476 lines
52 KiB
XML
476 lines
52 KiB
XML
<document ID-DOI="10.11646/zootaxa.5263.2.7" ID-ISSN="1175-5326" ID-Zenodo-Dep="7801139" ID-ZooBank="1D491798-9C50-4557-9906-F728B5B30F10" approvalRequired="1" approvalRequired_for_taxonomicNames="1" checkinTime="1680678503845" checkinUser="plazi" docAuthor="Özbek, Hasan Hüseyin" docDate="2023" docId="782353158610FF94FF36FC31631DE1D1" docLanguage="en" docName="zootaxa.5263.2.7.pdf" docOrigin="Zootaxa 5263 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5263.2.7" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Onchodellus circularis Özbek 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="291" masterDocId="841A2B6D8614FF92FFA1FFC76255E31C" masterDocTitle="The genus Onchodellus Berlese in Türkiye (Mesostigmata: Pachylaelapidae), with a new record and three new species" masterLastPageNumber="296" masterPageNumber="285" pageNumber="289" updateTime="1680678923019" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The genus Onchodellus Berlese in Türkiye (Mesostigmata: Pachylaelapidae), with a new record and three new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Özbek, Hasan Hüseyin</mods:namePart>
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<mods:affiliation>Faculty of Science and Arts, Erzincan Binali Yıldırım University, Erzincan, Türkiye</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-04-05</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>5263</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>2</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>285</mods:start>
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<mods:end>296</mods:end>
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</mods:extent>
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||
</mods:part>
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||
</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.11646/zootaxa.5263.2.7</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.11646/zootaxa.5263.2.7</mods:identifier>
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<mods:identifier type="ISSN">1175-5326</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7801139</mods:identifier>
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<mods:identifier type="ZooBank">1D491798-9C50-4557-9906-F728B5B30F10</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7801147" ID-Zenodo-Dep="7801147" LSID="urn:lsid:plazi:treatment:782353158610FF94FF36FC31631DE1D1" httpUri="http://treatment.plazi.org/id/782353158610FF94FF36FC31631DE1D1" lastPageId="6" lastPageNumber="291" pageId="4" pageNumber="289">
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||
<subSubSection box="[151,528,1014,1041]" pageId="4" pageNumber="289" type="nomenclature">
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||
<paragraph blockId="4.[151,764,1014,1077]" box="[151,528,1014,1041]" pageId="4" pageNumber="289">
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||
<heading bold="true" box="[151,528,1014,1041]" fontSize="11" level="1" pageId="4" pageNumber="289" reason="1">
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<emphasis bold="true" box="[151,528,1014,1041]" pageId="4" pageNumber="289">
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<taxonomicName authority="Özbek, 2023" authorityName="Özbek" authorityYear="2023" box="[151,428,1014,1041]" class="Arachnida" family="Pachylaelapidae" genus="Onchodellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="289" phylum="Arthropoda" rank="species" species="circularis" status="sp. nov.">
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<emphasis bold="true" box="[151,428,1014,1041]" italics="true" pageId="4" pageNumber="289">Onchodellus circularis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[435,528,1014,1040]" pageId="4" pageNumber="289" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[151,764,1050,1077]" pageId="4" pageNumber="289" type="description">
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<paragraph blockId="4.[151,764,1014,1077]" box="[151,764,1050,1077]" pageId="4" pageNumber="289">
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(
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<figureCitation box="[159,296,1050,1077]" captionStart="FIGURES 5–6" captionStartId="5.[152,270,1127,1151]" captionTargetBox="[156,1429,186,1102]" captionTargetId="figure-15@5.[151,1436,181,1103]" captionTargetPageId="5" captionText="FIGURES 5–6. Onchodellus circularis sp. nov., female. 5. Dorsal idiosoma; 6. Ventral idiosoma." figureDoi="http://doi.org/10.5281/zenodo.7801149" httpUri="https://zenodo.org/record/7801149/files/figure.png" pageId="4" pageNumber="289">Figures 5–6</figureCitation>
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,
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<figureCitation box="[308,505,1050,1076]" captionStart="FIGURES 9–23" captionStartId="8.[152,270,1894,1918]" captionTargetBox="[188,1394,395,1862]" captionTargetId="figure-138@8.[188,1398,381,1871]" captionTargetPageId="8" captionText="FIGURES 9–23. Onchodellus spp., female. 9–12. Epistome (O. karawaiewi, O. acrophilus, O.circularis, O. kosensis, respectively); 13–15. Spermathecal apparatus (O. acrophilus, O.circularis, O. kosensis, respectively); 16–19. Tarsus II (O. karawaiewi, O. acrophilus, O.circularis, respectively): O. acrophilus, male. 20. Tarsus II; 21. Femur, genu, tibia of Leg II: O.circularis, male. 22. Anterior part of holoventral shield; 23. Tarsus II." figureDoi="http://doi.org/10.5281/zenodo.7801155" httpUri="https://zenodo.org/record/7801155/files/figure.png" pageId="4" pageNumber="289">11, 14, 18, 22–23</figureCitation>
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,
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<figureCitation box="[516,546,1050,1076]" captionStart="FIGURES 24–29" captionStartId="9.[152,270,1542,1566]" captionTargetBox="[153,1433,181,1517]" captionTargetId="figure-15@9.[153,1433,181,1518]" captionTargetPageId="9" captionText="FIGURES 24–29. Onchodellus karawaiewi, 24. Chelicera of female; 25. of male. O. acrophilus, 26. Chelicera of female; 27. Chelicera of male. O.circularis, 28. Chelicera of male. O. kosensis, 29. Chelicera of female." figureDoi="http://doi.org/10.5281/zenodo.7801157" httpUri="https://zenodo.org/record/7801157/files/figure.png" pageId="4" pageNumber="289">28</figureCitation>
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,
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<figureCitation box="[558,631,1050,1077]" captionStart="FIGURES 30–35" captionStartId="10.[152,270,845,869]" captionTargetBox="[234,1353,181,821]" captionTargetId="figure-31@10.[234,1353,181,821]" captionTargetPageId="10" captionText="FIGURES 30–35. Onchodellus karawaiewi, male. 30.Tarsus II; 31. Femur, genu tibia of leg IV: O. acrophilus, male. 32.Tarsus II. 27: O.circularis, male. 33. Femur, genu, tibia, and tarsus of leg II; 34. Trochanter of leg IV; 35. Leg II." figureDoi="http://doi.org/10.5281/zenodo.7801159" httpUri="https://zenodo.org/record/7801159/files/figure.png" pageId="4" pageNumber="289">33–35</figureCitation>
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,
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<figureCitation box="[642,755,1050,1076]" captionStart="FIGURES 36–44" captionStartId="10.[152,270,1938,1962]" captionTargetBox="[230,1361,930,1914]" captionTargetId="figure-81@10.[227,1361,929,1914]" captionTargetPageId="10" captionText="FIGURES 36–44. Onchodellus karawaiewi, female. 36. Surface of ventral shield: O. acrophilus, female. 37. Surface of ventral shield: O. circularis sp. female. 38. Punctation of dorsal shield; 39. Surface of ventral shield: O. kosensis, female. 40. Surface of ventral shield:Onchodellus spp., female.41-44.Epistome (O.karawaiewi, O. acrophilus, O.circularis, O. kosensis, respectively)." figureDoi="http://doi.org/10.5281/zenodo.7801161" httpUri="https://zenodo.org/record/7801161/files/figure.png" pageId="4" pageNumber="289">38–39, 43</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="289" type="diagnosis">
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<paragraph blockId="4.[151,1437,1117,2007]" pageId="4" pageNumber="289">
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<emphasis bold="true" box="[151,266,1117,1143]" pageId="4" pageNumber="289">Diagnosis</emphasis>
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: Dorsal shield covered with distinct punctation; setae
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<emphasis box="[909,936,1117,1143]" italics="true" pageId="4" pageNumber="289">J5</emphasis>
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distinctly shorter than
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<emphasis box="[1211,1238,1117,1143]" italics="true" pageId="4" pageNumber="289">J4</emphasis>
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; soft integument of idiosoma with nine pairs of setae; anterior tips of peritremes extending to dorsal side of shield; one pair of posterolateral teat-like gland pores
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<emphasis box="[548,605,1189,1215]" italics="true" pageId="4" pageNumber="289">gdS4</emphasis>
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on shield. In male, cheliceral spermatodactyl leaf-like, its length half that of moveable digit; tarsus II with spur-like distal seta
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<emphasis box="[742,778,1225,1251]" italics="true" pageId="4" pageNumber="289">pl1</emphasis>
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and larger, well-developed opposite mediolateral process.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="5" lastPageNumber="290" pageId="4" pageNumber="289" type="materials_examined">
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<paragraph blockId="4.[151,1437,1117,2007]" pageId="4" pageNumber="289">
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<emphasis bold="true" box="[199,435,1261,1287]" pageId="4" pageNumber="289">Material examined:</emphasis>
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<materialsCitation collectingDate="2009-06-01" elevation="1455" latitude="40.65" location="Orumcek Forest" longLatPrecision="1161" longitude="39.016666" municipality="Kurtun Town" pageId="4" pageNumber="289" specimenCount="1" stateProvince="Gumushane" typeStatus="holotype">
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<typeStatus box="[442,549,1261,1287]" pageId="4" pageNumber="289">Holotype</typeStatus>
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, female,
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<collectingRegion box="[651,788,1261,1287]" country="Turkey" name="Gumushane" pageId="4" pageNumber="289">Gümüşhane</collectingRegion>
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,
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<collectingMunicipality box="[800,952,1261,1287]" pageId="4" pageNumber="289">Kürtün Town</collectingMunicipality>
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,
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<location LSID="urn:lsid:plazi:treatment:782353158610FF94FF36FC31631DE1D1:F555B4D88610FF96FC62FB296629E61B" box="[963,1148,1261,1289]" latitude="40.65" longLatPrecision="1161" longitude="39.016666" municipality="Kurtun Town" name="Orumcek Forest" pageId="4" pageNumber="289" stateProvince="Gumushane">Örümcek Forest</location>
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,
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<geoCoordinate box="[1159,1267,1261,1289]" degrees="40" direction="north" minutes="39" orientation="latitude" pageId="4" pageNumber="289" precision="925" value="40.65">40° 39′ N</geoCoordinate>
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,
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<geoCoordinate box="[1282,1389,1261,1289]" degrees="39" direction="east" minutes="01" orientation="longitude" pageId="4" pageNumber="289" precision="925" value="39.016666">39° 01′ E</geoCoordinate>
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, alt.
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<quantity box="[151,240,1297,1324]" metricMagnitude="3" metricUnit="m" metricValue="1.455" pageId="4" pageNumber="289" unit="m" value="1455.0">
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<elevation box="[151,240,1297,1324]" metricMagnitude="3" metricUnit="m" metricValue="1.455" pageId="4" pageNumber="289" unit="m" value="1455.0">1455 m</elevation>
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</quantity>
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,
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<date box="[253,394,1297,1323]" pageId="4" pageNumber="289" value="2009-06-01">
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<collectingDate box="[253,394,1297,1323]" pageId="4" pageNumber="289" value="2009-06-01">1 June 2009</collectingDate>
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</date>
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, litter under
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<taxonomicName box="[539,640,1297,1323]" class="Pinopsida" family="Pinaceae" genus="Picea" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="289" phylum="Tracheophyta" rank="species" species="undetermined">
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<emphasis box="[539,603,1298,1323]" italics="true" pageId="4" pageNumber="289">Picea</emphasis>
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sp
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</taxonomicName>
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</materialsCitation>
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||
.
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<materialsCitation box="[652,1150,1297,1323]" collectingDate="2009-06-01" elevation="1455" latitude="40.65" location="Orumcek Forest" longLatPrecision="1161" longitude="39.016666" municipality="Kurtun Town" pageId="4" pageNumber="289" specimenCount="1" specimenCount-male="1" stateProvince="Gumushane" typeStatus="paratype">
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<typeStatus box="[652,765,1297,1323]" pageId="4" pageNumber="289">Paratypes</typeStatus>
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:
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<specimenCount box="[777,883,1297,1323]" pageId="4" pageNumber="289" type="male">one male</specimenCount>
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, same data as holotype
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</materialsCitation>
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;
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<materialsCitation collectingDate="2013-11-24" elevation="1708" latitude="40.733334" location="Tahtamur Wold" longLatPrecision="1161" longitude="39.05" pageId="4" pageNumber="289" specimenCount="1" specimenCount-female="1" stateProvince="Gumushane" typeStatus="paratype">
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<specimenCount box="[1162,1290,1297,1323]" pageId="4" pageNumber="289" type="female">one female</specimenCount>
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,
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<collectingRegion country="Turkey" name="Gumushane" pageId="4" pageNumber="289">Gümüşhane Province</collectingRegion>
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,
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<location LSID="urn:lsid:plazi:treatment:782353158610FF94FF36FC31631DE1D1:F555B4D88610FF96FEA9FAF263ECE653" box="[264,441,1333,1359]" latitude="40.733334" longLatPrecision="1161" longitude="39.05" name="Tahtamur Wold" pageId="4" pageNumber="289" stateProvince="Gumushane">Tahtamur Wold</location>
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,
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<geoCoordinate box="[452,559,1333,1361]" degrees="40" direction="north" minutes="44" orientation="latitude" pageId="4" pageNumber="289" precision="925" value="40.733334">40° 44′ N</geoCoordinate>
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,
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<geoCoordinate box="[573,679,1333,1361]" degrees="39" direction="east" minutes="03" orientation="longitude" pageId="4" pageNumber="289" precision="925" value="39.05">39° 03′ E</geoCoordinate>
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, alt.
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<quantity box="[734,821,1333,1359]" metricMagnitude="3" metricUnit="m" metricValue="1.708" pageId="4" pageNumber="289" unit="m" value="1708.0">
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<elevation box="[734,821,1333,1359]" metricMagnitude="3" metricUnit="m" metricValue="1.708" pageId="4" pageNumber="289" unit="m" value="1708.0">1708 m</elevation>
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</quantity>
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,
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<date box="[832,1052,1333,1359]" pageId="4" pageNumber="289" value="2013-11-24">
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<collectingDate box="[832,1052,1333,1359]" pageId="4" pageNumber="289" value="2013-11-24">24 November 2013</collectingDate>
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</date>
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, in moss
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</materialsCitation>
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.
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</paragraph>
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<paragraph blockId="4.[151,1437,1117,2007]" box="[199,286,1369,1395]" pageId="4" pageNumber="289">
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<emphasis bold="true" box="[199,286,1369,1395]" pageId="4" pageNumber="289">Female</emphasis>
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</paragraph>
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<paragraph blockId="4.[151,1437,1117,2007]" pageId="4" pageNumber="289">
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<emphasis box="[199,289,1406,1431]" italics="true" pageId="4" pageNumber="289">Dorsum</emphasis>
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(
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<figureCitation box="[305,404,1405,1432]" captionStart="FIGURES 5–6" captionStartId="5.[152,270,1127,1151]" captionTargetBox="[156,1429,186,1102]" captionTargetId="figure-15@5.[151,1436,181,1103]" captionTargetPageId="5" captionText="FIGURES 5–6. Onchodellus circularis sp. nov., female. 5. Dorsal idiosoma; 6. Ventral idiosoma." figureDoi="http://doi.org/10.5281/zenodo.7801149" httpUri="https://zenodo.org/record/7801149/files/figure.png" pageId="4" pageNumber="289">Figure 5</figureCitation>
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). Dorsal shield well sclerotised, 530–560 long, 380–390 wide at widest point, length/width 1.35–1.47, oval, surface reticulate and punctate (
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<figureCitation box="[704,789,1441,1467]" captionStart="FIGURES 36–44" captionStartId="10.[152,270,1938,1962]" captionTargetBox="[230,1361,930,1914]" captionTargetId="figure-81@10.[227,1361,929,1914]" captionTargetPageId="10" captionText="FIGURES 36–44. Onchodellus karawaiewi, female. 36. Surface of ventral shield: O. acrophilus, female. 37. Surface of ventral shield: O. circularis sp. female. 38. Punctation of dorsal shield; 39. Surface of ventral shield: O. kosensis, female. 40. Surface of ventral shield:Onchodellus spp., female.41-44.Epistome (O.karawaiewi, O. acrophilus, O.circularis, O. kosensis, respectively)." figureDoi="http://doi.org/10.5281/zenodo.7801161" httpUri="https://zenodo.org/record/7801161/files/figure.png" pageId="4" pageNumber="289">Fig. 38</figureCitation>
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), with 30 pairs of smooth and pointed setae, clunal setae
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<emphasis box="[151,178,1477,1503]" italics="true" pageId="4" pageNumber="289">J5</emphasis>
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about 14 long,
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||
<emphasis box="[354,380,1477,1503]" italics="true" pageId="4" pageNumber="289">J4</emphasis>
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60–65, the ratio
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||
<emphasis box="[572,598,1477,1503]" italics="true" pageId="4" pageNumber="289">J4</emphasis>
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||
/
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||
<emphasis box="[606,632,1477,1503]" italics="true" pageId="4" pageNumber="289">J5</emphasis>
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||
4.28–4.33; a pair of posterolateral teat-like gland pores
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||
<emphasis box="[1264,1320,1477,1503]" italics="true" pageId="4" pageNumber="289">gdS4</emphasis>
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on shield,
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||
<emphasis box="[151,209,1513,1539]" italics="true" pageId="4" pageNumber="289">gdZ1</emphasis>
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subcircular and paraxial to setae
|
||
<emphasis box="[586,615,1513,1539]" italics="true" pageId="4" pageNumber="289">Z2</emphasis>
|
||
. Dorsal setae
|
||
<emphasis box="[776,797,1513,1539]" italics="true" pageId="4" pageNumber="289">j1</emphasis>
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||
about 20–24;
|
||
<emphasis box="[960,984,1513,1539]" italics="true" pageId="4" pageNumber="289">z1</emphasis>
|
||
about 14,
|
||
<emphasis box="[1104,1126,1513,1539]" italics="true" pageId="4" pageNumber="289">j4</emphasis>
|
||
,
|
||
<emphasis box="[1140,1162,1513,1539]" italics="true" pageId="4" pageNumber="289">j5</emphasis>
|
||
,
|
||
<emphasis box="[1177,1198,1513,1539]" italics="true" pageId="4" pageNumber="289">j6</emphasis>
|
||
,
|
||
<emphasis box="[1213,1238,1513,1539]" italics="true" pageId="4" pageNumber="289">z5</emphasis>
|
||
,
|
||
<emphasis box="[1252,1277,1513,1539]" italics="true" pageId="4" pageNumber="289">s6</emphasis>
|
||
,
|
||
<emphasis box="[1291,1316,1513,1539]" italics="true" pageId="4" pageNumber="289">r5</emphasis>
|
||
30–40,
|
||
<emphasis box="[1408,1436,1513,1539]" italics="true" pageId="4" pageNumber="289">S1</emphasis>
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||
and
|
||
<emphasis box="[199,226,1549,1575]" italics="true" pageId="4" pageNumber="289">S3</emphasis>
|
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20–30 long, other dorsal setae lengths between 50 and 68 long.
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||
</paragraph>
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<paragraph blockId="4.[151,1437,1117,2007]" pageId="4" pageNumber="289">
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||
<emphasis box="[199,271,1586,1611]" italics="true" pageId="4" pageNumber="289">Venter</emphasis>
|
||
(
|
||
<figureCitation box="[287,386,1585,1611]" captionStart="FIGURES 5–6" captionStartId="5.[152,270,1127,1151]" captionTargetBox="[156,1429,186,1102]" captionTargetId="figure-15@5.[151,1436,181,1103]" captionTargetPageId="5" captionText="FIGURES 5–6. Onchodellus circularis sp. nov., female. 5. Dorsal idiosoma; 6. Ventral idiosoma." figureDoi="http://doi.org/10.5281/zenodo.7801149" httpUri="https://zenodo.org/record/7801149/files/figure.png" pageId="4" pageNumber="289">Figure 6</figureCitation>
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). Sternal shield 170–180 long, 110–115 wide at level of coxae II, its surface decorated with reticulate pattern, bearing four pairs of pointed setae and three pairs of pores;
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<emphasis box="[1005,1038,1621,1647]" italics="true" pageId="4" pageNumber="289">st1</emphasis>
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55, slightly longer than
|
||
<emphasis box="[1305,1337,1621,1647]" italics="true" pageId="4" pageNumber="289">st2</emphasis>
|
||
(50);
|
||
<emphasis box="[1404,1437,1621,1647]" italics="true" pageId="4" pageNumber="289">st3</emphasis>
|
||
and
|
||
<emphasis box="[197,230,1657,1683]" italics="true" pageId="4" pageNumber="289">st4</emphasis>
|
||
equal in length (35). Genitiventral shield wider than long, length/width about 0.91, decorated with polygonal pattern, with two pairs of setae (
|
||
<figureCitation box="[522,608,1693,1719]" captionStart="FIGURES 36–44" captionStartId="10.[152,270,1938,1962]" captionTargetBox="[230,1361,930,1914]" captionTargetId="figure-81@10.[227,1361,929,1914]" captionTargetPageId="10" captionText="FIGURES 36–44. Onchodellus karawaiewi, female. 36. Surface of ventral shield: O. acrophilus, female. 37. Surface of ventral shield: O. circularis sp. female. 38. Punctation of dorsal shield; 39. Surface of ventral shield: O. kosensis, female. 40. Surface of ventral shield:Onchodellus spp., female.41-44.Epistome (O.karawaiewi, O. acrophilus, O.circularis, O. kosensis, respectively)." figureDoi="http://doi.org/10.5281/zenodo.7801161" httpUri="https://zenodo.org/record/7801161/files/figure.png" pageId="4" pageNumber="289">Fig. 39</figureCitation>
|
||
). Anal shield 70 long, 110–120 wide, length/width 0.58–0.59, triangular. Metapodal shields rounded. Lateral and opisthogastric soft integument with nine pairs of setae. Anterior tips of peritremes extend to dorsal side of idiosoma, between setae
|
||
<emphasis box="[825,849,1765,1791]" italics="true" pageId="4" pageNumber="289">z1</emphasis>
|
||
and
|
||
<emphasis box="[904,928,1765,1791]" italics="true" pageId="4" pageNumber="289">z2</emphasis>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="4.[151,1437,1117,2007]" box="[199,1436,1801,1827]" pageId="4" pageNumber="289">
|
||
<emphasis box="[199,473,1801,1827]" italics="true" pageId="4" pageNumber="289">Spermathecal apparatus</emphasis>
|
||
. Sperm access system associated with coxae III, tubes long, slender, and thin (
|
||
<figureCitation box="[1340,1423,1801,1827]" captionStart="FIGURES 9–23" captionStartId="8.[152,270,1894,1918]" captionTargetBox="[188,1394,395,1862]" captionTargetId="figure-138@8.[188,1398,381,1871]" captionTargetPageId="8" captionText="FIGURES 9–23. Onchodellus spp., female. 9–12. Epistome (O. karawaiewi, O. acrophilus, O.circularis, O. kosensis, respectively); 13–15. Spermathecal apparatus (O. acrophilus, O.circularis, O. kosensis, respectively); 16–19. Tarsus II (O. karawaiewi, O. acrophilus, O.circularis, respectively): O. acrophilus, male. 20. Tarsus II; 21. Femur, genu, tibia of Leg II: O.circularis, male. 22. Anterior part of holoventral shield; 23. Tarsus II." figureDoi="http://doi.org/10.5281/zenodo.7801155" httpUri="https://zenodo.org/record/7801155/files/figure.png" pageId="4" pageNumber="289">Fig. 14</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="4.[151,1437,1117,2007]" pageId="4" pageNumber="289">
|
||
<taxonomicName authorityName="Krefft" authorityYear="1871" box="[199,342,1837,1863]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="4" pageNumber="289" phylum="Nematoda" rank="genus">
|
||
<emphasis box="[199,342,1837,1863]" italics="true" pageId="4" pageNumber="289">Gnathosoma</emphasis>
|
||
</taxonomicName>
|
||
. Hypostome with seta
|
||
<emphasis box="[615,644,1837,1863]" italics="true" pageId="4" pageNumber="289">h3</emphasis>
|
||
longest,
|
||
<emphasis box="[753,781,1837,1863]" italics="true" pageId="4" pageNumber="289">h2</emphasis>
|
||
and palp coxal seta
|
||
<emphasis box="[1029,1055,1838,1863]" italics="true" pageId="4" pageNumber="289">pc</emphasis>
|
||
shortest and subequal in length; corniculi horn-like, deutosternal groove with six rows of denticles. Epistome with straight denticulate lateral margins and central projection with apical denticle (
|
||
<figureCitation box="[643,726,1909,1935]" captionStart="FIGURES 9–23" captionStartId="8.[152,270,1894,1918]" captionTargetBox="[188,1394,395,1862]" captionTargetId="figure-138@8.[188,1398,381,1871]" captionTargetPageId="8" captionText="FIGURES 9–23. Onchodellus spp., female. 9–12. Epistome (O. karawaiewi, O. acrophilus, O.circularis, O. kosensis, respectively); 13–15. Spermathecal apparatus (O. acrophilus, O.circularis, O. kosensis, respectively); 16–19. Tarsus II (O. karawaiewi, O. acrophilus, O.circularis, respectively): O. acrophilus, male. 20. Tarsus II; 21. Femur, genu, tibia of Leg II: O.circularis, male. 22. Anterior part of holoventral shield; 23. Tarsus II." figureDoi="http://doi.org/10.5281/zenodo.7801155" httpUri="https://zenodo.org/record/7801155/files/figure.png" pageId="4" pageNumber="289">Fig. 11</figureCitation>
|
||
), and its surface decorated with lines and punctation (
|
||
<figureCitation box="[1338,1424,1909,1935]" captionStart="FIGURES 36–44" captionStartId="10.[152,270,1938,1962]" captionTargetBox="[230,1361,930,1914]" captionTargetId="figure-81@10.[227,1361,929,1914]" captionTargetPageId="10" captionText="FIGURES 36–44. Onchodellus karawaiewi, female. 36. Surface of ventral shield: O. acrophilus, female. 37. Surface of ventral shield: O. circularis sp. female. 38. Punctation of dorsal shield; 39. Surface of ventral shield: O. kosensis, female. 40. Surface of ventral shield:Onchodellus spp., female.41-44.Epistome (O.karawaiewi, O. acrophilus, O.circularis, O. kosensis, respectively)." figureDoi="http://doi.org/10.5281/zenodo.7801161" httpUri="https://zenodo.org/record/7801161/files/figure.png" pageId="4" pageNumber="289">Fig. 43</figureCitation>
|
||
). Movable digit of chelicera 68 long, fixed digit about 70 long (to dorsal seta). Arthrodial brush short.
|
||
</paragraph>
|
||
<paragraph blockId="4.[151,1437,1117,2007]" box="[199,1387,1981,2007]" pageId="4" pageNumber="289">
|
||
<emphasis box="[199,252,1982,2007]" italics="true" pageId="4" pageNumber="289">Legs</emphasis>
|
||
. Chaetotaxy normal for the genus (
|
||
<bibRefCitation author="Masan, P." box="[648,799,1981,2007]" pageId="4" pageNumber="289" refId="ref7259" refString="Masan, P. (2007) A Review of the family Pachylaelapidae in Slovakia, with Systematics and Ecology of European Species (Acari: Mesostigmata: Eviphidoidea). Institute of Zoology, Slovak Academy of Sciences, Bratislava, 247 pp." type="book" year="2007">Mašán, 2007</bibRefCitation>
|
||
). Tarsus II with a thicker spur-like seta
|
||
<emphasis box="[1240,1275,1981,2007]" italics="true" pageId="4" pageNumber="289">pl1</emphasis>
|
||
(
|
||
<figureCitation box="[1291,1374,1981,2007]" captionStart="FIGURES 9–23" captionStartId="8.[152,270,1894,1918]" captionTargetBox="[188,1394,395,1862]" captionTargetId="figure-138@8.[188,1398,381,1871]" captionTargetPageId="8" captionText="FIGURES 9–23. Onchodellus spp., female. 9–12. Epistome (O. karawaiewi, O. acrophilus, O.circularis, O. kosensis, respectively); 13–15. Spermathecal apparatus (O. acrophilus, O.circularis, O. kosensis, respectively); 16–19. Tarsus II (O. karawaiewi, O. acrophilus, O.circularis, respectively): O. acrophilus, male. 20. Tarsus II; 21. Femur, genu, tibia of Leg II: O.circularis, male. 22. Anterior part of holoventral shield; 23. Tarsus II." figureDoi="http://doi.org/10.5281/zenodo.7801155" httpUri="https://zenodo.org/record/7801155/files/figure.png" pageId="4" pageNumber="289">Fig. 18</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7801149" ID-Zenodo-Dep="7801149" box="[152,1152,1127,1151]" httpUri="https://zenodo.org/record/7801149/files/figure.png" pageId="5" pageNumber="290" startId="5.[152,270,1127,1151]" targetBox="[156,1429,186,1102]" targetPageId="5">
|
||
<paragraph blockId="5.[152,1152,1127,1151]" box="[152,1152,1127,1151]" pageId="5" pageNumber="290">
|
||
<emphasis bold="true" box="[152,322,1127,1151]" pageId="5" pageNumber="290">FIGURES 5–6.</emphasis>
|
||
<taxonomicName authority="Özbek, 2023" authorityName="Özbek" authorityYear="2023" box="[329,561,1127,1151]" class="Arachnida" family="Pachylaelapidae" genus="Onchodellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="290" phylum="Arthropoda" rank="species" species="circularis" status="sp. nov.">
|
||
<emphasis box="[329,561,1127,1151]" italics="true" pageId="5" pageNumber="290">Onchodellus circularis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[567,648,1127,1151]" pageId="5" pageNumber="290">
|
||
<taxonomicNameLabel box="[567,648,1127,1151]" pageId="5" pageNumber="290" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, female. 5. Dorsal idiosoma; 6. Ventral idiosoma.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="5.[151,1437,1197,2016]" box="[199,260,1197,1223]" pageId="5" pageNumber="290">
|
||
<emphasis bold="true" box="[199,260,1197,1223]" pageId="5" pageNumber="290">Male</emphasis>
|
||
</paragraph>
|
||
<paragraph blockId="5.[151,1437,1197,2016]" box="[199,883,1233,1260]" pageId="5" pageNumber="290">
|
||
<emphasis box="[199,289,1234,1259]" italics="true" pageId="5" pageNumber="290">Dorsum</emphasis>
|
||
. Ornamentation and dorsal chaetotaxy as for female.
|
||
</paragraph>
|
||
<paragraph blockId="5.[151,1437,1197,2016]" pageId="5" pageNumber="290">
|
||
<emphasis box="[199,271,1270,1295]" italics="true" pageId="5" pageNumber="290">Venter</emphasis>
|
||
. Holoventral shield ornamented with the reticulate pattern and bearing 5 pairs of needle-like setae and 3 pairs of pores, and a pair well sclerotised ear-like process near
|
||
<emphasis box="[847,879,1305,1331]" italics="true" pageId="5" pageNumber="290">st1</emphasis>
|
||
(
|
||
<figureCitation box="[894,977,1305,1331]" captionStart="FIGURES 9–23" captionStartId="8.[152,270,1894,1918]" captionTargetBox="[188,1394,395,1862]" captionTargetId="figure-138@8.[188,1398,381,1871]" captionTargetPageId="8" captionText="FIGURES 9–23. Onchodellus spp., female. 9–12. Epistome (O. karawaiewi, O. acrophilus, O.circularis, O. kosensis, respectively); 13–15. Spermathecal apparatus (O. acrophilus, O.circularis, O. kosensis, respectively); 16–19. Tarsus II (O. karawaiewi, O. acrophilus, O.circularis, respectively): O. acrophilus, male. 20. Tarsus II; 21. Femur, genu, tibia of Leg II: O.circularis, male. 22. Anterior part of holoventral shield; 23. Tarsus II." figureDoi="http://doi.org/10.5281/zenodo.7801155" httpUri="https://zenodo.org/record/7801155/files/figure.png" pageId="5" pageNumber="290">Fig. 22</figureCitation>
|
||
). Opisthogastric region of the shield with three pairs of setae, three circum-anal setae.
|
||
</paragraph>
|
||
<paragraph blockId="5.[151,1437,1197,2016]" pageId="5" pageNumber="290">
|
||
<taxonomicName authorityName="Krefft" authorityYear="1871" box="[199,342,1377,1403]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="5" pageNumber="290" phylum="Nematoda" rank="genus">
|
||
<emphasis box="[199,342,1377,1403]" italics="true" pageId="5" pageNumber="290">Gnathosoma</emphasis>
|
||
</taxonomicName>
|
||
. Epistome as for female. Moveable digit of chelicerae 60 long, with a large subdistal tooth and a large terminal hook, spermatodactyl very short, leaf-shaped, 30 long, and its length half that of moveable digit (
|
||
<figureCitation captionStart="FIGURES 24–29" captionStartId="9.[152,270,1542,1566]" captionTargetBox="[153,1433,181,1517]" captionTargetId="figure-15@9.[153,1433,181,1518]" captionTargetPageId="9" captionText="FIGURES 24–29. Onchodellus karawaiewi, 24. Chelicera of female; 25. of male. O. acrophilus, 26. Chelicera of female; 27. Chelicera of male. O.circularis, 28. Chelicera of male. O. kosensis, 29. Chelicera of female." figureDoi="http://doi.org/10.5281/zenodo.7801157" httpUri="https://zenodo.org/record/7801157/files/figure.png" pageId="5" pageNumber="290">Fig. 28</figureCitation>
|
||
). Fixed digit with a subdistal tooth, a subdistal denticle and a large terminal hook, pilus dentilis clearly visible. Palptibiae without any processes. Arthrodial brush short.
|
||
</paragraph>
|
||
<paragraph blockId="5.[151,1437,1197,2016]" pageId="5" pageNumber="290">
|
||
<emphasis box="[199,252,1522,1547]" italics="true" pageId="5" pageNumber="290">Legs</emphasis>
|
||
. Tarsus II with a spur-like seta
|
||
<emphasis box="[603,642,1521,1547]" italics="true" pageId="5" pageNumber="290">p11</emphasis>
|
||
and strong opposite mediolateral apophysal spur; apophysal spur large and longer than
|
||
<emphasis box="[338,374,1557,1583]" italics="true" pageId="5" pageNumber="290">pl1</emphasis>
|
||
(
|
||
<figureCitation box="[390,479,1557,1583]" captionStart="FIGURES 9–23" captionStartId="8.[152,270,1894,1918]" captionTargetBox="[188,1394,395,1862]" captionTargetId="figure-138@8.[188,1398,381,1871]" captionTargetPageId="8" captionText="FIGURES 9–23. Onchodellus spp., female. 9–12. Epistome (O. karawaiewi, O. acrophilus, O.circularis, O. kosensis, respectively); 13–15. Spermathecal apparatus (O. acrophilus, O.circularis, O. kosensis, respectively); 16–19. Tarsus II (O. karawaiewi, O. acrophilus, O.circularis, respectively): O. acrophilus, male. 20. Tarsus II; 21. Femur, genu, tibia of Leg II: O.circularis, male. 22. Anterior part of holoventral shield; 23. Tarsus II." figureDoi="http://doi.org/10.5281/zenodo.7801155" httpUri="https://zenodo.org/record/7801155/files/figure.png" pageId="5" pageNumber="290">Figs 23</figureCitation>
|
||
,
|
||
<figureCitation box="[493,524,1557,1583]" captionStart="FIGURES 30–35" captionStartId="10.[152,270,845,869]" captionTargetBox="[234,1353,181,821]" captionTargetId="figure-31@10.[234,1353,181,821]" captionTargetPageId="10" captionText="FIGURES 30–35. Onchodellus karawaiewi, male. 30.Tarsus II; 31. Femur, genu tibia of leg IV: O. acrophilus, male. 32.Tarsus II. 27: O.circularis, male. 33. Femur, genu, tibia, and tarsus of leg II; 34. Trochanter of leg IV; 35. Leg II." figureDoi="http://doi.org/10.5281/zenodo.7801159" httpUri="https://zenodo.org/record/7801159/files/figure.png" pageId="5" pageNumber="290">33</figureCitation>
|
||
). Trochanter II with prominent median process; femur II with large, finger-like, spur-like process; genu and tibia each with small ventral process (
|
||
<figureCitation box="[876,1003,1593,1620]" captionStart="FIGURES 30–35" captionStartId="10.[152,270,845,869]" captionTargetBox="[234,1353,181,821]" captionTargetId="figure-31@10.[234,1353,181,821]" captionTargetPageId="10" captionText="FIGURES 30–35. Onchodellus karawaiewi, male. 30.Tarsus II; 31. Femur, genu tibia of leg IV: O. acrophilus, male. 32.Tarsus II. 27: O.circularis, male. 33. Femur, genu, tibia, and tarsus of leg II; 34. Trochanter of leg IV; 35. Leg II." figureDoi="http://doi.org/10.5281/zenodo.7801159" httpUri="https://zenodo.org/record/7801159/files/figure.png" pageId="5" pageNumber="290">Figs 33, 35</figureCitation>
|
||
). Trochanter IV with distal process and adjacent, well-developed, strong, and thicker setae (
|
||
<figureCitation box="[733,817,1629,1655]" captionStart="FIGURES 30–35" captionStartId="10.[152,270,845,869]" captionTargetBox="[234,1353,181,821]" captionTargetId="figure-31@10.[234,1353,181,821]" captionTargetPageId="10" captionText="FIGURES 30–35. Onchodellus karawaiewi, male. 30.Tarsus II; 31. Femur, genu tibia of leg IV: O. acrophilus, male. 32.Tarsus II. 27: O.circularis, male. 33. Femur, genu, tibia, and tarsus of leg II; 34. Trochanter of leg IV; 35. Leg II." figureDoi="http://doi.org/10.5281/zenodo.7801159" httpUri="https://zenodo.org/record/7801159/files/figure.png" pageId="5" pageNumber="290">Fig. 34</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="6" lastPageNumber="291" pageId="5" pageNumber="290" type="etymology">
|
||
<paragraph blockId="5.[151,1437,1197,2016]" box="[199,1437,1665,1692]" pageId="5" pageNumber="290">
|
||
<emphasis bold="true" box="[199,334,1665,1691]" pageId="5" pageNumber="290">Etymology.</emphasis>
|
||
The name of this species refers to the circular metapodal shield on the ventral idiososma of females.
|
||
</paragraph>
|
||
<paragraph blockId="5.[151,1437,1197,2016]" pageId="5" pageNumber="290">
|
||
<emphasis bold="true" box="[199,400,1701,1727]" pageId="5" pageNumber="290">Taxonomic notes</emphasis>
|
||
.
|
||
<taxonomicName authorityName="Özbek" authorityYear="2023" box="[415,671,1701,1727]" class="Arachnida" family="Pachylaelapidae" genus="Onchodellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="290" phylum="Arthropoda" rank="species" species="circularis">
|
||
<emphasis box="[415,671,1701,1727]" italics="true" pageId="5" pageNumber="290">Onchodellus circularis</emphasis>
|
||
</taxonomicName>
|
||
differs from most species of the genus by the presence of a specific leaf-like spermatodactyl and a well-developed mediolateral process on the tarsus II of the male, rounded metapodal shields, specifically shortened setae
|
||
<emphasis box="[554,582,1773,1799]" italics="true" pageId="5" pageNumber="290">S1</emphasis>
|
||
and
|
||
<emphasis box="[634,661,1773,1799]" italics="true" pageId="5" pageNumber="290">S3</emphasis>
|
||
, and a smaller number of setae on the soft integument of the idiosoma of the female, but can be compared with species of the
|
||
<taxonomicName baseAuthorityName="Hirschmann & Krauss" baseAuthorityYear="1965" box="[792,910,1809,1836]" class="Arachnida" family="Pachylaelapidae" genus="Onchodellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="290" phylum="Arthropoda" rank="species" species="falcifer">
|
||
<emphasis box="[792,812,1810,1836]" italics="true" pageId="5" pageNumber="290">O</emphasis>
|
||
.
|
||
<emphasis box="[829,910,1809,1835]" italics="true" pageId="5" pageNumber="290">falcifer</emphasis>
|
||
</taxonomicName>
|
||
group and the
|
||
<taxonomicName baseAuthorityName="Sellnick" baseAuthorityYear="1969" box="[1086,1237,1809,1836]" class="Arachnida" family="Pachylaelapidae" genus="Onchodellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="290" phylum="Arthropoda" rank="species" species="islandicus">
|
||
<emphasis box="[1086,1106,1810,1836]" italics="true" pageId="5" pageNumber="290">O</emphasis>
|
||
.
|
||
<emphasis box="[1123,1237,1809,1835]" italics="true" pageId="5" pageNumber="290">islandicus</emphasis>
|
||
</taxonomicName>
|
||
group as defined by
|
||
<bibRefCitation author="Masan, P." box="[189,351,1845,1871]" pageId="5" pageNumber="290" refId="ref7259" refString="Masan, P. (2007) A Review of the family Pachylaelapidae in Slovakia, with Systematics and Ecology of European Species (Acari: Mesostigmata: Eviphidoidea). Institute of Zoology, Slovak Academy of Sciences, Bratislava, 247 pp." type="book" year="2007">Mašán (2007)</bibRefCitation>
|
||
. These species groups share the features of a spermatodactyl that is shorter than the length of moveable digit and has 2–3 distal branches or is broadened like a leaf. This new species can be compared with the
|
||
<taxonomicName baseAuthorityName="Sellnick" baseAuthorityYear="1969" box="[151,299,1917,1944]" class="Arachnida" family="Pachylaelapidae" genus="Onchodellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="290" phylum="Arthropoda" rank="species" species="islandicus">
|
||
<emphasis box="[151,171,1918,1944]" italics="true" pageId="5" pageNumber="290">O</emphasis>
|
||
.
|
||
<emphasis box="[185,299,1917,1943]" italics="true" pageId="5" pageNumber="290">islandicus</emphasis>
|
||
</taxonomicName>
|
||
group, but there are important main differences in male and female specimens, as the anterior ends of the peritreme in
|
||
<taxonomicName authorityName="Özbek" authorityYear="2023" box="[331,472,1953,1980]" class="Arachnida" family="Pachylaelapidae" genus="Onchodellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="290" phylum="Arthropoda" rank="species" species="circularis">
|
||
<emphasis box="[331,351,1954,1980]" italics="true" pageId="5" pageNumber="290">O</emphasis>
|
||
.
|
||
<emphasis box="[364,472,1953,1979]" italics="true" pageId="5" pageNumber="290">circularis</emphasis>
|
||
</taxonomicName>
|
||
reach the dorsal surface of the shield, whereas in the species of the
|
||
<taxonomicName baseAuthorityName="Sellnick" baseAuthorityYear="1969" box="[1212,1359,1953,1980]" class="Arachnida" family="Pachylaelapidae" genus="Onchodellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="290" phylum="Arthropoda" rank="species" species="islandicus">
|
||
<emphasis box="[1212,1232,1954,1980]" italics="true" pageId="5" pageNumber="290">O</emphasis>
|
||
.
|
||
<emphasis box="[1245,1359,1953,1979]" italics="true" pageId="5" pageNumber="290">islandicus</emphasis>
|
||
</taxonomicName>
|
||
group, the peritreme is distinctly shortened and the anterior ends do not reach the dorsal surface of the shield.
|
||
</paragraph>
|
||
<paragraph blockId="6.[151,1437,151,717]" pageId="6" pageNumber="291">
|
||
It is also compared with species of the
|
||
<taxonomicName baseAuthorityName="Hirschmann & Krauss" baseAuthorityYear="1965" box="[663,782,151,177]" class="Arachnida" family="Pachylaelapidae" genus="Onchodellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="291" phylum="Arthropoda" rank="species" species="falcifer">
|
||
<emphasis box="[663,683,151,177]" italics="true" pageId="6" pageNumber="291">O</emphasis>
|
||
.
|
||
<emphasis box="[701,782,151,177]" italics="true" pageId="6" pageNumber="291">falcifer</emphasis>
|
||
</taxonomicName>
|
||
group, especially with male specimens. The male of
|
||
<taxonomicName authorityName="Özbek" authorityYear="2023" class="Arachnida" family="Pachylaelapidae" genus="Onchodellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="291" phylum="Arthropoda" rank="species" species="circularis">
|
||
<emphasis box="[1409,1429,151,177]" italics="true" pageId="6" pageNumber="291">O</emphasis>
|
||
.
|
||
<emphasis box="[151,259,187,213]" italics="true" pageId="6" pageNumber="291">circularis</emphasis>
|
||
</taxonomicName>
|
||
is clearly distinguished from all others by the presence of a larger, well-developed mediolateral process on the tarsus II, which is not present in males of this group. In addition, it differs from
|
||
<taxonomicName authority="(Hirschmann & Krauss, 1965)" baseAuthorityName="Hirschmann & Krauss" baseAuthorityYear="1965" class="Arachnida" family="Pachylaelapidae" genus="Onchodellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="291" phylum="Arthropoda" rank="species" species="falcifer">
|
||
<emphasis box="[1137,1157,223,249]" italics="true" pageId="6" pageNumber="291">O</emphasis>
|
||
.
|
||
<emphasis box="[1172,1253,223,249]" italics="true" pageId="6" pageNumber="291">falcifer</emphasis>
|
||
(
|
||
<bibRefCitation author="Hirschmann, W. & Krauss, W." pageId="6" pageNumber="291" pagination="1 - 5" refId="ref7059" refString="Hirschmann, W. & Krauss, W. (1965) Gangsystematik der Parasitiformes. Teil 8. Gamasiden. Bestimmungstafeln von 55 Pachylaelaps - Arten. Acarologie, Schriftenreihe fur vergleichende Milbenkunde, 7, 1 - 5." type="journal article" year="1965">Hirschmann & Krauss, 1965</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authority="(Moraza & Pena, 2005)" baseAuthorityName="Moraza & Pena" baseAuthorityYear="2005" box="[366,807,258,285]" class="Arachnida" family="Pachylaelapidae" genus="Onchodellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="291" phylum="Arthropoda" rank="species" species="canariensis">
|
||
<emphasis box="[366,386,259,285]" italics="true" pageId="6" pageNumber="291">O</emphasis>
|
||
.
|
||
<emphasis box="[402,531,259,284]" italics="true" pageId="6" pageNumber="291">canariensis</emphasis>
|
||
(Moraza & Peña, 2005)
|
||
</taxonomicName>
|
||
in having a non-branching, leaf-shaped spermatodactyl (two branches in
|
||
<taxonomicName baseAuthorityName="Moraza & Pena" baseAuthorityYear="2005" box="[345,508,295,321]" class="Arachnida" family="Pachylaelapidae" genus="Onchodellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="291" phylum="Arthropoda" rank="species" species="canariensis">
|
||
<emphasis box="[345,365,295,321]" italics="true" pageId="6" pageNumber="291">O</emphasis>
|
||
.
|
||
<emphasis box="[379,508,295,320]" italics="true" pageId="6" pageNumber="291">canariensis</emphasis>
|
||
</taxonomicName>
|
||
, three branches in
|
||
<taxonomicName baseAuthorityName="Hirschmann & Krauss" baseAuthorityYear="1965" box="[720,835,295,321]" class="Arachnida" family="Pachylaelapidae" genus="Onchodellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="291" phylum="Arthropoda" rank="species" species="falcifer">
|
||
<emphasis box="[720,740,295,321]" italics="true" pageId="6" pageNumber="291">O</emphasis>
|
||
.
|
||
<emphasis box="[754,835,295,321]" italics="true" pageId="6" pageNumber="291">falcifer</emphasis>
|
||
</taxonomicName>
|
||
) and differs from
|
||
<taxonomicName authority="(Koroleva, 1977)" baseAuthorityName="Koroleva" baseAuthorityYear="1977" box="[1040,1389,295,321]" class="Arachnida" family="Pachylaelapidae" genus="Onchodellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="291" phylum="Arthropoda" rank="species" species="nidicolens">
|
||
<emphasis box="[1040,1060,295,321]" italics="true" pageId="6" pageNumber="291">O</emphasis>
|
||
.
|
||
<emphasis box="[1074,1189,295,321]" italics="true" pageId="6" pageNumber="291">nidicolens</emphasis>
|
||
(
|
||
<bibRefCitation author="Koroleva, E. V." box="[1204,1380,295,321]" pageId="6" pageNumber="291" pagination="411 - 483" refId="ref7140" refString="Koroleva, E. V. (1977) Family Pachylaelaptidae Vitzthum, 1931. In: Ghilyarov, M. S. & Bregetova, N. G. (Eds.), Key to the Soil Inhabiting Mites. Mesostigmata, Nauka, Leningrad, pp. 411 - 483." type="book chapter" year="1977">Koroleva, 1977</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and O.
|
||
<emphasis box="[185,327,331,357]" italics="true" pageId="6" pageNumber="291">brevicrinitus</emphasis>
|
||
(
|
||
<bibRefCitation author="Hirschmann, W. & Krauss, W." box="[341,663,330,357]" pageId="6" pageNumber="291" pagination="1 - 5" refId="ref7059" refString="Hirschmann, W. & Krauss, W. (1965) Gangsystematik der Parasitiformes. Teil 8. Gamasiden. Bestimmungstafeln von 55 Pachylaelaps - Arten. Acarologie, Schriftenreihe fur vergleichende Milbenkunde, 7, 1 - 5." type="journal article" year="1965">Hirschmann & Krauss, 1965</bibRefCitation>
|
||
) by the specific shape of the spermatodactyl, only one process on the genu II and tibia II, and a distinct process on trochanter II. The female specimens are easily distinguished from the others in the genus because they have some specific character states, such as rounded metapodal shields, nine pairs of setae on the soft integument of the idiosoma, punctate surface of the dorsal shield. Ýt can be compared with
|
||
<taxonomicName authority="(Koroleva, 1977)" baseAuthorityName="Koroleva" baseAuthorityYear="1977" class="Arachnida" family="Pachylaelapidae" genus="Onchodellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="291" phylum="Arthropoda" rank="species" species="calcaratus">
|
||
<emphasis box="[1409,1429,439,465]" italics="true" pageId="6" pageNumber="291">O</emphasis>
|
||
.
|
||
<emphasis box="[151,269,475,501]" italics="true" pageId="6" pageNumber="291">calcaratus</emphasis>
|
||
(
|
||
<bibRefCitation author="Koroleva, E. V." box="[284,461,475,501]" pageId="6" pageNumber="291" pagination="411 - 483" refId="ref7140" refString="Koroleva, E. V. (1977) Family Pachylaelaptidae Vitzthum, 1931. In: Ghilyarov, M. S. & Bregetova, N. G. (Eds.), Key to the Soil Inhabiting Mites. Mesostigmata, Nauka, Leningrad, pp. 411 - 483." type="book chapter" year="1977">Koroleva, 1977</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
in that setae
|
||
<emphasis box="[617,644,475,501]" italics="true" pageId="6" pageNumber="291">J4</emphasis>
|
||
are longer than 50 and setae
|
||
<emphasis box="[971,998,475,501]" italics="true" pageId="6" pageNumber="291">J5</emphasis>
|
||
are shorter than 16, but they differ in a different
|
||
<emphasis box="[255,316,511,537]" italics="true" pageId="6" pageNumber="291">J4/J5</emphasis>
|
||
ratio (less than
|
||
<quantity box="[498,542,510,537]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="6" pageNumber="291" unit="in" value="5.0">5 in</quantity>
|
||
<taxonomicName authorityName="Özbek" authorityYear="2023" box="[549,693,511,537]" class="Arachnida" family="Pachylaelapidae" genus="Onchodellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="291" phylum="Arthropoda" rank="species" species="circularis">
|
||
<emphasis box="[549,569,511,537]" italics="true" pageId="6" pageNumber="291">O</emphasis>
|
||
.
|
||
<emphasis box="[585,693,511,537]" italics="true" pageId="6" pageNumber="291">circularis</emphasis>
|
||
</taxonomicName>
|
||
, more than
|
||
<quantity box="[829,873,510,537]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="6" pageNumber="291" unit="in" value="5.0">5 in</quantity>
|
||
<taxonomicName baseAuthorityName="Koroleva" baseAuthorityYear="1977" box="[880,1033,511,537]" class="Arachnida" family="Pachylaelapidae" genus="Onchodellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="291" phylum="Arthropoda" rank="species" species="calcaratus">
|
||
<emphasis box="[880,900,511,537]" italics="true" pageId="6" pageNumber="291">O</emphasis>
|
||
.
|
||
<emphasis box="[915,1033,511,537]" italics="true" pageId="6" pageNumber="291">calcaratus</emphasis>
|
||
</taxonomicName>
|
||
), the shape of the metapodal shield (not rounded in
|
||
<taxonomicName baseAuthorityName="Koroleva" baseAuthorityYear="1977" box="[333,486,547,573]" class="Arachnida" family="Pachylaelapidae" genus="Onchodellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="291" phylum="Arthropoda" rank="species" species="calcaratus">
|
||
<emphasis box="[333,353,547,573]" italics="true" pageId="6" pageNumber="291">O</emphasis>
|
||
.
|
||
<emphasis box="[368,486,547,573]" italics="true" pageId="6" pageNumber="291">calcaratus</emphasis>
|
||
</taxonomicName>
|
||
), the surface of the dorsal shield with distinct punctation, and diagnostic features in the males, such as the mediolateral process, which is shorter or the same length
|
||
<emphasis box="[1055,1091,583,609]" italics="true" pageId="6" pageNumber="291">
|
||
pl
|
||
<quantity box="[1080,1091,583,609]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="6" pageNumber="291" unit="in" value="1.0">1</quantity>
|
||
</emphasis>
|
||
in
|
||
<taxonomicName baseAuthorityName="Koroleva" baseAuthorityYear="1977" box="[1128,1281,583,609]" class="Arachnida" family="Pachylaelapidae" genus="Onchodellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="291" phylum="Arthropoda" rank="species" species="calcaratus">
|
||
<emphasis box="[1128,1148,583,609]" italics="true" pageId="6" pageNumber="291">O</emphasis>
|
||
.
|
||
<emphasis box="[1163,1281,583,609]" italics="true" pageId="6" pageNumber="291">calcaratus</emphasis>
|
||
</taxonomicName>
|
||
, significantly larger and longer than
|
||
<emphasis box="[405,441,619,645]" italics="true" pageId="6" pageNumber="291">
|
||
pl
|
||
<quantity box="[430,441,619,645]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="6" pageNumber="291" unit="in" value="1.0">1</quantity>
|
||
</emphasis>
|
||
in
|
||
<taxonomicName authorityName="Özbek" authorityYear="2023" box="[477,620,619,645]" class="Arachnida" family="Pachylaelapidae" genus="Onchodellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="291" phylum="Arthropoda" rank="species" species="circularis">
|
||
<emphasis box="[477,497,619,645]" italics="true" pageId="6" pageNumber="291">O</emphasis>
|
||
.
|
||
<emphasis box="[512,620,619,645]" italics="true" pageId="6" pageNumber="291">circularis</emphasis>
|
||
</taxonomicName>
|
||
, and the anterior part of the holoventral shield (a pair of well-sclerotised ear-like processes in
|
||
<taxonomicName authorityName="Özbek" authorityYear="2023" box="[397,543,655,681]" class="Arachnida" family="Pachylaelapidae" genus="Onchodellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="291" phylum="Arthropoda" rank="species" species="circularis">
|
||
<emphasis box="[397,417,655,681]" italics="true" pageId="6" pageNumber="291">O</emphasis>
|
||
.
|
||
<emphasis box="[435,543,655,681]" italics="true" pageId="6" pageNumber="291">circularis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation box="[562,649,655,681]" captionStart="FIGURES 9–23" captionStartId="8.[152,270,1894,1918]" captionTargetBox="[188,1394,395,1862]" captionTargetId="figure-138@8.[188,1398,381,1871]" captionTargetPageId="8" captionText="FIGURES 9–23. Onchodellus spp., female. 9–12. Epistome (O. karawaiewi, O. acrophilus, O.circularis, O. kosensis, respectively); 13–15. Spermathecal apparatus (O. acrophilus, O.circularis, O. kosensis, respectively); 16–19. Tarsus II (O. karawaiewi, O. acrophilus, O.circularis, respectively): O. acrophilus, male. 20. Tarsus II; 21. Femur, genu, tibia of Leg II: O.circularis, male. 22. Anterior part of holoventral shield; 23. Tarsus II." figureDoi="http://doi.org/10.5281/zenodo.7801155" httpUri="https://zenodo.org/record/7801155/files/figure.png" pageId="6" pageNumber="291">Fig. 22</figureCitation>
|
||
), which are absent in
|
||
<taxonomicName baseAuthorityName="Koroleva" baseAuthorityYear="1977" box="[912,1067,655,681]" class="Arachnida" family="Pachylaelapidae" genus="Onchodellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="291" phylum="Arthropoda" rank="species" species="calcaratus">
|
||
<emphasis box="[912,932,655,681]" italics="true" pageId="6" pageNumber="291">O</emphasis>
|
||
.
|
||
<emphasis box="[949,1067,655,681]" italics="true" pageId="6" pageNumber="291">calcaratus</emphasis>
|
||
</taxonomicName>
|
||
) (
|
||
<bibRefCitation author="Hirschmann, W. & Krauss, W." box="[1095,1431,654,681]" pageId="6" pageNumber="291" pagination="1 - 5" refId="ref7059" refString="Hirschmann, W. & Krauss, W. (1965) Gangsystematik der Parasitiformes. Teil 8. Gamasiden. Bestimmungstafeln von 55 Pachylaelaps - Arten. Acarologie, Schriftenreihe fur vergleichende Milbenkunde, 7, 1 - 5." type="journal article" year="1965">Hirschmann & Krauss, 1965</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Koroleva, E. V." box="[151,319,691,717]" pageId="6" pageNumber="291" pagination="411 - 483" refId="ref7140" refString="Koroleva, E. V. (1977) Family Pachylaelaptidae Vitzthum, 1931. In: Ghilyarov, M. S. & Bregetova, N. G. (Eds.), Key to the Soil Inhabiting Mites. Mesostigmata, Nauka, Leningrad, pp. 411 - 483." type="book chapter" year="1977">Koroleva 1977</bibRefCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |