD34C65B07FB4759F966" box="[301,326,1681,1705]" italics="true" pageId="6" pageNumber="283">ft’</emphasis>
|
||
,
|
||
<emphasis id="1BC44F67FE51BD34C62207FB476CF966" box="[340,371,1681,1705]" italics="true" pageId="6" pageNumber="283">ft”</emphasis>
|
||
,
|
||
<emphasis id="1BC44F67FE51BD34C6F407F847B9F966" box="[386,422,1682,1705]" italics="true" pageId="6" pageNumber="283">pv’</emphasis>
|
||
,
|
||
<emphasis id="1BC44F67FE51BD34C6C507F847C2F966" box="[435,477,1682,1705]" italics="true" pageId="6" pageNumber="283">pv”</emphasis>
|
||
,
|
||
<emphasis id="1BC44F67FE51BD34C69D07FB4415F965" box="[491,522,1681,1706]" italics="true" pageId="6" pageNumber="283">pl’</emphasis>
|
||
,
|
||
<emphasis id="1BC44F67FE51BD34C56E07FB4422F965" box="[536,573,1681,1706]" italics="true" pageId="6" pageNumber="283">pl”</emphasis>
|
||
,
|
||
<emphasis id="1BC44F67FE51BD34C53D07F84449F966" box="[587,598,1682,1705]" italics="true" pageId="6" pageNumber="283">s</emphasis>
|
||
,
|
||
<emphasis id="1BC44F67FE51BD34C51207F84467F966" box="[612,632,1682,1705]" italics="true" pageId="6" pageNumber="283">ω</emphasis>
|
||
1,
|
||
<emphasis id="1BC44F67FE51BD34C5F807F844BDF966" box="[654,674,1682,1705]" italics="true" pageId="6" pageNumber="283">ω</emphasis>
|
||
2,
|
||
<emphasis id="1BC44F67FE51BD34C5C107F844D9F966" box="[695,710,1682,1705]" italics="true" pageId="6" pageNumber="283">φ</emphasis>
|
||
1,
|
||
<emphasis id="1BC44F67FE51BD34C5AA07F844F4F966" box="[732,747,1682,1705]" italics="true" pageId="6" pageNumber="283">φ</emphasis>
|
||
2); leg II: Tr 1 (
|
||
<emphasis id="1BC44F67FE51BD34C4E307F845B4F966" box="[917,939,1682,1705]" italics="true" pageId="6" pageNumber="283">v’</emphasis>
|
||
), Fe 3 (
|
||
<emphasis id="1BC44F67FE51BD34C37507FB422FF965" box="[1027,1072,1681,1706]" italics="true" pageId="6" pageNumber="283">d, l’</emphasis>
|
||
,
|
||
<emphasis id="1BC44F67FE51BD34C34807F84245F966" box="[1086,1114,1682,1705]" italics="true" pageId="6" pageNumber="283">v”</emphasis>
|
||
), Ge 3 (
|
||
<emphasis id="1BC44F67FE51BD34C3CE07FB42D6F965" box="[1208,1225,1681,1706]" italics="true" pageId="6" pageNumber="283">l’</emphasis>
|
||
,
|
||
<emphasis id="1BC44F67FE51BD34C3A107FB42F1F965" box="[1239,1262,1681,1706]" italics="true" pageId="6" pageNumber="283">l”</emphasis>
|
||
,
|
||
<emphasis id="1BC44F67FE51BD34C38A07F8430EF966" box="[1276,1297,1682,1705]" italics="true" pageId="6" pageNumber="283">v’</emphasis>
|
||
), Ti 4(1) (
|
||
<emphasis id="1BC44F67FE51BD34C2F107FB438AF965" box="[1415,1429,1681,1706]" italics="true" pageId="6" pageNumber="283">d</emphasis>
|
||
,
|
||
<emphasis id="1BC44F67FE51BD34C7E107DD46B7F91F" box="[151,168,1719,1744]" italics="true" pageId="6" pageNumber="283">l’</emphasis>
|
||
,
|
||
<emphasis id="1BC44F67FE51BD34C7C007D246D3F900" box="[182,204,1720,1743]" italics="true" pageId="6" pageNumber="283">v’</emphasis>
|
||
,
|
||
<emphasis id="1BC44F67FE51BD34C7AD07D246E9F900" box="[219,246,1720,1743]" italics="true" pageId="6" pageNumber="283">v”</emphasis>
|
||
,
|
||
<emphasis id="1BC44F67FE51BD34C67307D3470BF91F" box="[261,276,1721,1744]" italics="true" pageId="6" pageNumber="283">φ</emphasis>
|
||
), ta 6(1) (
|
||
<emphasis id="1BC44F67FE51BD34C6FE07D247BAF900" box="[392,421,1720,1743]" italics="true" pageId="6" pageNumber="283">tс’</emphasis>
|
||
,
|
||
<emphasis id="1BC44F67FE51BD34C6C207D247C8F900" box="[436,471,1720,1743]" italics="true" pageId="6" pageNumber="283">tс”</emphasis>
|
||
,
|
||
<emphasis id="1BC44F67FE51BD34C69007DD4415F91F" box="[486,522,1719,1744]" italics="true" pageId="6" pageNumber="283">pl”</emphasis>
|
||
,
|
||
<emphasis id="1BC44F67FE51BD34C56C07D24422F900" box="[538,573,1720,1743]" italics="true" pageId="6" pageNumber="283">pv’</emphasis>
|
||
,
|
||
<emphasis id="1BC44F67FE51BD34C53A07D2446AF900" box="[588,629,1720,1743]" italics="true" pageId="6" pageNumber="283">pv”</emphasis>
|
||
,
|
||
<emphasis id="1BC44F67FE51BD34C5F207D24484F900" box="[644,667,1720,1743]" italics="true" pageId="6" pageNumber="283">u’</emphasis>
|
||
,
|
||
<emphasis id="1BC44F67FE51BD34C5DC07D344A1F91F" box="[682,702,1721,1744]" italics="true" pageId="6" pageNumber="283">ω</emphasis>
|
||
); leg III: Tr 1 (
|
||
<emphasis id="1BC44F67FE51BD34C41A07D2459DF900" box="[876,898,1720,1743]" italics="true" pageId="6" pageNumber="283">v’</emphasis>
|
||
), Fe 2 (
|
||
<emphasis id="1BC44F67FE51BD34C4AD07DD45F6F91F" box="[987,1001,1719,1744]" italics="true" pageId="6" pageNumber="283">d</emphasis>
|
||
,
|
||
<emphasis id="1BC44F67FE51BD34C48E07D24211F900" box="[1016,1038,1720,1743]" italics="true" pageId="6" pageNumber="283">v’</emphasis>
|
||
), Ge 2 (
|
||
<emphasis id="1BC44F67FE51BD34C31A07DD4262F91F" box="[1132,1149,1719,1744]" italics="true" pageId="6" pageNumber="283">l’</emphasis>
|
||
,
|
||
<emphasis id="1BC44F67FE51BD34C3FA07D242BEF900" box="[1164,1185,1720,1743]" italics="true" pageId="6" pageNumber="283">v’</emphasis>
|
||
), Ti 4(1) (
|
||
<emphasis id="1BC44F67FE51BD34C26E07DD4339F91F" box="[1304,1318,1719,1744]" italics="true" pageId="6" pageNumber="283">d</emphasis>
|
||
,
|
||
<emphasis id="1BC44F67FE51BD34C24307DD4359F91F" box="[1333,1350,1719,1744]" italics="true" pageId="6" pageNumber="283">l’</emphasis>
|
||
,
|
||
<emphasis id="1BC44F67FE51BD34C22207D24375F900" box="[1364,1386,1720,1743]" italics="true" pageId="6" pageNumber="283">v’</emphasis>
|
||
,
|
||
<emphasis id="1BC44F67FE51BD34C20F07D2438BF900" box="[1401,1428,1720,1743]" italics="true" pageId="6" pageNumber="283">v”</emphasis>
|
||
,
|
||
<emphasis id="1BC44F67FE51BD34C7E107B746B9F93B" box="[151,166,1757,1780]" italics="true" pageId="6" pageNumber="283">φ</emphasis>
|
||
), Ta 6 (
|
||
<emphasis id="1BC44F67FE51BD34C78A07B74706F93B" box="[252,281,1757,1780]" italics="true" pageId="6" pageNumber="283">tс’</emphasis>
|
||
,
|
||
<emphasis id="1BC44F67FE51BD34C65107B74754F93B" box="[295,331,1757,1780]" italics="true" pageId="6" pageNumber="283">tс”</emphasis>
|
||
,
|
||
<emphasis id="1BC44F67FE51BD34C62F07B64762F93A" box="[345,381,1756,1781]" italics="true" pageId="6" pageNumber="283">pl”</emphasis>
|
||
,
|
||
<emphasis id="1BC44F67FE51BD34C6FD07B747B0F93B" box="[395,431,1757,1780]" italics="true" pageId="6" pageNumber="283">pv’</emphasis>
|
||
,
|
||
<emphasis id="1BC44F67FE51BD34C6CA07B747FAF93B" box="[444,485,1757,1780]" italics="true" pageId="6" pageNumber="283">pv”</emphasis>
|
||
,
|
||
<emphasis id="1BC44F67FE51BD34C68207B74414F93B" box="[500,523,1757,1780]" italics="true" pageId="6" pageNumber="283">u’</emphasis>
|
||
); leg IV: Tr 1 (
|
||
<emphasis id="1BC44F67FE51BD34C5C407B744D8F93B" box="[690,711,1757,1780]" italics="true" pageId="6" pageNumber="283">v’</emphasis>
|
||
), F 2 (
|
||
<emphasis id="1BC44F67FE51BD34C46407B6453FF93A" box="[786,800,1756,1781]" italics="true" pageId="6" pageNumber="283">d</emphasis>
|
||
,
|
||
<emphasis id="1BC44F67FE51BD34C45B07B7455CF93B" box="[813,835,1757,1780]" italics="true" pageId="6" pageNumber="283">v’</emphasis>
|
||
), Ge 1 (
|
||
<emphasis id="1BC44F67FE51BD34C4E807B745ABF93B" box="[926,948,1757,1780]" italics="true" pageId="6" pageNumber="283">v’</emphasis>
|
||
), Ti 4(1) (
|
||
<emphasis id="1BC44F67FE51BD34C35107B6422AF93A" box="[1063,1077,1756,1781]" italics="true" pageId="6" pageNumber="283">d</emphasis>
|
||
,
|
||
<emphasis id="1BC44F67FE51BD34C33507B6424BF93A" box="[1091,1108,1756,1781]" italics="true" pageId="6" pageNumber="283">l’</emphasis>
|
||
,
|
||
<emphasis id="1BC44F67FE51BD34C31707B74268F93B" box="[1121,1143,1757,1780]" italics="true" pageId="6" pageNumber="283">v’</emphasis>
|
||
,
|
||
<emphasis id="1BC44F67FE51BD34C3F207B742BFF93B" box="[1156,1184,1757,1780]" italics="true" pageId="6" pageNumber="283">v”</emphasis>
|
||
,
|
||
<emphasis id="1BC44F67FE51BD34C3D807B742A2F93B" box="[1198,1213,1757,1780]" italics="true" pageId="6" pageNumber="283">φ</emphasis>
|
||
), Ta 6 (
|
||
<emphasis id="1BC44F67FE51BD34C26507B7432FF93B" box="[1299,1328,1757,1780]" italics="true" pageId="6" pageNumber="283">tс’</emphasis>
|
||
,
|
||
<emphasis id="1BC44F67FE51BD34C24807B7437DF93B" box="[1342,1378,1757,1780]" italics="true" pageId="6" pageNumber="283">tс”</emphasis>
|
||
,
|
||
<emphasis id="1BC44F67FE51BD34C21907B6438BF93A" box="[1391,1428,1756,1781]" italics="true" pageId="6" pageNumber="283">pl”</emphasis>
|
||
,
|
||
<emphasis id="1BC44F67FE51BD34C7E1066A46B1F8D8" box="[151,174,1792,1815]" italics="true" pageId="6" pageNumber="283">u’</emphasis>
|
||
,
|
||
<emphasis id="1BC44F67FE51BD34C7CA066A46FFF8D8" box="[188,224,1792,1815]" italics="true" pageId="6" pageNumber="283">pv’</emphasis>
|
||
,
|
||
<emphasis id="1BC44F67FE51BD34C798066A4707F8D8" box="[238,280,1792,1815]" italics="true" pageId="6" pageNumber="283">pv”</emphasis>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="290F9375FE51BD34C7B1064E45D7F83F" blockId="6.[151,1437,877,2032]" pageId="6" pageNumber="283">
|
||
<emphasis id="1BC44F67FE51BD34C7B1064E47D8F8F2" bold="true" box="[199,455,1828,1853]" pageId="6" pageNumber="283">Differential diagnosis</emphasis>
|
||
. The new genus is most similar to
|
||
<taxonomicName id="EEB0E8F6FE51BD34C42D064F426FF8F2" authority="Mahunka" authorityName="Mahunka" authorityYear="1968" box="[859,1136,1828,1853]" class="Arachnida" family="Pyemotidae" genus="Xystrorostrum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="283" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1BC44F67FE51BD34C42D064F45E4F8F3" box="[859,1019,1829,1852]" italics="true" pageId="6" pageNumber="283">Xystrorostrum</emphasis>
|
||
Mahunka
|
||
</taxonomicName>
|
||
, 19681 by the presence of extremely long palps. The new genus differs from
|
||
<taxonomicName id="EEB0E8F6FE51BD34C59E06234598F8AF" authorityName="Mahunka" authorityYear="1968" box="[744,903,1865,1888]" class="Arachnida" family="Pyemotidae" genus="Xystrorostrum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="283" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1BC44F67FE51BD34C59E06234598F8AF" box="[744,903,1865,1888]" italics="true" pageId="6" pageNumber="283">Xystrorostrum</emphasis>
|
||
</taxonomicName>
|
||
by the tripartite posterior margin of posterior sternal plate (vs. entire in
|
||
<taxonomicName id="EEB0E8F6FE51BD34C6B906074471F84B" authorityName="Mahunka" authorityYear="1968" box="[463,622,1901,1924]" class="Arachnida" family="Pyemotidae" genus="Xystrorostrum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="283" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1BC44F67FE51BD34C6B906074471F84B" box="[463,622,1901,1924]" italics="true" pageId="6" pageNumber="283">Xystrorostrum</emphasis>
|
||
</taxonomicName>
|
||
), presence of a well developed median genital sclerite (vs. absent in
|
||
<taxonomicName id="EEB0E8F6FE51BD34C7E106FB4728F867" authorityName="Mahunka" authorityYear="1968" box="[151,311,1937,1960]" class="Arachnida" family="Pyemotidae" genus="Xystrorostrum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="283" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1BC44F67FE51BD34C7E106FB4728F867" box="[151,311,1937,1960]" italics="true" pageId="6" pageNumber="283">Xystrorostrum</emphasis>
|
||
</taxonomicName>
|
||
), absence of the secondary transverse apodeme (vs. present in
|
||
<taxonomicName id="EEB0E8F6FE51BD34C36606FB42AFF867" authorityName="Mahunka" authorityYear="1968" box="[1040,1200,1937,1960]" class="Arachnida" family="Pyemotidae" genus="Xystrorostrum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="283" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1BC44F67FE51BD34C36606FB42AFF867" box="[1040,1200,1937,1960]" italics="true" pageId="6" pageNumber="283">Xystrorostrum</emphasis>
|
||
</taxonomicName>
|
||
), presence of lateral cuticular projections on the rostrum of the subcapitulum (vs. absent in
|
||
<taxonomicName id="EEB0E8F6FE51BD34C4DD06DF4254F803" authorityName="Mahunka" authorityYear="1968" box="[939,1099,1973,1996]" class="Arachnida" family="Pyemotidae" genus="Xystrorostrum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="283" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1BC44F67FE51BD34C4DD06DF4254F803" box="[939,1099,1973,1996]" italics="true" pageId="6" pageNumber="283">Xystrorostrum</emphasis>
|
||
</taxonomicName>
|
||
), and absence of an accessory setigenous structure and palpal solenidion (vs. present in
|
||
<taxonomicName id="EEB0E8F6FE51BD34C46E06B345A7F83F" authorityName="Mahunka" authorityYear="1968" box="[792,952,2009,2032]" class="Arachnida" family="Pyemotidae" genus="Xystrorostrum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="283" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1BC44F67FE51BD34C46E06B345A7F83F" box="[792,952,2009,2032]" italics="true" pageId="6" pageNumber="283">Xystrorostrum</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<caption id="7DCFC3FDFE50BD35C7E106664297F890" httpUri="https://zenodo.org/record/1303135/files/figure.png" pageId="7" pageNumber="284" startId="7.[151,250,1804,1826]" targetBox="[151,1436,180,1782]" targetPageId="7">
|
||
<paragraph id="290F9375FE50BD35C7E106664297F890" blockId="7.[151,1436,1804,1887]" pageId="7" pageNumber="284">
|
||
<emphasis id="1BC44F67FE50BD35C7E106664713F8EE" bold="true" box="[151,268,1804,1826]" pageId="7" pageNumber="284">FIGURE 5</emphasis>
|
||
. DIC micrographs of
|
||
<taxonomicName id="EEB0E8F6FE50BD35C69D06664535F8ED" authority="Khaustov & Frolov, 2018" authorityName="Khaustov & Frolov" authorityYear="2018" box="[491,810,1804,1826]" class="Arachnida" family="Pygmephoridae" genus="Spatulaphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="7" pageNumber="284" phylum="Arthropoda" rank="species" species="enoplotruporum" status="sp. nov.">
|
||
<emphasis id="1BC44F67FE50BD35C69D06664535F8ED" box="[491,810,1804,1826]" italics="true" pageId="7" pageNumber="284">Spatulaphorus enoplotruporum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="00F7F21CFE50BD35C4460666459FF8EE" box="[816,896,1804,1825]" pageId="7" pageNumber="284" rank="species">
|
||
<emphasis id="1BC44F67FE50BD35C4460666459FF8EE" bold="true" box="[816,896,1804,1825]" pageId="7" pageNumber="284">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, female: A—general view dorsally, B—general view ventrally, C—prosoma in dorsal view, D—prosoma in ventral view, E—metapodosoma in dorsal view, F—metapodosoma in ventral view, G—opisthosoma in dorsal view, H—opisthosoma in ventral view, I—genital sclerites.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="290F9375FE50BD35C7B106E64263F86B" blockId="7.[151,1436,1932,2029]" box="[199,1148,1932,1957]" pageId="7" pageNumber="284">
|
||
<emphasis id="1BC44F67FE50BD35C7B106E64793F86A" bold="true" box="[199,396,1932,1957]" pageId="7" pageNumber="284">Species included</emphasis>
|
||
. The genus
|
||
<taxonomicName id="EEB0E8F6FE50BD35C56206E64482F86A" authority="Khaustov & Frolov, 2018" authorityName="Khaustov & Frolov" authorityYear="2018" box="[532,669,1932,1957]" class="Arachnida" family="Pygmephoridae" genus="Nasutidania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="7" pageNumber="290" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="1BC44F67FE50BD35C56206E64482F86A" box="[532,669,1932,1957]" italics="true" pageId="7" pageNumber="284">Nasutidania</emphasis>
|
||
</taxonomicName>
|
||
includes one species
|
||
<taxonomicName id="EEB0E8F6FE50BD35C4E606E74203F86A" authority="Khaustov & Frolov, 2018" authorityName="Khaustov & Frolov" authorityYear="2018" box="[912,1052,1932,1957]" class="Arachnida" family="Pygmephoridae" genus="Nasutidania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="7" pageNumber="284" phylum="Arthropoda" rank="species" species="orientalis" status="sp. nov.">
|
||
<emphasis id="1BC44F67FE50BD35C4E606E745BCF86B" box="[912,931,1933,1956]" italics="true" pageId="7" pageNumber="284">N</emphasis>
|
||
.
|
||
<emphasis id="1BC44F67FE50BD35C4C706E64203F86A" box="[945,1052,1932,1957]" italics="true" pageId="7" pageNumber="284">orientalis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="00F7F21CFE50BD35C35506E64263F86B" box="[1059,1148,1932,1957]" pageId="7" pageNumber="284" rank="species">
|
||
<emphasis id="1BC44F67FE50BD35C35506E64269F86A" bold="true" box="[1059,1142,1932,1957]" pageId="7" pageNumber="284">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="61AAC0FEFE50BD35C7B106DA47A2F822" pageId="7" pageNumber="284" type="distribution">
|
||
<paragraph id="290F9375FE50BD35C7B106DA47A2F822" blockId="7.[151,1436,1932,2029]" pageId="7" pageNumber="284">
|
||
<emphasis id="1BC44F67FE50BD35C7B106DA4418F806" bold="true" box="[199,519,1968,1993]" pageId="7" pageNumber="284">Distribution and habitats</emphasis>
|
||
.
|
||
<taxonomicName id="EEB0E8F6FE50BD35C56806DA453CF806" authorityName="Khaustov & Frolov" authorityYear="2018" box="[542,803,1968,1993]" class="Arachnida" family="Pygmephoridae" genus="Nasutidania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="7" pageNumber="284" phylum="Arthropoda" rank="species" species="orientalis">
|
||
<emphasis id="1BC44F67FE50BD35C56806DA453CF806" box="[542,803,1968,1993]" italics="true" pageId="7" pageNumber="284">Nasutidania orientalis</emphasis>
|
||
</taxonomicName>
|
||
known only from the scarab beetles of the genus
|
||
<taxonomicName id="EEB0E8F6FE50BD35C7E106BE4731F822" authorityName="Lucas" authorityYear="1869" box="[151,302,2004,2029]" class="Insecta" family="Geotrupidae" genus="Enoplotrupes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="284" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1BC44F67FE50BD35C7E106BE4731F822" box="[151,302,2004,2029]" italics="true" pageId="7" pageNumber="284">Enoplotrupes</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<collectingCountry id="51A7D3E5FE50BD35C62706BE47A6F822" box="[337,441,2004,2029]" name="Thailand" pageId="7" pageNumber="284">Thailand</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="61AAC0FEFE5FBD3AC7B101FD426BFF1A" pageId="8" pageNumber="285" type="etymology">
|
||
<paragraph id="290F9375FE5FBD3AC7B101FD426BFF1A" blockId="8.[151,1436,151,213]" pageId="8" pageNumber="285">
|
||
<emphasis id="1BC44F67FE5FBD3AC7B101FD4759FF7F" bold="true" box="[199,326,151,176]" pageId="8" pageNumber="285">Etymology</emphasis>
|
||
. The generic name is a combination of two words: Latin
|
||
<emphasis id="1BC44F67FE5FBD3AC49201F34226FF7F" box="[996,1081,153,176]" italics="true" pageId="8" pageNumber="285">nasutus</emphasis>
|
||
meaning
|
||
<emphasis id="1BC44F67FE5FBD3AC3DA01FD42F2FF7F" box="[1196,1261,151,176]" italics="true" pageId="8" pageNumber="285">nosed</emphasis>
|
||
and
|
||
<emphasis id="1BC44F67FE5FBD3AC25F01FD436FFF7F" box="[1321,1392,151,176]" italics="true" pageId="8" pageNumber="285">dania,</emphasis>
|
||
the second part of the generic name
|
||
<taxonomicName id="EEB0E8F6FE5FBD3AC57501D6449AFF1A" authorityName="Sasa" authorityYear="1961" box="[515,645,188,213]" class="Arachnida" family="Pygmephoridae" genus="Bakerdania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="8" pageNumber="285" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1BC44F67FE5FBD3AC57501D6449AFF1A" box="[515,645,188,213]" italics="true" pageId="8" pageNumber="285">Bakerdania</emphasis>
|
||
</taxonomicName>
|
||
, the most abundant neopygmephorid genus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |